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<document id="ADF3B2DDE88163C17242C8F5F0DB19C6" ID-CLB-Dataset="28007" ID-DOI="10.11646/zootaxa.4514.3.6" ID-GBIF-Dataset="0973fa16-cbab-4b38-b71c-8eb6e086dd1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2608133" ID-ZooBank="39171900-3498-4BF1-BBF1-EE30467ADEB2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553590201797" checkinUser="plazi" docAuthor="Bayarsaikhan, Ulziijargal, Lee, Dong ˗ June &amp; Bae, Yang ˗ Seop" docDate="2018" docId="03E6421C8B62E17218A9FCA0C39BF81E" docLanguage="en" docName="zootaxa.4514.3.6.pdf" docOrigin="Zootaxa 4514 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diduga bispinosa Bayarsaikhan &amp; Lee &amp; Bae 2018, n. sp." docType="treatment" docVersion="9" lastPageNumber="417" masterDocId="FFDF3A648B64E174183EFFE4C059FF8D" masterDocTitle="Three new species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Cambodia" masterLastPageNumber="424" masterPageNumber="411" pageNumber="417" updateTime="1698713313983" updateUser="ExternalLinkService">
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<mods:title id="4B411F175C030F49A0E678482ED0B756">Three new species of Diduga Moore, [1887] (Lepidoptera, Erebidae, Arctiinae) from Cambodia</mods:title>
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<mods:namePart id="F427797C146F42A365D8F1C6214ED651">Bayarsaikhan, Ulziijargal</mods:namePart>
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<mods:namePart id="A2EB931BF5F489705C3FF0A6BA350729">Lee, Dong ˗ June</mods:namePart>
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<mods:namePart id="40E1F87858294BF5C991B2804AA945D3">Bae, Yang ˗ Seop</mods:namePart>
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<mods:date id="F2354B9061AB77CF9A1439B84205E770">2018</mods:date>
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<treatment id="03E6421C8B62E17218A9FCA0C39BF81E" ID-DOI="http://doi.org/10.5281/zenodo.5968074" ID-GBIF-Taxon="154496834" ID-Zenodo-Dep="5968074" LSID="urn:lsid:plazi:treatment:03E6421C8B62E17218A9FCA0C39BF81E" httpUri="http://treatment.plazi.org/id/03E6421C8B62E17218A9FCA0C39BF81E" lastPageNumber="417" pageId="6" pageNumber="417">
<subSubSection id="C355A0818B62E17218A9FCA0C29EFCD3" box="[151,711,836,862]" pageId="6" pageNumber="417" type="nomenclature">
<paragraph id="8BF0F30A8B62E17218A9FCA0C29EFCD3" blockId="6.[151,711,836,895]" box="[151,711,836,862]" pageId="6" pageNumber="417">
<heading id="D0B844668B62E17218A9FCA0C29EFCD3" bold="true" box="[151,711,836,862]" fontSize="11" level="1" pageId="6" pageNumber="417" reason="1">
<emphasis id="B93B2F188B62E17218A9FCA0C29EFCD3" bold="true" box="[151,711,836,862]" pageId="6" pageNumber="417">
<taxonomicName id="4C4F88898B62E17218A9FCA0C22EFCD3" ID-CoL="BDYW3" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Lee &amp; Bae" authorityYear="2018" box="[151,631,836,862]" class="Insecta" family="Erebidae" genus="Diduga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="bispinosa" status="sp. nov.">
<emphasis id="B93B2F188B62E17218A9FCA0C133FCD3" bold="true" box="[151,362,836,862]" italics="true" pageId="6" pageNumber="417">Diduga bispinosa</emphasis>
Bayarsaikhan &amp; Bae
</taxonomicName>
,
<taxonomicNameLabel id="A20892638B62E1721ABAFCA0C29EFCD3" box="[644,711,836,862]" pageId="6" pageNumber="417" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C355A0818B62E17218A9FC82C146FCF2" box="[151,287,870,895]" pageId="6" pageNumber="417" type="description">
<paragraph id="8BF0F30A8B62E17218A9FC82C146FCF2" blockId="6.[151,711,836,895]" box="[151,287,870,895]" pageId="6" pageNumber="417">
(
<figureCitation id="1374EF8F8B62E17218A1FC82C0B7FCF2" box="[159,238,870,895]" captionStart="FIGURES 16" captionStartId="3.[151,264,1908,1931]" captionTargetBox="[150,1436,223,1859]" captionTargetId="figure@3.[150,1436,223,1859]" captionTargetPageId="3" captionText="FIGURES 16. Adults of genus Diduga spp. from Cambodia. 1a. D. albicosta, male (INU˗1742C). 1b. ditto, female (INU˗1998C). 2a. D. dubatolovi n. sp., male (holotype, INU˗1775C). 2b. ditto, female (paratype, INU˗1193C). 3a. D. kohkongensis n. sp., male (holotype, INU˗1412C). 3b. ditto n. sp., female (paratype, INU˗1743C). 4. D. bispinosa n. sp., male (holotype, INU˗1415C). 5. D. barlowi, male (INU˗1410C). 6. D. amoenusa, male (INU˗1411C)." httpUri="https://zenodo.org/record/2608135/files/figure.png" pageId="6" pageNumber="417">Figs. 4</figureCitation>
,
<figureCitation id="1374EF8F8B62E17218C4FC83C14CFCF2" box="[250,277,871,895]" captionStart="FIGURES 1011" captionStartId="8.[151,264,1963,1986]" captionTargetBox="[174,1425,250,1869]" captionTargetId="figure@8.[150,1436,238,1874]" captionTargetPageId="8" captionText="FIGURES 1011. Male genitalia of genus Diduga spp. from Cambodia. 10. D. kohkongensis n. sp. (holotype, INU˗1412C). 11. D. bispinosa n. sp. (holotype, INU˗1415C). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2608141/files/figure.png" pageId="6" pageNumber="417">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C355A0818B62E17218A9FC4AC27BFBBE" pageId="6" pageNumber="417" type="materials_examined">
<paragraph id="8BF0F30A8B62E17218A9FC4AC27BFBBE" blockId="6.[151,1437,942,1939]" pageId="6" pageNumber="417">
<emphasis id="B93B2F188B62E17218A9FC4AC18EFC4A" bold="true" box="[151,471,942,967]" pageId="6" pageNumber="417">
Type materials.
<materialsCitation id="3B27F9578B62E172195FFC4AC18EFC4A" ID-GBIF-Occurrence="2028143357" box="[353,471,942,967]" collectingDate="2014-12-25" collectionCode="SM, JW" collectorName="YS Bae &amp; YD Ju &amp; MJ Qi &amp; U Bayarsaikhan &amp; BS Park &amp; Na &amp; Kim &amp; DJ Lee &amp; Gen. Slide No." location="Koh Kong" pageId="6" pageNumber="417" specimenCount="1" specimenCount-male="1" stateProvince="Koh Kong" typeStatus="holotype">
<typeStatus id="54F44DA88B62E172195FFC4AC18BFC4A" box="[353,466,942,967]" pageId="6" pageNumber="417" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D4938838B62E17219DAFC4BC24CFC4B" box="[484,533,943,967]" pageId="6" pageNumber="417" type="male">1 ♂</specimenCount>
,
<collectingRegion id="498B3DE88B62E1721A19FC4AC2FBFC4A" box="[551,674,942,967]" country="Cambodia" name="Koh Kong" pageId="6" pageNumber="417">Koh Kong</collectingRegion>
mountain area,
<collectingRegion id="498B3DE88B62E1721B58FC4AC3B8FC4A" box="[870,993,942,967]" country="Cambodia" name="Koh Kong" pageId="6" pageNumber="417">Koh Kong</collectingRegion>
Prov. (N11˚4144.9″, E103˚0651.8″),
<date id="FFF1D5CA8B62E17218A9FC36C146FC66" box="[151,287,978,1003]" pageId="6" pageNumber="417" value="2014-12-25">
<collectingDate id="EFB52C228B62E17218A9FC36C146FC66" box="[151,287,978,1003]" pageId="6" pageNumber="417" value="2014-12-25">25.XII.2014</collectingDate>
</date>
(
<collectorName id="26BA96DC8B62E172190DFC37C1CBFC67" box="[307,402,979,1003]" pageId="6" pageNumber="417">YS Bae</collectorName>
,
<collectorName id="26BA96DC8B62E172199CFC37C1A8FC67" box="[418,497,979,1002]" pageId="6" pageNumber="417">YD Ju</collectorName>
,
<collectorName id="26BA96DC8B62E1721A3CFC37C216FC66" box="[514,591,978,1003]" pageId="6" pageNumber="417">MJ Qi</collectorName>
,
<collectorName id="26BA96DC8B62E1721A5EFC37C342FC66" box="[608,795,978,1003]" pageId="6" pageNumber="417">U Bayarsaikhan</collectorName>
,
<collectorName id="26BA96DC8B62E1721B15FC37C3D6FC66" box="[811,911,978,1003]" pageId="6" pageNumber="417">BS Park</collectorName>
, SM
<collectorName id="26BA96DC8B62E1721BEDFC37C3AFFC67" box="[979,1014,979,1002]" pageId="6" pageNumber="417">Na</collectorName>
, JW
<collectorName id="26BA96DC8B62E1721C09FC36C432FC66" box="[1079,1131,978,1003]" pageId="6" pageNumber="417">Kim</collectorName>
,
<collectorName id="26BA96DC8B62E1721C42FC37C48DFC66" box="[1148,1236,979,1003]" pageId="6" pageNumber="417">DJ Lee</collectorName>
),
<collectorName id="26BA96DC8B62E1721CD2FC37C5C4FC67" box="[1260,1437,978,1003]" pageId="6" pageNumber="417">Gen. Slide No.</collectorName>
INU˗1415C.
<materialsCitation id="3B27F9578B62E1721912FC12C247FBBE" ID-GBIF-Occurrence="2028143354" collectingDate="2010-05-04" collectionCode="XV" collectorName="Prov. &amp; YS Bae &amp; Le &amp; DY Chae &amp; Gen. Slide No." location="Pursat, Pursat" pageId="6" pageNumber="417" specimenCount="1" specimenCount-male="1" stateProvince="Pursat, Pursat" typeStatus="paratype">
<emphasis id="B93B2F188B62E1721912FC12C1FBFB82" bold="true" box="[300,418,1014,1039]" pageId="6" pageNumber="417">
<typeStatus id="54F44DA88B62E1721912FC12C1C4FB82" box="[300,413,1014,1039]" pageId="6" pageNumber="417" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D4938838B62E1721995FC13C181FB83" box="[427,472,1015,1039]" pageId="6" pageNumber="417" type="male">1 ♂</specimenCount>
,
<collectingRegion id="498B3DE88B62E17219DBFC13C2D8FB83" box="[485,641,1015,1038]" isEnumeration="true" pageId="6" pageNumber="417">Pursat, Pursat</collectingRegion>
<collectorName id="26BA96DC8B62E1721AB7FC13C29DFB83" box="[649,708,1015,1038]" pageId="6" pageNumber="417">Prov.</collectorName>
(N12˚0534.2″, E103˚1110.1″),
<date id="FFF1D5CA8B62E1721C06FC13C4C5FB82" box="[1080,1180,1015,1039]" pageId="6" pageNumber="417" value="2010-05-04">
<collectingDate id="EFB52C228B62E1721C06FC13C4C5FB82" box="[1080,1180,1015,1039]" pageId="6" pageNumber="417" value="2010-05-04">4.V.2010</collectingDate>
</date>
(
<collectorName id="26BA96DC8B62E1721C93FC13C550FB83" box="[1197,1289,1015,1039]" pageId="6" pageNumber="417">YS Bae</collectorName>
,
<collectionCode id="ED5E6BCF8B62E1721D28FC13C566FB83" box="[1302,1343,1015,1038]" pageId="6" pageNumber="417">XV</collectionCode>
<collectorName id="26BA96DC8B62E1721D79FC13C53EFB83" box="[1351,1383,1015,1038]" pageId="6" pageNumber="417">Le</collectorName>
,
<collectorName id="26BA96DC8B62E1721D4AFC13C08CFBBE" pageId="6" pageNumber="417">DY Chae</collectorName>
),
<collectorName id="26BA96DC8B62E17218D6FBFFC1D6FBBF" box="[232,399,1050,1075]" pageId="6" pageNumber="417">Gen. Slide No.</collectorName>
INU˗1414C
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C355A0818B62E17218F9FBDAC131FAA2" pageId="6" pageNumber="417" type="diagnosis">
<paragraph id="8BF0F30A8B62E17218F9FBDAC131FAA2" blockId="6.[151,1437,942,1939]" pageId="6" pageNumber="417">
<emphasis id="B93B2F188B62E17218F9FBDAC118FBDA" bold="true" box="[199,321,1086,1111]" pageId="6" pageNumber="417">Diagnosis.</emphasis>
By the wing pattern, the new species is similar to
<taxonomicName id="4C4F88898B62E1721B50FBDAC481FBDA" authority="Holloway, 2001" authorityName="Holloway" authorityYear="2001" box="[878,1240,1086,1111]" class="Insecta" family="Erebidae" genus="Diduga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="barlowi">
<emphasis id="B93B2F188B62E1721B50FBDAC446FBDB" box="[878,1055,1086,1110]" italics="true" pageId="6" pageNumber="417">Diduga barlowi</emphasis>
<bibRefCitation id="EFDE8EFB8B62E1721C18FBDAC481FBDA" author="Holloway, J. D." box="[1062,1240,1086,1111]" pageId="6" pageNumber="417" pagination="279 - 486" refId="ref7161" refString="Holloway, J. D. (2001) The Moths of Borneo: Family Arctiidae, Subfamily Lithosiinae. Malayan Nature Journal, 7, 279 - 486." type="journal article" year="2001">Holloway, 2001</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFDE8EFB8B62E1721CD8FBDAC378FBF6" author="Holloway, J. D." pageId="6" pageNumber="417" pagination="279 - 486" refId="ref7161" refString="Holloway, J. D. (2001) The Moths of Borneo: Family Arctiidae, Subfamily Lithosiinae. Malayan Nature Journal, 7, 279 - 486." type="journal article" year="2001">Holloway 2001: 449˗450, pl. 8: 43, fig. 448; present paper: figs. 5, 14</bibRefCitation>
), but the male genitalia structure of both species are significantly different: apex of valva with an asymmetrical projection in lower angle; upper angle of valva roundly angled; saccus narrowly “U”˗shaped, almost same length with uncus; aedeagus with two slender spine˗shaped cornuti (
<figureCitation id="1374EF8F8B62E17218C6FB2AC115FB6A" box="[248,332,1230,1255]" captionStart="FIGURES 1011" captionStartId="8.[151,264,1963,1986]" captionTargetBox="[174,1425,250,1869]" captionTargetId="figure@8.[150,1436,238,1874]" captionTargetPageId="8" captionText="FIGURES 1011. Male genitalia of genus Diduga spp. from Cambodia. 10. D. kohkongensis n. sp. (holotype, INU˗1412C). 11. D. bispinosa n. sp. (holotype, INU˗1415C). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2608141/files/figure.png" pageId="6" pageNumber="417">Fig. 11</figureCitation>
). In
<taxonomicName id="4C4F88898B62E17219BAFB2BC259FB6B" authorityName="Holloway" authorityYear="2001" box="[388,512,1230,1254]" class="Insecta" family="Erebidae" genus="Diduga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="417" phylum="Arthropoda" rank="species" species="barlowi">
<emphasis id="B93B2F188B62E17219BAFB2BC259FB6B" box="[388,512,1230,1254]" italics="true" pageId="6" pageNumber="417">D. barlowi</emphasis>
</taxonomicName>
, apex of valva with large, less than asymmetrical projection in upper and lower angles; saccus broadly “U”˗shaped, almost two times shorter than uncus; aedeagus with a small spine˗shaped cornutus (
<figureCitation id="1374EF8F8B62E1721938FAF2C103FAA2" box="[262,346,1302,1327]" captionStart="FIGURES 1216" captionStartId="10.[151,264,1968,1991]" captionTargetBox="[154,1423,261,1860]" captionTargetId="figure@10.[150,1436,253,1889]" captionTargetPageId="10" captionText="FIGURES 1216. Male genitalia of genus Diduga spp. from Cambodia. 12. D. albicosta (INU˗1194C). 13. D. amoenusa (INU˗1411C). 14. D. barlowi (INU˗1410C). 15. D. alternota (INU˗1424C). 16. D. annulata (INU˗1420C). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2608143/files/figure.png" pageId="6" pageNumber="417">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C355A0818B62E17218F9FADEC25CF8AA" pageId="6" pageNumber="417" type="description">
<paragraph id="8BF0F30A8B62E17218F9FADEC140F98A" blockId="6.[151,1437,942,1939]" pageId="6" pageNumber="417">
<emphasis id="B93B2F188B62E17218F9FADEC100FADE" bold="true" box="[199,345,1338,1363]" pageId="6" pageNumber="417">Description.</emphasis>
<emphasis id="B93B2F188B62E172195DFADFC1C6FADE" box="[355,415,1339,1363]" italics="true" pageId="6" pageNumber="417">Adult</emphasis>
(
<figureCitation id="1374EF8F8B62E172198CFADEC1A3FADE" box="[434,506,1338,1363]" captionStart="FIGURES 16" captionStartId="3.[151,264,1908,1931]" captionTargetBox="[150,1436,223,1859]" captionTargetId="figure@3.[150,1436,223,1859]" captionTargetPageId="3" captionText="FIGURES 16. Adults of genus Diduga spp. from Cambodia. 1a. D. albicosta, male (INU˗1742C). 1b. ditto, female (INU˗1998C). 2a. D. dubatolovi n. sp., male (holotype, INU˗1775C). 2b. ditto, female (paratype, INU˗1193C). 3a. D. kohkongensis n. sp., male (holotype, INU˗1412C). 3b. ditto n. sp., female (paratype, INU˗1743C). 4. D. bispinosa n. sp., male (holotype, INU˗1415C). 5. D. barlowi, male (INU˗1410C). 6. D. amoenusa, male (INU˗1411C)." httpUri="https://zenodo.org/record/2608135/files/figure.png" pageId="6" pageNumber="417">Fig. 4</figureCitation>
). Length of forewing
<quantity id="4CB75EEF8B62E1721AC3FADEC301FADF" box="[765,856,1338,1363]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="6" pageNumber="417" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4˗5 mm</quantity>
in male. Head, patagium and tegula deep yellow. Thorax deep yellow, except dark brown second half. Forewing ground color dark brown (dark brown patch looks irregularly “Z” or “S”˗shaped), with deep yellow margin from basal to tornus, which hind border of margin broadly dentate in basal half of costal area and tornus; with several small, dark brown dots in deep yellow margin; cilia deep yellow. Hindwing ground color gray, tinged with pale brown; cilia pale brown. Abdomen brown, with deep yellow tuft.
</paragraph>
<paragraph id="8BF0F30A8B62E17218F9F9F6C0B7F88E" blockId="6.[151,1437,942,1939]" pageId="6" pageNumber="417">
Male genitalia (
<figureCitation id="1374EF8F8B62E1721947F9F6C192F9A6" box="[377,459,1554,1579]" captionStart="FIGURES 1011" captionStartId="8.[151,264,1963,1986]" captionTargetBox="[174,1425,250,1869]" captionTargetId="figure@8.[150,1436,238,1874]" captionTargetPageId="8" captionText="FIGURES 1011. Male genitalia of genus Diduga spp. from Cambodia. 10. D. kohkongensis n. sp. (holotype, INU˗1412C). 11. D. bispinosa n. sp. (holotype, INU˗1415C). Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/2608141/files/figure.png" pageId="6" pageNumber="417">Fig. 11</figureCitation>
). Uncus slender, waved, with strongly hook˗shaped apex. Tegumen triangular, weakly sclerotized. Valva short, rectangular, weakly sclerotized, with a stout, asymmetrical process (almost same length with valva) in lower angle of apex; process of left valva almost straight, tubular, with short, subapical and medial spines; process of right valva stout, weakly sickle˗shaped, with an apical spine and a large medial branch pointed, with an arch˗shaped process in inward. Juxta trapezoid, weakly sclerotized. Saccus “U”˗shaped, almost same length with uncus. Aedeagus slender, “S”˗shaped, weakly sclerotized and vesica with two slender spine˗shaped cornuti.
</paragraph>
<paragraph id="8BF0F30A8B62E17218F9F8EAC25CF8AA" blockId="6.[151,1437,942,1939]" box="[199,517,1806,1831]" pageId="6" pageNumber="417">Female genitalia. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C355A0818B62E17218F9F8D6C360F8C6" box="[199,825,1842,1867]" pageId="6" pageNumber="417" type="distribution">
<paragraph id="8BF0F30A8B62E17218F9F8D6C360F8C6" blockId="6.[151,1437,942,1939]" box="[199,825,1842,1867]" pageId="6" pageNumber="417">
<emphasis id="B93B2F188B62E17218F9F8D6C139F8C6" bold="true" box="[199,352,1842,1867]" pageId="6" pageNumber="417">Distribution.</emphasis>
<collectingCountry id="F358B39A8B62E1721959F8D6C183F8C6" box="[359,474,1842,1867]" name="Cambodia" pageId="6" pageNumber="417">Cambodia</collectingCountry>
(
<collectingRegion id="498B3DE88B62E17219D7F8D6C23AF8C6" box="[489,611,1842,1867]" country="Cambodia" name="Koh Kong" pageId="6" pageNumber="417">Koh Kong</collectingRegion>
,
<collectingRegion id="498B3DE88B62E1721A50F8D7C2EDF8C7" box="[622,692,1843,1866]" country="Cambodia" name="Pursat" pageId="6" pageNumber="417">Pursat</collectingRegion>
provinces).
</paragraph>
</subSubSection>
<subSubSection id="C355A0818B62E17218F9F8B2C39BF81E" pageId="6" pageNumber="417" type="etymology">
<paragraph id="8BF0F30A8B62E17218F9F8B2C39BF81E" blockId="6.[151,1437,942,1939]" pageId="6" pageNumber="417">
<emphasis id="B93B2F188B62E17218F9F8B2C117F8E2" bold="true" box="[199,334,1878,1903]" pageId="6" pageNumber="417">Etymology.</emphasis>
The specific epithet is derived from the Latin “
<emphasis id="B93B2F188B62E1721BA7F8B2C3F6F8E3" box="[921,943,1878,1902]" italics="true" pageId="6" pageNumber="417">bi</emphasis>
” meaning “twice” and “
<emphasis id="B93B2F188B62E1721CE4F8B3C57BF8E3" box="[1242,1314,1879,1902]" italics="true" pageId="6" pageNumber="417">spinos</emphasis>
” meaning “thorny”, refers to the two process of the left valva in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>