282 lines
36 KiB
XML
282 lines
36 KiB
XML
<document id="287259F6DA19C3BE62887246D4EC6C13" ID-DOI="10.11646/zootaxa.4007.3.6" ID-GBIF-Dataset="1c40eef0-6b7e-4a78-9d56-0e472e2b51a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244253" ID-ZooBank="8FF190D2-3ECA-4819-93D5-1EF29C3AB675" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461219579834" checkinUser="plazi" docAuthor="Suzuki, Toshiyuki, Greenfield, David W. & Motomura, Hiroyuki" docDate="2015" docId="03E587BACF26FFA3B1A0FAD57D24FC8A" docLanguage="en" docName="zt04007p408.pdf" docOrigin="Zootaxa 4007 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eviota rubrimaculata Suzuki, Greenfield & Motomura, 2015, n. sp." docType="treatment" docUuid="7B19C50F-EB22-47D3-8814-775AC256E33A" docUuidSource="ZooBank" docVersion="9" lastPageNumber="407" masterDocId="FFDCFFC2CF23FFABB137FFB07932FFCB" masterDocTitle="Two new dwarfgobies (Teleostei: Gobiidae) from the Ryukyu Islands, Japan: Eviota flavipinnata and Eviota rubrimaculata" masterLastPageNumber="408" masterPageNumber="399" pageNumber="404" updateTime="1698603715224" updateUser="plazi">
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<mods:mods id="A6742FB04F419D0BF8A8F5A11604BA02" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="439F6A48886CA298BB02C0D08D755DC2">
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<mods:title id="3B8B4812F0BD91A56D70A2D5BF0E5045">Two new dwarfgobies (Teleostei: Gobiidae) from the Ryukyu Islands, Japan: Eviota flavipinnata and Eviota rubrimaculata</mods:title>
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</mods:titleInfo>
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<mods:name id="6B7A1632A6073F405E8B28A21675E600" type="personal">
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<mods:role id="FFF4E2D4EA97E6546336C8D7A2667F5B">
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<mods:roleTerm id="0D8AA0C5452F2E73FD804216928EC5A4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4AE088642A5B5F0DDD9593F48CE63169">Suzuki, Toshiyuki</mods:namePart>
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<mods:name id="FD6D5BFD77B337123ACF38BAF0A19F48" type="personal">
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<mods:roleTerm id="620B817D9A0D70AB370D9C8FEBE70CBC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="75B2F8646E1CE10F08C76F6DCC4D5B69">Greenfield, David W.</mods:namePart>
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</mods:name>
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<mods:name id="9640EBAA012B5303524235E683DF31B9" type="personal">
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<mods:roleTerm id="C1F2902CD7BDFFA4396ACCDF890EA4D7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8F1550EB6DD00C2E67BB777648EDEB9D">Motomura, Hiroyuki</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="30A5DCD438204E7594AA15931B94C7CD">text</mods:typeOfResource>
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<mods:title id="67232F8C513D5B0FEF41D3664D6DADF2">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="54B524BE999EEA9491F779C8B373C000">
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<mods:date id="5B373FBE5A994B7202C974B938C6395F">2015</mods:date>
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<mods:detail id="DE8A4DF27B0B06DBB9C67B9E37E5341A" type="volume">
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<mods:number id="4F63B3B32999956E314F9CF2948E20FC">4007</mods:number>
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</mods:detail>
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<mods:detail id="FF9AF5B7D2C8E184F28764A331D5611F" type="issue">
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<mods:number id="C26EE664AC4277321DCF49EDB4C1FA5B">3</mods:number>
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<mods:end id="A83B777B4E925A96DC76D4DFD7151F9A">408</mods:end>
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<mods:classification id="C19A42AEB84697F94A3AE2EC1D46995B">journal article</mods:classification>
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<mods:identifier id="D2A2ADB4795F49D0E96DCCD2D7EDD1CE" type="DOI">10.11646/zootaxa.4007.3.6</mods:identifier>
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<mods:identifier id="2343715AA5023DAB9FD8EA2A3FD54022" type="GBIF-Dataset">1c40eef0-6b7e-4a78-9d56-0e472e2b51a8</mods:identifier>
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<mods:identifier id="E0E1589628795F77C5DFCB5461E9F3DA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="6C77293104780C2AD3BB736D64F61DEF" type="Zenodo-Dep">244253</mods:identifier>
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<mods:identifier id="1CEEED175F696B223C946F9592695BFD" type="ZooBank">8FF190D2-3ECA-4819-93D5-1EF29C3AB675</mods:identifier>
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</mods:mods>
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<treatment id="03E587BACF26FFA3B1A0FAD57D24FC8A" ID-DOI="http://doi.org/10.5281/zenodo.6102395" ID-GBIF-Taxon="119631384" ID-Zenodo-Dep="6102395" LSID="urn:lsid:zoobank.org:act:7B19C50F-EB22-47D3-8814-775AC256E33A" httpUri="http://treatment.plazi.org/id/03E587BACF26FFA3B1A0FAD57D24FC8A" lastPageId="8" lastPageNumber="407" pageId="5" pageNumber="404">
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<subSubSection id="C3566527CF26FFAEB1A0FAD57AB6F9E4" pageId="5" pageNumber="404" type="nomenclature">
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<paragraph id="8BF336ACCF26FFAEB1A0FAD578D1FAB4" blockId="5.[151,571,1381,1513]" box="[151,483,1381,1408]" pageId="5" pageNumber="404">
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<heading id="D0BB81C0CF26FFAEB1A0FAD578D1FAB4" bold="true" box="[151,483,1381,1408]" fontSize="11" level="1" pageId="5" pageNumber="404" reason="1">
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||
<emphasis id="B938EABECF26FFAEB1A0FAD578D1FAB4" bold="true" box="[151,483,1381,1408]" pageId="5" pageNumber="404">
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||
<taxonomicName id="4C4C4D2FCF26FFAEB1A0FAD578A8FAB4" ID-CoL="3DHNL" box="[151,410,1381,1407]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="404" phylum="Chordata" rank="species" species="rubrimaculata" status="sp. nov.">
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<emphasis id="B938EABECF26FFAEB1A0FAD578A8FAB4" bold="true" box="[151,410,1381,1407]" italics="true" pageId="5" pageNumber="404">Eviota rubrimaculata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B57C5CF26FFAEB096FAD678D1FAB4" box="[417,483,1381,1408]" pageId="5" pageNumber="404" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF336ACCF26FFAEB1A0FA38784BFA6A" blockId="5.[151,571,1381,1513]" box="[151,377,1416,1441]" pageId="5" pageNumber="404">Redspot Dwarfgoby</paragraph>
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<paragraph id="8BF336ACCF26FFAEB1A0FA1E7812FA22" blockId="5.[151,571,1381,1513]" pageId="5" pageNumber="404">
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New Japanese-name: Zakuro-Isohaze (
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<figureCitation id="13772A29CF26FFAEB1A8FA60782AFA22" box="[159,280,1488,1513]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStartId-0="6.[151,250,1184,1206]" captionStartId-1="6.[151,250,1735,1757]" captionStartId-2="7.[151,250,783,805]" captionStartId-3="7.[151,250,1349,1371]" captionTargetBox-0="[267,1355,824,1098]" captionTargetBox-1="[170,1416,1255,1714]" captionTargetBox-2="[169,1417,337,762]" captionTargetBox-3="[206,1363,942,1230]" captionTargetId-0="figure@6.[179,1408,733,1162]" captionTargetId-1="figure@6.[170,1417,1255,1714]" captionTargetId-2="figure@7.[169,1417,337,762]" captionTargetId-3="figure@7.[151,1436,886,1327]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionText-0="FIGURE 7. Eviota rubrimaculata, preserved holotype, KPM-NI 30770, light background. Photograph by Toshiyuki Suzuki." captionText-1="FIGURE 8. Eviota rubrimaculata, preserved holotype, KPM-NI 30770, dark background. Photograph by Toshiyuki Suzuki." captionText-2="FIGURE 9. Eviota rubrimaculata, freshly collected holotype, KPM-NI 30770, dark background. Photograph by Hiroshi Senou." captionText-3="FIGURE 10. Eviota rubrimaculata, freshly collected holotype, KPM-NI 30770, light background. Photograph by Hiroshi Senou." httpUri-0="https://zenodo.org/record/244260/files/figure.png" httpUri-1="https://zenodo.org/record/244261/files/figure.png" httpUri-2="https://zenodo.org/record/244262/files/figure.png" httpUri-3="https://zenodo.org/record/244263/files/figure.png" pageId="5" pageNumber="404">Figs. 7–10</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF336ACCF26FFAEB1A0F9AA7AB6F9E4" blockId="5.[151,900,1561,1583]" box="[151,900,1561,1583]" pageId="5" pageNumber="404">
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<taxonomicName id="4C4C4D2FCF26FFAEB1A0F9AA79EBF9E4" box="[151,217,1562,1583]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="404" phylum="Chordata" rank="genus">
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<emphasis id="B938EABECF26FFAEB1A0F9AA79EBF9E4" box="[151,217,1562,1583]" italics="true" pageId="5" pageNumber="404">Eviota</emphasis>
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</taxonomicName>
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sp. 4. Suzuki 2014: 517 (Yoron Island, the Ryukyu Islands,
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<collectingCountry id="F35B763CCF26FFAEB20CF9AA7A4BF9E4" box="[827,889,1562,1583]" name="Japan" pageId="5" pageNumber="404">Japan</collectingCountry>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3566527CF26FFAEB1A0F9EC7A74F953" pageId="5" pageNumber="404" type="materials_examined">
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<paragraph id="8BF336ACCF26FFAEB1A0F9EC7A74F953" blockId="5.[151,1436,1627,2015]" pageId="5" pageNumber="404">
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<emphasis id="B938EABECF26FFAEB1A0F9EC7839F9BF" bold="true" box="[151,267,1628,1652]" pageId="5" pageNumber="404">
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<typeStatus id="54F7880ECF26FFAEB1A0F9EC7834F9BF" box="[151,262,1628,1652]" pageId="5" pageNumber="404" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED5DAE69CF26FFAEB026F9EC7847F9B8" LSID="urn:lsid:biocol.org:col:35223" box="[273,373,1628,1651]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35223" name="Kanagawa Prefectural Museum of Natural History" pageId="5" pageNumber="404">KPM-NI</collectionCode>
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30770, 12.0 mm, female, off Maehama, Yoron Island, the Ryukyu Islands,
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<collectingCountry id="F35B763CCF26FFAEB5F7F9EC7C31F9B8" box="[1216,1283,1628,1651]" name="Japan" pageId="5" pageNumber="404">Japan</collectingCountry>
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,
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<quantity id="4CB49B49CF26FFAEB439F9EB7C62F9B8" box="[1294,1360,1627,1652]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="5" pageNumber="404" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">6-8 m</quantity>
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depth, 27°01ʹ13ʺN, 128°26ʹ26ʺE,
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<collectionCode id="ED5DAE69CF26FFAEB0F2F9307B28F953" box="[453,538,1664,1688]" httpUri="http://grbio.org/cool/r700-376n" name="Kagoshima University Museum" pageId="5" pageNumber="404">KAUM</collectionCode>
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Fish Team,
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<date id="FFF2106CCF26FFAEB392F9307A73F953" box="[677,833,1664,1688]" pageId="5" pageNumber="404" value="2012-08-14">14 Aug. 2012</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3566527CF26FFAEB1F0F9137B7CF8B8" pageId="5" pageNumber="404" type="diagnosis">
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<paragraph id="8BF336ACCF26FFAEB1F0F9137B7CF8B8" blockId="5.[151,1436,1627,2015]" pageId="5" pageNumber="404">
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<emphasis id="B938EABECF26FFAEB1F0F9137873F977" bold="true" box="[199,321,1699,1724]" pageId="5" pageNumber="404">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="4C4C4D2FCF26FFAEB0EAF9147B14F970" box="[477,550,1700,1723]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="404" phylum="Chordata" rank="genus">
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<emphasis id="B938EABECF26FFAEB0EAF9147B14F970" box="[477,550,1700,1723]" italics="true" pageId="5" pageNumber="404">Eviota</emphasis>
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</taxonomicName>
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with cephalic sensory-canal pore system pattern 2 (lacking only H [IT] pore); some pectoral-fin rays branched; dorsal/anal fin-ray formula 9/8; 5th pelvic-fin ray absent; no prominent distinct dark spots on pectoral-fin base; no caudal-peduncle spot over preural centrum; no postocular spots; no strong, dark spots on caudal fin. Two dark spots over anal-fin base; first dorsal fin clear with some light orange on anterodorsal portion and dark distal margin; second dorsal fin with red-orange spots on basal half, distal margin black and yellow; body with bright red markings.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3566527CF26FFA3B1F0F8CD7AB0FF33" lastPageId="8" lastPageNumber="407" pageId="5" pageNumber="404" type="description">
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<paragraph id="8BF336ACCF26FFADB1F0F8CD7A17FDD3" blockId="5.[151,1436,1627,2015]" lastBlockId="6.[151,1436,151,680]" lastPageId="6" lastPageNumber="405" pageId="5" pageNumber="404">
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<emphasis id="B938EABECF26FFAEB1F0F8CD786BF85D" bold="true" box="[199,345,1917,1942]" pageId="5" pageNumber="404">Description.</emphasis>
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Dorsal-fin rays VI-I,9, first dorsal fin triangular in shape, first spine longest and slightly elongated, but all spines not filamentous (not extending back to origin of second dorsal fin), all second dorsal-fin soft rays branched except first soft ray, the last ray branched to base; anal-fin rays I,8, all soft rays branched, the last ray branched to base; pectoral-fin rays 16, 11–15th rays branched, pectoral fin reaching to above third anal-fin soft ray base; pelvic fins without basal membrane, no pelvic frenum, pelvic-fin membrane reduced, pelvic-fin rays I,4, 4th pelvic-fin ray with 5 branches, 1-2 segments between branches, pelvic fins reaching to anal-fin origin; 13 branched and 17 segmented caudal-fin rays; lateral scale rows 23, transverse scale rows 5, no scales on head, nape, breast, pectoral-fin base, midline of belly and narrow areas along base of first dorsal fin, scales on body finely ctenoid except belly where they are cycloid; cephalic sensory-canal pore system pattern 2 (lacking only the H (IT) pore): anterior oculoscapular canal with pores B', single C, single D, E and F', two preopercular-canal pores, N' and O'; female urogenital papilla smooth, bulbous, with six short finger-like projections on end, not reaching to anal-fin origin; mouth oblique, both jaws equal, maxilla extending posterior to vertical at center of pupil; anterior nares tube not extending to upper lip and not dark in color; gill opening extending forward to below posteroventral edge of operculum, gill membranes attached anteriorly to isthmus.
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</paragraph>
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<paragraph id="8BF336ACCF25FFADB1F0FD947C47FD63" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="405">
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<emphasis id="B938EABECF25FFADB1F0FD947844FDF7" bold="true" box="[199,374,548,572]" pageId="6" pageNumber="405">Measurements</emphasis>
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. Standard length 12.0 mm; head length 31.7; origin of first dorsal fin 42.9, slightly behind above posterior end of pectoral-fin base; origin of second dorsal fin 62.7; origin of anal fin 64.3, at vertical through second soft ray base of second dorsal fin; caudal-peduncle length 23.0; caudal-peduncle depth 11.1; body depth 20.6; eye diameter 9.5; snout length 6.0; upper-jaw length 11.5; pectoral-fin length 31.7; pelvic-fin length 30.2.
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</paragraph>
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<caption id="DF336624CF25FFADB1A0FB107CB4FB7D" box="[151,1414,1184,1206]" httpUri="https://zenodo.org/record/244260/files/figure.png" pageId="6" pageNumber="405" targetBox="[267,1355,824,1098]" targetPageId="6">
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<paragraph id="8BF336ACCF25FFADB1A0FB107CB4FB7D" blockId="6.[151,1414,1184,1206]" box="[151,1414,1184,1206]" pageId="6" pageNumber="405">
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<emphasis id="B938EABECF25FFADB1A0FB107820FB7E" bold="true" box="[151,274,1184,1206]" pageId="6" pageNumber="405">FIGURE 7.</emphasis>
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<emphasis id="B938EABECF25FFADB02EFB1178C8FB7E" box="[281,506,1184,1206]" italics="true" pageId="6" pageNumber="405">
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<taxonomicName id="4C4C4D2FCF25FFADB02EFB1178C4FB7E" box="[281,502,1184,1206]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="405" phylum="Chordata" rank="species" species="rubrimaculata">Eviota rubrimaculata</taxonomicName>
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,
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</emphasis>
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preserved holotype, KPM-NI 30770, light background. Photograph by Toshiyuki Suzuki.
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</paragraph>
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</caption>
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<caption id="DF336624CF25FFADB1A0F9777CB7F916" box="[151,1413,1735,1757]" httpUri="https://zenodo.org/record/244261/files/figure.png" pageId="6" pageNumber="405" targetBox="[170,1416,1255,1714]" targetPageId="6">
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<paragraph id="8BF336ACCF25FFADB1A0F9777CB7F916" blockId="6.[151,1413,1735,1757]" box="[151,1413,1735,1757]" pageId="6" pageNumber="405">
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||
<emphasis id="B938EABECF25FFADB1A0F9777820F917" bold="true" box="[151,274,1735,1757]" pageId="6" pageNumber="405">FIGURE 8.</emphasis>
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<emphasis id="B938EABECF25FFADB02EF97878C8F916" box="[281,506,1736,1757]" italics="true" pageId="6" pageNumber="405">
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||
<taxonomicName id="4C4C4D2FCF25FFADB02EF97878C4F916" box="[281,502,1736,1757]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="405" phylum="Chordata" rank="species" species="rubrimaculata">Eviota rubrimaculata</taxonomicName>
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,
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</emphasis>
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preserved holotype, KPM-NI 30770, dark background. Photograph by Toshiyuki Suzuki.
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</paragraph>
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</caption>
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<paragraph id="8BF336ACCF25FFACB1F0F8B478ACFED6" blockId="6.[151,1436,1796,2038]" lastBlockId="7.[151,1437,151,285]" lastPageId="7" lastPageNumber="406" pageId="6" pageNumber="405">
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||
<emphasis id="B938EABECF25FFADB1F0F8B47B60F8D6" bold="true" box="[199,594,1796,1821]" pageId="6" pageNumber="405">
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||
Color in preservative of
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<typeStatus id="54F7880ECF25FFADB0DCF8B57B60F8D6" box="[491,594,1797,1821]" pageId="6" pageNumber="405" type="holotype">holotype</typeStatus>
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</emphasis>
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(
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<figureCitation id="13772A29CF25FFADB355F8B57B82F8D6" box="[610,688,1797,1822]" captionStart="FIGURE 7" captionStartId="6.[151,250,1184,1206]" captionTargetBox="[267,1355,824,1098]" captionTargetId="figure@6.[179,1408,733,1162]" captionTargetPageId="6" captionText="FIGURE 7. Eviota rubrimaculata, preserved holotype, KPM-NI 30770, light background. Photograph by Toshiyuki Suzuki." httpUri="https://zenodo.org/record/244260/files/figure.png" pageId="6" pageNumber="405">Figs. 7</figureCitation>
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&
|
||
<figureCitation id="13772A29CF25FFADB3E2F8B67BD7F8D5" box="[725,741,1798,1822]" captionStart="FIGURE 8" captionStartId="6.[151,250,1735,1757]" captionTargetBox="[170,1416,1255,1714]" captionTargetId="figure@6.[170,1417,1255,1714]" captionTargetPageId="6" captionText="FIGURE 8. Eviota rubrimaculata, preserved holotype, KPM-NI 30770, dark background. Photograph by Toshiyuki Suzuki." httpUri="https://zenodo.org/record/244261/files/figure.png" pageId="6" pageNumber="405">8</figureCitation>
|
||
). Background color of head and body translucent gray. Iris of eye black, pupil clear. A large brown-orange blotch behind upper half of eye and top of head. Cluster of dark chromatophores on snout above nares tube, another dark cluster at end of upper jaw. Scattered individual or clusters of dark chromatophores on side of head, more concentrated around eye, with a line of dark chromatophores extending from eye at five o’clock position to gular area. Operculum with scattered dark chromatophores and a pinkish wash extending to pectoral-fin base through cluster of dark chromatophores crossing center of base. Body with very few small dark chromatophores, mostly above midline. Edges of scales on dorsum under second dorsal fin dark. First dorsal fin mostly clear with a few dark chromatophores along basal half and more on distal margin of fin. Second dorsal fin with scattered dark chromatophores on rays and small dark chromatophores on distal margin. Caudal fin with a few scattered dark chromatophores on rays. Anal fin dusky, darker than other fins. Pectoral and pelvic fins immaculate.
|
||
</paragraph>
|
||
<caption id="DF336624CF24FFACB1A0FCBF79EFFC8F" httpUri="https://zenodo.org/record/244262/files/figure.png" pageId="7" pageNumber="406" targetBox="[169,1417,337,762]" targetPageId="7">
|
||
<paragraph id="8BF336ACCF24FFACB1A0FCBF79EFFC8F" blockId="7.[151,1436,783,836]" pageId="7" pageNumber="406">
|
||
<emphasis id="B938EABECF24FFACB1A0FCBF7825FCEF" bold="true" box="[151,279,783,805]" pageId="7" pageNumber="406">FIGURE 9.</emphasis>
|
||
<emphasis id="B938EABECF24FFACB015FCA07B3AFCEE" box="[290,520,784,805]" italics="true" pageId="7" pageNumber="406">
|
||
<taxonomicName id="4C4C4D2FCF24FFACB015FCA07B37FCEE" box="[290,517,784,805]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="406" phylum="Chordata" rank="species" species="rubrimaculata">Eviota rubrimaculata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
freshly collected holotype, KPM-NI 30770, dark background. Photograph by Hiroshi Senou.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF336624CF24FFACB1A0FAF579EFFAB2" httpUri="https://zenodo.org/record/244263/files/figure.png" pageId="7" pageNumber="406" targetBox="[206,1363,942,1230]" targetPageId="7">
|
||
<paragraph id="8BF336ACCF24FFACB1A0FAF579EFFAB2" blockId="7.[151,1436,1348,1401]" pageId="7" pageNumber="406">
|
||
<emphasis id="B938EABECF24FFACB1A0FAF57811FA91" bold="true" box="[151,291,1349,1371]" pageId="7" pageNumber="406">FIGURE 10.</emphasis>
|
||
<taxonomicName id="4C4C4D2FCF24FFACB01AFAF57B39FA91" box="[301,523,1349,1370]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="406" phylum="Chordata" rank="species" species="rubrimaculata">
|
||
<emphasis id="B938EABECF24FFACB01AFAF57B39FA91" box="[301,523,1349,1370]" italics="true" pageId="7" pageNumber="406">Eviota rubrimaculata</emphasis>
|
||
</taxonomicName>
|
||
, freshly collected holotype, KPM-NI 30770, light background. Photograph by Hiroshi Senou.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF336ACCF24FFA3B1F0FA107AB0FF33" blockId="7.[151,1436,1440,2006]" lastBlockId="8.[151,1436,151,833]" lastPageId="8" lastPageNumber="407" pageId="7" pageNumber="406">
|
||
<emphasis id="B938EABECF24FFACB1F0FA1078D6FA72" bold="true" box="[199,484,1440,1465]" pageId="7" pageNumber="406">
|
||
Color of fresh
|
||
<typeStatus id="54F7880ECF24FFACB04AFA1178D6FA72" box="[381,484,1441,1465]" pageId="7" pageNumber="406" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13772A29CF24FFACB0C1FA107B75FA72" box="[502,583,1440,1465]" captionStart="FIGURE 9" captionStartId="7.[151,250,783,805]" captionTargetBox="[169,1417,337,762]" captionTargetId="figure@7.[169,1417,337,762]" captionTargetPageId="7" captionText="FIGURE 9. Eviota rubrimaculata, freshly collected holotype, KPM-NI 30770, dark background. Photograph by Hiroshi Senou." httpUri="https://zenodo.org/record/244262/files/figure.png" pageId="7" pageNumber="406">Figs. 9</figureCitation>
|
||
&
|
||
<figureCitation id="13772A29CF24FFACB346FA117BA2FA72" box="[625,656,1441,1465]" captionStart="FIGURE 10" captionStartId="7.[151,250,1349,1371]" captionTargetBox="[206,1363,942,1230]" captionTargetId="figure@7.[151,1436,886,1327]" captionTargetPageId="7" captionText="FIGURE 10. Eviota rubrimaculata, freshly collected holotype, KPM-NI 30770, light background. Photograph by Hiroshi Senou." httpUri="https://zenodo.org/record/244263/files/figure.png" pageId="7" pageNumber="406">10</figureCitation>
|
||
). Background color of head and body translucent gray-white. Most obvious markings red spots on areas as described below. Pupil of eye black, iris silver with red spokes radiating out from the pupil. Red blotches around eye; two between eye and upper jaw, one at eight o’clock, a second at seven o’clock. Long red line extending from under eye at six o’clock down behind jaws onto gular area. Side of head with several scattered large red spots mixed with black chromatophores extending up to behind eye. Series of red lines extending back from lower edge of preoperculum and operculum across branchiostegals. Pectoral-fin base with two red bars, one dorsal and one ventral, separated by yellow area filed with small dark chromatophores. Top of head, nape and area under pectoral-fin base white, side of abdomen with two large red spots followed by smaller spot above anal opening. Lower part of abdomen white, both upper and lower areas heavily peppered with large dark chromatophores. Upper half of body translucent gray with just a few scattered scales with dark margins. Grayblue line along vertebral column. Ventral half of body with six red spots, two over anal-fin base and four more on ventral edge of caudal peduncle. Each of these spots, except last at caudal-fin base, with dark subcutaneous bar extending up to just below vertebral column. Series of small white spots, about half the size of pupil, on body; anteriormost behind second large red spot on abdomen, second above anal-fin origin, third above center of anal fin, fourth at end of anal fin, fifth and sixth on caudal peduncle near vertebral column, and seventh at top of caudal peduncle. First dorsal fin clear with some light orange on anterodorsal portion and a dark distal margin. Second dorsal fin with red-orange spots on basal half, distal margin black and yellow. Anal fin black, with two red areas extending from body spots onto basal portion of fin. Caudal fin with scattered orange spots on central rays and light gray spots on remaining rays. Pectoral and pelvic fins immaculate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566527CF2BFFA3B1F0FEB37D42FED6" box="[199,1136,259,285]" pageId="8" pageNumber="407" type="distribution">
|
||
<paragraph id="8BF336ACCF2BFFA3B1F0FEB37D42FED6" blockId="8.[151,1436,151,833]" box="[199,1136,259,285]" pageId="8" pageNumber="407">
|
||
<emphasis id="B938EABECF2BFFA3B1F0FEB37852FED7" bold="true" box="[199,352,259,284]" pageId="8" pageNumber="407">Distribution.</emphasis>
|
||
Yoron Island, the Ryukyu Islands,
|
||
<collectingCountry id="F35B763CCF2BFFA3B3DDFEB57A18FED7" box="[746,810,261,284]" name="Japan" pageId="8" pageNumber="407">Japan</collectingCountry>
|
||
(Suzuki 2014; in this study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566527CF2BFFA3B1F0FE987807FD41" pageId="8" pageNumber="407" type="etymology">
|
||
<paragraph id="8BF336ACCF2BFFA3B1F0FE987A17FEAE" blockId="8.[151,1436,151,833]" pageId="8" pageNumber="407">
|
||
<emphasis id="B938EABECF2BFFA3B1F0FE98787CFE8B" bold="true" box="[199,334,296,320]" pageId="8" pageNumber="407">Etymology.</emphasis>
|
||
The specific epithet is an adjective from the Latin
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B2B6FE987AF3FE8B" box="[897,961,296,320]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="rubra">
|
||
<emphasis id="B938EABECF2BFFA3B2B6FE987AF3FE8B" box="[897,961,296,320]" italics="true" pageId="8" pageNumber="407">rubra</emphasis>
|
||
</taxonomicName>
|
||
(red) and
|
||
<emphasis id="B938EABECF2BFFA3B504FE987DB4FE8B" box="[1075,1158,296,320]" italics="true" pageId="8" pageNumber="407">macula</emphasis>
|
||
(spot) in reference to the distinctive red spots on
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B096FEFD7B50FEAF" box="[417,610,332,356]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="rubrimaculata">
|
||
<emphasis id="B938EABECF2BFFA3B096FEFD7B50FEAF" box="[417,610,332,356]" italics="true" pageId="8" pageNumber="407">E. rubrimaculata</emphasis>
|
||
</taxonomicName>
|
||
’s head and body.
|
||
</paragraph>
|
||
<paragraph id="8BF336ACCF2BFFA3B1F0FEC27807FD41" blockId="8.[151,1436,151,833]" pageId="8" pageNumber="407">
|
||
<emphasis id="B938EABECF2BFFA3B1F0FEC2785FFE40" bold="true" box="[199,365,370,395]" pageId="8" pageNumber="407">Comparisons.</emphasis>
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B043FEC37B54FE40" box="[372,614,371,395]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="rubrimaculata">
|
||
<emphasis id="B938EABECF2BFFA3B043FEC37B54FE40" box="[372,614,371,395]" italics="true" pageId="8" pageNumber="407">Eviota rubrimaculata</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B201FEC37AEFFE40" box="[822,989,371,395]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="flavipinnata">
|
||
<emphasis id="B938EABECF2BFFA3B201FEC37AEFFE40" box="[822,989,371,395]" italics="true" pageId="8" pageNumber="407">E. flavipinnata</emphasis>
|
||
</taxonomicName>
|
||
, differing only in lacking the 5th pelvicfin ray, whereas it is 12% of the 4th ray in
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B354FE2A7A3AFE7A" box="[611,776,409,433]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="flavipinnata">
|
||
<emphasis id="B938EABECF2BFFA3B354FE2A7B46FE7A" box="[611,628,410,433]" italics="true" pageId="8" pageNumber="407">E</emphasis>
|
||
.
|
||
<emphasis id="B938EABECF2BFFA3B3B6FE297A3AFE7A" box="[641,776,409,433]" italics="true" pageId="8" pageNumber="407">flavipinnata</emphasis>
|
||
</taxonomicName>
|
||
, and in coloration, with the first dorsal fin mostly clear and anal fin black in
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B06AFE0E7B39FE1E" box="[349,523,445,469]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="rubrimacula">
|
||
<emphasis id="B938EABECF2BFFA3B06AFE0E7B39FE1E" box="[349,523,445,469]" italics="true" pageId="8" pageNumber="407">E. rubrimacula</emphasis>
|
||
</taxonomicName>
|
||
, whereas it is bright golden-yellow and anal fin orange in
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B580FE0E7C6DFE1E" box="[1207,1375,445,469]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="flavipinnata">
|
||
<emphasis id="B938EABECF2BFFA3B580FE0E7C6DFE1E" box="[1207,1375,445,469]" italics="true" pageId="8" pageNumber="407">E. flavipinnata</emphasis>
|
||
</taxonomicName>
|
||
. The specimens of
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B008FE5278DEFE32" box="[319,492,481,505]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="flavipinnata">
|
||
<emphasis id="B938EABECF2BFFA3B008FE5278DEFE32" box="[319,492,481,505]" italics="true" pageId="8" pageNumber="407">E. flavipinnata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B306FE527BCBFE32" box="[561,761,481,505]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="rubrimaculata">
|
||
<emphasis id="B938EABECF2BFFA3B306FE527BCBFE32" box="[561,761,481,505]" italics="true" pageId="8" pageNumber="407">E. rubrimaculata</emphasis>
|
||
</taxonomicName>
|
||
are both females, eliminating sexual dimorphism as an explanation of color differences between the two. The separation from other species of
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B59CFDB67DC6FDD6" box="[1195,1268,518,541]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EABECF2BFFA3B59CFDB67DC6FDD6" box="[1195,1268,518,541]" italics="true" pageId="8" pageNumber="407">Eviota</emphasis>
|
||
</taxonomicName>
|
||
with cephalic sensory-canal pore pattern 2 (lacking only the H [IT] pore) and branched pectoral-fin rays, is the same as for
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B4B3FD9A782DFDAE" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="flavipinnata">
|
||
<emphasis id="B938EABECF2BFFA3B4B3FD9A782DFDAE" italics="true" pageId="8" pageNumber="407">E. flavipinnata</emphasis>
|
||
</taxonomicName>
|
||
(Table 1), with the fresh coloration also differing from
|
||
<emphasis id="B938EABECF2BFFA3B29BFDFE7D8CFDAE" box="[940,1214,589,613]" italics="true" pageId="8" pageNumber="407">
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B29BFDFE7D3DFDAE" box="[940,1039,589,613]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="afelei">E. afelei</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B529FDFE7D8CFDAE" box="[1054,1214,589,613]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="bimaculata">E. bimaculata</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B435FDFE7CAEFDAE" box="[1282,1436,589,613]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="punctulata">
|
||
<emphasis id="B938EABECF2BFFA3B435FDFE7CAEFDAE" box="[1282,1436,589,613]" italics="true" pageId="8" pageNumber="407">E. punctulata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13772A29CF2BFFA3B1A8FDC179DCFD41" box="[159,238,625,650]" captionStart="FIGURE 4" captionStartId="4.[151,250,606,628]" captionTargetBox="[218,1346,193,584]" captionTargetId="figure@4.[169,1418,193,585]" captionTargetPageId="4" captionText="FIGURE 4. Eviota bimaculata, freshly collected specimen, Jurien Bay, Australia. Photograph by J. B. Hutchins." httpUri="https://zenodo.org/record/244257/files/figure.png" pageId="8" pageNumber="407">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="13772A29CF2BFFA3B1CDFDC17838FD41" box="[250,266,625,650]" captionStart="FIGURE 5" captionStartId="4.[151,250,1450,1472]" captionTargetBox="[150,1436,678,1428]" captionTargetId="figure@4.[150,1436,678,1428]" captionTargetPageId="4" captionText="FIGURE 5. Eviota punctulata, freshly collected specimen under water, Fiji. Photograph by J. E. Randall." httpUri="https://zenodo.org/record/244258/files/figure.png" pageId="8" pageNumber="407">5</figureCitation>
|
||
,
|
||
<figureCitation id="13772A29CF2BFFA3B021FDC17814FD41" box="[278,294,625,650]" captionStart="FIGURE 6" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[190,1392,1563,1886]" captionTargetId="figure@4.[169,1418,1522,1917]" captionTargetPageId="4" captionText="FIGURE 6. Eviota afelei, freshly collected specimen, Moorea. Photograph by J. T. Williams." httpUri="https://zenodo.org/record/244259/files/figure.png" pageId="8" pageNumber="407">6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3566527CF2BFFA3B1F0FD267D24FC8A" pageId="8" pageNumber="407" type="discussion">
|
||
<paragraph id="8BF336ACCF2BFFA3B1F0FD267D24FC8A" blockId="8.[151,1436,151,833]" pageId="8" pageNumber="407">
|
||
<emphasis id="B938EABECF2BFFA3B1F0FD267806FD65" bold="true" box="[199,308,662,686]" pageId="8" pageNumber="407">Remarks</emphasis>
|
||
. Both
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B0B6FD267B1AFD66" box="[385,552,661,685]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="flavipinnata">
|
||
<emphasis id="B938EABECF2BFFA3B0B6FD267B1AFD66" box="[385,552,661,685]" italics="true" pageId="8" pageNumber="407">E. flavipinnata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B357FD267A10FD66" box="[608,802,661,685]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="species" species="rubrimaculata">
|
||
<emphasis id="B938EABECF2BFFA3B357FD267A10FD66" box="[608,802,661,685]" italics="true" pageId="8" pageNumber="407">E. rubrimaculata</emphasis>
|
||
</taxonomicName>
|
||
share many characters in common, differing only in the development of the 5th pelvic-fin ray and major differences in fresh coloration. In their discussion of cryptic goby species within the Coral Triangle,
|
||
<bibRefCitation id="EFDD4B5DCF2BFFA3B377FD507A6CFD32" box="[576,862,736,761]" pageId="8" pageNumber="407" refString="Tornabene, L., Valdez, S., Erdmann, M. & Pezold, F. (2015) Support for a ' Center of Origin' in the Coral Triangle: cryptic diversity, recent speciation, and local endemism in a diverse lineage of reef fishes (Gobiidae: Eviota), Molecular Phylogenetics and Evolution, 82, 200 - 210." type="journal article">
|
||
Tornabene
|
||
<emphasis id="B938EABECF2BFFA3B3F3FD517A37FD33" box="[708,773,736,760]" italics="true" pageId="8" pageNumber="407">et al.</emphasis>
|
||
(2015)
|
||
</bibRefCitation>
|
||
demonstrated that coloration in various
|
||
<taxonomicName id="4C4C4D2FCF2BFFA3B464FD517CAEFD33" box="[1363,1436,737,760]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="407" phylum="Chordata" rank="genus">
|
||
<emphasis id="B938EABECF2BFFA3B464FD517CAEFD33" box="[1363,1436,737,760]" italics="true" pageId="8" pageNumber="407">Eviota</emphasis>
|
||
</taxonomicName>
|
||
complexes appears to evolve rapidly, before their genetic differences are reflected in their morphology. It is possible that this may be another example of species that have evolved recently.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |