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<document id="DA3B3FBFB5BFA5966548C770F4CEAD5C" ID-DOI="10.1080/00222930802610428" ID-ISSN="1464-5262" ID-Zenodo-Dep="4666701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617733898737" checkinUser="carolina" docAuthor="Korsós, Zoltán, Geoffroy, Jean-Jacques &amp; Mauriès, Jean-Paul" docDate="2009" docId="03E587B5FFA9BA66FE06D8A8FB0BFEDF" docLanguage="en" docName="JNATHIST.43.7-8.435-445.pdf" docOrigin="Journal of Natural History 43 (7 - 8)" docSource="http://dx.doi.org/10.1080/00222930802610428" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Hirudicryptus quintumelementum Korsós &amp; Geoffroy &amp; Mauriès 2009, sp. nov." docType="treatment" docVersion="3" lastPageNumber="442" masterDocId="FFDCFFCDFFAABA6CFF83DB22FF9AFF9D" masterDocTitle="The fifth element: reconnection of the disjunct distribution of the members of Siphonocryptida (Diplopoda) with the description of a new species from Nepal" masterLastPageNumber="445" masterPageNumber="435" pageNumber="436" updateTime="1698901179509" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="EB8ED20E301B27210C71AC5F0A1308AB">The fifth element: reconnection of the disjunct distribution of the members of Siphonocryptida (Diplopoda) with the description of a new species from Nepal</mods:title>
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<mods:namePart id="CC9C38EA8D0C4822251610FC5CA482E5">Korsós, Zoltán</mods:namePart>
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<mods:namePart id="FA78427BC367BDB7C7D2A8AFE12AD900">Geoffroy, Jean-Jacques</mods:namePart>
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<mods:namePart id="E82B40E2865CACFD0516BE46C5C32C26">Mauriès, Jean-Paul</mods:namePart>
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<mods:title id="956A7F0A638DB86675AFCC2E5007CD8C">Journal of Natural History</mods:title>
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<mods:date id="593943C422EC062D854B7C10B7FF90BB">2009</mods:date>
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<mods:number id="98DE1E2CDF65EE0A5FB3701124FD7FED">2009-02-28</mods:number>
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<mods:number id="13779019D28C201463EDAEDB9D4CE2FC">43</mods:number>
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<mods:number id="176FDC51C67746F79868D77E161659F7">7 - 8</mods:number>
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<subSubSection id="C3566528FFA9BA6FFE06D8A8FCC6FC3F" box="[389,860,906,930]" pageId="3" pageNumber="436" type="nomenclature">
<paragraph id="8BF336A3FFA9BA6FFE06D8A8FCC6FC3F" blockId="3.[389,860,906,963]" box="[389,860,906,930]" pageId="3" pageNumber="436">
<heading id="D0BB81CFFFA9BA6FFE06D8A8FCC6FC3F" box="[389,860,906,930]" centered="true" fontSize="36" level="2" pageId="3" pageNumber="436" reason="7">
<taxonomicName id="4C4C4D20FFA9BA6FFE06D8A8FD62FC3F" ID-CoL="3M4P3" authority="Korsós &amp; Geoffroy &amp; Mauriès, 2009" authorityName="Korsós &amp; Geoffroy &amp; Mauriès" authorityYear="2009" box="[389,760,906,930]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="3" pageNumber="436" phylum="Arthropoda" rank="species" species="quintumelementum" status="sp. nov.">
<emphasis id="B938EAB1FFA9BA6FFE06D8A8FD62FC3F" bold="true" box="[389,760,906,930]" italics="true" pageId="3" pageNumber="436">Hirudicryptus quintumelementum</emphasis>
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<taxonomicNameLabel id="A20B57CAFFA9BA6FFC82D8A8FCC6FC3F" box="[769,860,906,930]" pageId="3" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BF336A3FFA9BA6FFDA4D889FD23FC5E" blockId="3.[389,860,906,963]" box="[551,697,939,963]" pageId="3" pageNumber="436">
(
<figureCitation id="13772A26FFA9BA6FFDADD889FD2BFC5E" box="[558,689,939,963]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStartId-0="4.[213,280,735,757]" captionStartId-1="4.[213,280,1682,1704]" captionStartId-2="5.[146,213,834,856]" captionStartId-3="5.[146,213,1653,1675]" captionStartId-4="6.[213,280,682,704]" captionStartId-5="6.[213,280,1683,1705]" captionStartId-6="7.[146,213,496,518]" captionStartId-7="7.[146,213,1653,1675]" captionStartId-8="8.[213,280,733,755]" captionTargetBox-0="[292,1089,165,698]" captionTargetBox-1="[371,1010,1167,1646]" captionTargetBox-3="[305,945,1132,1616]" captionTargetBox-4="[371,1010,165,645]" captionTargetBox-5="[371,1010,1166,1645]" captionTargetBox-6="[305,945,165,459]" captionTargetBox-7="[305,945,994,1617]" captionTargetBox-8="[294,1090,165,697]" captionTargetId-0="figure-83@4.[292,1090,164,698]" captionTargetId-1="figure-103@4.[371,1011,1167,1646]" captionTargetId-3="figure-64@5.[305,945,1132,1616]" captionTargetId-4="figure-111@6.[371,1011,165,645]" captionTargetId-5="figure-132@6.[371,1011,1166,1645]" captionTargetId-6="figure-82@7.[305,945,165,459]" captionTargetId-7="figure-105@7.[305,945,994,1617]" captionTargetId-8="figure-272@8.[292,1092,165,697]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="6" captionTargetPageId-6="7" captionTargetPageId-7="7" captionTargetPageId-8="8" captionText-0="Figure 1. Body size relationships (max. body width plotted against number of terga) of four species of Siphonocryptida." captionText-1="Figure 2. Hirudicryptus quintumelementum sp. nov. Paratype (27 terga), lateral view." captionText-2="Figure 3. Midbody terga of paratype (27 terga), dorsal view." captionText-3="Figure 4. Head with collum and second tergum, fronto-lateral view. Scanning electron micrograph. Scale 0.1 mm." captionText-4="Figure 5. Midbody terga, showing enlarged tubercles at posterior margin in dorsal view. Scanning electron micrograph. Scale 0.1 mm." captionText-5="Figure 6. Last tergum, dorsal view. Scanning electron micrograph. Scale 0.1 mm." captionText-6="Figure 7. Hirudicryptus quintumelementum sp. nov. Holotype, anterior part of body, frontolateral view. Antennal setae have been omitted. Scale 0.1 mm." captionText-7="Figure 8. Hirudicryptus quintumelementum sp. nov. Holotype, anterior part of body, left lateral view. Scale 0.1 mm." captionText-8="Figure 9. Right second leg and vulva, ventro-caudal view. Notes: Op, operculum; vi-ve, internal and external valves; C, coxa; st, sternum. Scale 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4666703" figureDoi-1="http://doi.org/10.5281/zenodo.4666707" figureDoi-3="http://doi.org/10.5281/zenodo.4666711" figureDoi-4="http://doi.org/10.5281/zenodo.4666715" figureDoi-5="http://doi.org/10.5281/zenodo.4666717" figureDoi-6="http://doi.org/10.5281/zenodo.4666719" figureDoi-7="http://doi.org/10.5281/zenodo.4666723" figureDoi-8="http://doi.org/10.5281/zenodo.4666725" httpUri-0="https://zenodo.org/record/4666703/files/figure.png" httpUri-1="https://zenodo.org/record/4666707/files/figure.png" httpUri-3="https://zenodo.org/record/4666711/files/figure.png" httpUri-4="https://zenodo.org/record/4666715/files/figure.png" httpUri-5="https://zenodo.org/record/4666717/files/figure.png" httpUri-6="https://zenodo.org/record/4666719/files/figure.png" httpUri-7="https://zenodo.org/record/4666723/files/figure.png" httpUri-8="https://zenodo.org/record/4666725/files/figure.png" pageId="3" pageNumber="436">Figures 19</figureCitation>
)
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11D8DEFED1FB89" blockId="3.[146,331,1020,1044]" box="[146,331,1020,1044]" pageId="3" pageNumber="436">
<emphasis id="B938EAB1FFA9BA6FFF11D8DEFED1FB89" box="[146,331,1020,1044]" italics="true" pageId="3" pageNumber="436">Material studied</emphasis>
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DF07FE0CFB00" blockId="3.[146,1103,1060,1181]" pageId="3" pageNumber="436">
<specimenCount id="9D4AFD2AFFA9BA6FFF11DF07FF5FFBA0" box="[146,197,1060,1085]" pageId="3" pageNumber="436" type="female">3 ♀♀</specimenCount>
,
<collectingCountry id="F35B7633FFA9BA6FFF56DF07FE87FBA0" box="[213,285,1061,1085]" name="Nepal" pageId="3" pageNumber="436">Nepal</collectingCountry>
, Siwapuri Dara (5Sheopuri Lekh),
<quantity id="4CB49B46FFA9BA6FFD3CDF07FC99FBA0" box="[703,771,1061,1085]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="436" unit="km" value="15.0">15 km</quantity>
NE of Kathmandu,
<quantity id="4CB49B46FFA9BA6FFC7BDF07FBD0FBA0" box="[1016,1098,1061,1085]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="3" pageNumber="436" unit="m" value="2450.0">2450 m</quantity>
, dry mature oak forest; little brook in a shallow gulley, sifting of vegetation, fallen leaves and other debris on moist to wet habitats along the brook (No. 250),
<date id="FFF21063FFA9BA6FFC69DF47FF50FB00" pageId="3" pageNumber="436" value="1985-04-29">29. April 1985</date>
, leg. A. Smetana.
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DFF4FF63FB73" blockId="3.[146,249,1238,1262]" box="[146,249,1238,1262]" pageId="3" pageNumber="436">
<emphasis id="B938EAB1FFA9BA6FFF11DFF4FF63FB73" box="[146,249,1238,1262]" italics="true" pageId="3" pageNumber="436">
<typeStatus id="54F78801FFA9BA6FFF11DFF4FF63FB73" box="[146,249,1238,1262]" pageId="3" pageNumber="436" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DFDDFD28FA8A" blockId="3.[146,690,1279,1303]" box="[146,690,1279,1303]" pageId="3" pageNumber="436">Intact female specimen (34 segments), GMNH.</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DE72FE98FAF5" blockId="3.[146,258,1360,1384]" box="[146,258,1360,1384]" pageId="3" pageNumber="436">
<emphasis id="B938EAB1FFA9BA6FFF11DE72FE98FAF5" box="[146,258,1360,1384]" italics="true" pageId="3" pageNumber="436">
<typeStatus id="54F78801FFA9BA6FFF11DE72FE98FAF5" box="[146,258,1360,1384]" pageId="3" pageNumber="436" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DE5AFC72FA2D" blockId="3.[146,1103,1400,1456]" pageId="3" pageNumber="436">
<materialsCitation id="3B243CFEFFA9BA6FFF11DE5AFC7FFA2D" collectionCode="MNHN" pageId="3" pageNumber="436" specimenCode="BB001" specimenCount="1" typeStatus="paratype">
Female with 35 segments (dissected for temporary glycerine, and permanent SEM mounts); and another female with 27 segments,
<collectionCode id="ED5DAE66FFA9BA6FFD3EDEBAFC83FA2D" box="[701,793,1432,1456]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="436">MNHN</collectionCode>
[
<collectionCode id="ED5DAE66FFA9BA6FFCAADEBAFC1CFA2D" box="[809,902,1432,1456]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="436">MNHN</collectionCode>
<specimenCode id="DBEA9ED8FFA9BA6FFC0DDEBAFC47FA2D" box="[910,989,1432,1456]" collectionCode="BB" country="0" name="Buffalo Bill Museum" pageId="3" pageNumber="436">BB001</specimenCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DECBFE91F99C" blockId="3.[146,267,1513,1537]" box="[146,267,1513,1537]" pageId="3" pageNumber="436">
<emphasis id="B938EAB1FFA9BA6FFF11DECBFE91F99C" box="[146,267,1513,1537]" italics="true" pageId="3" pageNumber="436">Etymology</emphasis>
</paragraph>
<paragraph id="8BF336A3FFA9BA6FFF11DD30FD00F937" blockId="3.[146,1103,1554,1706]" pageId="3" pageNumber="436">
The species is named as the fifth member of the millipede order
<taxonomicName id="4C4C4D20FFA9BA6FFCDEDD30FB87F9B7" authorityName="Pocock" authorityYear="1894" box="[861,1053,1554,1578]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonocryptida" pageId="3" pageNumber="436" phylum="Arthropoda" rank="order">Siphonocryptida</taxonomicName>
; but also in honour of the alien custodians (called mondoshawans) in the movie by Luc Besson (Le cinquième élément 1997) who eventually saved the Earth from a catastrophe, and whom the head and the collum of the new species (and actually all members of the order) resemble superficially.
</paragraph>
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<paragraph id="8BF336A3FFAEBA68FF56D9FDFE6BFC8F" blockId="4.[213,1169,735,786]" pageId="4" pageNumber="437">
Figure 1. Body size relationships (max. body width plotted against number of terga) of four species of
<taxonomicName id="4C4C4D20FFAEBA68FEBCD9DEFE77FC8F" authorityName="Pocock" authorityYear="1894" box="[319,493,764,786]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonocryptida" pageId="4" pageNumber="437" phylum="Arthropoda" rank="order">Siphonocryptida</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF336A3FFAEBA68FF56D81CFED8FCCB" blockId="4.[213,322,830,854]" box="[213,322,830,854]" pageId="4" pageNumber="437">
<emphasis id="B938EAB1FFAEBA68FF56D81CFED8FCCB" box="[213,322,830,854]" italics="true" pageId="4" pageNumber="437">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF336A3FFAEBA68FF56D845FDA9FBDC" blockId="4.[213,1169,871,1089]" pageId="4" pageNumber="437">
Member of the genus
<taxonomicName id="4C4C4D20FFAEBA68FDB6D845FD55FCE2" authorityName=", Enghoff and Golovatch" authorityYear="1995" box="[565,719,871,895]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="4" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAB1FFAEBA68FDB6D845FD55FCE2" box="[565,719,871,895]" italics="true" pageId="4" pageNumber="437">Hirudicryptus</emphasis>
</taxonomicName>
(family
<taxonomicName id="4C4C4D20FFAEBA68FCE1D845FBB4FCE2" authorityName="Pocock" authorityYear="1894" box="[866,1070,871,895]" class="Diplopoda" family="Siphonocryptidae" kingdom="Animalia" order="Siphonocryptida" pageId="4" pageNumber="437" phylum="Arthropoda" rank="family">Siphonocryptidae</taxonomicName>
, order
<taxonomicName id="4C4C4D20FFAEBA68FF56D8A5FE0EFC02" authorityName="Pocock" authorityYear="1894" box="[213,404,903,927]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonocryptida" pageId="4" pageNumber="437" phylum="Arthropoda" rank="order">Siphonocryptida</taxonomicName>
) as defined by
<bibRefCitation id="EFDD4B52FFAEBA68FDC7D8A5FC32FC02" author="Enghoff H &amp; Golovatch SI" box="[580,936,903,927]" pageId="4" pageNumber="437" pagination="29 - 41" refId="ref3909" refString="Enghoff H, Golovatch SI. 1995. A revision of the Siphonocryptidae (Diplopoda, Polyzoniida). Zool Script. 24 (1): 29 - 41." type="journal article" year="1995">Enghoff and Golovatch (1995)</bibRefCitation>
, in having a narrow body (length/width ratio is.6) with midbody terga ca. four times wider than long, and a broad, crescent-shaped collum with no earlike lobes. Differs from other species of the genus in colouration and in the surface of the body terga being strongly granulate, with a transverse row of about 911 well-developed dark tubercles at the posterior margin on each side.
</paragraph>
<caption id="DF33662BFFAEBA68FF56DDB0FBDAF935" ID-DOI="http://doi.org/10.5281/zenodo.4666707" ID-Zenodo-Dep="4666707" box="[213,1088,1682,1704]" httpUri="https://zenodo.org/record/4666707/files/figure.png" pageId="4" pageNumber="437" startId="4.[213,280,1682,1704]" targetBox="[371,1010,1167,1646]" targetPageId="4">
<paragraph id="8BF336A3FFAEBA68FF56DDB0FBDAF935" blockId="4.[213,1088,1682,1704]" box="[213,1088,1682,1704]" pageId="4" pageNumber="437">
Figure 2.
<taxonomicName id="4C4C4D20FFAEBA68FEBDDDB1FD16F935" authority="Korsós &amp; Geoffroy &amp; Mauriès, 2009" authorityName="Korsós &amp; Geoffroy &amp; Mauriès" authorityYear="2009" box="[318,652,1683,1704]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="4" pageNumber="437" phylum="Arthropoda" rank="species" species="quintumelementum" status="sp. nov.">
<emphasis id="B938EAB1FFAEBA68FEBDDDB1FD16F935" box="[318,652,1683,1704]" italics="true" pageId="4" pageNumber="437">Hirudicryptus quintumelementum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57CAFFAEBA68FD17DDB0FD7CF935" box="[660,742,1682,1704]" pageId="4" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
Paratype (27 terga), lateral view.
</paragraph>
</caption>
<caption id="DF33662BFFAFBA69FF11D860FC96FCC5" box="[146,780,834,856]" pageId="5" pageNumber="438">
<paragraph id="8BF336A3FFAFBA69FF11D860FC96FCC5" blockId="5.[146,780,834,856]" box="[146,780,834,856]" pageId="5" pageNumber="438">Figure 3. Midbody terga of paratype (27 terga), dorsal view.</paragraph>
</caption>
<paragraph id="8BF336A3FFAFBA69FF11D859FE8BFC0E" blockId="5.[146,273,891,915]" box="[146,273,891,915]" pageId="5" pageNumber="438">
<emphasis id="B938EAB1FFAFBA69FF11D859FE8BFC0E" box="[146,273,891,915]" italics="true" pageId="5" pageNumber="438">Description</emphasis>
</paragraph>
<paragraph id="8BF336A3FFAFBA69FF11D881FD85FB80" blockId="5.[146,1103,931,1053]" pageId="5" pageNumber="438">
<specimenCount id="9D4AFD2AFFAFBA69FF11D881FF3AFC21" box="[146,160,931,956]" pageId="5" pageNumber="438" type="female"></specimenCount>
(
<typeStatus id="54F78801FFAFBA69FF31D886FE82FC21" box="[178,280,932,956]" pageId="5" pageNumber="438" type="holotype">holotype</typeStatus>
) 34 segments (incl. collum, without telson), body length
<quantity id="4CB49B46FFAFBA69FC30D886FB93FC21" box="[947,1033,932,956]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="438" unit="mm" value="6.2">6.2 mm</quantity>
, max. width
<quantity id="4CB49B46FFAFBA69FF5AD8E6FEB4FC41" box="[217,302,964,988]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="438" unit="mm" value="0.9">0.9 mm</quantity>
;
<typeStatus id="54F78801FFAFBA69FEBAD8E6FE3DFC41" box="[313,423,964,988]" pageId="5" pageNumber="438" type="paratype">paratypes</typeStatus>
(
<specimenCount id="9D4AFD2AFFAFBA69FE34D8E1FE49FC41" box="[439,467,963,988]" pageId="5" pageNumber="438" type="female">♀♀</specimenCount>
): 35 and 27 segments, body length 7.7 and
<quantity id="4CB49B46FFAFBA69FC34D8E6FB96FC41" box="[951,1036,964,988]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="438" unit="mm" value="4.8">4.8 mm</quantity>
, max. width 1.1 and
<quantity id="4CB49B46FFAFBA69FEBED8C7FE08FC60" box="[317,402,997,1021]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="438" unit="mm" value="0.8">0.8 mm</quantity>
, respectively. Body length/width ratio: 6.9, 7, and 6, for the three specimens. See also
<figureCitation id="13772A26FFAFBA69FE38DF27FD80FB80" box="[443,538,1029,1053]" captionStart="Figure 1" captionStartId="4.[213,280,735,757]" captionTargetBox="[292,1089,165,698]" captionTargetId="figure-83@4.[292,1090,164,698]" captionTargetPageId="4" captionText="Figure 1. Body size relationships (max. body width plotted against number of terga) of four species of Siphonocryptida." figureDoi="http://doi.org/10.5281/zenodo.4666703" httpUri="https://zenodo.org/record/4666703/files/figure.png" pageId="5" pageNumber="438">Figure 1</figureCitation>
.
</paragraph>
<caption id="DF33662BFFAFBA69FF11DD57FE3FF935" ID-DOI="http://doi.org/10.5281/zenodo.4666711" ID-Zenodo-Dep="4666711" httpUri="https://zenodo.org/record/4666711/files/figure.png" pageId="5" pageNumber="438" startId="5.[146,213,1653,1675]" targetBox="[305,945,1132,1616]" targetPageId="5">
<paragraph id="8BF336A3FFAFBA69FF11DD57FE3FF935" blockId="5.[146,1102,1653,1704]" pageId="5" pageNumber="438">Figure 4. Head with collum and second tergum, fronto-lateral view. Scanning electron micrograph. Scale 0.1 mm.</paragraph>
</caption>
<caption id="DF33662BFFACBA6AFF56D988FD3CFD40" ID-DOI="http://doi.org/10.5281/zenodo.4666715" ID-Zenodo-Dep="4666715" httpUri="https://zenodo.org/record/4666715/files/figure.png" pageId="6" pageNumber="439" startId="6.[213,280,682,704]" targetBox="[371,1010,165,645]" targetPageId="6">
<paragraph id="8BF336A3FFACBA6AFF56D988FD3CFD40" blockId="6.[213,1170,682,733]" pageId="6" pageNumber="439">Figure 5. Midbody terga, showing enlarged tubercles at posterior margin in dorsal view. Scanning electron micrograph. Scale 0.1 mm.</paragraph>
</caption>
<paragraph id="8BF336A3FFACBA6AFF56D824FE18FC7C" blockId="6.[213,1169,774,993]" pageId="6" pageNumber="439">
<emphasis id="B938EAB1FFACBA6AFF56D824FE62FC83" box="[213,504,774,798]" italics="true" pageId="6" pageNumber="439">
Colour (
<figureCitation id="13772A26FFACBA6AFEB6D824FE04FC83" box="[309,414,774,798]" captionStart="Figure 2" captionStartId="4.[213,280,1682,1704]" captionTargetBox="[371,1010,1167,1646]" captionTargetId="figure-103@4.[371,1011,1167,1646]" captionTargetPageId="4" captionText="Figure 2. Hirudicryptus quintumelementum sp. nov. Paratype (27 terga), lateral view." figureDoi="http://doi.org/10.5281/zenodo.4666707" httpUri="https://zenodo.org/record/4666707/files/figure.png" pageId="6" pageNumber="439">Figures 2</figureCitation>
and
<figureCitation id="13772A26FFACBA6AFE5ED824FE74FC83" box="[477,494,774,798]" captionStart="Figure 3" captionStartId="5.[146,213,834,856]" captionText="Figure 3. Midbody terga of paratype (27 terga), dorsal view." pageId="6" pageNumber="439">3</figureCitation>
)
</emphasis>
. A light yellow mid-dorsal and two dark brown lateral stripes make the animal appear to have three equal stripes. This pattern is unique in the order except for
<taxonomicName id="4C4C4D20FFACBA6AFE46D865FDB8FCC2" authorityName="Enghoff and Golovatch" authorityYear="1995" box="[453,546,839,863]" class="Diplopoda" family="Siphonocryptidae" genus="Siphonocryptus" kingdom="Animalia" order="Siphonocryptida" pageId="6" pageNumber="439" phylum="Arthropoda" rank="species" species="latior">
<emphasis id="B938EAB1FFACBA6AFE46D865FDB8FCC2" box="[453,546,839,863]" italics="true" pageId="6" pageNumber="439">S. latior</emphasis>
</taxonomicName>
where the light median stripe is less pronounced and also narrower (
<bibRefCitation id="EFDD4B52FFACBA6AFE07D845FD40FCE2" author="Enghoff H &amp; Golovatch SI" box="[388,730,871,895]" pageId="6" pageNumber="439" pagination="29 - 41" refId="ref3909" refString="Enghoff H, Golovatch SI. 1995. A revision of the Siphonocryptidae (Diplopoda, Polyzoniida). Zool Script. 24 (1): 29 - 41." type="journal article" year="1995">Enghoff and Golovatch 1995</bibRefCitation>
). Tubercles at the posterior margin of the marbled paraterga are dark brown. Antennal segments distally, and sides of head are darker brown, eyespots black in dark patches. Underside, legs, telson uniformly light yellowish.
</paragraph>
<paragraph id="8BF336A3FFACBA6BFF56DF38FC4EFD27" blockId="6.[213,1170,1049,1106]" lastBlockId="7.[146,1102,609,698]" lastPageId="7" lastPageNumber="440" pageId="6" pageNumber="439">
<emphasis id="B938EAB1FFACBA6AFF56DF38FE55FBAF" box="[213,463,1050,1074]" italics="true" pageId="6" pageNumber="439">
Head (
<figureCitation id="13772A26FFACBA6AFEA1DF38FE17FBAF" box="[290,397,1050,1074]" captionStart="Figure 4" captionStartId="5.[146,213,1653,1675]" captionTargetBox="[305,945,1132,1616]" captionTargetId="figure-64@5.[305,945,1132,1616]" captionTargetPageId="5" captionText="Figure 4. Head with collum and second tergum, fronto-lateral view. Scanning electron micrograph. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666711" httpUri="https://zenodo.org/record/4666711/files/figure.png" pageId="6" pageNumber="439">Figures 4</figureCitation>
,
<figureCitation id="13772A26FFACBA6AFE19DF38FE5CFBAF" box="[410,454,1050,1074]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStartId-0="7.[146,213,496,518]" captionStartId-1="7.[146,213,1653,1675]" captionTargetBox-0="[305,945,165,459]" captionTargetBox-1="[305,945,994,1617]" captionTargetId-0="figure-82@7.[305,945,165,459]" captionTargetId-1="figure-105@7.[305,945,994,1617]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figure 7. Hirudicryptus quintumelementum sp. nov. Holotype, anterior part of body, frontolateral view. Antennal setae have been omitted. Scale 0.1 mm." captionText-1="Figure 8. Hirudicryptus quintumelementum sp. nov. Holotype, anterior part of body, left lateral view. Scale 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4666719" figureDoi-1="http://doi.org/10.5281/zenodo.4666723" httpUri-0="https://zenodo.org/record/4666719/files/figure.png" httpUri-1="https://zenodo.org/record/4666723/files/figure.png" pageId="6" pageNumber="439">78</figureCitation>
)
</emphasis>
. Elongated, proboscoid, sharply pointed triangular, about as long as wide. In dorsal view completely hidden under collum, only antennae visible. Antennae stout, slightly bent ventrad, with six antennomeres subequal in length, the last (seventh) much shorter, its length about one-third of the sixth. Ocelli 2+2, black, posterior bigger than anterior ones. A pair of long frontal setae present.
</paragraph>
<caption id="DF33662BFFACBA6AFF56DDB1FB85F934" ID-DOI="http://doi.org/10.5281/zenodo.4666717" ID-Zenodo-Dep="4666717" box="[213,1055,1683,1705]" httpUri="https://zenodo.org/record/4666717/files/figure.png" pageId="6" pageNumber="439" startId="6.[213,280,1683,1705]" targetBox="[371,1010,1166,1645]" targetPageId="6">
<paragraph id="8BF336A3FFACBA6AFF56DDB1FB85F934" blockId="6.[213,1055,1683,1705]" box="[213,1055,1683,1705]" pageId="6" pageNumber="439">Figure 6. Last tergum, dorsal view. Scanning electron micrograph. Scale 0.1 mm.</paragraph>
</caption>
<caption id="DF33662BFFADBA6BFF11DAD2FC88FDBE" ID-DOI="http://doi.org/10.5281/zenodo.4666719" ID-Zenodo-Dep="4666719" httpUri="https://zenodo.org/record/4666719/files/figure.png" pageId="7" pageNumber="440" startId="7.[146,213,496,518]" targetBox="[305,945,165,459]" targetPageId="7">
<paragraph id="8BF336A3FFADBA6BFF11DAD2FC88FDBE" blockId="7.[146,1103,496,547]" pageId="7" pageNumber="440">
Figure 7.
<taxonomicName id="4C4C4D20FFADBA6BFF7FDAD2FDD3FD98" authority="Korsós &amp; Geoffroy &amp; Mauriès, 2009" authorityName="Korsós &amp; Geoffroy &amp; Mauriès" authorityYear="2009" box="[252,585,496,517]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="quintumelementum" status="sp. nov.">
<emphasis id="B938EAB1FFADBA6BFF7FDAD2FDD3FD98" box="[252,585,496,517]" italics="true" pageId="7" pageNumber="440">Hirudicryptus quintumelementum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57CAFFADBA6BFDD1DAD2FD3EFD9B" box="[594,676,496,518]" pageId="7" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, anterior part of body, frontolateral view. Antennal setae have been omitted. Scale 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF336A3FFADBA6BFF11D9D1FC70FCF1" blockId="7.[146,1103,755,876]" pageId="7" pageNumber="440">
<emphasis id="B938EAB1FFADBA6BFF11D9D1FE3CFC96" box="[146,422,755,779]" italics="true" pageId="7" pageNumber="440">
Collum (
<figureCitation id="13772A26FFADBA6BFF75D9D1FEF8FC96" box="[246,354,755,779]" captionStart="Figure 4" captionStartId="5.[146,213,1653,1675]" captionTargetBox="[305,945,1132,1616]" captionTargetId="figure-64@5.[305,945,1132,1616]" captionTargetPageId="5" captionText="Figure 4. Head with collum and second tergum, fronto-lateral view. Scanning electron micrograph. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666711" httpUri="https://zenodo.org/record/4666711/files/figure.png" pageId="7" pageNumber="440">Figures 4</figureCitation>
,
<figureCitation id="13772A26FFADBA6BFEF1D9D1FE04FC96" box="[370,414,755,779]" captionStart-0="Figure 7" captionStart-1="Figure 8" captionStartId-0="7.[146,213,496,518]" captionStartId-1="7.[146,213,1653,1675]" captionTargetBox-0="[305,945,165,459]" captionTargetBox-1="[305,945,994,1617]" captionTargetId-0="figure-82@7.[305,945,165,459]" captionTargetId-1="figure-105@7.[305,945,994,1617]" captionTargetPageId-0="7" captionTargetPageId-1="7" captionText-0="Figure 7. Hirudicryptus quintumelementum sp. nov. Holotype, anterior part of body, frontolateral view. Antennal setae have been omitted. Scale 0.1 mm." captionText-1="Figure 8. Hirudicryptus quintumelementum sp. nov. Holotype, anterior part of body, left lateral view. Scale 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4666719" figureDoi-1="http://doi.org/10.5281/zenodo.4666723" httpUri-0="https://zenodo.org/record/4666719/files/figure.png" httpUri-1="https://zenodo.org/record/4666723/files/figure.png" pageId="7" pageNumber="440">78</figureCitation>
)
</emphasis>
. Distinctly separate from second tergum, dorsally and in anterior view crescent-shaped, forming two smooth subrectangular lobes, without ventro-lateral ears. In lateral view dorsal part highly elevated and protruded forwards, casque or helmet-shaped, widely separated from second tergum.
</paragraph>
<caption id="DF33662BFFADBA6BFF11DD57FE3FF935" ID-DOI="http://doi.org/10.5281/zenodo.4666723" ID-Zenodo-Dep="4666723" httpUri="https://zenodo.org/record/4666723/files/figure.png" pageId="7" pageNumber="440" startId="7.[146,213,1653,1675]" targetBox="[305,945,994,1617]" targetPageId="7">
<paragraph id="8BF336A3FFADBA6BFF11DD57FE3FF935" blockId="7.[146,1103,1653,1704]" pageId="7" pageNumber="440">
Figure 8.
<taxonomicName id="4C4C4D20FFADBA6BFE83DD57FDCBF917" authority="Korsós &amp; Geoffroy &amp; Mauriès, 2009" authorityName="Korsós &amp; Geoffroy &amp; Mauriès" authorityYear="2009" box="[256,593,1653,1674]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="7" pageNumber="440" phylum="Arthropoda" rank="species" species="quintumelementum" status="sp. nov.">
<emphasis id="B938EAB1FFADBA6BFE83DD57FDCBF917" box="[256,593,1653,1674]" italics="true" pageId="7" pageNumber="440">Hirudicryptus quintumelementum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57CAFFADBA6BFDDDDD57FD2EF916" box="[606,692,1653,1675]" pageId="7" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, anterior part of body, left lateral view. Scale 0.1 mm.
</paragraph>
</caption>
<caption id="DF33662BFFA2BA64FF56D9FFFCF6FC8C" ID-DOI="http://doi.org/10.5281/zenodo.4666725" ID-Zenodo-Dep="4666725" httpUri="https://zenodo.org/record/4666725/files/figure.png" pageId="8" pageNumber="441" startId="8.[213,280,733,755]" targetBox="[294,1090,165,697]" targetPageId="8">
<paragraph id="8BF336A3FFA2BA64FF56D9FFFCF6FC8C" blockId="8.[213,1169,733,785]" pageId="8" pageNumber="441">Figure 9. Right second leg and vulva, ventro-caudal view. Notes: Op, operculum; vi-ve, internal and external valves; C, coxa; st, sternum. Scale 0.1 mm.</paragraph>
</caption>
<paragraph id="8BF336A3FFA2BA64FF56D814FCE5FB71" blockId="8.[213,1169,821,1260]" pageId="8" pageNumber="441">
<emphasis id="B938EAB1FFA2BA64FF56D814FECAFCD3" box="[213,336,822,846]" italics="true" pageId="8" pageNumber="441">Body terga</emphasis>
. Midbody terga (
<figureCitation id="13772A26FFA2BA64FD9ED817FD1EFCD0" box="[541,644,821,845]" captionStart="Figure 3" captionStartId="5.[146,213,834,856]" captionText="Figure 3. Midbody terga of paratype (27 terga), dorsal view." pageId="8" pageNumber="441">Figures 3</figureCitation>
and
<figureCitation id="13772A26FFA2BA64FD42D817FD55FCD0" box="[705,719,821,845]" captionStart="Figure 5" captionStartId="6.[213,280,682,704]" captionTargetBox="[371,1010,165,645]" captionTargetId="figure-111@6.[371,1011,165,645]" captionTargetPageId="6" captionText="Figure 5. Midbody terga, showing enlarged tubercles at posterior margin in dorsal view. Scanning electron micrograph. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666715" httpUri="https://zenodo.org/record/4666715/files/figure.png" pageId="8" pageNumber="441">5</figureCitation>
) about four to five times broader than long, slightly tapering towards both ends, with deep, distinct middorsal suture along entire length. Large, dark brown tubercles from the fourth tergum onwards, 911 on each half, uneven, partly paired and distributed at the posterior margin (1222221222121+1212221222221 is a usual arrangement). The 1+1 median ones usually the largest, and most conspicuous with their dark brown colour against the light median dorsal stripe. Postero-lateral projections on each tergum from third onwards strong, but not passing posterior tergal contour, also coloured dark brown. Arrangement of paraterga normal, 25 not overlying the preceding ones. Lateral margin of paraterga forming strong peritreme from fifth onwards, occupying almost entire length of paraterga, large ozopore on the fifth anteriorly, from the sixth onwards more or less in median position. Last tergum (
<figureCitation id="13772A26FFA2BA64FF5FDF96FEA1FB51" box="[220,315,1204,1228]" captionStart="Figure 6" captionStartId="6.[213,280,1683,1705]" captionTargetBox="[371,1010,1166,1645]" captionTargetId="figure-132@6.[371,1011,1166,1645]" captionTargetPageId="6" captionText="Figure 6. Last tergum, dorsal view. Scanning electron micrograph. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666717" httpUri="https://zenodo.org/record/4666717/files/figure.png" pageId="8" pageNumber="441">Figure 6</figureCitation>
) rounded or subtrapeziform, with a median longitudinal suture, but without posterior lobes, and no indication of (diplo)fusion.
</paragraph>
<paragraph id="8BF336A3FFA2BA64FF56DE06FBFAFAA6" blockId="8.[213,1120,1315,1340]" box="[213,1120,1315,1340]" pageId="8" pageNumber="441">
<taxonomicName id="4C4C4D20FFA2BA64FF56DE06FE84FAA1" box="[213,286,1316,1340]" class="Aves" family="Laridae" genus="Sterna" kingdom="Animalia" order="Charadriiformes" pageId="8" pageNumber="441" phylum="Chordata" rank="genus">
<emphasis id="B938EAB1FFA2BA64FF56DE06FE84FAA1" box="[213,286,1316,1340]" italics="true" pageId="8" pageNumber="441">Sterna</emphasis>
</taxonomicName>
. At midbody much wider than long, about two times wider than coxae.
</paragraph>
<paragraph id="8BF336A3FFA2BA64FF56DE51FD74FA36" blockId="8.[213,1169,1395,1451]" pageId="8" pageNumber="441">
<emphasis id="B938EAB1FFA2BA64FF56DE51FE84FA16" box="[213,286,1395,1419]" italics="true" pageId="8" pageNumber="441">Telson</emphasis>
. Very small, completely hidden under last tergum, with two strong setae. Anal valves with two smaller setae directed ventrad.
</paragraph>
<paragraph id="8BF336A3FFA2BA64FF56DEC1FDF5F9A6" blockId="8.[213,1169,1507,1595]" pageId="8" pageNumber="441">
<emphasis id="B938EAB1FFA2BA64FF56DEC1FE90FA66" box="[213,266,1507,1531]" italics="true" pageId="8" pageNumber="441">Legs</emphasis>
. Long, slender, in normal position not surpassing body margin (except under posteriormost terga). Coxa widely separated from each other. Telopodite comprising five podomeres. No accessory claw.
</paragraph>
<paragraph id="8BF336A3FFA2BA64FF56DD50FDACF937" blockId="8.[213,1169,1650,1706]" pageId="8" pageNumber="441">
<emphasis id="B938EAB1FFA2BA64FF56DD50FE4FF917" box="[213,469,1650,1674]" italics="true" pageId="8" pageNumber="441">Male sexual characters</emphasis>
. In the absence of males in the present sample, no description of gonopods can be presented.
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF11DB8BFB8AFEBF" blockId="9.[146,1103,168,744]" pageId="9" pageNumber="442">
<emphasis id="B938EAB1FFA3BA65FF11DB8BFDAAFF5C" box="[146,560,169,193]" italics="true" pageId="9" pageNumber="442">
Female sexual characters (
<figureCitation id="13772A26FFA3BA65FE47DB8BFDBCFF5C" box="[452,550,169,193]" captionStart="Figure 9" captionStartId="8.[213,280,733,755]" captionTargetBox="[294,1090,165,697]" captionTargetId="figure-272@8.[292,1092,165,697]" captionTargetPageId="8" captionText="Figure 9. Right second leg and vulva, ventro-caudal view. Notes: Op, operculum; vi-ve, internal and external valves; C, coxa; st, sternum. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666725" httpUri="https://zenodo.org/record/4666725/files/figure.png" pageId="9" pageNumber="442">Figure 9</figureCitation>
)
</emphasis>
. It should be emphasized that this is the first time that the vulvae of a female siphonocryptid have been properly studied (i.e. after removal, under a compound microscope). The following description is essentially based on polyzoniid terminology, and compared with members of that order.
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF3EDA08FED7FD75" blockId="9.[146,1103,168,744]" pageId="9" pageNumber="442">
Coxae of second legpair hypertrophied and very close to each other due to the fact they use the space usually occupied by the coxal sacs of other subsequent legpairs. Vulval cavity (termed lucarne by
<bibRefCitation id="EFDD4B52FFA3BA65FD16DA49FCE4FE1E" author="Brolemann HW" box="[661,894,363,387]" pageId="9" pageNumber="442" refId="ref3838" refString="Brolemann HW. 1935. Myriapodes Diplopodes (Chilognathes I). Faune de France. Vol. 29. Paris (France): P. Lechevalier. 369 p." type="book" year="1935">Brölemann 1935: 50</bibRefCitation>
), very large, using mesally more than half of posterior part of coxa, in a similar way as in Polyzoniida. In contrast, however, each vulva provided with a strongly developed operculum, entirely closing the vulval cavity. Operculum simple, rectangular plate, apically rounded and slightly concave. No setae or hairs could be observed at all, unlike most other diplopod vulvae.
<collectingRegion id="4988F841FFA3BA65FE39D92FFE66FDB8" box="[442,508,525,549]" country="Turkey" name="Bursa" pageId="9" pageNumber="442">Bursa</collectingRegion>
more or less globular as in Polyzoniida, with prominent vertical cleft dividing it into two subequal and poorly chitinized halves. In analogy with the structures generally encountered in other diplopod vulvae these two parts could be termed as internal and external valves. Under a compound microscope, in continuation of medial cleft the outline of an apodematic tube can be distinguished (see dotted line in
<figureCitation id="13772A26FFA3BA65FD97D98DFDE8FD5A" box="[532,626,687,711]" captionStart="Figure 9" captionStartId="8.[213,280,733,755]" captionTargetBox="[294,1090,165,697]" captionTargetId="figure-272@8.[292,1092,165,697]" captionTargetPageId="8" captionText="Figure 9. Right second leg and vulva, ventro-caudal view. Notes: Op, operculum; vi-ve, internal and external valves; C, coxa; st, sternum. Scale 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4666725" httpUri="https://zenodo.org/record/4666725/files/figure.png" pageId="9" pageNumber="442">Figure 9</figureCitation>
), together with a spherical
<emphasis id="B938EAB1FFA3BA65FC3FD992FF78FD75" italics="true" pageId="9" pageNumber="442">receptaculum seminis</emphasis>
beneath.
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF11D803FD57FCA4" blockId="9.[146,717,801,825]" box="[146,717,801,825]" pageId="9" pageNumber="442">
<emphasis id="B938EAB1FFA3BA65FF11D803FD57FCA4" bold="true" box="[146,717,801,825]" pageId="9" pageNumber="442">
Updated key to the five species of
<taxonomicName id="4C4C4D20FFA3BA65FD9BD803FD57FCA4" authorityName="Pocock" authorityYear="1894" box="[536,717,801,825]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="order">Siphonocryptida</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF16D87BFBD4FC4E" blockId="9.[149,1103,857,979]" pageId="9" pageNumber="442">
1. Body very broad, length/width ratio of body 1.93.0, midbody terga ca. six times wider than long. First five paraterga overlying preceding paraterga, collum anteriorly with two extra, subrectangular lobes (ears). No frontal setae...................
<taxonomicName id="4C4C4D20FFA3BA65FDCFD899FD6DFC4E" authorityName="Pocock" authorityYear="1894" box="[588,759,955,979]" class="Diplopoda" family="Siphonocryptidae" genus="Siphonocryptus" kingdom="Animalia" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAB1FFA3BA65FDCFD899FD6DFC4E" box="[588,759,955,979]" italics="true" pageId="9" pageNumber="442">Siphonocryptus</emphasis>
</taxonomicName>
.................... 2
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF49D8C3FBD4FBC7" blockId="9.[202,1103,993,1114]" pageId="9" pageNumber="442">
Body much narrower, length/width ratio.4 (except in very small juveniles), midbody terga ca. four times wider than long. Anterior paraterga not overlapping reversely, collum simple crescent-shaped, with broad lobes in anterior view. Two long frontal setae..............
<taxonomicName id="4C4C4D20FFA3BA65FD39DF60FCC9FBC7" authorityName=", Enghoff and Golovatch" authorityYear="1995" box="[698,851,1090,1114]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAB1FFA3BA65FD39DF60FCC9FBC7" box="[698,851,1090,1114]" italics="true" pageId="9" pageNumber="442">Hirudicryptus</emphasis>
</taxonomicName>
............... 3
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF16DF51FBD4FB70" blockId="9.[149,1103,1139,1261]" pageId="9" pageNumber="442">
2. With a dorsal colour pattern of three dark and two light longitudinal stripes, body narrow, paratergal projections surpassing posterior contour of tergum, apical part of terminal podomere of posterior gonopods longer. (Sumatra)..............................................
<taxonomicName id="4C4C4D20FFA3BA65FC39DFF7FBD4FB70" authorityName="Pocock" authorityYear="1894" box="[954,1102,1237,1261]" class="Diplopoda" family="Siphonocryptidae" genus="Siphonocryptus" kingdom="Animalia" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="species" species="compactus">
<emphasis id="B938EAB1FFA3BA65FC39DFF7FBD4FB70" box="[954,1102,1237,1261]" italics="true" pageId="9" pageNumber="442">S. compactus</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF49DFDAFBD4FACC" blockId="9.[202,1102,1272,1361]" pageId="9" pageNumber="442">
A mid-dorsal light stripe (or body uniformly pale), body broad, paratergal projections not surpassing posterior contour of tergum, apical part of terminal podomere of posterior gonopod shorter. (Malaya).............
<taxonomicName id="4C4C4D20FFA3BA65FC71DE1BFBD4FACC" authorityName="Enghoff and Golovatch" authorityYear="1995" box="[1010,1102,1337,1361]" class="Diplopoda" family="Siphonocryptidae" genus="Siphonocryptus" kingdom="Animalia" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="species" species="latior">
<emphasis id="B938EAB1FFA3BA65FC71DE1BFBD4FACC" box="[1010,1102,1337,1361]" italics="true" pageId="9" pageNumber="442">S. latior</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF16DE48FBD4FA3F" blockId="9.[149,1103,1386,1442]" pageId="9" pageNumber="442">3. With a dorsal colour pattern of five longitudinal stripes: one median and two lateral dark ones, with two light ones between...................... 4</paragraph>
<paragraph id="8BF336A3FFA3BA65FF49DE8CFBD4FA7B" blockId="9.[202,1103,1454,1543]" pageId="9" pageNumber="442">
Dorsal colour pattern different: only three longitudinal stripes of equal width, one median light and two lateral dark ones. (
<collectingCountry id="F35B7633FFA3BA65FC78DEECFBDDFA7B" box="[1019,1095,1486,1510]" name="Nepal" pageId="9" pageNumber="442">Nepal</collectingCountry>
)
</paragraph>
<paragraph id="8BF336A3FFA3BA65FF54DECCFBD5F99B" blockId="9.[202,1103,1454,1543]" box="[215,1103,1518,1543]" pageId="9" pageNumber="442">
.................................
<taxonomicName id="4C4C4D20FFA3BA65FD77DECDFC70F99A" authority="Korsós &amp; Geoffroy &amp; Mauriès, 2009" authorityName="Korsós &amp; Geoffroy &amp; Mauriès" authorityYear="2009" box="[756,1002,1519,1543]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="species" species="quintumelementum" status="sp. nov.">
<emphasis id="B938EAB1FFA3BA65FD77DECDFC70F99A" box="[756,1002,1519,1543]" italics="true" pageId="9" pageNumber="442">H. quintumelementum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57CAFFA3BA65FC77DECCFBD5F99B" box="[1012,1103,1518,1542]" pageId="9" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BF336A3FFA3BA66FF16DD3DFB0BFEDF" blockId="9.[149,1103,1567,1689]" lastBlockId="10.[268,1169,168,322]" lastPageId="10" lastPageNumber="443" pageId="9" pageNumber="442">
4. Body very narrow, adult specimens even with more than 50 paraterga are only
<quantity id="4CB49B46FFA3BA65FF49DD62FE86F9C5" box="[202,284,1600,1624]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="9" pageNumber="442" unit="mm" value="1.2">1.2 mm</quantity>
wide. Number of tubercles on posterior margin of each half of midbody paraterga 1415. Terminal podomere of male posterior gonopod long, slender, simple. (Macaronesian islands).......................
<taxonomicName id="4C4C4D20FFA3BA66FC2CDDA3FED2FF5D" authority="Body" authorityName="Body" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" lastPageId="10" lastPageNumber="443" order="Siphonocryptida" pageId="9" pageNumber="442" phylum="Arthropoda" rank="species" species="canariensis">
<emphasis id="B938EAB1FFA3BA65FC2CDDA3FBD5F904" box="[943,1103,1665,1689]" italics="true" pageId="9" pageNumber="442">H. canariensis</emphasis>
Body
</taxonomicName>
less narrow, adult specimens with approximately 50 paraterga are 2.0
<quantity id="4CB49B46FFA0BA66FE8FDBEBFEC5FF7C" box="[268,351,201,225]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="443" unit="mm" value="2.5">2.5 mm</quantity>
wide (almost like
<taxonomicName id="4C4C4D20FFA0BA66FDB0DBEBFD15FF7C" authorityName="Enghoff and Golovatch" authorityYear="1995" box="[563,655,201,225]" class="Diplopoda" family="Siphonocryptidae" genus="Siphonocryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="443" phylum="Arthropoda" rank="species" species="latior">
<emphasis id="B938EAB1FFA0BA66FDB0DBEBFD15FF7C" box="[563,655,201,225]" italics="true" pageId="10" pageNumber="443">S. latior</emphasis>
</taxonomicName>
). Tubercles on posterior margin of midbody paraterga large, the number on each half only 68. Terminal podomere of male posterior gonopod with a clearly visible, distinct shoulder at its proximal third. (
<collectingRegion id="4988F841FFA0BA66FE97DA08FEE9FEDF" box="[276,371,298,322]" country="China" name="Taiwan)" pageId="10" pageNumber="443">Taiwan)</collectingRegion>
.......................................
<taxonomicName id="4C4C4D20FFA0BA66FC6EDA08FB0BFEDF" authorityName="Korsos, Enghoff &amp; Chang" authorityYear="2008" box="[1005,1169,298,322]" class="Diplopoda" family="Siphonocryptidae" genus="Hirudicryptus" kingdom="Animalia" order="Siphonocryptida" pageId="10" pageNumber="443" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="B938EAB1FFA0BA66FC6EDA08FB0BFEDF" box="[1005,1169,298,322]" italics="true" pageId="10" pageNumber="443">H. taiwanensis</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>