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<document id="B380554B151FB34C67152CD1D2DA8E55" ID-DOI="10.26028/cybium/2020-444-003" ID-ISSN="2101-0315" ID-Zenodo-Dep="12712019" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720646339926" checkinUser="felipe" docAuthor="Keith, Philippe, Mennesson, Marion I., Sauri, Sopian, Busson, Frédéric, Delrieu-Trottin, Erwan, Limmon, Gino, Nurjirana, Dahruddin, Hadi &amp; Hubert, Nicolas" docDate="2020" docId="039E87F5E23BFF9CFE8939D75352FA96" docLanguage="en" docName="Cybium.44.4.317-329.pdf" docOrigin="Cybium 44 (4)" docSource="http://dx.doi.org/10.26028/cybium/2020-444-003" docStyle="DocumentStyle:EAB95795642928CB986F0923DE58F0B7.1:Cybium.2019-.journal_article" docStyleId="EAB95795642928CB986F0923DE58F0B7" docStyleName="Cybium.2019-.journal_article" docStyleVersion="1" docTitle="Giuris tolsoni" docType="treatment" docVersion="2" lastPageNumber="325" masterDocId="FFA7FF8DE23CFF94FFAA3F6F5724FFA4" masterDocTitle="Giuris (Teleostei: Eleotridae) from Indonesia, with description of a new species" masterLastPageNumber="329" masterPageNumber="317" pageNumber="324" updateTime="1720650568442" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="86D015B0A5FBF2914382CE4EBC270774">Giuris (Teleostei: Eleotridae) from Indonesia, with description of a new species</mods:title>
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<mods:namePart id="03B0C9F310383036797A3AEF1C4FC8B1">Keith, Philippe</mods:namePart>
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<mods:namePart id="860B9029E222FDEB402D0F372FCE0774">Mennesson, Marion I.</mods:namePart>
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<mods:namePart id="6A081E5ED0E92A0D9A32AC54BC21BB86">Sauri, Sopian</mods:namePart>
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<mods:namePart id="2313ECA15F79C92FB6B7E24368F5FD28">Busson, Frédéric</mods:namePart>
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<mods:namePart id="66B74FDCF5E16A3CB004A5C7A436195F">Delrieu-Trottin, Erwan</mods:namePart>
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<mods:namePart id="CC7B659FF5BF039A0EEEA7555314EF2E">Limmon, Gino</mods:namePart>
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<mods:namePart id="62BB50F74BAA194D93564B0DD01015DC">Nurjirana</mods:namePart>
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<mods:namePart id="B881483DFD9D9C15C1A081A5B6034C33">Dahruddin, Hadi</mods:namePart>
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<mods:namePart id="DC5EF3E9365D0195B52D025FF82CB0C6">Hubert, Nicolas</mods:namePart>
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<mods:title id="D04A9E634DE91571B7D765FCC7374330">Cybium</mods:title>
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<mods:date id="C9E549310C0C8E6E09675D374B40B9D6">2020</mods:date>
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<mods:classification id="2F98C4224017C992AF09E1D137A4CF7A">journal article</mods:classification>
<mods:identifier id="44C27454F7EA65CB0B16DC2529375E13" type="DOI">10.26028/cybium/2020-444-003</mods:identifier>
<mods:identifier id="8F5D0AED6C2299BB92C4E78DDACD33CF" type="ISSN">2101-0315</mods:identifier>
<mods:identifier id="66A5E07BAC174A59E57989313CB8002C" type="Zenodo-Dep">12712019</mods:identifier>
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<treatment id="039E87F5E23BFF9CFE8939D75352FA96" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96" httpUri="http://treatment.plazi.org/id/039E87F5E23BFF9CFE8939D75352FA96" lastPageId="8" lastPageNumber="325" pageId="7" pageNumber="324">
<subSubSection id="C32D6568E23BFF93FE8939D75543F974" box="[291,615,1719,1744]" pageId="7" pageNumber="324" type="nomenclature">
<paragraph id="8B8836E3E23BFF93FE8939D75543F974" blockId="7.[291,615,1719,1744]" box="[291,615,1719,1744]" pageId="7" pageNumber="324">
<heading id="D0C0818FE23BFF93FE8939D75543F974" bold="true" box="[291,615,1719,1744]" centered="true" fontSize="9" level="2" pageId="7" pageNumber="324" reason="2">
<taxonomicName id="4C374D60E23BFF93FE8939D75543F974" authority="(Bleeker, 1854)" baseAuthorityName="Bleeker" baseAuthorityYear="1854" box="[291,615,1719,1744]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="324" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B943EAF1E23BFF93FE8939D75543F974" bold="true" box="[291,615,1719,1744]" pageId="7" pageNumber="324">
<emphasis id="B943EAF1E23BFF93FE8939D7569DF96B" bold="true" box="[291,441,1719,1744]" italics="true" pageId="7" pageNumber="324">Giuris tolsoni</emphasis>
(Bleeker, 1854)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D6568E23BFF93FEF839B6551DF955" box="[338,569,1753,1777]" pageId="7" pageNumber="324" type="description">
<paragraph id="8B8836E3E23BFF93FEF839B6551DF955" blockId="7.[338,569,1753,1777]" box="[338,569,1753,1777]" pageId="7" pageNumber="324">
(
<figureCitation id="130C2A66E23BFF93FEF039B656B8F955" box="[346,412,1753,1777]" captionStart="Figure 1" captionStartId="4.[132,195,1051,1073]" captionTargetBox="[133,1455,215,1031]" captionTargetId="figure-394@4.[132,1455,215,1034]" captionTargetPageId="4" captionText="Figure 1. Most likely ML tree inferred using the K80+I model (lnL = 1548.01608, Transition/tranversion ratio = 20.137, I = 0.776). Barcode Index Numbers (BIN) are given for each lineages." figureDoi="http://doi.org/10.5281/zenodo.12712021" httpUri="https://zenodo.org/record/12712021/files/figure.png" pageId="7" pageNumber="324">Figs 1</figureCitation>
,
<figureCitation id="130C2A66E23BFF93FE0339B6569CF955" box="[425,440,1753,1777]" captionStart="Figure 3" captionStartId="7.[813,879,1789,1811]" captionTargetBox="[813,1456,1065,1775]" captionTargetId="figure-699@7.[813,1456,1065,1775]" captionTargetPageId="7" captionText="Figure 3. A: Giuris tolsoni, RMNH 5180, holotype, male; SL 61.5 mm (Photo RMNH, P. Keith). B: G. tolsoni, MZB.25328, male, Bali; BIF 2283; SL 66 mm (Photo N. Hubert). C: G. tolsoni, in MZB.25329, female, Bali; BIF 2848; SL 34 mm (Photo N. Hubert)." figureDoi="http://doi.org/10.5281/zenodo.12712027" httpUri="https://zenodo.org/record/12712027/files/figure.png" pageId="7" pageNumber="324">3</figureCitation>
; Tabs II-III)
</paragraph>
</subSubSection>
<caption id="DF48666BE23BFF93FC873992545AF8D7" ID-DOI="http://doi.org/10.5281/zenodo.12712027" ID-Zenodo-Dep="12712027" httpUri="https://zenodo.org/record/12712027/files/figure.png" pageId="7" pageNumber="324" startId="7.[813,879,1789,1811]" targetBox="[813,1456,1065,1775]" targetPageId="7" targetType="figure">
<paragraph id="8B8836E3E23BFF93FC873992545AF8D7" blockId="7.[813,1456,1789,1907]" pageId="7" pageNumber="324">
Figure 3.
<emphasis id="B943EAF1E23BFF93FC0939925490F8B6" bold="true" box="[931,948,1789,1810]" pageId="7" pageNumber="324">A</emphasis>
:
<taxonomicName id="4C374D60E23BFF93FC6939925383F8B6" authority=", RMNH" authorityName="RMNH" box="[963,1191,1789,1810]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="324" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B943EAF1E23BFF93FC6939925372F8B6" box="[963,1110,1789,1810]" italics="true" pageId="7" pageNumber="324">Giuris tolsoni,</emphasis>
RMNH
</taxonomicName>
5180, holotype, male; SL 61.5 mm (Photo RMNH, P. Keith).
<emphasis id="B943EAF1E23BFF93FB30387A538EF88E" bold="true" box="[1178,1194,1813,1834]" pageId="7" pageNumber="324">B</emphasis>
:
<taxonomicName id="4C374D60E23BFF93FB13387A524EF88F" authority=", MZB." authorityName="MZB." box="[1209,1386,1813,1835]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="324" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B943EAF1E23BFF93FB13387A5201F88E" box="[1209,1317,1813,1834]" italics="true" pageId="7" pageNumber="324">G. tolsoni,</emphasis>
MZB.
</taxonomicName>
25328, male, Bali; BIF 2283; SL 66 mm (Photo N. Hubert).
<emphasis id="B943EAF1E23BFF93FA933842526EF8E7" bold="true" box="[1337,1354,1837,1859]" pageId="7" pageNumber="324">C</emphasis>
:
<emphasis id="B943EAF1E23BFF93FAF238425444F8FE" italics="true" pageId="7" pageNumber="324">
<taxonomicName id="4C374D60E23BFF93FAF238425466F8FE" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="324" phylum="Chordata" rank="species" species="tolsoni">G. tolsoni</taxonomicName>
, in
</emphasis>
MZB.25329, female, Bali; BIF 2848; SL 34 mm (Photo N. Hubert).
</paragraph>
</caption>
<subSubSection id="C32D6568E23BFF93FF2E38765490FDB5" pageId="7" pageNumber="324" type="materials_examined">
<paragraph id="8B8836E3E23BFF93FF2E38765678F895" blockId="7.[132,774,1817,1907]" box="[132,348,1817,1841]" pageId="7" pageNumber="324">
<emphasis id="B943EAF1E23BFF93FF2E38765678F895" bold="true" box="[132,348,1817,1841]" pageId="7" pageNumber="324">Material examined</emphasis>
</paragraph>
<paragraph id="8B8836E3E23BFF93FF0038555551F8D7" blockId="7.[132,774,1817,1907]" pageId="7" pageNumber="324">
<materialsCitation id="3B5F3CBEE23BFF93FF0038555551F8D7" collectionCode="RMNH" country="Indonesia" location="Near Djung-Kulon" pageId="7" pageNumber="324" specimenCode="RMNH 5180" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<emphasis id="B943EAF1E23BFF93FF003855562BF8F6" box="[170,271,1850,1874]" italics="true" pageId="7" pageNumber="324">
<typeStatus id="548C8841E23BFF93FF003855562BF8F6" box="[170,271,1850,1874]" pageId="7" pageNumber="324" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="DB919E98E23BFF93FE98385456E6F8F6" box="[306,450,1850,1874]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="324" type="Museum">RMNH 5180</specimenCode>
(1 of 17),
<location id="8EE86038E23BFF93FD9238545422F8F6" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FD9238545422F8F6" box="[568,774,1850,1874]" country="Indonesia" name="Near Djung-Kulon" pageId="7" pageNumber="324" stateProvince="Jawa Timur">Near Djung-Kulon</location>
(Djunkulon), extreme western
<collectingRegion id="49F3F801E23BFF93FE60383456D8F8D6" box="[458,508,1883,1906]" country="Indonesia" name="Jawa Timur" pageId="7" pageNumber="324">Java</collectingRegion>
,
<collectingCountry id="F3207673E23BFF93FDAC38345555F8D7" box="[518,625,1883,1907]" name="Indonesia" pageId="7" pageNumber="324">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8B8836E3E23BFF93FCF93FB65490FDB5" blockId="7.[813,1455,217,529]" pageId="7" pageNumber="324">
<emphasis id="B943EAF1E23BFF93FCF93FB654BAFF55" box="[851,926,217,241]" italics="true" pageId="7" pageNumber="324">Others</emphasis>
.
<materialsCitation id="3B5F3CBEE23BFF93FC6A3FB652B2FEB5" collectingDate="2014-04-15" collectionCode="MZB" collectorName="West &amp; Kab Jembrana &amp; Nbang" location="Hubert" pageId="7" pageNumber="324" specimenCount="1" stateProvince="Bali">
<collectionCode id="ED26AE26E23BFF93FC6A3FB654DAFF55" box="[960,1022,217,241]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="324" type="Museum">MZB</collectionCode>
.25328,
<collectingRegion id="49F3F801E23BFF93FBFF3FB653A2FF55" box="[1109,1158,217,241]" country="Indonesia" name="Bali" pageId="7" pageNumber="324">Bali</collectingRegion>
,
<collectorName id="26C25335E23BFF93FB3B3FB553E3FF55" box="[1169,1223,218,241]" pageId="7" pageNumber="324">West</collectorName>
<collectingRegion id="49F3F801E23BFF93FB643FB653DBFF55" box="[1230,1279,217,241]" country="Indonesia" name="Bali" pageId="7" pageNumber="324">Bali</collectingRegion>
,
<collectorName id="26C25335E23BFF93FAA13FB6528FFF55" box="[1291,1451,217,241]" pageId="7" pageNumber="324">Kab Jembrana</collectorName>
,
<collectorName id="26C25335E23BFF93FC873F965459FEB5" box="[813,893,249,273]" pageId="7" pageNumber="324">Nbang</collectorName>
,
<date id="FF891023E23BFF93FC213F96530EFEB5" box="[907,1066,249,273]" pageId="7" pageNumber="324" value="2014-04-15">
<collectingDate id="EFCDE9CBE23BFF93FC213F96530EFEB5" box="[907,1066,249,273]" pageId="7" pageNumber="324" value="2014-04-15">15 Apr. 2014</collectingDate>
</date>
,
<location id="8EE86038E23BFF93FB923F9653ACFEB5" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FB923F9653ACFEB5" box="[1080,1160,249,273]" name="Hubert" pageId="7" pageNumber="324" stateProvince="Bali">Hubert</location>
<emphasis id="B943EAF1E23BFF93FB393F9553EBFEB5" box="[1171,1231,249,273]" italics="true" pageId="7" pageNumber="324">et al.</emphasis>
coll.; BIF 2283
</materialsCitation>
.
<materialsCitation id="3B5F3CBEE23BFF93FC873E7652B3FEF5" collectingDate="2014-04-22" collectionCode="MZB" collectorName="West &amp; Kab Kelungkung &amp; Tukad Unda" location="Hubert" pageId="7" pageNumber="324" specimenCount="1" stateProvince="Bali">
<collectionCode id="ED26AE26E23BFF93FC873E76544DFE95" box="[813,873,281,305]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="324" type="Museum">MZB</collectionCode>
.25329,
<collectingRegion id="49F3F801E23BFF93FC163E7654CFFE95" box="[956,1003,281,305]" country="Indonesia" name="Bali" pageId="7" pageNumber="324">Bali</collectingRegion>
,
<collectorName id="26C25335E23BFF93FC5E3E75530DFE95" box="[1012,1065,282,305]" pageId="7" pageNumber="324">West</collectorName>
<collectingRegion id="49F3F801E23BFF93FB873E765378FE95" box="[1069,1116,281,305]" country="Indonesia" name="Bali" pageId="7" pageNumber="324">Bali</collectingRegion>
,
<collectorName id="26C25335E23BFF93FBCC3E76523AFE95" box="[1126,1310,281,305]" pageId="7" pageNumber="324">Kab Kelungkung</collectorName>
,
<collectorName id="26C25335E23BFF93FA8C3E76528FFE95" box="[1318,1451,281,305]" pageId="7" pageNumber="324">Tukad Unda</collectorName>
,
<date id="FF891023E23BFF93FC873E56549BFEF5" box="[813,959,313,337]" pageId="7" pageNumber="324" value="2014-04-22">
<collectingDate id="EFCDE9CBE23BFF93FC873E56549BFEF5" box="[813,959,313,337]" pageId="7" pageNumber="324" value="2014-04-22">22 Apr. 2014</collectingDate>
</date>
,
<location id="8EE86038E23BFF93FC613E565333FEF5" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FC613E565333FEF5" box="[971,1047,313,337]" name="Hubert" pageId="7" pageNumber="324" stateProvince="Bali">Hubert</location>
<emphasis id="B943EAF1E23BFF93FBB53E555373FEF5" box="[1055,1111,313,337]" italics="true" pageId="7" pageNumber="324">et al.</emphasis>
coll.; BIF 2846 to BIF 2850
</materialsCitation>
.
<materialsCitation id="3B5F3CBEE23BFF93FC873E365221FE35" collectingDate="2014-04-24" collectionCode="MZB" county="West" location="Tukad Balian" municipality="Kab Tabanan" pageId="7" pageNumber="324" specimenCount="1" stateProvince="Bali">
<collectionCode id="ED26AE26E23BFF93FC873E36544FFED5" box="[813,875,345,369]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="324" type="Museum">MZB</collectionCode>
.25330,
<date id="FF891023E23BFF93FC683E365371FED5" box="[962,1109,345,369]" pageId="7" pageNumber="324" value="2014-04-24">
<collectingDate id="EFCDE9CBE23BFF93FC683E365371FED5" box="[962,1109,345,369]" pageId="7" pageNumber="324" value="2014-04-24">24 Apr. 2014</collectingDate>
</date>
,
<collectingRegion id="49F3F801E23BFF93FBCB3E3653B6FED5" box="[1121,1170,345,369]" country="Indonesia" name="Bali" pageId="7" pageNumber="324">Bali</collectingRegion>
,
<collectingCounty id="62E94E6FE23BFF93FB373E3553F0FED5" box="[1181,1236,346,369]" pageId="7" pageNumber="324">West</collectingCounty>
<collectingRegion id="49F3F801E23BFF93FB703E36522FFED5" box="[1242,1291,345,369]" country="Indonesia" name="Bali" pageId="7" pageNumber="324">Bali</collectingRegion>
,
<collectingMunicipality id="6BECAC99E23BFF93FABD3E36528FFED5" box="[1303,1451,345,369]" pageId="7" pageNumber="324">Kab Tabanan</collectingMunicipality>
,
<location id="8EE86038E23BFF93FC873E1654E6FE35" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FC873E1654E6FE35" box="[813,962,377,401]" county="West" municipality="Kab Tabanan" name="Tukad Balian" pageId="7" pageNumber="324" stateProvince="Bali">Tukad Balian</location>
,
<location id="8EE86038E23BFF93FC673E16533CFE35" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FC673E16533CFE35" box="[973,1048,377,401]" county="West" municipality="Kab Tabanan" name="Hubert" pageId="7" pageNumber="324" stateProvince="Bali">Hubert</location>
<emphasis id="B943EAF1E23BFF93FBB53E155372FE35" box="[1055,1110,377,401]" italics="true" pageId="7" pageNumber="324">et al.</emphasis>
coll.; BIF 2962
</materialsCitation>
.
<materialsCitation id="3B5F3CBEE23BFF93FA893E165301FE75" collectingDate="2015-04-02" collectionCode="MZB" collectorName="Lombok &amp; Lombok Utara &amp; Kali Sidutan" location="Hubert" pageId="7" pageNumber="324" specimenCount="1" stateProvince="Bali">
<collectionCode id="ED26AE26E23BFF93FA893E165245FE35" box="[1315,1377,377,401]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="324" type="Museum">MZB</collectionCode>
.25331,
<collectorName id="26C25335E23BFF93FC873EF654A8FE15" box="[813,908,409,433]" pageId="7" pageNumber="324">Lombok</collectorName>
,
<collectorName id="26C25335E23BFF93FC3C3EF65313FE15" box="[918,1079,409,433]" pageId="7" pageNumber="324">Lombok Utara</collectorName>
,
<collectorName id="26C25335E23BFF93FBE83EF653E9FE15" box="[1090,1229,409,433]" pageId="7" pageNumber="324">Kali Sidutan</collectorName>
,
<date id="FF891023E23BFF93FB723EF6527DFE15" box="[1240,1369,409,433]" pageId="7" pageNumber="324" value="2015-04-02">
<collectingDate id="EFCDE9CBE23BFF93FB723EF6527DFE15" box="[1240,1369,409,433]" pageId="7" pageNumber="324" value="2015-04-02">2 Apr. 2015</collectingDate>
</date>
,
<location id="8EE86038E23BFF93FACE3EF6528BFE15" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FACE3EF6528BFE15" box="[1380,1455,409,433]" name="Hubert" pageId="7" pageNumber="324" stateProvince="Bali">Hubert</location>
<emphasis id="B943EAF1E23BFF93FC873ED55443FE75" box="[813,871,441,465]" italics="true" pageId="7" pageNumber="324">et al.</emphasis>
coll.; BIF 4040
</materialsCitation>
.
<materialsCitation id="3B5F3CBEE23BFF93FBEC3ED65494FDB5" collectingDate="2015-03-28" collectionCode="MZB" collectorName="Lombok &amp; Lombok Utara &amp; Sidutan" location="Hubert" pageId="7" pageNumber="324" specimenCount="1" stateProvince="Bali">
<collectionCode id="ED26AE26E23BFF93FBEC3ED653A2FE75" box="[1094,1158,441,465]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="7" pageNumber="324" type="Museum">MZB</collectionCode>
.25332,
<collectorName id="26C25335E23BFF93FB4B3ED65260FE75" box="[1249,1348,441,465]" pageId="7" pageNumber="324">Lombok</collectorName>
,
<collectorName id="26C25335E23BFF93FAFA3ED6544FFE55" pageId="7" pageNumber="324">Lombok Utara</collectorName>
,
<collectorName id="26C25335E23BFF93FCDF3EB654EDFE55" box="[885,969,473,497]" pageId="7" pageNumber="324">Sidutan</collectorName>
,
<date id="FF891023E23BFF93FC793EB6535CFE55" box="[979,1144,473,497]" pageId="7" pageNumber="324" value="2015-03-28">
<collectingDate id="EFCDE9CBE23BFF93FC793EB6535CFE55" box="[979,1144,473,497]" pageId="7" pageNumber="324" value="2015-03-28">28 March 2015</collectingDate>
</date>
,
<location id="8EE86038E23BFF93FB283EB653E8FE55" LSID="urn:lsid:plazi:treatment:039E87F5E23BFF9CFE8939D75352FA96:8EE86038E23BFF93FB283EB653E8FE55" box="[1154,1228,473,497]" name="Hubert" pageId="7" pageNumber="324" stateProvince="Bali">Hubert</location>
<emphasis id="B943EAF1E23BFF93FB793EB5522CFE55" box="[1235,1288,473,497]" italics="true" pageId="7" pageNumber="324">et al.</emphasis>
coll.; BIF 3790 &amp; BIF 3792
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D6568E23BFF9CFC873D5654D2FE15" lastPageId="8" lastPageNumber="325" pageId="7" pageNumber="324" type="description">
<paragraph id="8B8836E3E23BFF93FC873D565495FDF5" blockId="7.[813,1456,569,1010]" box="[813,945,569,593]" pageId="7" pageNumber="324">
<emphasis id="B943EAF1E23BFF93FC873D565495FDF5" bold="true" box="[813,945,569,593]" pageId="7" pageNumber="324">Description</emphasis>
</paragraph>
<paragraph id="8B8836E3E23BFF93FCF93D36537DFD35" blockId="7.[813,1456,569,1010]" pageId="7" pageNumber="324">The scale and ray counts are given in table II and morphomeristic data in table III.</paragraph>
<paragraph id="8B8836E3E23BFF93FCF93DF654BDFC95" blockId="7.[813,1456,569,1010]" pageId="7" pageNumber="324">
The body is more ovoid than elongated. The body depth at anus is 16-22 (% SL), at first dorsal fin 20-24 (% SL), and the caudal peduncle depth is 13-15 (% SL). Predorsal length 39-47 (% SL) and preanal length 59-68 (% SL). Size: up to
<quantity id="4CCF9B06E23BFF93FC873C765449FC95" box="[813,877,793,817]" metricMagnitude="-1" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="324" unit="cm" value="11.0">11 cm</quantity>
SL.
</paragraph>
<paragraph id="8B8836E3E23BFF9CFCF93C5655B1FEB5" blockId="7.[813,1456,569,1010]" lastBlockId="8.[132,776,217,914]" lastPageId="8" lastPageNumber="325" pageId="7" pageNumber="324">The head (31-37% SL) is depressed, the snout is convex. The anterior nostrils are short, not reaching upper margin of upper lip. Mouth and jaw length (9-12% SL) small. Inwardly curved teeth set on both jaws and larger on outer row. Posterior end of maxillary extending to below around anterior margin of eye. Lips with numerous internal short papillae, large and bifid in several specimens. Eye diameter 6-7 (% SL), and interorbital length small (9-12% SL).</paragraph>
<paragraph id="8B8836E3E234FF9CFF003E76569CFD16" blockId="8.[132,776,217,914]" pageId="8" pageNumber="325">Dorsal fins VI-I,8 without filamentous rays. The first dorsal fin is with second, third and fourth rays longer. Anal fin I,9 directly opposite to the second dorsal fin. The caudal fin is with 13-15 branched rays and its posterior margin is rounded. Pelvic fins separate, I,5. Pectoral fins 14, with the posterior margin rounded. Lateral scales 29-31, with ctenoid scales on flanks and caudal peduncle. Cycloid scales from snout to top of head and anterior part of D1, on operculum, on base of pectoral fins and on belly extending to anus. Scales of top of head and back greater than those of belly. Scales in transverse back series 10-11, in transverse forward 13-16, in predorsal 14-16 and in zigzag 8-10. 6-9 scales around the eye. 25 vertebrae.</paragraph>
<paragraph id="8B8836E3E234FF9CFF003DD55425FD56" blockId="8.[132,776,217,914]" pageId="8" pageNumber="325">Absence of oculoscapular canal and supratemporals and presence of a short preopercular canal with pores N and O.</paragraph>
<paragraph id="8B8836E3E234FF9CFF003D9556A0FC96" blockId="8.[132,776,217,914]" pageId="8" pageNumber="325">
Cephalic sensory papillae system developed as described by
<bibRefCitation id="EFA64B12E234FF9CFF0E3C7556A4FC96" author="AKIHITO &amp; HAYASHI M. &amp; YOSHINO T." box="[164,384,794,818]" pageId="8" pageNumber="325" pagination="236 - 289" refId="ref11110" refString="AKIHITO, HAYASHI M. &amp; YOSHINO T., 1988. - Suborder Gobioidei. In: The Fishes of the Japanese Archipelago, 2 nd Edit. (Masuda H., Amaoko K., Araga C., Uyeno T. &amp; Yoshino T., eds), pp. 236 - 289, 445. Tokai University Press: Tokyo." type="journal article" year="1988">
Akihito
<emphasis id="B943EAF1E234FF9CFF513C75560DFC96" box="[251,297,794,818]" italics="true" pageId="8" pageNumber="325">et al</emphasis>
. (1988)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B8836E3E234FF9CFF003C5555AEFC36" blockId="8.[132,776,217,914]" pageId="8" pageNumber="325">Males with a rounded/triangular urogenital papilla with distal tip rounded. Females have a bulbous urogenital papilla with fimbriate projections around distal opening.</paragraph>
<paragraph id="8B8836E3E234FF9CFF2E3CD556CAFC76" blockId="8.[132,775,954,1202]" box="[132,494,954,978]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFF2E3CD556A1FC76" bold="true" box="[132,389,954,978]" pageId="8" pageNumber="325">Colour in preservation</emphasis>
(
<figureCitation id="130C2A66E234FF9CFE393CD556C1FC76" box="[403,485,954,978]" captionStart="Figure 3" captionStartId="7.[813,879,1789,1811]" captionTargetBox="[813,1456,1065,1775]" captionTargetId="figure-699@7.[813,1456,1065,1775]" captionTargetPageId="7" captionText="Figure 3. A: Giuris tolsoni, RMNH 5180, holotype, male; SL 61.5 mm (Photo RMNH, P. Keith). B: G. tolsoni, MZB.25328, male, Bali; BIF 2283; SL 66 mm (Photo N. Hubert). C: G. tolsoni, in MZB.25329, female, Bali; BIF 2848; SL 34 mm (Photo N. Hubert)." figureDoi="http://doi.org/10.5281/zenodo.12712027" httpUri="https://zenodo.org/record/12712027/files/figure.png" pageId="8" pageNumber="325">Fig. 3A</figureCitation>
)
</paragraph>
<paragraph id="8B8836E3E234FF9CFF003CB5567FFB16" blockId="8.[132,775,954,1202]" pageId="8" pageNumber="325">Male and female similar. Background of body and head brown to beige. Lateral part brownish with, for several specimens, 1-3 brown stripes radiating from the eye to the cheeks and operculum and four to five longitudinal parallel rows of aligned small brownish dots, or a dark longitudinal and medium brown band from pectoral to hypural base. The fins are greyish to beige.</paragraph>
<paragraph id="8B8836E3E234FF9CFF2E3BB5563EFB56" blockId="8.[132,776,1242,1907]" box="[132,282,1242,1266]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFF2E3BB5563EFB56" bold="true" box="[132,282,1242,1266]" pageId="8" pageNumber="325">Colour in life</emphasis>
</paragraph>
<paragraph id="8B8836E3E234FF9CFF003B955790F916" blockId="8.[132,776,1242,1907]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFF003B9557C9FAB6" box="[170,237,1274,1298]" italics="true" pageId="8" pageNumber="325">Males</emphasis>
(
<figureCitation id="130C2A66E234FF9CFF563B95566BFAB6" box="[252,335,1274,1298]" captionStart="Figure 3" captionStartId="7.[813,879,1789,1811]" captionTargetBox="[813,1456,1065,1775]" captionTargetId="figure-699@7.[813,1456,1065,1775]" captionTargetPageId="7" captionText="Figure 3. A: Giuris tolsoni, RMNH 5180, holotype, male; SL 61.5 mm (Photo RMNH, P. Keith). B: G. tolsoni, MZB.25328, male, Bali; BIF 2283; SL 66 mm (Photo N. Hubert). C: G. tolsoni, in MZB.25329, female, Bali; BIF 2848; SL 34 mm (Photo N. Hubert)." figureDoi="http://doi.org/10.5281/zenodo.12712027" httpUri="https://zenodo.org/record/12712027/files/figure.png" pageId="8" pageNumber="325">Fig. 3B</figureCitation>
).
<emphasis id="B943EAF1E234FF9CFECC3B945657FAB6" box="[358,371,1275,1298]" italics="true" pageId="8" pageNumber="325"></emphasis>
Male with background of body light brown on the back to bright beige and yellow on the flanks. Belly white. Top of head brownish, with several small reddish dots, lateral part greyish to bluish with 3 thin red stripes radiating from the eye to the cheeks and operculum, the first or two highest continuing on the pectoral base. Four to five longitudinal parallel rows of aligned small red dots, alternating with blue and yellow parts. Sometimes the medium row forms a continuous line from pectoral base to hypural base. The first dorsal fin is translucent with yellow stripes in the medium part. The second dorsal and anal fins are yellowish at the base and hyaline at the distal half with a thin white stripe at the distal tip. Pectoral, pelvic and caudal fins hyaline.
</paragraph>
<paragraph id="8B8836E3E234FF9CFF0039D554D2FE15" blockId="8.[132,776,1242,1907]" lastBlockId="8.[813,1456,217,433]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFF0039D55621F976" box="[170,261,1722,1746]" italics="true" pageId="8" pageNumber="325">Females</emphasis>
(
<figureCitation id="130C2A66E234FF9CFEB939D55640F977" box="[275,356,1722,1747]" captionStart="Figure 3" captionStartId="7.[813,879,1789,1811]" captionTargetBox="[813,1456,1065,1775]" captionTargetId="figure-699@7.[813,1456,1065,1775]" captionTargetPageId="7" captionText="Figure 3. A: Giuris tolsoni, RMNH 5180, holotype, male; SL 61.5 mm (Photo RMNH, P. Keith). B: G. tolsoni, MZB.25328, male, Bali; BIF 2283; SL 66 mm (Photo N. Hubert). C: G. tolsoni, in MZB.25329, female, Bali; BIF 2848; SL 34 mm (Photo N. Hubert)." figureDoi="http://doi.org/10.5281/zenodo.12712027" httpUri="https://zenodo.org/record/12712027/files/figure.png" pageId="8" pageNumber="325">Fig. 3C</figureCitation>
).
<emphasis id="B943EAF1E234FF9CFED039D456A3F976" box="[378,391,1723,1746]" italics="true" pageId="8" pageNumber="325"></emphasis>
background of body light brown on the back to bright beige on the flanks. Top of head brownish, lateral part greyish to bluish with 3 thin red stripes radiating from the eye to the cheeks and operculum, the first or two highest continuing on the pectoral base. Small alternating reddish to bluish rounded patches along the flanks from pectoral base to hypural base, forming a medium straight or sinuous band. Belly whitish as the lower part of the flanks. The first dorsal fin is translucent. The second dorsal fin is slightly greyish with a thin white stripe at the distal tip. Pectoral fins hyaline. Pelvic fins hyaline with a white distal margin. The anal fin is yellow or grey with a distal white stripe. Caudal fin hyaline.
</paragraph>
</subSubSection>
<subSubSection id="C32D6568E234FF9CFC873EB654F0FD15" pageId="8" pageNumber="325" type="biology_ecology">
<paragraph id="8B8836E3E234FF9CFC873EB654A3FE55" blockId="8.[813,1456,473,689]" box="[813,903,473,497]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFC873EB654A3FE55" bold="true" box="[813,903,473,497]" pageId="8" pageNumber="325">Ecology</emphasis>
</paragraph>
<paragraph id="8B8836E3E234FF9CFCF83E9554F0FD15" blockId="8.[813,1456,473,689]" pageId="8" pageNumber="325">
<taxonomicName id="4C374D60E234FF9CFCF83E9554D7FDB5" box="[850,1011,505,529]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B943EAF1E234FF9CFCF83E9554D7FDB5" box="[850,1011,505,529]" italics="true" pageId="8" pageNumber="325">Giuris tolsoni</emphasis>
</taxonomicName>
occurs in estuaries and lower parts of coastal streams, usually in riverbank vegetation. It is mostly carnivorous and feeds on small shrimps, insects and fish. It is thought to be amphidromous. In
<collectingCountry id="F3207673E234FF9CFB0B3D36522DFDD5" box="[1185,1289,601,625]" name="Indonesia" pageId="8" pageNumber="325">Indonesia</collectingCountry>
it is sometimes found in sympatry with
<taxonomicName id="4C374D60E234FF9CFB843D1553CFFD35" authorityName=", MNHN" authorityYear="2020" box="[1070,1259,634,657]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="margaritaceus">
<emphasis id="B943EAF1E234FF9CFB843D1553CFFD35" box="[1070,1259,634,657]" italics="true" pageId="8" pageNumber="325">G. margaritaceus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C374D60E234FF9CFAA43D15524BFD35" authorityName="Keith &amp; Mennesson &amp; Sauri &amp; Busson &amp; Delrieu-Trottin &amp; Limmon &amp; Nurjirana &amp; Dahruddin &amp; Hubert" authorityYear="2020" box="[1294,1391,634,657]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="viator" status="n. sp.">
<emphasis id="B943EAF1E234FF9CFAA43D15524BFD35" box="[1294,1391,634,657]" italics="true" pageId="8" pageNumber="325">G. viator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A270578AE234FF9CFADC3D15528BFD35" box="[1398,1455,634,657]" pageId="8" pageNumber="325" rank="species">n. sp.</taxonomicNameLabel>
(present paper).
</paragraph>
</subSubSection>
<subSubSection id="C32D6568E234FF9CFC873DB6522BFB56" pageId="8" pageNumber="325" type="distribution">
<paragraph id="8B8836E3E234FF9CFC873DB6549CFD55" blockId="8.[813,1455,729,817]" box="[813,952,729,753]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFC873DB6549CFD55" bold="true" box="[813,952,729,753]" pageId="8" pageNumber="325">Distribution</emphasis>
</paragraph>
<paragraph id="8B8836E3E234FF9CFCF93D955243FC95" blockId="8.[813,1455,729,817]" pageId="8" pageNumber="325">
<taxonomicName id="4C374D60E234FF9CFCF93D9554CEFCB5" box="[851,1002,761,785]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B943EAF1E234FF9CFCF93D9554CEFCB5" box="[851,1002,761,785]" italics="true" pageId="8" pageNumber="325">Giuris tolsoni</emphasis>
</taxonomicName>
is known from
<collectingCountry id="F3207673E234FF9CFB3D3D965226FCB5" box="[1175,1282,761,785]" name="Indonesia" pageId="8" pageNumber="325">Indonesia</collectingCountry>
(
<collectingRegion id="49F3F801E234FF9CFABA3D965265FCB5" box="[1296,1345,761,785]" country="Indonesia" name="Bali" pageId="8" pageNumber="325">Bali</collectingRegion>
, Lombok, Halmahera,
<collectingRegion id="49F3F801E234FF9CFC043C765336FC95" box="[942,1042,793,817]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="325">Sulawesi</collectingRegion>
),
<collectingCountry id="F3207673E234FF9CFB8E3C7653FBFC95" box="[1060,1247,793,817]" name="Philippines Japan" pageId="8" pageNumber="325">Philippines Japan</collectingCountry>
and
<collectingCountry id="F3207673E234FF9CFABB3C765247FC95" box="[1297,1379,793,817]" name="Taiwan" pageId="8" pageNumber="325">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="8B8836E3E234FF9CFC873C36549EFCD5" blockId="8.[813,1456,857,1266]" box="[813,954,857,881]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFC873C36549EFCD5" bold="true" box="[813,954,857,881]" pageId="8" pageNumber="325">Comparison</emphasis>
</paragraph>
<paragraph id="8B8836E3E234FF9CFCF93C15522BFB56" blockId="8.[813,1456,857,1266]" pageId="8" pageNumber="325">
<taxonomicName id="4C374D60E234FF9CFCF93C1554CFFC35" box="[851,1003,889,913]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="tolsoni">
<emphasis id="B943EAF1E234FF9CFCF93C1554CFFC35" box="[851,1003,889,913]" italics="true" pageId="8" pageNumber="325">Giuris tolsoni</emphasis>
</taxonomicName>
differs from the other species sequenced and present in the area by displaying reciprocal monophyly from its closest relatives and high K80+I genetic distances to its relatives at COI gene (0.0556 to its closest relative
<taxonomicName id="4C374D60E234FF9CFC873C9554D0FBB5" authorityName=", MNHN" authorityYear="2020" box="[813,1012,1018,1041]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="margaritaceus">
<emphasis id="B943EAF1E234FF9CFC873C9554D0FBB5" box="[813,1012,1018,1041]" italics="true" pageId="8" pageNumber="325">G. margaritaceus</emphasis>
</taxonomicName>
). Moreover, it differs from
<taxonomicName id="4C374D60E234FF9CFA993C955469FB95" authorityName="Keith &amp; Mennesson &amp; Sauri &amp; Busson &amp; Delrieu-Trottin &amp; Limmon &amp; Nurjirana &amp; Dahruddin &amp; Hubert" authorityYear="2020" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="viator" status="sp. nov.">
<emphasis id="B943EAF1E234FF9CFA993C955469FB95" italics="true" pageId="8" pageNumber="325">Giuris viator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A270578AE234FF9CFCF93B7554AAFB95" box="[851,910,1050,1073]" pageId="8" pageNumber="325" rank="species">n. sp.</taxonomicNameLabel>
(present paper) in having fewer scales around the eye (6-9
<emphasis id="B943EAF1E234FF9CFC393B555496FBF5" box="[915,946,1082,1105]" italics="true" pageId="8" pageNumber="325">vs.</emphasis>
10-13), a smaller body depth at anus (16-22
<emphasis id="B943EAF1E234FF9CFC873B35546EFBD5" box="[813,842,1114,1137]" italics="true" pageId="8" pageNumber="325">vs.</emphasis>
21-24% SL) and by the colourful pattern of the male. It differs from
<emphasis id="B943EAF1E234FF9CFC133B15538BFB35" box="[953,1199,1146,1169]" italics="true" pageId="8" pageNumber="325">
<taxonomicName id="4C374D60E234FF9CFC133B155388FB35" authorityName=", MNHN" authorityYear="2020" box="[953,1196,1146,1169]" family="Eleotridae" genus="Giuris" kingdom="Animalia" order="Perciformes" pageId="8" pageNumber="325" phylum="Chordata" rank="species" species="margaritaceus">Giuris margaritaceus</taxonomicName>
,
</emphasis>
in having fewer scales around the eye (6-8
<emphasis id="B943EAF1E234FF9CFBA63BF5530EFB15" box="[1036,1066,1178,1201]" italics="true" pageId="8" pageNumber="325">vs.</emphasis>
8-11), a smaller interorbital length (9-12
<emphasis id="B943EAF1E234FF9CFCC43BD554AFFB75" box="[878,907,1210,1233]" italics="true" pageId="8" pageNumber="325">vs.</emphasis>
12-14% SL), always 14 pectorals rays
<emphasis id="B943EAF1E234FF9CFA983BD5526BFB75" box="[1330,1359,1210,1233]" italics="true" pageId="8" pageNumber="325">vs.</emphasis>
14 or 15 rays, and by the colourful pattern of the male.
</paragraph>
</subSubSection>
<subSubSection id="C32D6568E234FF9CFC873A755352FA96" box="[813,1142,1305,1330]" pageId="8" pageNumber="325" type="description">
<paragraph id="8B8836E3E234FF9CFC873A755352FA96" blockId="8.[813,1142,1305,1330]" box="[813,1142,1305,1330]" pageId="8" pageNumber="325">
<emphasis id="B943EAF1E234FF9CFC873A755352FA96" bold="true" box="[813,1142,1305,1330]" italics="true" pageId="8" pageNumber="325">Description of the new species</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>