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<mods:title id="D03231454BAA2A1A43B51F825B053D89">Morphological diagnoses of higher taxa in Ophiuroidea (Echinodermata) in support of a new classification</mods:title>
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<treatment id="039E87E81F67302DFDD1FC4F7D4EFA02" ID-DOI="http://doi.org/10.5281/zenodo.3816345" ID-GBIF-Taxon="163971278" ID-Zenodo-Dep="3816345" LSID="urn:lsid:zoobank.org:act:67BE8BBA-D959-40E4-9E02-73A9F483A18E" httpUri="http://treatment.plazi.org/id/039E87E81F67302DFDD1FC4F7D4EFA02" lastPageNumber="8" pageId="7" pageNumber="8">
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Family
<taxonomicName id="4C374D7D1F67302DFD0CFC4E7C54FC5A" ID-CoL="7NK7H" authorityName="O'Hara, Stohr, Hugall, Thuy &amp; Martynov" authorityYear="2018" box="[670,892,908,934]" class="Ophiuroidea" family="Ophiosphalmidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="family" status="fam. nov.">
<emphasis id="B943EAEC1F67302DFD0CFC4E7C54FC5A" bold="true" box="[670,892,908,934]" pageId="7" pageNumber="8">Ophiosphalmidae</emphasis>
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.
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<heading id="D0C081921F67302DFF2FFC387E62FBE8" bold="true" box="[189,330,1018,1044]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
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genus
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<taxonomicName id="4C374D7D1F67302DFF2FFBEF7D68FBBB" authority="H. L. Clark, 1941" authorityName="H. L. Clark" authorityYear="1941" box="[189,576,1069,1096]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F67302DFF2FFBEF7E42FBBB" box="[189,362,1069,1095]" italics="true" pageId="7" pageNumber="8">Ophiosphalma</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F67302DFE21FBEF7D68FBBB" author="Clark" box="[435,576,1069,1095]" firstAuthor="Clark" pageId="7" pageNumber="8" pagination="1 - 154" refId="ref15378" refString="Clark H. L. 1941. Reports on the scientific results of the Atlantis expeditions to the West Indies, under the joint auspices of the University of Havana and Harvard University. The echinoderms (other than Holothurians). Memorias de la Sociedad cubana de Historia natural, &quot; Felipe Poey &quot; 15: 1 - 154." type="journal article" year="1941">Clark, 1941</bibRefCitation>
</taxonomicName>
(
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species:
<taxonomicName id="4C374D7D1F67302DFD64FBEF7B15FBB4" authority="(Lyman, 1878 b)" baseAuthorityName="Lyman" baseAuthorityYear="1878" box="[758,1085,1069,1096]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="planum">
<emphasis id="B943EAEC1F67302DFD64FBEF7C23FBBB" box="[758,779,1069,1095]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="B943EAEC1F67302DFC8EFBEF7C5CFBBB" box="[796,884,1069,1095]" italics="true" pageId="7" pageNumber="8">planum</emphasis>
(
<bibRefCitation id="EFA64B0F1F67302DFC14FBEF7B1DFBB4" author="Lyman" box="[902,1077,1069,1096]" firstAuthor="Lyman" pageId="7" pageNumber="8" pagination="217 - 238" refId="ref17435" refString="Lyman T. 1878 b. Ophiurans and Astrophytons. Reports on the dredging operations of the U. S. coast survey Str. &quot; Blake &quot;. Bulletin of the Museum of Comparative Zoology at Harvard 5: 217 - 238, 3 pls." type="book chapter" year="1878" yearSuffix="b">Lyman, 1878b</bibRefCitation>
)
</taxonomicName>
=
<taxonomicName id="4C374D7D1F67302DFBF0FBEF7E34FB93" authority="(Lyman, 1878 a))" authorityName="" baseAuthorityName="Lyman" baseAuthorityYear="1878" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiosphalma" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="armigerum">
<emphasis id="B943EAEC1F67302DFBF0FBEF7B5FFBBB" box="[1122,1143,1069,1095]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="B943EAEC1F67302DFB1AFBEC7A22FBBB" box="[1160,1290,1070,1095]" italics="true" pageId="7" pageNumber="8">armigerum</emphasis>
(
<bibRefCitation id="EFA64B0F1F67302DFA8EFBEF7E20FB93" author="Lyman" firstAuthor="Lyman" pageId="7" pageNumber="8" pagination="65 - 168" refId="ref17401" refString="Lyman T. 1878 a. Ophiuridae and Astrophytidae of the &quot; Challenger &quot; expedition. Part I. Bulletin of the Museum of Comparative Zoology at Harvard 5: 65 - 168." type="journal article" year="1878" yearSuffix="a">Lyman, 1878a</bibRefCitation>
))
</taxonomicName>
.
</paragraph>
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<heading id="D0C081921F67302DFF2FFB587E4CFB49" bold="true" box="[189,356,1178,1205]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B943EAEC1F67302DFF2FFB587E4CFB49" bold="true" box="[189,356,1178,1205]" pageId="7" pageNumber="8">Other genera</emphasis>
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<taxonomicName id="4C374D7D1F67302DFF2FFB0F7EDBFB1B" authority="Lyman, 1878 b" authorityName="Lyman" authorityYear="1878" box="[189,499,1229,1256]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiolipus" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F67302DFF2FFB0F7E17FB1B" box="[189,319,1229,1255]" italics="true" pageId="7" pageNumber="8">Ophiolipus</emphasis>
<bibRefCitation id="EFA64B0F1F67302DFED5FB0C7EDBFB1B" author="Lyman" box="[327,499,1229,1256]" firstAuthor="Lyman" pageId="7" pageNumber="8" pagination="217 - 238" refId="ref17435" refString="Lyman T. 1878 b. Ophiurans and Astrophytons. Reports on the dredging operations of the U. S. coast survey Str. &quot; Blake &quot;. Bulletin of the Museum of Comparative Zoology at Harvard 5: 217 - 238, 3 pls." type="book chapter" year="1878" yearSuffix="b">Lyman, 1878b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F67302DFE6DFB0F7C6EFB1B" authority="Lyman, 1869" authorityName="Lyman" authorityYear="1869" box="[511,838,1229,1256]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F67302DFE6DFB0F7D8CFB1B" box="[511,676,1229,1255]" italics="true" pageId="7" pageNumber="8">Ophiomusium</emphasis>
<bibRefCitation id="EFA64B0F1F67302DFD39FB0C7C6EFB1B" author="Lyman" box="[683,838,1229,1256]" firstAuthor="Lyman" pageId="7" pageNumber="8" pagination="309 - 354" refId="ref17290" refString="Lyman T. 1869. Preliminary report on the Ophiuridae and Astrophytidae dredged in deep water between Cuba and Florida Reef. Bulletin of the Museum of Comparative Zoology 1: 309 - 354." type="journal article" year="1869">Lyman, 1869</bibRefCitation>
</taxonomicName>
(only species
<taxonomicName id="4C374D7D1F67302DFC7CFB0C7A06FB14" authority="Lyman, 1869" authorityName="Lyman" authorityYear="1869" box="[1006,1326,1229,1256]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomusium" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="species" species="eburneum">
<emphasis id="B943EAEC1F67302DFC7CFB0C7B2BFB14" box="[1006,1027,1230,1256]" italics="true" pageId="7" pageNumber="8">O</emphasis>
.
<emphasis id="B943EAEC1F67302DFB80FB0F7BAFFB1B" box="[1042,1159,1229,1255]" italics="true" pageId="7" pageNumber="8">eburneum</emphasis>
<bibRefCitation id="EFA64B0F1F67302DFB1DFB0C7A06FB14" author="Lyman" box="[1167,1326,1229,1256]" firstAuthor="Lyman" pageId="7" pageNumber="8" pagination="309 - 354" refId="ref17290" refString="Lyman T. 1869. Preliminary report on the Ophiuridae and Astrophytidae dredged in deep water between Cuba and Florida Reef. Bulletin of the Museum of Comparative Zoology 1: 309 - 354." type="journal article" year="1869">Lyman, 1869</bibRefCitation>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C32D65751F67302DFF2FFAD17D4EFA02" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8B8836FE1F67302DFF2FFAD17E1EFAD1" blockId="7.[189,310,1299,1325]" box="[189,310,1299,1325]" pageId="7" pageNumber="8">
<heading id="D0C081921F67302DFF2FFAD17E1EFAD1" bold="true" box="[189,310,1299,1325]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B943EAEC1F67302DFF2FFAD17E1EFAD1" bold="true" box="[189,310,1299,1325]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F67302DFF2FFA857D4EFA02" blockId="7.[189,1399,1350,1535]" pageId="7" pageNumber="8">
Numerous disc scales, small primary plates. Dorsal and ventral arm plates present along most of the arm. Lateral arm plate with a finely meshed proximal band that lacks spurs. Longer genital slits than
<taxonomicName id="4C374D7D1F67302DFF2FFA577E6CFA53" authorityName="Hertz" authorityYear="1927" box="[189,324,1429,1455]" class="Ophiuroidea" family="Ophiomusaidae" genus="Ophiomusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F67302DFF2FFA577E6CFA53" box="[189,324,1429,1455]" italics="true" pageId="7" pageNumber="8">Ophiomusa</emphasis>
</taxonomicName>
. Three (rarely 25) visible proximal pairs of tentacle pores. Frequently with small disc spines distal to the genital slit around the disc margin. In our molecular analysis, the genus
<taxonomicName id="4C374D7D1F67302DFB66FA7F7A5EFA2B" authorityName="Lyman" authorityYear="1878" box="[1268,1398,1469,1495]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiolipus" kingdom="Animalia" order="Ophiurida" pageId="7" pageNumber="8" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F67302DFB66FA7F7A5EFA2B" box="[1268,1398,1469,1495]" italics="true" pageId="7" pageNumber="8">Ophiolipus</emphasis>
</taxonomicName>
(lacking disc plates) is polyphyletic.
</paragraph>
</subSubSection>
</treatment>
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