157 lines
19 KiB
XML
157 lines
19 KiB
XML
<document id="D2CAF7546572FF7803B0B0BBD6225C1F" ID-CLB-Dataset="46646" ID-DOI="10.5281/zenodo.206513" ID-GBIF-Dataset="3c536076-5504-4715-9a96-930397f8e6ce" ID-ISSN="1175-5326" ID-Zenodo-Dep="206513" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460370589621" checkinUser="plazi" docAuthor="Cruz, Estevam L., Silva, Da & Lise, Arno A." docDate="2011" docId="039E87E0D37AD1481CABFAB7CD88363B" docLanguage="en" docName="zt02919p068.pdf" docOrigin="Zootaxa 2919" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Enna triste Cruz, Silva & Lise, 2011, new species" docType="treatment" docVersion="9" lastPageNumber="64" masterDocId="FFA7FF98D379D14C1C3CFFE3CC283704" masterDocTitle="Seven new species of Enna (Araneae: Trechaleidae) from Central and South America" masterLastPageNumber="68" masterPageNumber="60" pageNumber="63" updateTime="1698295673398" updateUser="ExternalLinkService">
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<mods:mods id="9C59CF2D2CABD0B1E179E46C7AB011E6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="724259F8E0EF6B64934F6342049D1D2D">
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<mods:title id="6CEE8205402DD398184EB4EEF4F516CF">Seven new species of Enna (Araneae: Trechaleidae) from Central and South America</mods:title>
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</mods:titleInfo>
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<mods:name id="EAB45613EFC94E0C302904A1DC081E26" type="personal">
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<mods:roleTerm id="7E86601E2125FC248DC96F989EB6FB1E">Author</mods:roleTerm>
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<mods:namePart id="1BFD27C1605A08F59976C18417D90F10">Cruz, Estevam L.</mods:namePart>
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<mods:name id="9F04E7EBD564554FDA83AAA1CCEE4DB8" type="personal">
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<mods:roleTerm id="2A023746A3998340C8108258AE1DA529">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F208639E1F5F0F584A92BEEA87C7680E">Silva, Da</mods:namePart>
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</mods:name>
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<mods:name id="884982DD3DEBFEBA2E3AB653D445BE56" type="personal">
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<mods:role id="01A74E362B2747A8950365DBF8B89039">
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<mods:roleTerm id="89DF466E79F01B77F4A2618C8E964D23">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F9E9047B27218C103AFDEBA76928009">Lise, Arno A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="1D1E453A61F89BF83090874C8046E7DA">text</mods:typeOfResource>
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<mods:titleInfo id="3DA2C7B274530E2DA3E0ED96D093E5BF">
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<mods:title id="31C5397FECD1EF6340405E2B8FE11836">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="B808EF797580F20D0A00AE17A8FE41BE">
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<mods:date id="042123559BDCA734CB52C094F9E08E22">2011</mods:date>
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<mods:detail id="9F833A997A5BEB0CE00026A1F6BCD24B" type="volume">
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<mods:number id="F397C6B70BEC9F8B805907C41DFD7AC0">2919</mods:number>
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<mods:start id="10D2B48EE225ACDE09384CB29DD41553">60</mods:start>
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<mods:end id="29C8676DA0FD18A94E0DD315C07A9F77">68</mods:end>
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<mods:classification id="A4D8D699E08EBA80CEE6760C5B9DC9CB">journal article</mods:classification>
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<mods:identifier id="33C0FA218D116F45B024B8A44716851F" type="CLB-Dataset">46646</mods:identifier>
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<mods:identifier id="0D649901D3693E130EAA25DF5C2C147B" type="DOI">10.5281/zenodo.206513</mods:identifier>
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<mods:identifier id="911E73877D6E5E2653DE470A81834513" type="GBIF-Dataset">3c536076-5504-4715-9a96-930397f8e6ce</mods:identifier>
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<mods:identifier id="A594CBA0D50BB264ECE5973C72ABF774" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D7BF9C94F582E352D9450CAF33028EEF" type="Zenodo-Dep">206513</mods:identifier>
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<treatment id="039E87E0D37AD1481CABFAB7CD88363B" ID-DOI="http://doi.org/10.5281/zenodo.6184199" ID-GBIF-Taxon="119411832" ID-Zenodo-Dep="6184199" LSID="urn:lsid:plazi:treatment:039E87E0D37AD1481CABFAB7CD88363B" httpUri="http://treatment.plazi.org/id/039E87E0D37AD1481CABFAB7CD88363B" lastPageId="4" lastPageNumber="64" pageId="3" pageNumber="63">
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<subSubSection id="C32D657DD37AD14F1CABFAB7CCD0328B" pageId="3" pageNumber="63" type="nomenclature">
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<paragraph id="8B8836F6D37AD14F1CABFAB7CD9C326A" blockId="3.[151,436,1364,1423]" box="[151,436,1364,1390]" pageId="3" pageNumber="63">
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<heading id="D0C0819AD37AD14F1CABFAB7CD9C326A" bold="true" box="[151,436,1364,1390]" fontSize="11" level="1" pageId="3" pageNumber="63" reason="1">
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<emphasis id="B943EAE4D37AD14F1CABFAB7CD9C326A" bold="true" box="[151,436,1364,1390]" pageId="3" pageNumber="63">
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<taxonomicName id="4C374D75D37AD14F1CABFAB7CD35326A" ID-CoL="39VL2" box="[151,285,1364,1390]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="triste" status="sp. nov.">
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<emphasis id="B943EAE4D37AD14F1CABFAB7CD35326A" bold="true" box="[151,285,1364,1390]" italics="true" pageId="3" pageNumber="63">Enna triste</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A270579FD37AD14F1D18FAB7CD9C326A" box="[292,436,1364,1390]" pageId="3" pageNumber="63" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8836F6D37AD14F1CABFA94CCD0328B" blockId="3.[151,436,1364,1423]" box="[151,248,1399,1423]" pageId="3" pageNumber="63">
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<figureCitation id="130C2A73D37AD14F1CABFA94CCD0328B" box="[151,248,1399,1423]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[204,1377,204,1842]" captionTargetId="figure@2.[204,1380,193,1843]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Enna carinata sp. nov. 1, 2. male palpus (1 ventral, 2 retrolateral). Enna triste sp. nov. 3, 4. male palpus (3 ventral, 4 retrolateral). Enna caricoi sp. nov. 5, 6. male palpus (5 ventral, 6 retrolateral). Enna venezuelana sp. nov. 7, 8. female epigynum (7 ventral, 8 dorsal). (DD, dorsal division of the median apophysis; ECD, ectal division of the retrolateral tibial apophysis; FD, fertilizations ducts; HS, head of spermathecae; LL, lateral lobe; LLA, lateral lamella; MF, middle field of epigynum; ST, subtegulum; T, tegulum; VD, ventral division of the median apophysis)." httpUri="https://zenodo.org/record/206514/files/figure.png" pageId="3" pageNumber="63">Figs 3, 4</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32D657DD37AD14F1CABFA5CCD2F32FF" pageId="3" pageNumber="63" type="materials_examined">
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<paragraph id="8B8836F6D37AD14F1CABFA5CCD2F32FF" blockId="3.[151,1437,1471,2035]" pageId="3" pageNumber="63">
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<emphasis id="B943EAE4D37AD14F1CABFA5CCCFE32D3" bold="true" box="[151,214,1471,1495]" pageId="3" pageNumber="63">
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<typeStatus id="548C8854D37AD14F1CABFA5CCCF832D3" box="[151,208,1471,1495]" pageId="3" pageNumber="63">Type</typeStatus>
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.
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</emphasis>
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Male
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<typeStatus id="548C8854D37AD14F1D1FFA5CCDAD32D3" box="[291,389,1471,1495]" pageId="3" pageNumber="63" type="holotype">holotype</typeStatus>
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from Golfo
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<taxonomicName id="4C374D75D37AD14F1E29FA5CCE7032D3" box="[533,600,1471,1495]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="triste" status="sp. nov.">Triste</taxonomicName>
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,
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<collectingCountry id="F3207666D37AD14F1E5AFA5CCEF132D3" box="[614,729,1471,1495]" name="Venezuela" pageId="3" pageNumber="63">Venezuela</collectingCountry>
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[
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<geoCoordinate id="EE035031D37AD14F1ED4FA5CCFB132D3" box="[744,921,1471,1495]" direction="north" orientation="latitude" pageId="3" pageNumber="63" precision="1" value="10.006014">10°00’21.65”N</geoCoordinate>
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, 66°55’49.44’W],
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<date id="FF891036D37AD14F1852FA5CC8D832D3" box="[1134,1264,1471,1495]" pageId="3" pageNumber="63" value="1942-09-06">06.IX.1942</date>
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, W. M. Beeber (
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<collectionCode id="ED26AE33D37AD14F1CA3FA00CCD332FF" LSID="urn:lsid:biocol.org:col:34925" box="[159,251,1507,1531]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="63">AMNH</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D657DD37AD14F1CF9F9E4CFC8311B" box="[197,992,1543,1568]" pageId="3" pageNumber="63" type="etymology">
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<paragraph id="8B8836F6D37AD14F1CF9F9E4CFC8311B" blockId="3.[151,1437,1471,2035]" box="[197,992,1543,1568]" pageId="3" pageNumber="63">
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<emphasis id="B943EAE4D37AD14F1CF9F9E4CD63311B" bold="true" box="[197,331,1543,1567]" pageId="3" pageNumber="63">Etymology.</emphasis>
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The specific name is a noun and refers to the
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<typeStatus id="548C8854D37AD14F1F73F9EBCF573124" box="[847,895,1544,1568]" pageId="3" pageNumber="63">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D657DD37AD14F1CF9F9C8C86731AB" pageId="3" pageNumber="63" type="diagnosis">
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<paragraph id="8B8836F6D37AD14F1CF9F9C8C86731AB" blockId="3.[151,1437,1471,2035]" pageId="3" pageNumber="63">
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<emphasis id="B943EAE4D37AD14F1CF9F9C8CD173140" bold="true" box="[197,319,1579,1604]" pageId="3" pageNumber="63">Diagnosis.</emphasis>
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The male of
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<taxonomicName id="4C374D75D37AD14F1DE8F9CFCE053147" box="[468,557,1580,1603]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="triste" status="sp. nov.">
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<emphasis id="B943EAE4D37AD14F1DE8F9CFCE053147" box="[468,557,1580,1603]" italics="true" pageId="3" pageNumber="63">E. triste</emphasis>
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</taxonomicName>
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<emphasis id="B943EAE4D37AD14F1E08F9CFCEA53140" bold="true" box="[564,653,1580,1604]" pageId="3" pageNumber="63">
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<taxonomicNameLabel id="A270579FD37AD14F1E08F9CFCEA53140" box="[564,653,1580,1604]" pageId="3" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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resembles those of
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<taxonomicName id="4C374D75D37AD14F1F50F9CFC93F3147" authority="Silva, Lise & Carico 2008" authorityName="Silva, Lise & Carico" authorityYear="2008" box="[876,1303,1579,1603]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="huanuco">
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<emphasis id="B943EAE4D37AD14F1F50F9CFCFC33147" box="[876,1003,1579,1603]" italics="true" pageId="3" pageNumber="63">E. huanuco</emphasis>
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<bibRefCitation id="EFA64B07D37AD14F1FCEF9C8C93F3147" author="Silva" box="[1010,1303,1579,1603]" pageId="3" pageNumber="63" refString="Silva, E. L. C., Lise, A. A. & Carico, J. E. (2008) Revision of the Neotropical spider genus Enna (Araneae, Lycosoidea, Trechaleidae). Journal of Arachnology, 36, 76 - 110." type="journal article" year="2008">Silva, Lise & Carico 2008</bibRefCitation>
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</taxonomicName>
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by the general shape of the dorsal division (DD) of the median apophysis (
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<bibRefCitation id="EFA64B07D37AD14F1F55F9ACC80E3163" author="Silva" box="[873,1062,1615,1639]" pageId="3" pageNumber="63" refString="Silva, E. L. C., Lise, A. A. & Carico, J. E. (2008) Revision of the Neotropical spider genus Enna (Araneae, Lycosoidea, Trechaleidae). Journal of Arachnology, 36, 76 - 110." type="journal article" year="2008">
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Silva
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<emphasis id="B943EAE4D37AD14F1F97F9B3CFF43163" box="[939,988,1615,1639]" italics="true" pageId="3" pageNumber="63">et al</emphasis>
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. 2008
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</bibRefCitation>
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, p. 91, fig. 73), but can be distinguished by the larger lateral lamella (LLA) on the dorsal division of the median apophysis (
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<figureCitation id="130C2A73D37AD14F18F4F990C926318F" box="[1224,1294,1651,1675]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[204,1377,204,1842]" captionTargetId="figure@2.[204,1380,193,1843]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Enna carinata sp. nov. 1, 2. male palpus (1 ventral, 2 retrolateral). Enna triste sp. nov. 3, 4. male palpus (3 ventral, 4 retrolateral). Enna caricoi sp. nov. 5, 6. male palpus (5 ventral, 6 retrolateral). Enna venezuelana sp. nov. 7, 8. female epigynum (7 ventral, 8 dorsal). (DD, dorsal division of the median apophysis; ECD, ectal division of the retrolateral tibial apophysis; FD, fertilizations ducts; HS, head of spermathecae; LL, lateral lobe; LLA, lateral lamella; MF, middle field of epigynum; ST, subtegulum; T, tegulum; VD, ventral division of the median apophysis)." httpUri="https://zenodo.org/record/206514/files/figure.png" pageId="3" pageNumber="63">Fig. 3</figureCitation>
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) and by the slender, acute ectal division (ECD) of the retrolateral tibial apophysis (RTA) (
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<figureCitation id="130C2A73D37AD14F1FC0F974C86A31AB" box="[1020,1090,1687,1711]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[204,1377,204,1842]" captionTargetId="figure@2.[204,1380,193,1843]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Enna carinata sp. nov. 1, 2. male palpus (1 ventral, 2 retrolateral). Enna triste sp. nov. 3, 4. male palpus (3 ventral, 4 retrolateral). Enna caricoi sp. nov. 5, 6. male palpus (5 ventral, 6 retrolateral). Enna venezuelana sp. nov. 7, 8. female epigynum (7 ventral, 8 dorsal). (DD, dorsal division of the median apophysis; ECD, ectal division of the retrolateral tibial apophysis; FD, fertilizations ducts; HS, head of spermathecae; LL, lateral lobe; LLA, lateral lamella; MF, middle field of epigynum; ST, subtegulum; T, tegulum; VD, ventral division of the median apophysis)." httpUri="https://zenodo.org/record/206514/files/figure.png" pageId="3" pageNumber="63">Fig. 4</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32D657DD37AD1481CF9F958CF0637F3" lastPageId="4" lastPageNumber="64" pageId="3" pageNumber="63" type="description">
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<paragraph id="8B8836F6D37AD1481CF9F958CF0637F3" blockId="3.[151,1437,1471,2035]" lastBlockId="4.[151,1436,151,320]" lastPageId="4" lastPageNumber="64" pageId="3" pageNumber="63">
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<emphasis id="B943EAE4D37AD14F1CF9F958CD7F31D0" bold="true" box="[197,343,1723,1748]" pageId="3" pageNumber="63">Description.</emphasis>
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Male (
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<typeStatus id="548C8854D37AD14F1D95F958CE3D31D7" box="[425,533,1723,1747]" pageId="3" pageNumber="63" type="holotype">Holotype</typeStatus>
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, Golfo
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<taxonomicName id="4C374D75D37AD14F1E51F958CE9831D7" box="[621,688,1723,1747]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="63" phylum="Arthropoda" rank="species" species="triste" status="sp. nov.">Triste</taxonomicName>
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,
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<collectingCountry id="F3207666D37AD14F1E81F958CF1B31D7" box="[701,819,1723,1747]" name="Venezuela" pageId="3" pageNumber="63">Venezuela</collectingCountry>
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, AMNH). Total length 6.22. Carapace 4.15 long, 3.48 wide, dark brown. Clypeus dark brown, 0.28 high. Anterior eye row straight, 0.94 wide; posterior 1.70 wide. Eye diameters, interdistances, and median ocular quadrangle: AME 0.18, ALE 0.14, PME 0.22, PLE 0.26; AME-AME 0.12, AME-ALE 0.07, PME-PME 0.40, PME-PLE 0.34, OQA 0.50, OQP 0.90, OQH 0.64. Chelicerae dark brown, swollen distally, glabrous, with lateral carina; promargin and retromargin of fang furrow with three teeth equidistant and equal in size. Sternum light brown, darker laterally, scattered setae; 1.49 long, 1.99 wide. Labium dark brown, light brown distally, 0.41 long, 0.66 wide. Legs dark brown, unmarked, relative length: II-IV-III. Leg measurements: I—femur 6.22/ tibia-patella 7.71/ metatarsus 6.30/ tarsus 2.98/ total 23.21; II—6.25/ 8.72/ 7.55/ 2.57/ 25.09; III—5.06/ 6.22/ 5.56/ 1.85/ 18.69. Ventral pairs of macrosetae on tibiae:
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<date id="FF891036D37AD14F1831F838C81A30F7" box="[1037,1074,2011,2035]" pageId="3" pageNumber="63">I-4</date>
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;
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<date id="FF891036D37AD14F1801F838C84230F7" box="[1085,1130,2011,2035]" pageId="3" pageNumber="63">II-4</date>
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;
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<date id="FF891036D37AD14F1849F838C89830F7" box="[1141,1200,2011,2035]" pageId="3" pageNumber="63">III-3.</date>
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Abdomen 2.90 long, grayish, with irregular white patches on dorsum; three pairs of sigilla. Venter light gray, unmarked. Palpus with enlarged lateral lamella (LLA) on the dorsal division of the median apophysis (
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<figureCitation id="130C2A73D37DD1481834FF5FC86437D0" box="[1032,1100,188,212]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[204,1377,204,1842]" captionTargetId="figure@2.[204,1380,193,1843]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Enna carinata sp. nov. 1, 2. male palpus (1 ventral, 2 retrolateral). Enna triste sp. nov. 3, 4. male palpus (3 ventral, 4 retrolateral). Enna caricoi sp. nov. 5, 6. male palpus (5 ventral, 6 retrolateral). Enna venezuelana sp. nov. 7, 8. female epigynum (7 ventral, 8 dorsal). (DD, dorsal division of the median apophysis; ECD, ectal division of the retrolateral tibial apophysis; FD, fertilizations ducts; HS, head of spermathecae; LL, lateral lobe; LLA, lateral lamella; MF, middle field of epigynum; ST, subtegulum; T, tegulum; VD, ventral division of the median apophysis)." httpUri="https://zenodo.org/record/206514/files/figure.png" pageId="4" pageNumber="64">Fig. 3</figureCitation>
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). Retrolateral tibial apophysis with a prominent and slender ectal division (ECD) (
|
||
<figureCitation id="130C2A73D37DD1481EE0FF3CCF0937F3" box="[732,801,223,247]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1880,1903]" captionTargetBox="[204,1377,204,1842]" captionTargetId="figure@2.[204,1380,193,1843]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Enna carinata sp. nov. 1, 2. male palpus (1 ventral, 2 retrolateral). Enna triste sp. nov. 3, 4. male palpus (3 ventral, 4 retrolateral). Enna caricoi sp. nov. 5, 6. male palpus (5 ventral, 6 retrolateral). Enna venezuelana sp. nov. 7, 8. female epigynum (7 ventral, 8 dorsal). (DD, dorsal division of the median apophysis; ECD, ectal division of the retrolateral tibial apophysis; FD, fertilizations ducts; HS, head of spermathecae; LL, lateral lobe; LLA, lateral lamella; MF, middle field of epigynum; ST, subtegulum; T, tegulum; VD, ventral division of the median apophysis)." httpUri="https://zenodo.org/record/206514/files/figure.png" pageId="4" pageNumber="64">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32D657DD37DD1481CF9FEE0CD88363B" pageId="4" pageNumber="64" type="distribution">
|
||
<paragraph id="8B8836F6D37DD1481CF9FEE0CD88363B" blockId="4.[151,1436,151,320]" pageId="4" pageNumber="64">
|
||
<emphasis id="B943EAE4D37DD1481CF9FEE0CD763618" bold="true" box="[197,350,259,284]" pageId="4" pageNumber="64">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="548C8854D37DD1481E62FEE6CEA63619" box="[606,654,261,285]" pageId="4" pageNumber="64">type</typeStatus>
|
||
locality (Golfo
|
||
<taxonomicName id="4C374D75D37DD1481F7EFEE7CFAD3618" box="[834,901,260,284]" class="Arachnida" family="Trechaleidae" genus="Enna" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="triste" status="sp. nov.">Triste</taxonomicName>
|
||
,
|
||
<collectingCountry id="F3207666D37DD1481FACFEE7C8213618" box="[912,1033,260,284]" name="Venezuela" pageId="4" pageNumber="64">Venezuela</collectingCountry>
|
||
).
|
||
<emphasis id="B943EAE4D37DD1481CF9FECBCD0B3644" bold="true" box="[197,291,296,320]" pageId="4" pageNumber="64">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |