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<document ID-DOI="10.3853/j.0067-1975.55.2003.1387" ID-ISSN="2201-4349" approvalRequired="85" approvalRequired_for_document="1" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="34" approvalRequired_for_textStreams="45" approvalRequired_for_treatments="4" checkinTime="1629578461805" checkinUser="felipe" docAuthor="Rickards, R. B., Chapman, A. J., Wright, A. J. &amp; Packham, G. H" docDate="2003" docId="039E87A3F92AFFBD73B84030996AE59C" docLanguage="en" docName="RecAustMus.55.3.305-330.pdf" docOrigin="Records of the Australian Museum 55 (3)" docSource="https://journals.australian.museum/rickards-et-al-2003-rec-aust-mus-553-305330/" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.1:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="1" docTitle="Cyclograptus Spencer 1884" docType="treatment" docVersion="3" lastPageNumber="328" masterDocId="FFA7FFDBF93DFFAA7009443B9D31E021" masterDocTitle="Dendroid and Tuboid Graptolites from the Llandovery (Silurian) of the Four Mile Creek Area, New South Wales" masterLastPageNumber="330" masterPageNumber="305" pageNumber="328" updateTime="1630661534800" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Dendroid and Tuboid Graptolites from the Llandovery (Silurian) of the Four Mile Creek Area, New South Wales</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Rickards, R. B.</mods:namePart>
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<mods:namePart>Chapman, A. J.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wright, A. J.</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Packham, G. H</mods:namePart>
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<mods:title>Records of the Australian Museum</mods:title>
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<mods:date>2003</mods:date>
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<mods:number>2003-12-10</mods:number>
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<mods:number>55</mods:number>
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<taxonomicName authority="Spencer, 1884" authorityName="Spencer" authorityYear="1884" box="[945,1270,1034,1059]" class="Graptolithina" family="Cyclograptidae" genus="Cyclograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tuboidea" pageId="23" pageNumber="328" phylum="Hemichordata" rank="genus">
<emphasis bold="true" box="[945,1270,1034,1059]" pageId="23" pageNumber="328">
<emphasis bold="true" box="[945,1098,1035,1059]" italics="true" pageId="23" pageNumber="328">Cyclograptus</emphasis>
<bibRefCitation author="Spencer" box="[1106,1270,1034,1058]" firstAuthor="Spencer" pageId="23" pageNumber="328" pagination="1 - 43" refId="ref19572" refString="Spencer, J. W., 1884. Graptolites of the Upper Silurian System. University of Missouri Museum Bulletin 1: 1 - 43." type="journal article" year="1884">Spencer, 1884</bibRefCitation>
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<subSubSection pageId="23" pageNumber="328" type="type_taxon">
<paragraph blockId="23.[794,1421,1079,1131]" pageId="23" pageNumber="328">
<emphasis bold="true" box="[794,940,1079,1103]" pageId="23" pageNumber="328">
<typeStatus box="[794,851,1079,1103]" pageId="23" pageNumber="328">Type</typeStatus>
species
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.
<taxonomicName authority="Spencer, 1884, p. 42" authorityName="Spencer" authorityPageNumber="42" authorityYear="1884" class="Graptolithina" family="Cyclograptidae" genus="Cyclograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tuboidea" pageId="23" pageNumber="328" phylum="Hemichordata" rank="species" species="rotadentatus">
<emphasis box="[957,1248,1079,1102]" italics="true" pageId="23" pageNumber="328">Cyclograptus rotadentatus</emphasis>
<bibRefCitation author="Spencer" box="[1258,1417,1079,1102]" firstAuthor="Spencer" pageId="23" pageNumber="328" pagination="1 - 43" refId="ref19572" refString="Spencer, J. W., 1884. Graptolites of the Upper Silurian System. University of Missouri Museum Bulletin 1: 1 - 43." type="journal article" year="1884">Spencer, 1884</bibRefCitation>
, p. 42
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; by original designation.
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<subSubSection pageId="23" pageNumber="328" type="discussion">
<paragraph blockId="23.[794,1422,1152,1469]" pageId="23" pageNumber="328">
<emphasis bold="true" box="[794,897,1152,1176]" pageId="23" pageNumber="328">Remarks</emphasis>
.
<bibRefCitation author="Bulman" box="[908,1068,1153,1176]" firstAuthor="Bulman" pageId="23" pageNumber="328" pagination="1 - 163" refId="ref17600" refString="Bulman, O. M. B., 1970. Graptolithina with sections on Entero - pneusta and Pterobranchia. In Treatise on Invertebrate Paleontology, ed. C. Teichert, pt. V, (2 nd edition), i - xxxii + 1 - 163." type="book chapter" year="1970">Bulman (1970)</bibRefCitation>
referred to the branching of stipes at their mid-length in his revised diagnosis of the genus. However, the original material is not quite so consistent as this, as pointed out by
<bibRefCitation author="Boueek" box="[1039,1200,1241,1264]" firstAuthor="Boueek" pageId="23" pageNumber="328" pagination="1 - 294" refId="ref17380" refString="Boueek, B., 1957. The dendroid graptolites of the Silurian of Bohemia. Ustredniho Ustavu Geologicky Rozpravy 23: 1 - 294." type="journal article" year="1957">Bouěek (1957)</bibRefCitation>
who recorded much longer stipes in some species than in the
<typeStatus box="[1250,1296,1270,1293]" pageId="23" pageNumber="328">type</typeStatus>
species. In Bouěeks species (
<taxonomicName authorityName="Boueek" authorityYear="1957" box="[994,1140,1300,1323]" class="Graptolithina" family="Cyclograptidae" genus="Cyclograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tuboidea" pageId="23" pageNumber="328" phylum="Hemichordata" rank="species" species="irregularis">
<emphasis box="[994,1140,1300,1323]" italics="true" pageId="23" pageNumber="328">C. irregularis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Boueek" authorityYear="1957" box="[1191,1367,1300,1323]" class="Graptolithina" family="Cyclograptidae" genus="Cyclograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tuboidea" pageId="23" pageNumber="328" phylum="Hemichordata" rank="species" species="multithecatus">
<emphasis box="[1191,1367,1300,1323]" italics="true" pageId="23" pageNumber="328">C. multithecatus</emphasis>
</taxonomicName>
) and
<taxonomicName authority="(Goppert, 1860)" baseAuthorityName="Goppert" baseAuthorityYear="1860" box="[794,1126,1329,1352]" class="Graptolithina" family="Cyclograptidae" genus="Cyclograptus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tuboidea" pageId="23" pageNumber="328" phylum="Hemichordata" rank="species" species="scharyanus">
<emphasis box="[794,947,1329,1352]" italics="true" pageId="23" pageNumber="328">C. scharyanus</emphasis>
(
<bibRefCitation author="Goppert" box="[960,1118,1329,1352]" firstAuthor="Goppert" pageId="23" pageNumber="328" pagination="426 - 606" refId="ref17922" refString="Goppert, H. R., 1860. Ueber die fossile Flora der Silurischen, der Devonischen und unteren Kohlen Formation oder des sogenannten Uebergangs Gebirges. Nova Acta Academiae Caesarae Lepoldino Carolinae Germanicae Naturae Curio - sorum 27: 426 - 606." type="journal article" year="1860">Göppert, 1860</bibRefCitation>
)
</taxonomicName>
the stipes divide at roughly mid-length, but with additional bifurcations occurring.
<bibRefCitation author="Boueek" box="[794,949,1387,1411]" firstAuthor="Boueek" pageId="23" pageNumber="328" pagination="1 - 294" refId="ref17380" refString="Boueek, B., 1957. The dendroid graptolites of the Silurian of Bohemia. Ustredniho Ustavu Geologicky Rozpravy 23: 1 - 294." type="journal article" year="1957">Bouěek (1957</bibRefCitation>
: p. 141) also questioned whether, given such long stipes, all the autothecae must originate in the thecorhiza: it seems unlikely.
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