treatments-xml/data/03/9E/87/039E8793F333FFAE6ACBF996FAB6792A.xml

<document id="E39DFB9F8E1CB399B020435F4DEE94E1" ID-CLB-Dataset="7747" ID-DOI="10.11646/zootaxa.4938.4.5" ID-GBIF-Dataset="c9ee5fe0-60c0-4b08-9dc2-3467b51a997a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4574948" ID-ZooBank="F3741DEB-5690-4BD4-9B98-18A4EA2A57CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614752491017" checkinUser="plazi" docAuthor="Sun, Jing, Jiang, Ji-Bao, Wu, Juzhen, Yuan, Zhu &amp; Qiu, Jiang-Ping" docDate="2021" docId="039E8793F333FFAE6ACBF996FAB6792A" docLanguage="en" docName="zootaxa.4938.4.5.pdf" docOrigin="Zootaxa 4938 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas yuanjiangensis Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="467" masterDocId="FFA7FFEBF334FFA46A5CFFCCFFAF7F07" masterDocTitle="Three new widely distributed and polymorphic species of Amynthas earthworms (Oligochaeta, Clitellata, Megascolecidae) from South China" masterLastPageNumber="474" masterPageNumber="457" pageNumber="464" updateTime="1698931910685" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="96047C1FC050863A6D9526CE90C5B4DA">Three new widely distributed and polymorphic species of Amynthas earthworms (Oligochaeta, Clitellata, Megascolecidae) from South China</mods:title>
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<mods:namePart id="229CA1E2B7500F4E465C1D5FB3559A84">Sun, Jing</mods:namePart>
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<mods:affiliation id="02C8FB4797A0BA42B3C54D670B24AA7E">School of Materials and Environmental Engineering, Chengdu Technological University, Chengdu, 610031, China. &amp; sun _ jing. 2005 @ aliyun. com; https: // orcid. org / 0000 - 0003 - 1326 - 1886</mods:affiliation>
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<mods:namePart id="654C25492B86EDC6C1D173011D49F05E">Jiang, Ji-Bao</mods:namePart>
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<mods:affiliation id="D04F11925967E08C1CC0F0CDB7D7C414">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, 200240, China &amp; jibao @ sjtu. edu. cn; https: // orcid. org / 0000 - 0002 - 5137 - 6097</mods:affiliation>
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<mods:namePart id="CD01827D9E2DF4D67CE39B4F4C1F252E">Wu, Juzhen</mods:namePart>
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<mods:affiliation id="1EE52BC7766F5E6ACB0A348FC5497F06">School of Materials and Environmental Engineering, Chengdu Technological University, Chengdu, 610031, China. &amp; 403141272 @ qq. com; https: // orcid. org / 0000 - 0001 - 7002 - 8023</mods:affiliation>
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<mods:namePart id="86642FDCC97C8D7C16B677ED0C82A0E3">Yuan, Zhu</mods:namePart>
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<mods:affiliation id="5D95B4312EDEF8A196E1D252E73080E0">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, 200240, China &amp; zhuyuan 0452 @ jibao @ sjtu. edu. cn; https: // orcid. org / 0000 - 0003 - 2631 - 6495</mods:affiliation>
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<mods:namePart id="5B874DAF9F763654D7A01422492D2491">Qiu, Jiang-Ping</mods:namePart>
<mods:affiliation id="716BF711B19E53C9762EF05CE3978CD7">School of Agriculture and Biology, Shanghai Jiao Tong University, Shanghai, 200240, China</mods:affiliation>
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<taxonomicName id="4C374D06F333FFA36ACBF996FDCF7973" authority="Sun &amp; Qiu" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[151,608,1626,1653]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="7" pageNumber="464" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F333FFA36ACBF996FE7F7973" bold="true" box="[151,464,1626,1653]" italics="true" pageId="7" pageNumber="464">Amynthas yuanjiangensis</emphasis>
Sun &amp; Qiu
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, 
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(
<figureCitation id="130C2A00F333FFA36AFCF9B2FF4C799F" box="[160,227,1662,1688]" captionStart="FIGURE 4" captionStartId="8.[151,250,1925,1949]" captionTargetBox="[151,1429,203,1899]" captionTargetId="figure-21@8.[151,1436,179,1900]" captionTargetPageId="8" captionText="FIGURE 4. Amynthas yuanjiangensis Sun &amp; Qiu sp. nov., holotype. (a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="7" pageNumber="464">Fig. 4</figureCitation>
)
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<paragraph id="8B883685F333FFA36ACBF90AFC0E782F" blockId="7.[151,1437,1734,2013]" pageId="7" pageNumber="464">
<emphasis id="B943EA97F333FFA36ACBF90AFEEB79E7" bold="true" box="[151,324,1734,1760]" pageId="7" pageNumber="464">Type material.</emphasis>
<materialsCitation id="3B5F3CD8F333FFA36B17F90AFC0E782F" ID-GBIF-Occurrence="3053067335" collectingDate="2011-07-16" collectorName="J. B. Jiang &amp; J. Sun &amp; X. D. Lei &amp; H. W. Feng" country="China" elevation="2096" latitude="23.664999" location="Yuanjiang National Nature Reserve" longLatPrecision="21" longitude="101.77473" municipality="One" pageId="7" pageNumber="464" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B943EA97F333FFA36B17F90AFE1D79E7" box="[331,434,1734,1760]" italics="true" pageId="7" pageNumber="464">
<typeStatus id="548C8827F333FFA36B17F90AFE1D79E7" box="[331,434,1734,1760]" pageId="7" pageNumber="464" type="holotype">Holotype</typeStatus>
</emphasis>
: 
<collectingMunicipality id="6BECACFFF333FFA36B9CF90AFE4079E7" box="[448,495,1734,1760]" pageId="7" pageNumber="464">One</collectingMunicipality>
clitellate specimen (C-YN201102-01), 
<collectingCountry id="F3207615F333FFA369F5F90AFC4079E7" box="[937,1007,1734,1760]" name="China" pageId="7" pageNumber="464">China</collectingCountry>
, 
<collectingRegion id="49F3F867F333FFA369A4F90BFB1579E7" box="[1016,1210,1734,1761]" country="China" name="Yunnan" pageId="7" pageNumber="464">Yunnan Province</collectingRegion>
, 
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(
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, 
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), 
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asl
</elevation>
, by the side of a ditch, under chestnut tree, yellow cinnamon soil; 
<date id="FF891045F333FFA36A92F8C2FECE782F" box="[206,353,1806,1832]" pageId="7" pageNumber="464" value="2011-07-16">
<collectingDate id="EFCDE9ADF333FFA36A92F8C2FECE782F" box="[206,353,1806,1832]" pageId="7" pageNumber="464" value="2011-07-16">Jul. 16, 2011</collectingDate>
</date>
; 
<collectorName id="26C25353F333FFA36B31F8C3FE4B782F" box="[365,484,1806,1833]" pageId="7" pageNumber="464">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F333FFA36BB3F8C3FD99782F" box="[495,566,1806,1833]" pageId="7" pageNumber="464">J. Sun</collectorName>
, 
<collectorName id="26C25353F333FFA3681EF8C3FD01782F" box="[578,686,1806,1833]" pageId="7" pageNumber="464">X. D. Lei</collectorName>
, and 
<collectorName id="26C25353F333FFA368B5F8C3FCC6782E" box="[745,873,1807,1833]" pageId="7" pageNumber="464">H. W. Feng</collectorName>
coll.
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</paragraph>
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<materialsCitation id="3B5F3CD8F333FFA36A9BF8FFFD067893" ID-GBIF-Occurrence="3053067336" collectingDate="2017-08-02" collectorName="One" country="China" elevation="1451" latitude="27.205444" location="Yulong Snow Mountain Provincial Nature Reserve" longLatPrecision="1" longitude="100.27992" pageId="7" pageNumber="464" specimenCount="55" stateProvince="Yunnan" typeStatus="paratype">
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<typeStatus id="548C8827F333FFA36A9BF8FFFE96784B" box="[199,313,1843,1868]" pageId="7" pageNumber="464" type="paratype">Paratypes</typeStatus>
</emphasis>
: A total of 
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, as follows: 
<collectorName id="26C25353F333FFA368A5F8FEFC87784B" box="[761,808,1842,1868]" pageId="7" pageNumber="464">One</collectorName>
clitellate (C-YN201703-01), 
<collectingCountry id="F3207615F333FFA36E22F8FEFB6B784B" box="[1150,1220,1842,1868]" name="China" pageId="7" pageNumber="464">China</collectingCountry>
, 
<collectingRegion id="49F3F867F333FFA36E8EF8FFFA37784B" box="[1234,1432,1842,1869]" country="China" name="Yunnan" pageId="7" pageNumber="464">Yunnan Province</collectingRegion>
, 
<location id="8EE8605EF333FFA36ACBF89AFD737876" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF333FFA36ACBF89AFD737876" box="[151,732,1878,1905]" country="China" latitude="27.205444" longLatPrecision="1" longitude="100.27992" name="Yulong Snow Mountain Provincial Nature Reserve" pageId="7" pageNumber="464" stateProvince="Yunnan">Yulong Snow Mountain Provincial Nature Reserve</location>
(
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, 
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), 
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<quantity id="4CCF9B60F333FFA36E08F89AFB037876" box="[1108,1196,1878,1905]" metricMagnitude="3" metricUnit="m" metricValue="1.451" pageId="7" pageNumber="464" unit="m" value="1451.0">1451 m</quantity>
asl
</elevation>
; 
<date id="FF891045F333FFA36EBFF89AFAD37877" box="[1251,1404,1878,1904]" pageId="7" pageNumber="464" value="2017-08-02">
<collectingDate id="EFCDE9ADF333FFA36EBFF89AFAD37877" box="[1251,1404,1878,1904]" pageId="7" pageNumber="464" value="2017-08-02">Aug. 2, 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F333FFA36FD6F89BFF577893" pageId="7" pageNumber="464">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F333FFA36B59F8B7FEC87892" box="[261,359,1915,1941]" pageId="7" pageNumber="464">Y. Dong</collectorName>
, 
<collectorName id="26C25353F333FFA36B28F8B6FE7A7893" box="[372,469,1914,1940]" pageId="7" pageNumber="464">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F333FFA368EEF8B6FE3078DB" ID-GBIF-Occurrence="3053067306" collectingDate="2017-08-05" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Nine" country="China" elevation="2158" latitude="24.773027" location="Wuliangshan Nature Reserve" longLatPrecision="1" longitude="100.52011" municipality="Dali City" pageId="7" pageNumber="464" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F333FFA368EEF8B6FD477893" box="[690,744,1914,1940]" pageId="7" pageNumber="464">Nine</collectorName>
clitellates (C-YN201713-06), 
<collectingCountry id="F3207615F333FFA36E1BF8B6FB227893" box="[1095,1165,1914,1940]" name="China" pageId="7" pageNumber="464">China</collectingCountry>
, 
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, 
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(
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, 
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), 
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<quantity id="4CCF9B60F333FFA36962F852FC3878BE" box="[830,919,1950,1977]" metricMagnitude="3" metricUnit="m" metricValue="2.158" pageId="7" pageNumber="464" unit="m" value="2158.0">2158 m</quantity>
asl
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; 
<date id="FF891045F333FFA3698EF852FBDF78BF" box="[978,1136,1950,1977]" pageId="7" pageNumber="464" value="2017-08-05">
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</date>
; 
<collectorName id="26C25353F333FFA36EDCF853FAAF78BF" box="[1152,1280,1950,1977]" pageId="7" pageNumber="464">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F333FFA36F52F853FADD78BE" box="[1294,1394,1951,1977]" pageId="7" pageNumber="464">Y. Dong</collectorName>
, 
<collectorName id="26C25353F333FFA36FDDF852FF7C78DB" pageId="7" pageNumber="464">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F333FFAD6BF9F80EFEA47FD2" ID-GBIF-Occurrence="3053067323" collectingDate="2017-08-06" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Nine" country="China" elevation="2364" lastPageId="9" lastPageNumber="466" latitude="25.781973" location="Cangshan National Nature Reserve" longLatPrecision="1" longitude="100.09095" municipality="Dali City" pageId="7" pageNumber="464" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F333FFA36BF9F80EFE7478DB" box="[421,475,1986,2012]" pageId="7" pageNumber="464">Nine</collectorName>
clitellates (C-YN201719-01), 
<collectingCountry id="F3207615F333FFA36968F80EFCD578DB" box="[820,890,1986,2012]" name="China" pageId="7" pageNumber="464">China</collectingCountry>
, 
<collectingRegion id="49F3F867F333FFA369D8F80FFBE878DB" box="[900,1095,1986,2013]" country="China" name="Yunnan" pageId="7" pageNumber="464">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF333FFA36E0EF80EFB1478DB" box="[1106,1211,1986,2012]" pageId="7" pageNumber="464">Dali City</collectingMunicipality>
, 
<location id="8EE8605EF333FFAD6E9AF80EFEE97FB6" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF333FFAD6E9AF80EFEE97FB6" country="China" lastPageId="9" lastPageNumber="466" latitude="25.781973" longLatPrecision="1" longitude="100.09095" municipality="Dali City" name="Cangshan National Nature Reserve" pageId="7" pageNumber="464" stateProvince="Yunnan">Cangshan National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6B0BFF54FE5B7FB4" box="[343,500,152,179]" degrees="25" direction="north" minutes="46" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="55.1" value="25.781973">25°46ʹ55.1ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD685DFF54FD077FB4" box="[513,680,152,179]" degrees="100" direction="east" minutes="05" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="27.4" value="100.09095">100°05ʹ27.4ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD68E1FF5BFC907FB6" box="[701,831,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.364" pageId="9" pageNumber="466" unit="m" value="2364.0">
<quantity id="4CCF9B60F33DFFAD68E1FF5BFCBB7FB6" box="[701,788,151,177]" metricMagnitude="3" metricUnit="m" metricValue="2.364" pageId="9" pageNumber="466" unit="m" value="2364.0">2364 m</quantity>
asl
</elevation>
; 
<date id="FF891045F33DFFAD6917FF5BFC4C7FB6" box="[843,995,151,177]" pageId="9" pageNumber="466" value="2017-08-06">
<collectingDate id="EFCDE9ADF33DFFAD6917FF5BFC4C7FB6" box="[843,995,151,177]" pageId="9" pageNumber="466" value="2017-08-06">Aug. 6, 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F33DFFAD69ADFF5BFBC37FB6" box="[1009,1132,151,177]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6E24FF5BFB767FB6" box="[1144,1241,151,177]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, 
<collectorName id="26C25353F33DFFAD6EBAFF5BFAE97FB6" box="[1254,1350,151,177]" pageId="9" pageNumber="466">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6B48FF77FF667E1A" ID-GBIF-Occurrence="3053067317" collectingDate="2017-08-07" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Four" country="China" county="Dehong Dai" elevation="2621" latitude="25.861694" location="Yunlong Tianchi National Nature Reserve" longLatPrecision="1" longitude="99.28352" municipality="Dali City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6B48FF77FEE67FD2" box="[276,329,187,213]" pageId="9" pageNumber="466">Four</collectorName>
clitellates (C-YN201722-01), 
<collectingCountry id="F3207615F33DFFAD68FBFF77FD427FD2" box="[679,749,187,213]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD68A6FF77FC117FD2" box="[762,958,187,213]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD6990FF77FB987FD2" box="[972,1079,187,213]" pageId="9" pageNumber="466">Dali City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6E1FFF77FE8B7FFE" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6E1FFF77FE8B7FFE" country="China" latitude="25.861694" longLatPrecision="1" longitude="99.28352" municipality="Dali City" name="Yunlong Tianchi National Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Yunlong Tianchi National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6B68FF2CFE7E7FFC" box="[308,465,224,251]" degrees="25" direction="north" minutes="51" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="42.1" value="25.861694">25°51ʹ42.1ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD6B81FF2CFDD87FFC" box="[477,631,224,251]" degrees="99" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="00.7" value="99.28352">99°17ʹ00.7ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD68D7FF13FCA57FFE" box="[651,778,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.621" pageId="9" pageNumber="466" unit="m" value="2621.0">
<quantity id="4CCF9B60F33DFFAD68D7FF13FD4E7FFE" box="[651,737,223,249]" metricMagnitude="3" metricUnit="m" metricValue="2.621" pageId="9" pageNumber="466" unit="m" value="2621.0">2621 m</quantity>
asl
</elevation>
; 
<date id="FF891045F33DFFAD6949FF13FC037FFE" box="[789,940,223,249]" pageId="9" pageNumber="466" value="2017-08-07">
<collectingDate id="EFCDE9ADF33DFFAD6949FF13FC037FFE" box="[789,940,223,249]" pageId="9" pageNumber="466" value="2017-08-07">Aug. 7, 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F33DFFAD69E4FF13FB9E7FFE" box="[952,1073,223,249]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6E60FF13FB327FFE" box="[1084,1181,223,249]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, 
<collectorName id="26C25353F33DFFAD6EF5FF13FAA87FFE" box="[1193,1287,223,249]" pageId="9" pageNumber="466">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6A93FECFFDB77E62" ID-GBIF-Occurrence="3053067320" collectingDate="2017-08-11" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Two" country="China" county="Dehong Dai" elevation="957" latitude="24.148972" location="Tongbiguan Provincial Nature Reserve" longLatPrecision="1" longitude="98.02528" municipality="Jingpo Autonomous Prefecture" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6A93FECFFEAF7E1A" box="[207,256,259,285]" pageId="9" pageNumber="466">Two</collectorName>
clitellates (C-YN201735-08), 
<collectingCountry id="F3207615F33DFFAD6806FECFFD0F7E1A" box="[602,672,259,285]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD68F6FECFFCC27E1A" box="[682,877,259,285]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingCounty id="62E94E09F33DFFAD6925FECFFBAE7E1A" box="[889,1025,259,285]" pageId="9" pageNumber="466">Dehong Dai</collectingCounty>
and 
<collectingMunicipality id="6BECACFFF33DFFAD6E65FECFFA377E1A" box="[1081,1432,259,285]" pageId="9" pageNumber="466">Jingpo Autonomous Prefecture</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6ACBFEEBFDFB7E46" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6ACBFEEBFDFB7E46" box="[151,596,295,321]" country="China" county="Dehong Dai" latitude="24.148972" longLatPrecision="1" longitude="98.02528" municipality="Jingpo Autonomous Prefecture" name="Tongbiguan Provincial Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Tongbiguan Provincial Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD683AFEE4FCAC7E44" box="[614,771,296,323]" degrees="24" direction="north" minutes="08" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="56.3" value="24.148972">24°08ʹ56.3ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD694DFEE4FC047E44" box="[785,939,296,323]" degrees="98" direction="east" minutes="01" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="31.0" value="98.02528">98°01ʹ31.0ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD699DFEEAFB997E46" box="[961,1078,294,321]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="9" pageNumber="466" unit="m" value="957.0">
<quantity id="4CCF9B60F33DFFAD699DFEEAFBA57E46" box="[961,1034,294,321]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="9" pageNumber="466" unit="m" value="957.0">957 m</quantity>
asl
</elevation>
; 
<date id="FF891045F33DFFAD6E1FFEEBFB447E46" box="[1091,1259,295,321]" pageId="9" pageNumber="466" value="2017-08-11">
<collectingDate id="EFCDE9ADF33DFFAD6E1FFEEBFB447E46" box="[1091,1259,295,321]" pageId="9" pageNumber="466" value="2017-08-11">Aug. 11, 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F33DFFAD6EA6FEEBFAD77E46" box="[1274,1400,295,321]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6FD9FEEBFF777E62" pageId="9" pageNumber="466">Y. Dong</collectorName>
, 
<collectorName id="26C25353F33DFFAD6AB9FE87FEEA7E62" box="[229,325,331,357]" pageId="9" pageNumber="466">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6843FE87FEA37EAA" ID-GBIF-Occurrence="3053067324" collectingDate="2017-08-14" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Two" country="China" elevation="2122" latitude="25.15086" location="Yongdedaxueshan Nature Reserve" longLatPrecision="1" longitude="99.70872" municipality="Lincang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6843FE87FDFF7E62" box="[543,592,331,357]" pageId="9" pageNumber="466">Two</collectorName>
clitellates (C-YN201749-01), 
<collectingCountry id="F3207615F33DFFAD69F2FE87FC5B7E62" box="[942,1012,331,357]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD69A3FE87FB6B7E62" box="[1023,1220,331,357]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<location id="8EE8605EF33DFFAD6E8CFE87FEE97E8E" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6E8CFE87FEE97E8E" country="China" latitude="25.15086" longLatPrecision="1" longitude="99.70872" name="Yongdedaxueshan Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Yongdedaxueshan Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6B04FEBCFE5A7E8C" box="[344,501,368,395]" degrees="25" direction="north" minutes="09" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="03.1" value="25.15086">25°09ʹ03.1ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD685EFEBCFD337E8C" box="[514,668,368,395]" degrees="99" direction="east" minutes="42" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="31.4" value="99.70872">99°42ʹ31.4ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD68EDFEA3FC9C7E8E" box="[689,819,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.122" pageId="9" pageNumber="466" unit="m" value="2122.0">
<quantity id="4CCF9B60F33DFFAD68EDFEA3FCA77E8E" box="[689,776,367,393]" metricMagnitude="3" metricUnit="m" metricValue="2.122" pageId="9" pageNumber="466" unit="m" value="2122.0">2122 m</quantity>
asl
</elevation>
, 
<date id="FF891045F33DFFAD691CFEA3FC707E8E" box="[832,991,367,393]" pageId="9" pageNumber="466" value="2017-08-14">
<collectingDate id="EFCDE9ADF33DFFAD691CFEA3FC707E8E" box="[832,991,367,393]" pageId="9" pageNumber="466" value="2017-08-14">14 Aug. 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F33DFFAD69B1FEA3FBC67E8E" box="[1005,1129,367,393]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6E29FEA3FB787E8E" box="[1141,1239,367,393]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, 
<collectorName id="26C25353F33DFFAD6EB8FEA3FAEA7E8E" box="[1252,1349,367,393]" pageId="9" pageNumber="466">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6B4AFE5FFF667EF2" ID-GBIF-Occurrence="3053067304" collectingDate="2017-08-16" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; One" country="China" elevation="1429" latitude="23.649529" location="Nangunhe National Nature Reserve" longLatPrecision="1" longitude="99.342476" municipality="Lincang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6B4AFE5FFEEA7EAA" box="[278,325,403,429]" pageId="9" pageNumber="466">One</collectorName>
clitellate (C-YN201756-04), 
<collectingCountry id="F3207615F33DFFAD68C7FE5FFD4E7EAA" box="[667,737,403,429]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD68B2FE5FFC1B7EAA" box="[750,948,403,429]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD699FFE5FFBF57EAA" box="[963,1114,403,429]" pageId="9" pageNumber="466">Lincang City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6E34FE5FFF5E7ED6" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6E34FE5FFF5E7ED6" country="China" latitude="23.649529" longLatPrecision="1" longitude="99.342476" municipality="Lincang City" name="Nangunhe National Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Nangunhe National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6B5FFE74FE0F7ED4" box="[259,416,440,467]" degrees="23" direction="north" minutes="38" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="58.3" value="23.649529">23°38ʹ58.3ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD6BF2FE74FDE77ED4" box="[430,584,440,467]" degrees="99" direction="east" minutes="20" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="32.9" value="99.342476">99°20ʹ32.9ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6802FE7BFD4E7ED6" box="[606,737,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.429" pageId="9" pageNumber="466" unit="m" value="1429.0">
<quantity id="4CCF9B60F33DFFAD6802FE7BFD197ED6" box="[606,694,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.429" pageId="9" pageNumber="466" unit="m" value="1429.0">1429 m</quantity>
asl
</elevation>
; 
<date id="FF891045F33DFFAD68B2FE7BFC387ED6" box="[750,919,439,465]" pageId="9" pageNumber="466" value="2017-08-16">
<collectingDate id="EFCDE9ADF33DFFAD68B2FE7BFC387ED6" box="[750,919,439,465]" pageId="9" pageNumber="466" value="2017-08-16">Aug. 16, 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F33DFFAD69FAFE7BFB8C7ED6" box="[934,1059,439,465]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6E6CFE7BFB3D7ED6" box="[1072,1170,439,465]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, 
<collectorName id="26C25353F33DFFAD6EFCFE7BFAAE7ED6" box="[1184,1281,439,465]" pageId="9" pageNumber="466">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6A92FE17FAF17D1E" ID-GBIF-Occurrence="3053067303" collectingDate="2017-08-17" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Two" country="China" county="Jingdong Yi Autonomous County" elevation="2129" latitude="23.315248" location="Nangunhe National Nature Reserve" longLatPrecision="1" longitude="99.220665" municipality="Lincang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6A92FE17FF507EF2" box="[206,255,475,501]" pageId="9" pageNumber="466">Two</collectorName>
clitellates (C-YN201762-05), 
<collectingCountry id="F3207615F33DFFAD6809FE17FD347EF2" box="[597,667,475,501]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD68F8FE17FCC97EF2" box="[676,870,475,501]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD692DFE17FBAB7EF2" box="[881,1028,475,501]" pageId="9" pageNumber="466">Lincang City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6E53FE17FA337EF2" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6E53FE17FA337EF2" box="[1039,1436,475,501]" country="China" latitude="23.315248" longLatPrecision="1" longitude="99.220665" municipality="Lincang City" name="Nangunhe National Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Nangunhe National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6AC3FDCCFE937D1C" box="[159,316,512,539]" degrees="23" direction="north" minutes="18" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="54.9" value="23.315248">23°18ʹ54.9ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD6B15FDCCFE4D7D1C" box="[329,482,512,539]" degrees="99" direction="east" minutes="13" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="14.4" value="99.220665">99°13ʹ14.4ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6BABFE33FDD77D1E" box="[503,632,511,537]" metricMagnitude="3" metricUnit="m" metricValue="2.129" pageId="9" pageNumber="466" unit="m" value="2129.0">
<quantity id="4CCF9B60F33DFFAD6BABFE33FDE17D1E" box="[503,590,511,537]" metricMagnitude="3" metricUnit="m" metricValue="2.129" pageId="9" pageNumber="466" unit="m" value="2129.0">2129 m</quantity>
asl
</elevation>
; 
<date id="FF891045F33DFFAD68D8FE33FC847D1E" box="[644,811,511,537]" pageId="9" pageNumber="466" value="2017-08-17">
<collectingDate id="EFCDE9ADF33DFFAD68D8FE33FC847D1E" box="[644,811,511,537]" pageId="9" pageNumber="466" value="2017-08-17">Aug. 17, 2017</collectingDate>
</date>
; 
<collectorName id="26C25353F33DFFAD6964FE33FC1C7D1E" box="[824,947,511,537]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD69E3FE33FB8F7D1E" box="[959,1056,511,537]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, 
<collectorName id="26C25353F33DFFAD6E71FE33FB237D1E" box="[1069,1164,511,537]" pageId="9" pageNumber="466">Q. Zhao</collectorName>
, and Z. Yuan coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6F3AFE33FECF7D82" ID-GBIF-Occurrence="3053067331" collectingDate="2016-07-30" collectorName="J. B. Jiang &amp; Y. Dong &amp; Q. Zhao &amp; Nine" country="China" county="Jingdong Yi Autonomous County" elevation="1860" latitude="24.269306" location="Pu'er City" longLatPrecision="1" longitude="100.76036" municipality="Lijiang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6F3AFE33FA337D1E" box="[1382,1436,511,537]" pageId="9" pageNumber="466">Nine</collectorName>
clitellates and four aclitellates (C-YN201627-02), 
<collectingCountry id="F3207615F33DFFAD6889FDEFFCB47D3A" box="[725,795,547,573]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD697AFDEFFC457D3A" box="[806,1002,547,573]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<location id="8EE8605EF33DFFAD69A4FDEFFBC17D3A" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD69A4FDEFFBC17D3A" box="[1016,1134,547,573]" country="China" county="Jingdong Yi Autonomous County" latitude="24.269306" longLatPrecision="1" longitude="100.76036" name="Pu'er City" pageId="9" pageNumber="466" stateProvince="Yunnan">Pu’er City</location>
, 
<collectingCounty id="62E94E09F33DFFAD6E27FDEFFF467D66" pageId="9" pageNumber="466">Jingdong Yi Autonomous County</collectingCounty>
(
<geoCoordinate id="EE035042F33DFFAD6AA5FD84FE327D66" box="[249,413,583,611]" degrees="24" direction="north" minutes="16" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="09.5" value="24.269306">24°16ʹ09.5ʺ N</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD6BF5FD84FDF87D66" box="[425,599,583,611]" degrees="100" direction="east" minutes="45" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="37.3" value="100.76036">100°45ʹ37.3ʺ E</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6830FD8BFD437D66" box="[620,748,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="9" pageNumber="466" unit="m" value="1860.0">
<quantity id="4CCF9B60F33DFFAD6830FD8BFD6D7D66" box="[620,706,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="9" pageNumber="466" unit="m" value="1860.0">1860 m</quantity>
asl
</elevation>
, in black sandy soil; 
<date id="FF891045F33DFFAD6987FD8BFBDD7D66" box="[987,1138,583,609]" pageId="9" pageNumber="466" value="2016-07-30">
<collectingDate id="EFCDE9ADF33DFFAD6987FD8BFBDD7D66" box="[987,1138,583,609]" pageId="9" pageNumber="466" value="2016-07-30">Jul. 30, 2016</collectingDate>
</date>
; X. Gao, Y. F. Lu, J. Z. Jiang, 
<collectorName id="26C25353F33DFFAD6A8CFDA7FE8A7D82" box="[208,293,619,645]" pageId="9" pageNumber="466">J. Long</collectorName>
coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6B34FDA7FD437DCA" ID-GBIF-Occurrence="3053067319" collectingDate="2017-08-02" collectorName="J. Long" country="China" elevation="3140" latitude="27.202002" location="Yulong Snow Mountain Provincial Nature Reserve" longLatPrecision="1" longitude="100.28109" municipality="Lijiang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
Two clitellates and three aclitellates (C-YN201703-01), 
<collectingCountry id="F3207615F33DFFAD69B5FDA7FB807D82" box="[1001,1071,619,645]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD6E67FDA7FAAF7D82" box="[1083,1280,619,645]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD6F51FDA7FA377D82" box="[1293,1432,619,645]" pageId="9" pageNumber="466">Lijiang City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6ACBFD43FD637DAE" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6ACBFD43FD637DAE" box="[151,716,655,681]" country="China" latitude="27.202002" longLatPrecision="1" longitude="100.28109" municipality="Lijiang City" name="Yulong Snow Mountain Provincial Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Yulong Snow Mountain Provincial Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6887FD5CFCD77DAC" box="[731,888,656,683]" degrees="27" direction="north" minutes="12" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="07.2" value="27.202002">27°12ʹ07.2ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD69DEFD5CFB857DAC" box="[898,1066,656,683]" degrees="100" direction="east" minutes="16" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="51.9" value="100.28109">100°16ʹ51.9ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6E60FD43FB177DAE" box="[1084,1208,655,681]" metricMagnitude="3" metricUnit="m" metricValue="3.1399999999999997" pageId="9" pageNumber="466" unit="m" value="3140.0">
<quantity id="4CCF9B60F33DFFAD6E60FD43FB3F7DAE" box="[1084,1168,655,681]" metricMagnitude="3" metricUnit="m" metricValue="3.1399999999999997" pageId="9" pageNumber="466" unit="m" value="3140.0">3140 m</quantity>
asl
</elevation>
, in yellow soil; 
<date id="FF891045F33DFFAD6F39FD43FF407DCA" pageId="9" pageNumber="466" value="2017-08-02">
<collectingDate id="EFCDE9ADF33DFFAD6F39FD43FF407DCA" pageId="9" pageNumber="466" value="2017-08-02">Aug. 2, 2017</collectingDate>
</date>
; Z. 
<collectorName id="26C25353F33DFFAD6B47FD7FFEF77DCA" box="[283,344,691,717]" pageId="9" pageNumber="466">Yuan</collectorName>
, 
<collectorName id="26C25353F33DFFAD6B38FD7FFE727DCA" box="[356,477,691,717]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6BB5FD7FFDE67DCA" box="[489,585,691,717]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, Q. Zhao coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD68AFFD7FFD417C12" ID-GBIF-Occurrence="3053067341" collectingDate="2017-08-07" collectorName="Yuan &amp; J. B. Jiang &amp; Y. Dong &amp; Four" country="China" elevation="2621" latitude="25.858528" location="Yunlong Tianchi National Nature Reserve" longLatPrecision="1" longitude="99.28436" municipality="Dali City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD68AFFD7FFC877DCA" box="[755,808,691,717]" pageId="9" pageNumber="466">Four</collectorName>
clitellates (C-YN201722-01), 
<collectingCountry id="F3207615F33DFFAD6EDFFD7FFB667DCA" box="[1155,1225,691,717]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD6E88FD7FFA377DCA" box="[1236,1432,691,717]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD6ACBFD1BFF507DF6" box="[151,255,727,753]" pageId="9" pageNumber="466">Dali City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6B55FD1BFD737DF6" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6B55FD1BFD737DF6" box="[265,732,727,753]" country="China" latitude="25.858528" longLatPrecision="1" longitude="99.28436" municipality="Dali City" name="Yunlong Tianchi National Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Yunlong Tianchi National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD68B7FD14FC287DF4" box="[747,903,728,755]" degrees="25" direction="north" minutes="51" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="30.7" value="25.858528">25°51ʹ30.7ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD69CEFD14FB837DF4" box="[914,1068,728,755]" degrees="99" direction="east" minutes="17" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="03.7" value="99.28436">99°17ʹ03.7ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6E63FD1BFB147DF6" box="[1087,1211,727,753]" metricMagnitude="3" metricUnit="m" metricValue="2.621" pageId="9" pageNumber="466" unit="m" value="2621.0">
<quantity id="4CCF9B60F33DFFAD6E63FD1BFB3C7DF6" box="[1087,1171,727,753]" metricMagnitude="3" metricUnit="m" metricValue="2.621" pageId="9" pageNumber="466" unit="m" value="2621.0">2621 m</quantity>
asl
</elevation>
, in brown soil; 
<date id="FF891045F33DFFAD6F39FD1BFF407C12" pageId="9" pageNumber="466" value="2017-08-07">
<collectingDate id="EFCDE9ADF33DFFAD6F39FD1BFF407C12" pageId="9" pageNumber="466" value="2017-08-07">Aug. 7, 2017</collectingDate>
</date>
; Z. 
<collectorName id="26C25353F33DFFAD6B40FD37FEF67C12" box="[284,345,763,789]" pageId="9" pageNumber="466">Yuan</collectorName>
, 
<collectorName id="26C25353F33DFFAD6B39FD37FE707C12" box="[357,479,763,789]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD6BB6FD37FDE47C12" box="[490,587,763,789]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, Q. Zhao coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD68AAFD37FCC97C5A" ID-GBIF-Occurrence="3053067340" collectingDate="2017-08-16" collectorName="Yuan &amp; J. B. Jiang &amp; Y. Dong &amp; Six" country="China" county="Dehong Dai" elevation="1429" latitude="23.646944" location="Nangunhe National Nature Reserve" longLatPrecision="1" longitude="99.343506" municipality="Lincang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD68AAFD37FCB47C12" box="[758,795,763,789]" pageId="9" pageNumber="466">Six</collectorName>
aclitellates (C-YN201756-04), 
<collectingCountry id="F3207615F33DFFAD6EDFFD37FB667C12" box="[1155,1225,763,789]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD6E88FD37FA377C12" box="[1236,1432,763,789]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD6ACBFCD3FE837C3E" box="[151,300,799,825]" pageId="9" pageNumber="466">Lincang City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6B64FCD3FD667C3E" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6B64FCD3FD667C3E" box="[312,713,799,825]" country="China" latitude="23.646944" longLatPrecision="1" longitude="99.343506" municipality="Lincang City" name="Nangunhe National Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Nangunhe National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6885FCECFCD87C3C" box="[729,887,800,827]" degrees="23" direction="north" minutes="38" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="49.0" value="23.646944">23°38ʹ49.0ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD69DFFCECFBB37C3C" box="[899,1052,800,827]" degrees="99" direction="east" minutes="20" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="36.6" value="99.343506">99°20ʹ36.6ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6E6CFCD3FB1F7C3E" box="[1072,1200,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.429" pageId="9" pageNumber="466" unit="m" value="1429.0">
<quantity id="4CCF9B60F33DFFAD6E6CFCD3FB297C3E" box="[1072,1158,799,825]" metricMagnitude="3" metricUnit="m" metricValue="1.429" pageId="9" pageNumber="466" unit="m" value="1429.0">1429 m</quantity>
asl
</elevation>
, in yellow cinnamon soil; 
<date id="FF891045F33DFFAD6A91FC8FFEDE7C5A" box="[205,369,835,861]" pageId="9" pageNumber="466" value="2017-08-16">
<collectingDate id="EFCDE9ADF33DFFAD6A91FC8FFEDE7C5A" box="[205,369,835,861]" pageId="9" pageNumber="466" value="2017-08-16">Aug. 16, 2017</collectingDate>
</date>
; Z. 
<collectorName id="26C25353F33DFFAD6BC7FC8FFE777C5A" box="[411,472,835,861]" pageId="9" pageNumber="466">Yuan</collectorName>
, 
<collectorName id="26C25353F33DFFAD6BBFFC8FFDF37C5A" box="[483,604,835,861]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD683AFC8FFD697C5A" box="[614,710,835,861]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, Q. Zhao coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6931FC8FFB5C7CA2" ID-GBIF-Occurrence="3053067333" collectingDate="2017-08-11" collectorName="Yuan &amp; J. B. Jiang &amp; Y. Dong &amp; Two" country="China" county="Dehong Dai" elevation="957" latitude="24.14614" location="Tongbiguan Provincial Nature Reserve" longLatPrecision="1" longitude="98.02564" municipality="Jingpo Autonomous Prefecture" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6931FC8FFC317C5A" box="[877,926,835,861]" pageId="9" pageNumber="466">Two</collectorName>
clitellates (C-YN201735-08), 
<collectingCountry id="F3207615F33DFFAD6EABFC8FFA927C5A" box="[1271,1341,835,861]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD6F1BFC8FFF527C86" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingCounty id="62E94E09F33DFFAD6B56FCABFE3D7C86" box="[266,402,871,897]" pageId="9" pageNumber="466">Dehong Dai</collectingCounty>
and 
<collectingMunicipality id="6BECACFFF33DFFAD6B96FCABFC857C86" box="[458,810,871,897]" pageId="9" pageNumber="466">Jingpo Autonomous Prefecture</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6969FCABFB447C86" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6969FCABFB447C86" box="[821,1259,871,897]" country="China" county="Dehong Dai" latitude="24.14614" longLatPrecision="1" longitude="98.02564" municipality="Jingpo Autonomous Prefecture" name="Tongbiguan Provincial Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Tongbiguan Provincial Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6EA7FCA4FA377C84" box="[1275,1432,872,899]" degrees="24" direction="north" minutes="08" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="46.1" value="24.14614">24°08ʹ46.1ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD6ACBFC40FE9E7CA0" box="[151,305,908,935]" degrees="98" direction="east" minutes="01" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="32.3" value="98.02564">98°01ʹ32.3ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6B18FC46FE1B7CA2" box="[324,436,906,933]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="9" pageNumber="466" unit="m" value="957.0">
<quantity id="4CCF9B60F33DFFAD6B18FC46FE247CA2" box="[324,395,906,933]" metricMagnitude="2" metricUnit="m" metricValue="9.57" pageId="9" pageNumber="466" unit="m" value="957.0">957 m</quantity>
asl
</elevation>
, in brown soil; 
<date id="FF891045F33DFFAD6803FC47FCAE7CA2" box="[607,769,907,933]" pageId="9" pageNumber="466" value="2017-08-11">
<collectingDate id="EFCDE9ADF33DFFAD6803FC47FCAE7CA2" box="[607,769,907,933]" pageId="9" pageNumber="466" value="2017-08-11">Aug. 11, 2017</collectingDate>
</date>
; Z. 
<collectorName id="26C25353F33DFFAD6977FC47FCC77CA2" box="[811,872,907,933]" pageId="9" pageNumber="466">Yuan</collectorName>
, 
<collectorName id="26C25353F33DFFAD692FFC47FC457CA2" box="[883,1002,907,933]" pageId="9" pageNumber="466">J. B. Jiang</collectorName>
, 
<collectorName id="26C25353F33DFFAD69A8FC47FBFB7CA2" box="[1012,1108,907,933]" pageId="9" pageNumber="466">Y. Dong</collectorName>
, Q. Zhao coll.
</materialsCitation>
<materialsCitation id="3B5F3CD8F33DFFAD6EA5FC47FD2A7B16" ID-GBIF-Occurrence="3053067302" collectingDate="2017-08-17" collectorName="Yuan &amp; J. B. Jiang &amp; Y. Dong &amp; Two" country="China" elevation="2129" latitude="23.312555" location="Nangunhe National Nature Reserve" longLatPrecision="1" longitude="99.221504" municipality="Lincang City" pageId="9" pageNumber="466" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
<collectorName id="26C25353F33DFFAD6EA5FC47FA857CA2" box="[1273,1322,907,933]" pageId="9" pageNumber="466">Two</collectorName>
clitellates (C-YN201762-05), 
<collectingCountry id="F3207615F33DFFAD6B26FC63FE6F7CCE" box="[378,448,943,969]" name="China" pageId="9" pageNumber="466">China</collectingCountry>
, 
<collectingRegion id="49F3F867F33DFFAD6B92FC63FD3A7CCE" box="[462,661,943,969]" country="China" name="Yunnan" pageId="9" pageNumber="466">Yunnan Province</collectingRegion>
, 
<collectingMunicipality id="6BECACFFF33DFFAD68F9FC63FC927CCE" box="[677,829,943,969]" pageId="9" pageNumber="466">Lincang City</collectingMunicipality>
, 
<location id="8EE8605EF33DFFAD6910FC63FB477CCE" LSID="urn:lsid:plazi:treatment:039E8793F333FFAE6ACBF996FAB6792A:8EE8605EF33DFFAD6910FC63FB477CCE" box="[844,1256,943,969]" country="China" latitude="23.312555" longLatPrecision="1" longitude="99.221504" municipality="Lincang City" name="Nangunhe National Nature Reserve" pageId="9" pageNumber="466" stateProvince="Yunnan">Nangunhe National Nature Reserve</location>
(
<geoCoordinate id="EE035042F33DFFAD6EA7FC7CFA377CCC" box="[1275,1432,944,971]" degrees="23" direction="north" minutes="18" orientation="latitude" pageId="9" pageNumber="466" precision="1" seconds="45.2" value="23.312555">23°18ʹ45.2ʺN</geoCoordinate>
, 
<geoCoordinate id="EE035042F33DFFAD6ACBFC18FE9F7CE8" box="[151,304,980,1007]" degrees="99" direction="east" minutes="13" orientation="longitude" pageId="9" pageNumber="466" precision="1" seconds="17.4" value="99.221504">99°13ʹ17.4ʺE</geoCoordinate>
), 
<elevation id="001AD1B6F33DFFAD6B19FC1FFE6C7CEA" box="[325,451,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.129" pageId="9" pageNumber="466" unit="m" value="2129.0">
<quantity id="4CCF9B60F33DFFAD6B19FC1FFE357CEA" box="[325,410,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.129" pageId="9" pageNumber="466" unit="m" value="2129.0">2129 m</quantity>
asl
</elevation>
, in brown soil; 
<date id="FF891045F33DFFAD682DFC1FFCBA7CEA" box="[625,789,979,1005]" pageId="9" pageNumber="466" value="2017-08-17">
<collectingDate id="EFCDE9ADF33DFFAD682DFC1FFCBA7CEA" box="[625,789,979,1005]" pageId="9" pageNumber="466" value="2017-08-17">Aug. 17, 2017</collectingDate>
</date>
; Z. Yuan, J. B. Jiang, Y. Dong, Q. Zhao coll. The habitats of the 
<typeStatus id="548C8827F33DFFAD6A83FC3BFEE57B16" box="[223,330,1015,1041]" pageId="9" pageNumber="466" type="paratype">paratypes</typeStatus>
were not recorded in detail.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF48660DF33CFFAC6ACBF849FB7678C6" ID-DOI="http://doi.org/10.5281/zenodo.4574956" ID-Zenodo-Dep="4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="8" pageNumber="465" startId="8.[151,250,1925,1949]" targetBox="[151,1429,203,1899]" targetPageId="8">
<paragraph id="8B883685F33CFFAC6ACBF849FB7678C6" blockId="8.[151,1437,1925,1986]" pageId="8" pageNumber="465">
<emphasis id="B943EA97F33CFFAC6ACBF849FCC07899" bold="true" box="[151,879,1925,1950]" pageId="8" pageNumber="465">
FIGURE 4. 
<taxonomicName id="4C374D06F33CFFAC6B47F849FD09789A" authority="Sun &amp; Qiu" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[283,678,1925,1949]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="8" pageNumber="465" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F33CFFAC6B47F849FD86789A" bold="true" box="[283,553,1925,1949]" italics="true" pageId="8" pageNumber="465">Amynthas yuanjiangensis</emphasis>
Sun &amp; Qiu
</taxonomicName>
<taxonomicNameLabel id="A27057ECF33CFFAC68F1F84AFCAE7899" box="[685,769,1926,1950]" pageId="8" pageNumber="465" rank="species">sp. nov.</taxonomicNameLabel>
, holotype.
</emphasis>
(a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C32D650EF33DFFAD6A9BFBD7FC1B7B51" pageId="9" pageNumber="466" type="etymology">
<paragraph id="8B883685F33DFFAD6A9BFBD7FC1B7B51" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="466">
<emphasis id="B943EA97F33DFFAD6A9BFBD7FEE17B32" bold="true" box="[199,334,1051,1077]" pageId="9" pageNumber="466">Etymology.</emphasis>
The name 
<taxonomicName id="4C374D06F33DFFAD6B90FBD7FDDB7B33" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[460,628,1051,1076]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="466" phylum="Annelida" rank="species" species="yuanjiangensis">
<emphasis id="B943EA97F33DFFAD6B90FBD7FDDB7B33" box="[460,628,1051,1076]" italics="true" pageId="9" pageNumber="466">yuanjiangensis</emphasis>
</taxonomicName>
is derived from the collection site (Yuanjiang National Nature Reserve) of the 
<typeStatus id="548C8827F33DFFAD6A83FBF3FEEA7B5E" box="[223,325,1087,1113]" pageId="9" pageNumber="466" type="holotype">holotype</typeStatus>
. In Chinese, 
<taxonomicName id="4C374D06F33DFFAD6B89FBF3FD5D7B5F" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[469,754,1087,1113]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="9" pageNumber="466" phylum="Annelida" rank="species" species="yuanjiangensis">
<emphasis id="B943EA97F33DFFAD6B89FBF3FD5D7B5F" box="[469,754,1087,1113]" italics="true" pageId="9" pageNumber="466">Amynthas yuanjiangensis</emphasis>
</taxonomicName>
is 
<collectingRegion id="49F3F867F33DFFAD6946FBF3FCD77B51" box="[794,888,1087,1110]" country="Vietnam" name="Ca Mau" pageId="9" pageNumber="466">Kmũ</collectingRegion>
Ḃḋ.
</paragraph>
</subSubSection>
<subSubSection id="C32D650EF33DFFAD6A9BFBAEFCBB7BEE" pageId="9" pageNumber="466" type="diagnosis">
<paragraph id="8B883685F33DFFAD6A9BFBAEFCBB7BEE" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="466">
<emphasis id="B943EA97F33DFFAD6A9BFBAEFEED7B7B" bold="true" box="[199,322,1122,1148]" pageId="9" pageNumber="466">Diagnosis.</emphasis>
Medium-sized; four pairs of spermathecal pores in 5/6–8/9; each male pore on the top of a large raised pulvinate pad; no genital papillae in the spermathecal pore and male pore regions; prostate gland partially vestigial in 1/2 XVII–1/2 XIX, or developed; spermathecal diverticulum about ¾ as long as the main spermathecal axis, distal U dilated into a rod-shaped seminal chamber.
</paragraph>
</subSubSection>
<subSubSection id="C32D650EF33DFFAE6A9BFB3EFEA67D82" lastPageId="10" lastPageNumber="467" pageId="9" pageNumber="466" type="description">
<paragraph id="8B883685F33DFFAD6A9BFB3EFA8E7956" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="466">
<emphasis id="B943EA97F33DFFAD6A9BFB3EFEF67A0B" bold="true" box="[199,345,1266,1292]" pageId="9" pageNumber="466">Description.</emphasis>
<emphasis id="B943EA97F33DFFAD6B3CFB3FFDEF7A0A" box="[352,576,1267,1293]" italics="true" pageId="9" pageNumber="466">External characters</emphasis>
: Preserved specimens yellowish-brown on dorsum, lacking pigment on ventrum. Mid-dorsal line pigmented. Dimensions 
<quantity id="4CCF9B60F33DFFAD68C4FADAFCB27A36" box="[664,797,1302,1329]" metricMagnitude="-1" metricUnit="m" metricValue="1.075" metricValueMax="1.35" metricValueMin="0.8" pageId="9" pageNumber="466" unit="mm" value="107.5" valueMax="135.0" valueMin="80.0">80–135 mm</quantity>
by 3.0–5.0 mm at clitellum; segments 82–110. Secondary annulations present in VI–XIII. Prostomium ¾ epilobous. First dorsal pore in 11/12. Clitellum annular, in XIV–XVI, gray-brown, smooth, swollen, no setae on the ventral side; dorsal pores absent but with vestiges on clitellum. Setae uniformly distributed, 22–30 at III, 30–36 at V, 32–36 at VIII, 38–48 at XX, 46–62 at XXV; 7–9/VII, 7–14/VIII between spermathecal pores, 7–12 between male pores; setal formula AA=1.0–1.2AB, ZZ=1.2–1.3ZY. Spermathecal pores four pairs in 5/6–8/9 (
<figureCitation id="130C2A00F33DFFAD6BAFFA07FDEC7AE2" box="[499,579,1483,1509]" captionStart="FIGURE 4" captionStartId="8.[151,250,1925,1949]" captionTargetBox="[151,1429,203,1899]" captionTargetId="figure-21@8.[151,1436,179,1900]" captionTargetPageId="8" captionText="FIGURE 4. Amynthas yuanjiangensis Sun &amp; Qiu sp. nov., holotype. (a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="9" pageNumber="466">Fig. 4a</figureCitation>
, sp.p), ventrally separated by slightly more than 0.33 of circumference. Male pores in XVIII, ventrally separated by 0.25 of circumference, each on top of a large raised pulvinate pad, surrounded by two skin folds medially and one rhombus-shaped skin fold laterally in the 
<typeStatus id="548C8827F33DFFAD69B7F9DFFBE2792A" box="[1003,1101,1555,1581]" pageId="9" pageNumber="466" type="holotype">holotype</typeStatus>
(
<figureCitation id="130C2A00F33DFFAD6E07F9DFFB05792A" box="[1115,1194,1555,1581]" captionStart="FIGURE 4" captionStartId="8.[151,250,1925,1949]" captionTargetBox="[151,1429,203,1899]" captionTargetId="figure-21@8.[151,1436,179,1900]" captionTargetPageId="8" captionText="FIGURE 4. Amynthas yuanjiangensis Sun &amp; Qiu sp. nov., holotype. (a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="9" pageNumber="466">Fig. 4a</figureCitation>
, mp), but no skin fold in 
<typeStatus id="548C8827F33DFFAD6AE8F9FBFE8C7956" box="[180,291,1591,1617]" pageId="9" pageNumber="466" type="paratype">paratypes</typeStatus>
. No genital papillae in male pore region. Female pore single, mid-ventral in XIV, elliptical.
</paragraph>
<paragraph id="8B883685F33DFFAE6A9BF997FBB87E61" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,1581]" lastPageId="10" lastPageNumber="467" pageId="9" pageNumber="466">
<emphasis id="B943EA97F33DFFAD6A9BF997FE0C7972" box="[199,419,1627,1653]" italics="true" pageId="9" pageNumber="466">Internal characters</emphasis>
: Septa 8/9–9/10 absent, 5/6–7/8 thickened and muscular, 10/11–14/15 thicker than those following. Dorsal blood vessel single, continuous onto pharynx; esophageal hearts 4 pairs in X–XIII, all developed. Gizzard bucket-like, in VIII–X; intestinal swelling in XV. Intestinal caeca between simple and complex, originating in XXVII and extending forward to 1/2XXIII, horn-shaped sacs; tiny incisions deeper on dorsal margin than on ventral margin in 
<typeStatus id="548C8827F33DFFAD6BD1F927FE407802" box="[397,495,1771,1797]" pageId="9" pageNumber="466" type="holotype">holotype</typeStatus>
(
<figureCitation id="130C2A00F33DFFAD685EF927FDF67801" box="[514,601,1771,1798]" captionStart="FIGURE 4" captionStartId="8.[151,250,1925,1949]" captionTargetBox="[151,1429,203,1899]" captionTargetId="figure-21@8.[151,1436,179,1900]" captionTargetPageId="8" captionText="FIGURE 4. Amynthas yuanjiangensis Sun &amp; Qiu sp. nov., holotype. (a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="9" pageNumber="466">Fig. 4b</figureCitation>
); tiny incisions only on dorsal margin in 
<typeStatus id="548C8827F33DFFAD6E60F927FB087802" box="[1084,1191,1771,1797]" pageId="9" pageNumber="466" type="paratype">paratypes</typeStatus>
C-YN201703-01, CYN201703-01, C-YN201719-01, and C-YN201735-08. Male organs: testis sacs in X and XI; oval, developed, two lobes connected ventromedially in the first pair and separated ventromedially in the second pair; two pairs of seminal vesicles in XI and XII, first pair separated ventromedially, second pair connected with a slim tube; in 
<typeStatus id="548C8827F33DFFAD6F6FF89BFA377876" box="[1331,1432,1879,1905]" pageId="9" pageNumber="466" type="holotype">holotype</typeStatus>
, left prostate gland located in 1/2 XVII–1/2 XIX and degenerated into a small and compact lobe, right prostate gland pachytic and developed (
<figureCitation id="130C2A00F33DFFAD6BF1F853FE5178BE" box="[429,510,1951,1977]" captionStart="FIGURE 4" captionStartId="8.[151,250,1925,1949]" captionTargetBox="[151,1429,203,1899]" captionTargetId="figure-21@8.[151,1436,179,1900]" captionTargetPageId="8" captionText="FIGURE 4. Amynthas yuanjiangensis Sun &amp; Qiu sp. nov., holotype. (a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="9" pageNumber="466">Fig. 4c</figureCitation>
), its duct in XVIII, U-shaped, stout ventrally; prostate glands vestigial in 
<typeStatus id="548C8827F33DFFAD6F6DF853FA3378BE" box="[1329,1436,1951,1977]" pageId="9" pageNumber="466" type="paratype">paratypes</typeStatus>
C-YN201703-01, C-YN201719-0, and C-YN201735-08, but developed in 
<typeStatus id="548C8827F33DFFAD69AEF80FFBFD78DA" box="[1010,1106,1987,2013]" pageId="9" pageNumber="466" type="paratype">paratype</typeStatus>
C-YN201713-06; no visible accessory glands on XVIII. Spermathecae paired in VI–IX, about 
<quantity id="4CCF9B60F33EFFAE692EFF5BFC687FB6" box="[882,967,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="10" pageNumber="467" unit="mm" value="2.7">2.7 mm</quantity>
long; spermathecal ampulla heart-shaped, 
<quantity id="4CCF9B60F33EFFAE6ACBFF77FF437FD2" box="[151,236,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="10" pageNumber="467" unit="mm" value="2.1">2.1 mm</quantity>
long; in 
<typeStatus id="548C8827F33EFFAE6B11FF77FE1D7FD2" box="[333,434,187,213]" pageId="10" pageNumber="467" type="holotype">holotype</typeStatus>
, diverticulum about 3/4 as long as the main spermathecal axis, distal 2/5 dilated into a rodshaped seminal chamber; in 
<typeStatus id="548C8827F33EFFAE6B88FF13FD907FFE" box="[468,575,223,249]" pageId="10" pageNumber="467" type="paratype">paratypes</typeStatus>
C-YN201703-01, C-YN201713-06, C-YN201722-01, and C-YN201735-08, diverticulum about 1/2 as long as the main spermathecal axis, distal 1/2–1/3 dilated into rod-shaped seminal chamber; in 
<typeStatus id="548C8827F33EFFAE6AB4FEEBFEE77E46" box="[232,328,295,321]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
C-YN201719-01, diverticulum about 2/5 as long as the main spermathecal axis, distal 1/3 dilated into rod-shaped seminal chamber; no nephridia on spermathecal ducts (
<figureCitation id="130C2A00F33EFFAE69EAFE87FBA47E61" box="[950,1035,331,358]" captionStart="FIGURE 4" captionStartId="8.[151,250,1925,1949]" captionTargetBox="[151,1429,203,1899]" captionTargetId="figure-21@8.[151,1436,179,1900]" captionTargetPageId="8" captionText="FIGURE 4. Amynthas yuanjiangensis Sun &amp; Qiu sp. nov., holotype. (a) Ventral view showing spermathecal pores (sp) and male pores (mp); (b) do., intestinal cecum; (c) do., prostate gland; (d) do., spermathecae. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4574956" httpUri="https://zenodo.org/record/4574956/files/figure.png" pageId="10" pageNumber="467">Fig. 4d</figureCitation>
).
</paragraph>
<paragraph id="8B883685F33EFFAE6A9BFEA2FC107E8E" blockId="10.[151,1437,151,1581]" box="[199,959,366,393]" pageId="10" pageNumber="467">
<emphasis id="B943EA97F33EFFAE6A9BFEA2FE907E8F" bold="true" box="[199,319,366,392]" pageId="10" pageNumber="467">Variation.</emphasis>
The prostate glands are either degenerated or developed.
</paragraph>
<paragraph id="8B883685F33EFFAE6A9BFE5EFEA67D82" blockId="10.[151,1437,151,1581]" pageId="10" pageNumber="467">
<emphasis id="B943EA97F33EFFAE6A9BFE5EFE2A7EAA" bold="true" box="[199,389,402,429]" pageId="10" pageNumber="467">DNA barcodes.</emphasis>
<materialsCitation id="3B5F3CD8F33EFFAE6BC9FE5FFEAA7D82" ID-GBIF-Occurrence="3053067307" accessionNumber="KF205466, MH845539, YN201703, MH845531, MH845522, MH845514, MH845487, MH845470, MH845460, MH845451, MH837679" collectionCode="MH" pageId="10" pageNumber="467" specimenCount="15" typeStatus="holotype">
GenBank accession numbers 
<accessionNumber id="9464AB66F33EFFAE68A0FE5EFCDB7EAA" box="[764,884,402,429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KF205466" pageId="10" pageNumber="467">KF205466</accessionNumber>
(C-YN201102-01, 
<typeStatus id="548C8827F33EFFAE6E3EFE5FFB677EA9" box="[1122,1224,403,430]" pageId="10" pageNumber="467" type="holotype">holotype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6EB9FE5EFAC97EAA" box="[1253,1382,402,429]" httpUri="https://www.ncbi.nlm.nih.gov/nuccore/MH845539" pageId="10" pageNumber="467" type="GenBank">
<collectionCode id="ED26AE40F33EFFAE6EB9FE5EFABD7EAA" box="[1253,1298,402,429]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845539
</accessionNumber>
(CYN201703-01, 
<typeStatus id="548C8827F33EFFAE6B13FE7BFE1B7ED6" box="[335,436,439,465]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6B93FE7AFDFF7ED6" box="[463,592,438,465]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845531" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6B93FE7AFE527ED6" box="[463,509,438,465]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845531
</accessionNumber>
(C-YN201713-06, 
<typeStatus id="548C8827F33EFFAE6961FE7BFC0D7ED6" box="[829,930,439,465]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE69E2FE7AFB907ED6" box="[958,1087,438,465]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845522" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE69E2FE7AFC447ED6" box="[958,1003,438,465]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845522
</accessionNumber>
(C-YN201719-01, 
<typeStatus id="548C8827F33EFFAE6F70FE7BFA3E7ED6" box="[1324,1425,439,465]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6ACBFE16FEB77EF2" box="[151,280,474,501]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845514" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6ACBFE16FF6B7EF2" box="[151,196,474,501]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845514
</accessionNumber>
(C-YN201722-01, 
<typeStatus id="548C8827F33EFFAE685CFE17FDCA7EF2" box="[512,613,475,501]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6821FE16FD517EF2" box="[637,766,474,501]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845487" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6821FE16FD057EF2" box="[637,682,474,501]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845487
</accessionNumber>
(C-YN201735-08, 
<typeStatus id="548C8827F33EFFAE69BAFE17FBE47EF2" box="[998,1099,475,501]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6E3FFE16FB4B7EF2" box="[1123,1252,474,501]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845470" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6E3FFE16FB3F7EF2" box="[1123,1168,474,501]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845470
</accessionNumber>
(C-YN201749- 01, 
<typeStatus id="548C8827F33EFFAE6A94FE33FE827D1E" box="[200,301,511,537]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6B1BFE32FE677D1E" box="[327,456,510,537]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845460" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6B1BFE32FEDB7D1E" box="[327,372,510,537]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845460
</accessionNumber>
(C-YN201756-04, 
<typeStatus id="548C8827F33EFFAE68EEFE33FCB87D1E" box="[690,791,511,537]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE696DFE32FC1D7D1E" box="[817,946,510,537]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845451" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE696DFE32FCF07D1E" box="[817,863,510,537]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845451
</accessionNumber>
(C-YN201762-05, 
<typeStatus id="548C8827F33EFFAE6EC0FE33FAAE7D1E" box="[1180,1281,511,537]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6F47FE33FA337D1E" box="[1307,1436,511,537]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH837679" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6F47FE33FAE77D1E" box="[1307,1352,511,537]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
837679
</accessionNumber>
(C-YN201627-02, 
<typeStatus id="548C8827F33EFFAE6B33FDEFFE7B7D3A" box="[367,468,547,573]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6BB6FDEEFDC47D3A" box="[490,619,546,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845539" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6BB6FDEEFDB87D3A" box="[490,535,546,573]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845539
</accessionNumber>
(C-YN201703-01, 
<typeStatus id="548C8827F33EFFAE6911FDEFFC1D7D3A" box="[845,946,547,573]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6994FDEEFBE67D3A" box="[968,1097,546,573]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845514" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6994FDEEFC5A7D3A" box="[968,1013,546,573]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845514
</accessionNumber>
(C-YN201722-01, 
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), 
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<collectionCode id="ED26AE40F33EFFAE6ACBFD8AFF6B7D66" box="[151,196,582,609]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845460
</accessionNumber>
(C-YN201756-04, 
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), 
<accessionNumber id="9464AB66F33EFFAE6832FD8AFD5F7D66" box="[622,752,582,609]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845487" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6832FD8AFD347D66" box="[622,667,582,609]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845487
</accessionNumber>
(C-YN201735-08, 
<typeStatus id="548C8827F33EFFAE6991FD8BFB9D7D66" box="[973,1074,583,609]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
), 
<accessionNumber id="9464AB66F33EFFAE6E19FD8AFB697D66" box="[1093,1222,582,609]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MH845451" pageId="10" pageNumber="467" type="EnaNcbi">
<collectionCode id="ED26AE40F33EFFAE6E19FD8AFBDC7D66" box="[1093,1139,582,609]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="10" pageNumber="467">MH</collectionCode>
845451
</accessionNumber>
(C-YN201762-05, 
<typeStatus id="548C8827F33EFFAE6ACBFDA7FF527D82" box="[151,253,619,645]" pageId="10" pageNumber="467" type="paratype">paratype</typeStatus>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D650EF33EFFAE6A9BFD43FAB6792A" pageId="10" pageNumber="467" type="discussion">
<paragraph id="8B883685F33EFFAE6A9BFD43FF4F7C86" blockId="10.[151,1437,151,1581]" pageId="10" pageNumber="467">
<emphasis id="B943EA97F33EFFAE6A9BFD43FE947DAE" bold="true" box="[199,315,655,681]" pageId="10" pageNumber="467">Remarks.</emphasis>
<taxonomicName id="4C374D06F33EFFAE6B1FFD43FDA37DAF" authority="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu, 2021" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[323,524,655,681]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F33EFFAE6B1FFD43FDA37DAF" box="[323,524,655,681]" italics="true" pageId="10" pageNumber="467">A. yuanjiangensis</emphasis>
</taxonomicName>
<emphasis id="B943EA97F33EFFAE6848FD43FDC07DAE" bold="true" box="[532,623,655,681]" pageId="10" pageNumber="467">
<taxonomicNameLabel id="A27057ECF33EFFAE6848FD43FDC07DAE" box="[532,623,655,681]" pageId="10" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
keys to the 
<taxonomicName id="4C374D06F33EFFAE68A7FD43FC697DAF" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[763,966,655,681]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="corticis">
<emphasis id="B943EA97F33EFFAE68A7FD43FC697DAF" box="[763,966,655,681]" italics="true" pageId="10" pageNumber="467">Amynthas corticis</emphasis>
</taxonomicName>
-group (
<bibRefCitation id="EFA64B74F33EFFAE6E7DFD43FAB77DAE" author="Sims, R. W. &amp; Easton, E. G." box="[1057,1304,655,681]" pageId="10" pageNumber="467" pagination="169 - 268" refId="ref13169" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biological Journal of the Linnean Society, 4 (3), 169 - 268. https: // doi. org / 10.1111 / j. 1095 - 8312.1972. tb 00694. x" type="journal article" year="1972">Sims &amp; Easton, 1972</bibRefCitation>
), characterized by four spermathecal pores located intersegmentally in 5/6–8/9 and by holandry. By now, there are 109 species belonging into 
<taxonomicName id="4C374D06F33EFFAE6B1AFD1BFE147DF7" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[326,443,727,753]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="corticis">
<emphasis id="B943EA97F33EFFAE6B1AFD1BFE147DF7" box="[326,443,727,753]" italics="true" pageId="10" pageNumber="467">A. corticis</emphasis>
</taxonomicName>
-group totally (
<bibRefCitation id="EFA64B74F33EFFAE6834FD1BFCF57DF6" author="Nguyen, T. T. &amp; Lam, D. H. &amp; Trinh, B. T. K. &amp; Nguyen, A. D." box="[616,858,727,753]" pageId="10" pageNumber="467" pagination="1 - 25" refId="ref13018" refString="Nguyen, T. T., Lam, D. H., Trinh, B. T. K. &amp; Nguyen, A. D. (2020 a) The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys, 932, 1 - 25. https: // doi. org / 10.3897 / zookeys. 932.50314" type="journal article" year="2020">
Nguyen 
<emphasis id="B943EA97F33EFFAE6890FD1BFCA87DF6" box="[716,775,727,753]" italics="true" pageId="10" pageNumber="467">et al.</emphasis>
2020a
</bibRefCitation>
). The combined characters of medium-sized body, male pore on the top of a large raised pulvinate pad, no genital papillae in the spermathecal pore and male pore regions, intestinal caeca between simple and complex, prostate gland partially vestigial or developed, and rod-shaped spermathecal seminal chamber make this new species different from the species reported from 
<collectingCountry id="F3207615F33EFFAE6EE6FC8FFB527C5A" box="[1210,1277,835,861]" name="China" pageId="10" pageNumber="467">China</collectingCountry>
in 
<taxonomicName id="4C374D06F33EFFAE6F7CFC8FFA3C7C5B" baseAuthorityName="Kinberg" baseAuthorityYear="1867" box="[1312,1427,835,861]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="corticis">
<emphasis id="B943EA97F33EFFAE6F7CFC8FFA3C7C5B" box="[1312,1427,835,861]" italics="true" pageId="10" pageNumber="467">A. corticis</emphasis>
</taxonomicName>
-group.
</paragraph>
<paragraph id="8B883685F33EFFAE6A9BFC47FAFD7B32" blockId="10.[151,1437,151,1581]" pageId="10" pageNumber="467">
This new species is similar to 
<taxonomicName id="4C374D06F33EFFAE6847FC47FC0E7CA2" authority="(Gates, 1935)" baseAuthorityName="Gates" baseAuthorityYear="1935" box="[539,929,906,933]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="fornicates">
<emphasis id="B943EA97F33EFFAE6847FC47FCAE7CA2" box="[539,769,907,933]" italics="true" pageId="10" pageNumber="467">Amynthas fornicates</emphasis>
(Gates, 1935)
</taxonomicName>
in medium-sized body, no genital papillae in the male pore region, and the arrangement of spermathecal pores. However, in 
<taxonomicName id="4C374D06F33EFFAE6E45FC63FB4E7CCF" authority="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu, 2021" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[1049,1249,943,969]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F33EFFAE6E45FC63FB4E7CCF" box="[1049,1249,943,969]" italics="true" pageId="10" pageNumber="467">A. yuanjiangensis</emphasis>
</taxonomicName>
<emphasis id="B943EA97F33EFFAE6EB5FC63FAEB7CCE" bold="true" box="[1257,1348,943,969]" pageId="10" pageNumber="467">
<taxonomicNameLabel id="A27057ECF33EFFAE6EB5FC63FAEB7CCE" box="[1257,1348,943,969]" pageId="10" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the intestinal caeca are between simple and complex, the prostate glands are partially degenerated, and the spermathecal diverticulum shorter than the main spermathecal axis; however, in 
<taxonomicName id="4C374D06F33EFFAE69DCFC3BFBA07B16" baseAuthorityName="Gates" baseAuthorityYear="1935" box="[896,1039,1015,1041]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="fornicates">
<emphasis id="B943EA97F33EFFAE69DCFC3BFBA07B16" box="[896,1039,1015,1041]" italics="true" pageId="10" pageNumber="467">A. fornicates</emphasis>
</taxonomicName>
, the intestinal caeca are simple, the prostate glands are developed, and the spermathecal diverticulum is longer than the main spermathecal axis.
</paragraph>
<paragraph id="8B883685F33EFFAE6A9BFBF3FC7C7A0A" blockId="10.[151,1437,151,1581]" pageId="10" pageNumber="467">
<taxonomicName id="4C374D06F33EFFAE6A9BFBF3FE227B5F" authority="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu, 2021" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[199,397,1087,1113]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F33EFFAE6A9BFBF3FE227B5F" box="[199,397,1087,1113]" italics="true" pageId="10" pageNumber="467">A. yuanjiangensis</emphasis>
</taxonomicName>
<emphasis id="B943EA97F33EFFAE6BCFFBF3FE437B5E" bold="true" box="[403,492,1087,1113]" pageId="10" pageNumber="467">
<taxonomicNameLabel id="A27057ECF33EFFAE6BCFFBF3FE437B5E" box="[403,492,1087,1113]" pageId="10" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is resemble 
<taxonomicName id="4C374D06F33EFFAE682AFBF3FBB87B5E" authority="(Chen, 1938)" baseAuthorityName="Chen" baseAuthorityYear="1938" box="[630,1047,1087,1113]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="homochaetus">
<emphasis id="B943EA97F33EFFAE682AFBF3FCD07B5E" box="[630,895,1087,1113]" italics="true" pageId="10" pageNumber="467">Amynthas homochaetus</emphasis>
(Chen, 1938)
</taxonomicName>
in the large raised pulvinate pad of male pore area, no genital papillae on spermathecal pore region and male pore region especially. But 
<taxonomicName id="4C374D06F33EFFAE6F4DFBAFFF727BA7" authority="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu, 2021" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F33EFFAE6F4DFBAFFF727BA7" italics="true" pageId="10" pageNumber="467">A. yuanjiangensis</emphasis>
</taxonomicName>
<emphasis id="B943EA97F33EFFAE6AB9FB4BFE907BA6" bold="true" box="[229,319,1159,1185]" pageId="10" pageNumber="467">
<taxonomicNameLabel id="A27057ECF33EFFAE6AB9FB4BFE907BA6" box="[229,319,1159,1185]" pageId="10" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is clearly distinguished from 
<taxonomicName id="4C374D06F33EFFAE68D2FB4BFCED7BA6" baseAuthorityName="Chen" baseAuthorityYear="1938" box="[654,834,1159,1185]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="homochaetus">
<emphasis id="B943EA97F33EFFAE68D2FB4BFCED7BA6" box="[654,834,1159,1185]" italics="true" pageId="10" pageNumber="467">A. homochaetus</emphasis>
</taxonomicName>
by the following characters: 1) the intestinal caeca in 
<taxonomicName id="4C374D06F33EFFAE6ACBFB67FEE57BC2" baseAuthorityName="Chen" baseAuthorityYear="1938" box="[151,330,1195,1221]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="homochaetus">
<emphasis id="B943EA97F33EFFAE6ACBFB67FEE57BC2" box="[151,330,1195,1221]" italics="true" pageId="10" pageNumber="467">A. homochaetus</emphasis>
</taxonomicName>
are simple, but in this new species, they are between simple and complex; 2) the prostate glands in 
<taxonomicName id="4C374D06F33EFFAE6ACBFB03FEE57BEE" baseAuthorityName="Chen" baseAuthorityYear="1938" box="[151,330,1231,1257]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="homochaetus">
<emphasis id="B943EA97F33EFFAE6ACBFB03FEE57BEE" box="[151,330,1231,1257]" italics="true" pageId="10" pageNumber="467">A. homochaetus</emphasis>
</taxonomicName>
are well developed, but in the new species, they are partially degenerated; 3) the seminal chambers in 
<taxonomicName id="4C374D06F33EFFAE6AE8FB3FFEC87A0A" baseAuthorityName="Chen" baseAuthorityYear="1938" box="[180,359,1267,1293]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="homochaetus">
<emphasis id="B943EA97F33EFFAE6AE8FB3FFEC87A0A" box="[180,359,1267,1293]" italics="true" pageId="10" pageNumber="467">A. homochaetus</emphasis>
</taxonomicName>
are ovoid, but those in the new species are rod-shaped.
</paragraph>
<paragraph id="8B883685F33EFFAE6A9BFADBFAB6792A" blockId="10.[151,1437,151,1581]" pageId="10" pageNumber="467">
<taxonomicName id="4C374D06F33EFFAE6A9BFADBFDC77A36" authority="Sun &amp; Qiu, 2018" authorityName="Sun &amp; Qiu" authorityYear="2018" box="[199,616,1303,1329]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="disperses">
<emphasis id="B943EA97F33EFFAE6A9BFADBFE0B7A36" box="[199,420,1303,1329]" italics="true" pageId="10" pageNumber="467">Amynthas disperses</emphasis>
<bibRefCitation id="EFA64B74F33EFFAE6BF7FADBFDC77A36" author="Sun, J. &amp; Jiang, J. &amp; Bartlam, S. &amp; Qiu, J. &amp; Hu, F." box="[427,616,1303,1329]" pageId="10" pageNumber="467" pagination="287 - 301" refId="ref13318" refString="Sun, J., Jiang, J., Bartlam, S., Qiu, J. &amp; Hu, F. (2018) Four new Amynthas and Metaphire earthworm species from nine provinces in southern China. Zootaxa, 4496 (1), 287 - 301. https: // doi. org / 10.11646 / zootaxa. 4496.1.24" type="journal article" year="2018">Sun &amp; Qiu, 2018</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C374D06F33EFFAE68C1FADBFCCC7A37" authority="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu, 2021" authorityName="Sun &amp; Jiang &amp; Wu &amp; Yuan &amp; Qiu" authorityYear="2021" box="[669,867,1303,1329]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="yuanjiangensis" status="sp. nov.">
<emphasis id="B943EA97F33EFFAE68C1FADBFCCC7A37" box="[669,867,1303,1329]" italics="true" pageId="10" pageNumber="467">A. yuanjiangensis</emphasis>
</taxonomicName>
<emphasis id="B943EA97F33EFFAE6936FADBFC6C7A36" bold="true" box="[874,963,1303,1329]" pageId="10" pageNumber="467">
<taxonomicNameLabel id="A27057ECF33EFFAE6936FADBFC6C7A36" box="[874,963,1303,1329]" pageId="10" pageNumber="467" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
have both been collected in several localities in South 
<collectingCountry id="F3207615F33EFFAE6B76FAF7FEDF7A52" box="[298,368,1339,1365]" name="China" pageId="10" pageNumber="467">China</collectingCountry>
, and share some characters (e.g. megium-sized body, the area of male pore porophore is bigger, partially vestigial prostate gland, and band- or rod-shaped seminal chamber), but they are different from each other in the following ways: 1) the first dorsal pore is located at 10/11 or 11/12 on 
<taxonomicName id="4C374D06F33EFFAE6E54FA4FFB3E7A9A" authorityName="Sun &amp; Qiu" authorityYear="2018" box="[1032,1169,1411,1437]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="disperses">
<emphasis id="B943EA97F33EFFAE6E54FA4FFB3E7A9A" box="[1032,1169,1411,1437]" italics="true" pageId="10" pageNumber="467">A. disperses</emphasis>
</taxonomicName>
, but always at 11/ 
<quantity id="4CCF9B60F33EFFAE6F3EFA4FFA337A9A" box="[1378,1436,1411,1437]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="10" pageNumber="467" unit="in" value="12.0">12 in</quantity>
the new species; 2) small genital papillae are always present in the spermathecal pore and male pore region in 
<taxonomicName id="4C374D06F33EFFAE6FD8FA6BFF507AE2" authorityName="Sun &amp; Qiu" authorityYear="2018" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="disperses">
<emphasis id="B943EA97F33EFFAE6FD8FA6BFF507AE2" italics="true" pageId="10" pageNumber="467">A. disperses</emphasis>
</taxonomicName>
, but there are no genital papillae in the new species; 3) the intestinal caeca in 
<taxonomicName id="4C374D06F33EFFAE6E34FA07FB407AE2" authorityName="Sun &amp; Qiu" authorityYear="2018" box="[1128,1263,1483,1509]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="disperses">
<emphasis id="B943EA97F33EFFAE6E34FA07FB407AE2" box="[1128,1263,1483,1509]" italics="true" pageId="10" pageNumber="467">A. disperses</emphasis>
</taxonomicName>
are simple, but those in this new species are between simple and complex; 4) the spermathecal diverticulum is about as long as the main spermathecal axis in 
<taxonomicName id="4C374D06F33EFFAE6B9EF9DFFDE6792A" authorityName="Sun &amp; Qiu" authorityYear="2018" box="[450,585,1555,1581]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="10" pageNumber="467" phylum="Annelida" rank="species" species="disperses">
<emphasis id="B943EA97F33EFFAE6B9EF9DFFDE6792A" box="[450,585,1555,1581]" italics="true" pageId="10" pageNumber="467">A. disperses</emphasis>
</taxonomicName>
, but shorter than the main spermathecal axis in the new species.
</paragraph>
</subSubSection>
</treatment>
</document>