351 lines
42 KiB
XML
351 lines
42 KiB
XML
<document id="DEA76BCD84FDAC830325FCD8E47F6E23" ID-CLB-Dataset="24795" ID-DOI="10.11646/zootaxa.4701.1.3" ID-GBIF-Dataset="1fb77063-018a-4222-8733-9450729ac222" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557803" ID-ZooBank="668205B0-B6E9-48F3-9E21-87C534CDC837" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575043522256" checkinUser="plazi" docAuthor="Huang, Sunbin, Tú, Výõng Tân, Phú, Ph ạm Văn, Tian, Mingyi & Faille, Arnaud" docDate="2019" docId="0399DA2B7F32A80CFF33FD61FAB0F210" docLanguage="en" docName="zootaxa.4701.1.3.pdf" docOrigin="Zootaxa 4701 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonkinaphaenops anthonyi Huang & Tú & Phú & Tian & Faille 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="52" masterDocId="FFA0A2537F3FA81DFFA4FF9CFFD8F02F" masterDocTitle="Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="53" masterPageNumber="35" pageNumber="48" updateTime="1698769588363" updateUser="ExternalLinkService">
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<mods:mods id="CE820D8E0A9265318D611FC818EDB922" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="788BEB387956089E337869CB66F18AAA">
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<mods:title id="0415242351A981265204C9D0C949CD0B">Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="ED78A5CB7E9A51C9C44C3DAAE61EE19A">Huang, Sunbin</mods:namePart>
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<mods:roleTerm id="9E7411421749F9B6EE4F8748BEFF2739">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="240CD6E1BBEF9EDBE0C954FA8FC15960">Tú, Výõng Tân</mods:namePart>
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<mods:roleTerm id="628F48B0887E24590A894DDA9ADFAB15">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="961EB02F284A73EF32F50B5D931B50CD">Phú, Ph ạm Văn</mods:namePart>
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<mods:roleTerm id="1E6BF05BE62697AE1FDC549F9C88914F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B41FBA8F0CA5F94BA0EAA154974C7695">Tian, Mingyi</mods:namePart>
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<mods:roleTerm id="DDFCD69E09071CF2ABE5B8C54BD5E495">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9AB657FA24FF2048DD1B247106F6DC2D">Faille, Arnaud</mods:namePart>
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<mods:title id="51E4FD9A860128BA3D0BF890668A2F51">Zootaxa</mods:title>
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<mods:part id="901CAD3648C861A2CE403AE5CD36317B">
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<mods:date id="B7A013124935FB3C982B5417601E8324">2019</mods:date>
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<mods:number id="A116DEDC9FAE7F4C750BE200978BDC84">2019-11-26</mods:number>
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<mods:number id="407B955B4E010F3785815E5D189A9154">4701</mods:number>
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<mods:classification id="EE08F2BE60C14F03C1D6AC9DD41DDF12">journal article</mods:classification>
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<mods:identifier id="2AB8A6FA59D70F8E2F90F35EEC15EBF4" type="DOI">10.11646/zootaxa.4701.1.3</mods:identifier>
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<treatment id="0399DA2B7F32A80CFF33FD61FAB0F210" ID-DOI="http://doi.org/10.5281/zenodo.5929274" ID-GBIF-Taxon="160605351" ID-Zenodo-Dep="5929274" LSID="urn:lsid:plazi:treatment:0399DA2B7F32A80CFF33FD61FAB0F210" httpUri="http://treatment.plazi.org/id/0399DA2B7F32A80CFF33FD61FAB0F210" lastPageId="17" lastPageNumber="52" pageId="13" pageNumber="48">
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<subSubSection id="C32A38B67F32A810FF33FD61FD24F338" box="[151,764,765,792]" pageId="13" pageNumber="48" type="nomenclature">
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<paragraph id="8B8F6B3D7F32A810FF33FD61FD24F338" blockId="13.[151,764,765,827]" box="[151,764,765,792]" pageId="13" pageNumber="48">
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<heading id="D0C7DC517F32A810FF33FD61FD24F338" bold="true" box="[151,764,765,792]" fontSize="11" level="1" pageId="13" pageNumber="48" reason="1">
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<emphasis id="B944B72F7F32A810FF33FD61FD24F338" bold="true" box="[151,764,765,792]" pageId="13" pageNumber="48">
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<taxonomicName id="4C3010BE7F32A810FF33FD61FD4AF338" ID-CoL="8MLSG" authority="Faille & Tian" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[151,658,765,792]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="anthonyi" status="sp. nov.">
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||
<emphasis id="B944B72F7F32A810FF33FD61FE3AF337" bold="true" box="[151,482,765,792]" italics="true" pageId="13" pageNumber="48">Tonkinaphaenops anthonyi</emphasis>
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||
Faille & Tian
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||
</taxonomicName>
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||
,
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||
<taxonomicNameLabel id="A2770A547F32A810FD3BFD61FD24F338" box="[671,764,765,791]" pageId="13" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32A38B67F32A810FF33FCBDFE5AF314" box="[151,386,801,827]" pageId="13" pageNumber="48" type="description">
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<paragraph id="8B8F6B3D7F32A810FF33FCBDFE5AF314" blockId="13.[151,764,765,827]" box="[151,386,801,827]" pageId="13" pageNumber="48">
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(
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<figureCitation id="130B77B87F32A810FF3BFCBDFF35F314" box="[159,237,801,827]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1826]" captionTargetBox="[159,1428,877,1779]" captionTargetId="figure@1.[151,1436,877,1781]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Tonkinaphaenops Deuve, 2013" figureDoi="http://doi.org/10.5281/zenodo.3557805" httpUri="https://zenodo.org/record/3557805/files/figure.png" pageId="13" pageNumber="48">Figs. 1</figureCitation>
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,
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<figureCitation id="130B77B87F32A810FF5FFCBDFE9AF314" box="[251,322,801,827]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="14.[151,250,1995,2020]" captionStartId-1="15.[151,250,911,936]" captionStartId-2="15.[151,250,1996,2021]" captionTargetBox-1="[175,1422,184,876]" captionTargetBox-2="[151,1436,984,1948]" captionTargetId-0="figure@14.[403,1185,171,1972]" captionTargetId-1="figure@15.[151,1435,174,883]" captionTargetId-2="figure@15.[151,1436,984,1948]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="15" captionText-0="FIGURE 11. Tonkinaphaenops anthonyi sp. nov., habitus of holotype, male." captionText-1="FIGURE 12. Male genitalia of Tonkinaphaenops anthonyi sp. nov. (A. median lobe and parameres, lateral view; B. apical lobe, dorsal view)." captionText-2="FIGURE 13. Entrance of Hang Dõi cave, type locality of Tonkinaphaenops anthonyi sp. nov." figureDoi-0="http://doi.org/10.5281/zenodo.3614848" figureDoi-1="http://doi.org/10.5281/zenodo.3557819" figureDoi-2="http://doi.org/10.5281/zenodo.3557821" httpUri-0="https://zenodo.org/record/3614848/files/figure.png" httpUri-1="https://zenodo.org/record/3557819/files/figure.png" httpUri-2="https://zenodo.org/record/3557821/files/figure.png" pageId="13" pageNumber="48">11–13</figureCitation>
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,
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||
<figureCitation id="130B77B87F32A810FEEAFCBDFEA2F314" box="[334,378,801,827]" captionStart="FIGURE 14" captionStartId="16.[151,250,1912,1937]" captionTargetBox="[214,1391,181,1869]" captionTargetId="figure@16.[189,1398,181,1887]" captionTargetPageId="16" captionText="FIGURE 14. Chaetotaxal pattern of Tonkinaphaenops (A. Tonkinaphaenops marinae Deuve, 2013, holotype, male from cave TH20; B, C. Tonkinaphaenops yinquanicus sp. nov.: B. male from Yinquan Dong, C. female from Yinquan Dong; D. Tonkinaphaenops impunctatus sp. nov., male from Hang Rắn cave; E. Tonkinaphaenops jingxicus sp. nov., female from Nianluo Dong; F. Tonkinaphaenops anthonyi sp. nov., male from Hang Dõi cave)." figureDoi="http://doi.org/10.5281/zenodo.3557823" httpUri="https://zenodo.org/record/3557823/files/figure.png" pageId="13" pageNumber="48">14F</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32A38B67F32A810FF33FCF5FE68F3C0" pageId="13" pageNumber="48" type="materials_examined">
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<paragraph id="8B8F6B3D7F32A810FF33FCF5FE68F3C0" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">
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<materialsCitation id="3B5861607F32A810FF33FCF5FAEFF388" ID-GBIF-Occurrence="2460169374" collectingDate="2018-10-20" collectionCode="MNHN" collectorName="Cao B & A. Faille & Faille, P. V & Phu, V. T" country="Vietnam" county="Hang Doi" elevation="386" latitude="22.746584" location="Lang district" longLatPrecision="1" longitude="106.7433" municipality="Lung Rum" pageId="13" pageNumber="48" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<emphasis id="B944B72F7F32A810FF33FCF5FED3F3AC" bold="true" box="[151,267,873,899]" pageId="13" pageNumber="48">
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<typeStatus id="548BD59F7F32A810FF33FCF5FEDEF3AC" box="[151,262,873,899]" pageId="13" pageNumber="48" type="holotype">Holotype</typeStatus>
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.
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||
</emphasis>
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||
<specimenCount id="9D36A0B47F32A810FEB0FCF5FEF4F3AC" box="[276,300,873,899]" pageId="13" pageNumber="48" type="male">♂</specimenCount>
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;
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<collectingCountry id="F3272BAD7F32A810FE9EFCF5FE45F3AC" box="[314,413,873,899]" name="Vietnam" pageId="13" pageNumber="48">Vietnam</collectingCountry>
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:
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<collectorName id="26C50EEB7F32A810FE0EFCF5FE2AF3AC" box="[426,498,872,899]" pageId="13" pageNumber="48">Cao B</collectorName>
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ằng province: Hạ
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<location id="8EEF3DE67F32A810FD66FCF5FC8DF3AC" LSID="urn:lsid:plazi:treatment:0399DA2B7F32A80CFF33FD61FAB0F210:8EEF3DE67F32A810FD66FCF5FC8DF3AC" box="[706,853,873,899]" country="Vietnam" county="Hang Doi" latitude="22.746584" longLatPrecision="1" longitude="106.7433" municipality="Lung Rum" name="Lang district" pageId="13" pageNumber="48">Lang district</location>
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||
, Đồng
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<location id="8EEF3DE67F32A810FC0EFCF5FC3CF3AC" LSID="urn:lsid:plazi:treatment:0399DA2B7F32A80CFF33FD61FAB0F210:8EEF3DE67F32A810FC0EFCF5FC3CF3AC" box="[938,996,873,899]" country="Vietnam" county="Hang Doi" latitude="22.746584" longLatPrecision="1" longitude="106.7433" municipality="Lung Rum" name="Loan" pageId="13" pageNumber="48">Loan</location>
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commune,
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<collectingMunicipality id="6BEBF1477F32A810FBCFFCF5FB33F3AC" box="[1131,1259,873,899]" pageId="13" pageNumber="48">Lũng Rúm</collectingMunicipality>
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:
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<collectingCounty id="62EE13B17F32A810FB53FCF5FABFF3AC" box="[1271,1383,873,899]" pageId="13" pageNumber="48">Hang Dõi</collectingCounty>
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(bat cave);
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<geoCoordinate id="EE040DFA7F32A810FF47FC11FE5AF388" box="[227,386,909,935]" degrees="22" direction="north" minutes="44" orientation="latitude" pageId="13" pageNumber="48" precision="1" seconds="47.7" value="22.746584">22°44’47.7”N</geoCoordinate>
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<geoCoordinate id="EE040DFA7F32A810FE2DFC10FDEDF388" box="[393,565,908,935]" degrees="106" direction="east" minutes="44" orientation="longitude" pageId="13" pageNumber="48" precision="1" seconds="35.9" value="106.7433">106°44’35.9”E</geoCoordinate>
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,
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<quantity id="4CC8C6D87F32A810FDE5FC11FD54F388" box="[577,652,909,935]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="13" pageNumber="48" unit="m" value="386.0">
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<elevation id="001D8C0E7F32A810FDE5FC11FD54F388" box="[577,652,909,935]" metricMagnitude="2" metricUnit="m" metricValue="3.86" pageId="13" pageNumber="48" unit="m" value="386.0">386 m</elevation>
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</quantity>
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;
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<date id="FF8E4DFD7F32A810FD3CFC11FC81F388" box="[664,857,909,935]" pageId="13" pageNumber="48" value="2018-10-20">
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<collectingDate id="EFCAB4157F32A810FD3CFC11FC81F388" box="[664,857,909,935]" pageId="13" pageNumber="48" value="2018-10-20">20 October 2018</collectingDate>
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</date>
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||
; coll.
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||
<collectorName id="26C50EEB7F32A810FC39FC11FBDBF388" box="[925,1027,909,935]" pageId="13" pageNumber="48">A. Faille</collectorName>
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||
, P.V.
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||
<collectorName id="26C50EEB7F32A810FBE5FC11FB7EF388" box="[1089,1190,909,935]" pageId="13" pageNumber="48">Phu, V.T</collectorName>
|
||
. Tu;
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||
<collectionCode id="ED21F3F87F32A810FB7AFC11FAEFF388" box="[1246,1335,909,935]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="48">MNHN</collectionCode>
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||
</materialsCitation>
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||
.
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||
<materialsCitation id="3B5861607F32A810FAE6FC11FD04F3E4" ID-GBIF-Occurrence="2460169371" collectionCode="SMNS" country="Vietnam" pageId="13" pageNumber="48" specimenCode="L1791" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<emphasis id="B944B72F7F32A810FAE6FC11FF66F3E4" bold="true" pageId="13" pageNumber="48">
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<typeStatus id="548BD59F7F32A810FAE6FC11FF66F3E4" pageId="13" pageNumber="48" type="paratype">Paratypes</typeStatus>
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</emphasis>
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.
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<specimenCount id="9D36A0B47F32A810FF68FC2DFF2DF3E4" box="[204,245,945,971]" pageId="13" pageNumber="48" type="male">1 ♂</specimenCount>
|
||
(DNA extraction code:
|
||
<collectionCode id="ED21F3F87F32A810FDA5FC2DFD97F3E4" box="[513,591,945,971]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="13" pageNumber="48">SMNS</collectionCode>
|
||
_
|
||
<specimenCode id="DB96C3467F32A810FDFAFC2DFD7EF3E4" box="[606,678,945,971]" collectionCode="SMNS" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="13" pageNumber="48">L1791</specimenCode>
|
||
) and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5861607F32A810FD47FC2DFBB9F3E4" ID-GBIF-Occurrence="2460169380" box="[739,1121,945,971]" collectingDate="2018-10-20" collectionCode="IEBR" collectorName="Cao B & A. Faille & Faille, P. V & Phu, V. T" country="Vietnam" county="Hang Doi" elevation="386" latitude="22.746584" location="Lang district" longLatPrecision="1" longitude="106.7433" municipality="Lung Rum" pageId="13" pageNumber="48" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D36A0B47F32A810FD47FC2DFCD5F3E4" box="[739,781,945,971]" pageId="13" pageNumber="48" type="female">1 ♀</specimenCount>
|
||
; same data as holotype;
|
||
<collectionCode id="ED21F3F87F32A810FBBAFC2DFBB9F3E4" box="[1054,1121,945,971]" pageId="13" pageNumber="48">IEBR</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B5861607F32A810FBC8FC2DFE74F3C0" ID-GBIF-Occurrence="2460169376" collectingDate="2018-10-20" collectionCode="SMNS" collectorName="Cao B & A. Faille & Faille, P. V & Phu, V. T" country="Vietnam" county="Hang Doi" elevation="386" latitude="22.746584" location="Lang district" longLatPrecision="1" longitude="106.7433" municipality="Lung Rum" pageId="13" pageNumber="48" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
|
||
<specimenCount id="9D36A0B47F32A810FBC8FC2DFB4EF3E4" box="[1132,1174,945,971]" pageId="13" pageNumber="48" type="female">1 ♀</specimenCount>
|
||
; spare in ethanol; same data as holotype;
|
||
<collectionCode id="ED21F3F87F32A810FEF8FC49FE74F3C0" box="[348,428,981,1007]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="13" pageNumber="48">SMNS</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A38B67F32A810FF63FC64FBA9F7F0" pageId="13" pageNumber="48" type="description">
|
||
<paragraph id="8B8F6B3D7F32A810FF63FC64FF12F450" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">
|
||
<emphasis id="B944B72F7F32A810FF63FC64FE81F43C" bold="true" box="[199,345,1016,1043]" pageId="13" pageNumber="48">Description.</emphasis>
|
||
Habitus as in
|
||
<figureCitation id="130B77B87F32A810FE5DFC65FD92F43C" box="[505,586,1017,1043]" captionStart="FIGURE 11" captionStartId="14.[151,250,1995,2020]" captionTargetId="figure@14.[403,1185,171,1972]" captionTargetPageId="14" captionText="FIGURE 11. Tonkinaphaenops anthonyi sp. nov., habitus of holotype, male." figureDoi="http://doi.org/10.5281/zenodo.3614848" httpUri="https://zenodo.org/record/3614848/files/figure.png" pageId="13" pageNumber="48">Fig. 11</figureCitation>
|
||
. Length: 6.0 mm (from the middle of the labrum to the apex of the elytron). Anophthalmous. Body elongate, elytra long and subparallel. Narrow fore-body: elytra wider than pronotum, which is wider than head. EL:
|
||
<quantity id="4CC8C6D87F32A810FE09FBDDFDDFF474" box="[429,519,1089,1115]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="48" unit="mm" value="3.4">3.4 mm</quantity>
|
||
, EW:
|
||
<quantity id="4CC8C6D87F32A810FDF4FBDDFD73F474" box="[592,683,1089,1115]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="48" unit="mm" value="1.9">1.9 mm</quantity>
|
||
. PL:
|
||
<quantity id="4CC8C6D87F32A810FD4DFBDDFC9BF474" box="[745,835,1089,1115]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="13" pageNumber="48" unit="mm" value="1.4">1.4 mm</quantity>
|
||
, PW: 1.0 mm. HL: 1.0 mm, HW:
|
||
<quantity id="4CC8C6D87F32A810FB6BFBDDFAE0F474" box="[1231,1336,1089,1115]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="13" pageNumber="48" unit="mm" value="0.76">0.76 mm</quantity>
|
||
. AL:
|
||
<quantity id="4CC8C6D87F32A810FADDFBDCFF1EF450" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="48" unit="mm" value="4.5">4.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8F6B3D7F32A810FF63FB15FD28F48C" blockId="13.[151,1437,872,2015]" box="[199,752,1161,1187]" pageId="13" pageNumber="48">Color: body dark-reddish brown, concolor, shiny.</paragraph>
|
||
<paragraph id="8B8F6B3D7F32A810FF63FB31FC6FF4C4" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">Microsculpture fine but distinct across dorsal surface of pronotum and most of head. On head, only the vertex lacks micro-sculpture. Elytra with very fine polygonal micro-sculpture.</paragraph>
|
||
<paragraph id="8B8F6B3D7F32A810FF63FB69FF15F51C" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">Glabrous on dorsal surface of pronotum, sparsely pubescent on the head and pubescent on elytra, except on the disk.</paragraph>
|
||
<paragraph id="8B8F6B3D7F32A810FF63FAA1FC85F6B4" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">
|
||
Head (without mandibles) longer than wide, HLl/HW = 1.3; frontal furrows incomplete, deep and long, slightly and regularly curved, convergent in anterior portion to their mid-length, then slightly divergent; labrum bisinuate, six-setose. Clypeus quadrisetose, without additional setae; right mandible bidentate; eyes absent, two pairs of supraorbital pores present; mentum bisetose, base widely concave, mental tooth simple, slightly protruding; lateral lobe sharp and pointed at apex; labial suture invisible; submentum 6-setose; ligula slightly protruding at apex, multi-setose; penultimate article of the labial palps about 1.5 times longer than the last, quadrisetose, with two setae located on inner and two on outer margin. penultimate article of the maxillary palps much longer than the last one; antennae extending to about the last third of elytra; scape with a few setae; the 2
|
||
<superScript id="7C45C6757F32A810FC40F9A4FBDDF67C" attach="none" box="[996,1029,1592,1619]" fontSize="6" pageId="13" pageNumber="48">nd–</superScript>
|
||
11
|
||
<superScript id="7C45C6757F32A810FBBAF9A4FBF3F669" attach="left" box="[1054,1067,1592,1606]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
|
||
antennomeres pubescent; the 2
|
||
<superScript id="7C45C6757F32A810FA28F9A4FA44F669" attach="none" box="[1420,1436,1592,1606]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
|
||
antennomere a little longer than scape; antennomere 3–7 the longest, subequal, then other antennomeres decreasing in length posteriorly until the 10
|
||
<superScript id="7C45C6757F32A810FDA5F91CFDD6F6A1" attach="left" box="[513,526,1664,1678]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
|
||
which is shorter than the 11
|
||
<superScript id="7C45C6757F32A810FCEEF91CFC8FF6A1" attach="none" box="[842,855,1664,1678]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8F6B3D7F32A810FF63F939FDB5F728" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">Pronotum longer than wide, PW/PL = 0.7, wider than head, PW/HW = 1.3, front wider than base, PbW/PfW = 0.9; lateral margins slightly bordered throughout; two pairs of marginal setae present, at the level of 1/5 from front and a little before hind angle respectively.</paragraph>
|
||
<paragraph id="8B8F6B3D7F32A810FF63F88DFBA9F7F0" blockId="13.[151,1437,872,2015]" pageId="13" pageNumber="48">
|
||
Elytra elongate, much longer than wide, EL/EW = 1.8, subparallel, widest at about 1/3 from the base, much longer than pronotum, EL/PL = 3.4, wider than pronotum, EW/PW = 1.9; chaetotaxy as in
|
||
<figureCitation id="130B77B87F32A810FBFAF8A9FB67F760" box="[1118,1215,1845,1871]" captionStart="FIGURE 14" captionStartId="16.[151,250,1912,1937]" captionTargetBox="[214,1391,181,1869]" captionTargetId="figure@16.[189,1398,181,1887]" captionTargetPageId="16" captionText="FIGURE 14. Chaetotaxal pattern of Tonkinaphaenops (A. Tonkinaphaenops marinae Deuve, 2013, holotype, male from cave TH20; B, C. Tonkinaphaenops yinquanicus sp. nov.: B. male from Yinquan Dong, C. female from Yinquan Dong; D. Tonkinaphaenops impunctatus sp. nov., male from Hang Rắn cave; E. Tonkinaphaenops jingxicus sp. nov., female from Nianluo Dong; F. Tonkinaphaenops anthonyi sp. nov., male from Hang Dõi cave)." figureDoi="http://doi.org/10.5281/zenodo.3557823" httpUri="https://zenodo.org/record/3557823/files/figure.png" pageId="13" pageNumber="48">Fig. 14F</figureCitation>
|
||
: basal pores present at the beginning of the 2
|
||
<superScript id="7C45C6757F32A810FE09F8C4FE65F749" attach="left" box="[429,445,1880,1894]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
|
||
stria; 4 dorsal setiferous pores present in 3
|
||
<superScript id="7C45C6757F32A810FC02F8C4FC6CF749" attach="left" box="[934,948,1880,1894]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
|
||
stria; the first pore of the umbilicate series shifted to level of the 7
|
||
<superScript id="7C45C6757F32A810FE3CF8E0FE7DF7A5" attach="left" box="[408,421,1916,1930]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
|
||
stria (hardly discernible) and a little before the 2
|
||
<superScript id="7C45C6757F32A810FC61F8E0FC0DF7A5" attach="left" box="[965,981,1916,1930]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
|
||
pore; the 3
|
||
<superScript id="7C45C6757F32A810FBF6F8E0FBB8F7A5" attach="left" box="[1106,1120,1916,1930]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
|
||
pore of marginal umbilicate series much closer to 2nd than to 4
|
||
<superScript id="7C45C6757F32A810FDBBF83CFDF4F781" attach="none" box="[543,556,1952,1966]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
|
||
, making distance between 3
|
||
<superScript id="7C45C6757F32A810FCCEF83CFCA0F781" attach="left" box="[874,888,1952,1966]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
|
||
and 4
|
||
<superScript id="7C45C6757F32A810FC19F83CFC12F781" attach="left" box="[957,970,1952,1966]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
|
||
twice as that between 2
|
||
<superScript id="7C45C6757F32A810FB73F83CFB3FF781" attach="left" box="[1239,1255,1952,1966]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
|
||
and 3
|
||
<superScript id="7C45C6757F32A810FA88F83CFAE2F781" attach="none" box="[1324,1338,1952,1966]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
|
||
; the two setae of the middle group at about the half of elytra from base; two apical pores present.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4F3BB57F31A813FF33F857FC7AF7CB" ID-DOI="http://doi.org/10.5281/zenodo.3614848" ID-Zenodo-Dep="3614848" box="[151,930,1995,2020]" httpUri="https://zenodo.org/record/3614848/files/figure.png" pageId="14" pageNumber="49" targetBox="[423,1176,189,1960]" targetPageId="14">
|
||
<paragraph id="8B8F6B3D7F31A813FF33F857FC7AF7CB" blockId="14.[151,930,1995,2020]" box="[151,930,1995,2020]" pageId="14" pageNumber="49">
|
||
<emphasis id="B944B72F7F31A813FF33F857FEC6F7CB" bold="true" box="[151,286,1995,2020]" pageId="14" pageNumber="49">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C3010BE7F31A813FE81F850FDEEF7CB" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[293,566,1996,2020]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="anthonyi" status="sp. nov.">
|
||
<emphasis id="B944B72F7F31A813FE81F850FDEEF7CB" box="[293,566,1996,2020]" italics="true" pageId="14" pageNumber="49">Tonkinaphaenops anthonyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944B72F7F31A813FD98F850FD54F7CB" bold="true" box="[572,652,1996,2020]" pageId="14" pageNumber="49">
|
||
<taxonomicNameLabel id="A2770A547F31A813FD98F850FD54F7CB" box="[572,652,1996,2020]" pageId="14" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus of holotype, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F3BB57F30A812FF33FC13FEC4F3E3" ID-DOI="http://doi.org/10.5281/zenodo.3557819" ID-Zenodo-Dep="3557819" httpUri="https://zenodo.org/record/3557819/files/figure.png" pageId="15" pageNumber="50" startId="15.[151,250,911,936]" targetBox="[175,1422,184,876]" targetPageId="15">
|
||
<paragraph id="8B8F6B3D7F30A812FF33FC13FEC4F3E3" blockId="15.[151,1436,911,972]" pageId="15" pageNumber="50">
|
||
<emphasis id="B944B72F7F30A812FF33FC13FEC7F387" bold="true" box="[151,287,911,936]" pageId="15" pageNumber="50">FIGURE 12.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C3010BE7F30A812FE72FC0CFD3FF387" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[470,743,912,936]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="50" phylum="Arthropoda" rank="species" species="anthonyi" status="sp. nov.">
|
||
<emphasis id="B944B72F7F30A812FE72FC0CFD3FF387" box="[470,743,912,936]" italics="true" pageId="15" pageNumber="50">Tonkinaphaenops anthonyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944B72F7F30A812FD48FC0CFCE4F387" bold="true" box="[748,828,912,936]" pageId="15" pageNumber="50">
|
||
<taxonomicNameLabel id="A2770A547F30A812FD48FC0CFCE4F387" box="[748,828,912,936]" pageId="15" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A. median lobe and parameres, lateral view; B. apical lobe, dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F3BB57F30A812FF33F850FB96F7CA" ID-DOI="http://doi.org/10.5281/zenodo.3557821" ID-Zenodo-Dep="3557821" box="[151,1102,1996,2022]" httpUri="https://zenodo.org/record/3557821/files/figure.png" pageId="15" pageNumber="50" startId="15.[151,250,1996,2021]" targetBox="[151,1436,984,1948]" targetPageId="15">
|
||
<paragraph id="8B8F6B3D7F30A812FF33F850FB96F7CA" blockId="15.[151,1102,1996,2022]" box="[151,1102,1996,2022]" pageId="15" pageNumber="50">
|
||
<emphasis id="B944B72F7F30A812FF33F850FEF8F7CA" bold="true" box="[151,288,1996,2021]" pageId="15" pageNumber="50">FIGURE 13.</emphasis>
|
||
Entrance of Hang Dõi cave, type locality of
|
||
<taxonomicName id="4C3010BE7F30A812FD42F851FC2FF7CA" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[742,1015,1997,2021]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="50" phylum="Arthropoda" rank="species" species="anthonyi" status="sp. nov.">
|
||
<emphasis id="B944B72F7F30A812FD42F851FC2FF7CA" box="[742,1015,1997,2021]" italics="true" pageId="15" pageNumber="50">Tonkinaphaenops anthonyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2770A547F30A812FC59F851FB96F7CA" box="[1021,1102,1997,2021]" pageId="15" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4F3BB57F2FA80DFF33F8E4FC59F7D2" ID-DOI="http://doi.org/10.5281/zenodo.3557823" ID-Zenodo-Dep="3557823" httpUri="https://zenodo.org/record/3557823/files/figure.png" pageId="16" pageNumber="51" startId="16.[151,250,1912,1937]" targetBox="[214,1391,181,1869]" targetPageId="16">
|
||
<paragraph id="8B8F6B3D7F2FA80DFF33F8E4FC59F7D2" blockId="16.[151,1437,1912,2045]" pageId="16" pageNumber="51">
|
||
<emphasis id="B944B72F7F2FA80DFF33F8E4FEF9F7BE" bold="true" box="[151,289,1912,1937]" pageId="16" pageNumber="51">FIGURE 14.</emphasis>
|
||
Chaetotaxal pattern of
|
||
<taxonomicName id="4C3010BE7F2FA80DFDB6F8E5FD1CF7BE" authorityName="Deuve" authorityYear="2013" box="[530,708,1913,1937]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944B72F7F2FA80DFDB6F8E5FD1CF7BE" box="[530,708,1913,1937]" italics="true" pageId="16" pageNumber="51">Tonkinaphaenops</emphasis>
|
||
</taxonomicName>
|
||
(A.
|
||
<taxonomicName id="4C3010BE7F2FA80DFD57F8E5FB54F7BE" authority="Deuve, 2013" authorityName="Deuve" authorityYear="2013" box="[755,1164,1913,1937]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="marinae">
|
||
<emphasis id="B944B72F7F2FA80DFD57F8E5FBD8F7BF" box="[755,1024,1913,1937]" italics="true" pageId="16" pageNumber="51">Tonkinaphaenops marinae</emphasis>
|
||
<bibRefCitation id="EFA116CC7F2FA80DFBA3F8E5FB54F7BE" author="Deuve, T." box="[1031,1164,1913,1937]" pageId="16" pageNumber="51" pagination="209 - 214" refId="ref6294" refString="Deuve, T. (2013) Un nouveau Trechini troglobie du Vietnam septentrional (Coleoptera, Caraboidea, Trechidae). Bulletin de la Societe entomologique de France, 118 (2), 209 - 214." type="journal article" year="2013">Deuve, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, male from cave TH20; B, C.
|
||
<taxonomicName id="4C3010BE7F2FA80DFE84F801FD8BF79B" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[288,595,1949,1973]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="yinquanicus" status="sp. nov.">
|
||
<emphasis id="B944B72F7F2FA80DFE84F801FD8BF79B" box="[288,595,1949,1973]" italics="true" pageId="16" pageNumber="51">Tonkinaphaenops yinquanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944B72F7F2FA80DFDF8F801FD76F79A" bold="true" box="[604,686,1949,1973]" pageId="16" pageNumber="51">
|
||
<taxonomicNameLabel id="A2770A547F2FA80DFDF8F801FD76F79A" box="[604,686,1949,1973]" pageId="16" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: B. male from Yinquan Dong, C. female from Yinquan Dong; D.
|
||
<taxonomicName id="4C3010BE7F2FA80DFAF8F801FE4EF7F7" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="impunctatus" status="sp. nov.">
|
||
<emphasis id="B944B72F7F2FA80DFAF8F801FE4EF7F7" italics="true" pageId="16" pageNumber="51">Tonkinaphaenops impunctatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944B72F7F2FA80DFE3BF85DFE2AF7F6" bold="true" box="[415,498,1985,2009]" pageId="16" pageNumber="51">
|
||
<taxonomicNameLabel id="A2770A547F2FA80DFE3BF85DFE2AF7F6" box="[415,498,1985,2009]" pageId="16" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male from Hang Rắn cave; E.
|
||
<taxonomicName id="4C3010BE7F2FA80DFCE6F85DFB80F7F7" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[834,1112,1985,2009]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="jingxicus" status="sp. nov.">
|
||
<emphasis id="B944B72F7F2FA80DFCE6F85DFB80F7F7" box="[834,1112,1985,2009]" italics="true" pageId="16" pageNumber="51">Tonkinaphaenops jingxicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944B72F7F2FA80DFBC5F85DFB6CF7F6" bold="true" box="[1121,1204,1985,2009]" pageId="16" pageNumber="51">
|
||
<taxonomicNameLabel id="A2770A547F2FA80DFBC5F85DFB6CF7F6" box="[1121,1204,1985,2009]" pageId="16" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, female from Nianluo Dong; F.
|
||
<taxonomicName id="4C3010BE7F2FA80DFF52F879FDDFF7D2" authority="Huang & Tú & Phú & Tian & Faille, 2019" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[246,519,2021,2045]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="anthonyi" status="sp. nov.">
|
||
<emphasis id="B944B72F7F2FA80DFF52F879FDDFF7D2" box="[246,519,2021,2045]" italics="true" pageId="16" pageNumber="51">Tonkinaphaenops anthonyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B944B72F7F2FA80DFDA9F879FD85F7D2" bold="true" box="[525,605,2021,2045]" pageId="16" pageNumber="51">
|
||
<taxonomicNameLabel id="A2770A547F2FA80DFDA9F879FD85F7D2" box="[525,605,2021,2045]" pageId="16" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male from Hang Dõi cave).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32A38B67F2EA80CFF63FF05FE76F0D5" pageId="17" pageNumber="52" type="etymology">
|
||
<paragraph id="8B8F6B3D7F2EA80CFF63FF05FE76F0D5" blockId="17.[151,1437,152,575]" pageId="17" pageNumber="52">
|
||
<emphasis id="B944B72F7F2EA80CFF63FF05FE96F09C" bold="true" box="[199,334,153,179]" pageId="17" pageNumber="52">Etymology.</emphasis>
|
||
The specific name is a genitive proper noun in honor of the late Anthony « Tony » Whitten for his contribution to the protection of karstic areas of Asia, and his understanding of the importance of taxonomy in the protection of cave fauna.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A38B67F2EA80CFF63FE99FF28F235" pageId="17" pageNumber="52" type="discussion">
|
||
<paragraph id="8B8F6B3D7F2EA80CFF63FE99FD39F16D" blockId="17.[151,1437,152,575]" pageId="17" pageNumber="52">
|
||
<emphasis id="B944B72F7F2EA80CFF63FE99FEE3F130" bold="true" box="[199,315,261,287]" pageId="17" pageNumber="52">Remarks.</emphasis>
|
||
This new species shares the characters of the genus
|
||
<taxonomicName id="4C3010BE7F2EA80CFC06FE98FBBFF131" authorityName="Deuve" authorityYear="2013" box="[930,1127,260,286]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944B72F7F2EA80CFC06FE98FBBFF131" box="[930,1127,260,286]" italics="true" pageId="17" pageNumber="52">Tonkinaphaenops</emphasis>
|
||
</taxonomicName>
|
||
: protibiae without anterior grooves, mentum and submentum completely fused.
|
||
</paragraph>
|
||
<paragraph id="8B8F6B3D7F2EA80CFF63FED1FF28F235" blockId="17.[151,1437,152,575]" pageId="17" pageNumber="52">
|
||
From
|
||
<taxonomicName id="4C3010BE7F2EA80CFEAAFED1FE52F149" authorityName="Deuve" authorityYear="2013" box="[270,394,333,358]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="marinae">
|
||
<emphasis id="B944B72F7F2EA80CFEAAFED1FE52F149" box="[270,394,333,358]" italics="true" pageId="17" pageNumber="52">T. marinae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3010BE7F2EA80CFE6CFED1FDB6F149" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[456,622,333,358]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="yinquanicus">
|
||
<emphasis id="B944B72F7F2EA80CFE6CFED1FDB6F149" box="[456,622,333,358]" italics="true" pageId="17" pageNumber="52">T. yinquanicus</emphasis>
|
||
</taxonomicName>
|
||
, this species differs in its greater size, less convex, longer and nearly parallel elytra, its head with only a few setae whereas pubescent in
|
||
<taxonomicName id="4C3010BE7F2EA80CFC35FEEDFBD2F1A5" authorityName="Deuve" authorityYear="2013" box="[913,1034,369,394]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="marinae">
|
||
<emphasis id="B944B72F7F2EA80CFC35FEEDFBD2F1A5" box="[913,1034,369,394]" italics="true" pageId="17" pageNumber="52">T. marinae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3010BE7F2EA80CFBE6FEEDFB3DF1A5" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[1090,1253,369,394]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="yinquanicus">
|
||
<emphasis id="B944B72F7F2EA80CFBE6FEEDFB3DF1A5" box="[1090,1253,369,394]" italics="true" pageId="17" pageNumber="52">T. yinquanicus</emphasis>
|
||
</taxonomicName>
|
||
, and depression along the suture in the first quarter of the elytra. Punctation of elytra less marked, with striae reaching nearly the apex, whereas the striae are ending at the third of the elytra in
|
||
<emphasis id="B944B72F7F2EA80CFCF9FE25FB6CF1FD" box="[861,1204,440,466]" italics="true" pageId="17" pageNumber="52">
|
||
<taxonomicName id="4C3010BE7F2EA80CFCF9FE25FC0FF1FD" authorityName="Deuve" authorityYear="2013" box="[861,983,441,466]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="marinae">T. marinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3010BE7F2EA80CFBB5FE25FB6CF1FD" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[1041,1204,441,466]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="yinquanicus">T. yinquanicus</taxonomicName>
|
||
</emphasis>
|
||
. Finally, it differs at the first glance from
|
||
<taxonomicName id="4C3010BE7F2EA80CFE24FE41FDD8F1D9" authorityName="Huang & Tú & Phú & Tian & Faille" authorityYear="2019" box="[384,512,477,502]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="52" phylum="Arthropoda" rank="species" species="jingxicus">
|
||
<emphasis id="B944B72F7F2EA80CFE24FE41FDD8F1D9" box="[384,512,477,502]" italics="true" pageId="17" pageNumber="52">T. jingxicus</emphasis>
|
||
</taxonomicName>
|
||
by the shape of elytra, parallel with prominent shoulders whereas ovoid in this last species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A38B67F2EA80CFF63FDB8FAB0F210" box="[199,1384,548,575]" pageId="17" pageNumber="52" type="distribution">
|
||
<paragraph id="8B8F6B3D7F2EA80CFF63FDB8FAB0F210" blockId="17.[151,1437,152,575]" box="[199,1384,548,575]" pageId="17" pageNumber="52">
|
||
<emphasis id="B944B72F7F2EA80CFF63FDB8FEB9F210" bold="true" box="[199,353,548,575]" pageId="17" pageNumber="52">Distribution.</emphasis>
|
||
Known only from the four
|
||
<typeStatus id="548BD59F7F2EA80CFD32FDB9FD1EF210" box="[662,710,549,575]" pageId="17" pageNumber="52">type</typeStatus>
|
||
specimens collected in Hang Dõi cave, Đồng Loan (
|
||
<figureCitation id="130B77B87F2EA80CFAB2FDB8FA8FF210" box="[1302,1367,548,575]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1826]" captionTargetBox="[159,1428,877,1779]" captionTargetId="figure@1.[151,1436,877,1781]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Tonkinaphaenops Deuve, 2013" figureDoi="http://doi.org/10.5281/zenodo.3557805" httpUri="https://zenodo.org/record/3557805/files/figure.png" pageId="17" pageNumber="52">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |