258 lines
32 KiB
XML
258 lines
32 KiB
XML
<document id="7885FD356699C4CCD3B4428085ED2B34" ID-DOI="10.5281/zenodo.183532" ID-GBIF-Dataset="90941c20-aece-4da0-b4bf-19e800996840" ID-ISSN="1175-5326" ID-Zenodo-Dep="183532" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920883928" checkinUser="plazi" docAuthor="Short, Andrew Edward Z." docDate="2008" docId="03993C2AFFAE3204B8ACFCB2FD005A7C" docLanguage="en" docName="zt01855p055.pdf" docOrigin="Zootaxa 1855" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tritonus complanatus Short, 2008, sp.n." docType="treatment" docVersion="7" lastPageNumber="48" masterDocId="FFA04452FFAA3203B83BFF8DFFC45B06" masterDocTitle="Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)" masterLastPageNumber="55" masterPageNumber="41" pageNumber="45" updateTime="1698229214647" updateUser="plazi">
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<mods:titleInfo id="7B5D8D5BBE5F13E309B117D4C322CE77">
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<mods:title id="9F6C22A0BC160B089378DCCFA07D59D3">Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="CFC8A762C5FD5878FC65FE654A679AEA" type="personal">
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<mods:roleTerm id="4E0EB21F004BE9161451983C215921E5">Author</mods:roleTerm>
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<mods:namePart id="2D3E5A3E9FCEEA811E410690CB031A52">Short, Andrew Edward Z.</mods:namePart>
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<mods:typeOfResource id="EB453DD840C4759F697774923107AF6F">text</mods:typeOfResource>
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<mods:title id="6274F579E67E5554FBC9EB63BF6C7982">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A534B3CF16ED4E3E3BAFB5C76EC1BAF3">
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<mods:date id="A846A516CC5C25EE4566400563DA4CC3">2008</mods:date>
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<mods:detail id="57C198F9B2B5F5F244E73D696B24BA0B" type="volume">
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<mods:number id="C6A0B5F71487A64AC4DC88DEC48A7D00">1855</mods:number>
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<mods:start id="B0828EF6B64E2330B9AD27389F27AC87">41</mods:start>
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<mods:end id="35405CAB7DC37E8F0FBC972CA87E8E2A">55</mods:end>
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<mods:classification id="C82492B8F5321E41DDFB7F0D3EA4EB92">journal article</mods:classification>
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<mods:identifier id="37459A42AE9DC737A3AFDCE54307EACC" type="DOI">10.5281/zenodo.183532</mods:identifier>
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<mods:identifier id="C5B2A5CFDE8C17834C21CFBF588548E3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5F8A216A1EA34ACBB6557CE847FC3BEC" type="Zenodo-Dep">183532</mods:identifier>
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<treatment id="03993C2AFFAE3204B8ACFCB2FD005A7C" ID-DOI="http://doi.org/10.5281/zenodo.6232621" ID-GBIF-Taxon="119373675" ID-Zenodo-Dep="6232621" LSID="urn:lsid:plazi:treatment:03993C2AFFAE3204B8ACFCB2FD005A7C" httpUri="http://treatment.plazi.org/id/03993C2AFFAE3204B8ACFCB2FD005A7C" lastPageId="7" lastPageNumber="48" pageId="4" pageNumber="45">
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<subSubSection id="C32ADEB7FFAE3207B8ACFCB2FDD4587A" pageId="4" pageNumber="45" type="nomenclature">
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<paragraph id="8B8F8D3CFFAE3207B8ACFCB2FE26585F" blockId="4.[151,528,831,892]" box="[151,482,831,857]" pageId="4" pageNumber="45">
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<heading id="D0C73A50FFAE3207B8ACFCB2FE26585F" bold="true" box="[151,482,831,857]" fontSize="11" level="1" pageId="4" pageNumber="45" reason="1">
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<emphasis id="B944512EFFAE3207B8ACFCB2FE26585F" bold="true" box="[151,482,831,857]" pageId="4" pageNumber="45">
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<taxonomicName id="4C30F6BFFFAE3207B8ACFCB2FE64585F" ID-CoL="595BT" box="[151,416,831,857]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="complanatus" status="sp. nov.">
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<emphasis id="B944512EFFAE3207B8ACFCB2FE64585F" bold="true" box="[151,416,831,857]" italics="true" pageId="4" pageNumber="45">Tritonus complanatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277EC55FFAE3207B99CFCB2FE26585F" box="[423,482,831,857]" pageId="4" pageNumber="45" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B8F8D3CFFAE3207B8ACFCEFFDD4587A" blockId="4.[151,528,831,892]" box="[151,528,866,892]" pageId="4" pageNumber="45">
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(
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<figureCitation id="130B91B9FFAE3207B8A4FCEFFED4587A" box="[159,272,866,892]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 3" captionStartId-0="5.[151,255,1947,1971]" captionStartId-1="6.[151,269,1008,1032]" captionTargetBox-0="[187,1400,208,1905]" captionTargetBox-1="[175,1411,194,983]" captionTargetId-0="figure@5.[175,1411,197,1921]" captionTargetId-1="figure@6.[175,1411,194,984]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 1. Tritonus complanatus paratype, dorsal and lateral habitus." captionText-1="FIGURES 2 – 3. SEMs of head, dorsal and ventral views; 2) T. complanatus; 3) T. riambavy. Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/183533/files/figure.png" httpUri-1="https://zenodo.org/record/183534/files/figure.png" pageId="4" pageNumber="45">Figs. 1–2</figureCitation>
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,
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<figureCitation id="130B91B9FFAE3207B927FCEFFEE9587A" box="[284,301,866,892]" captionStart="FIGURES 4 – 5" captionStartId="7.[151,269,1587,1611]" captionTargetBox="[151,1435,418,1568]" captionTargetId="figure@7.[151,1436,414,1569]" captionTargetPageId="7" captionText="FIGURES 4 – 5. SEMs of elytra; 4) T. complanatus; 5) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183535/files/figure.png" pageId="4" pageNumber="45">4</figureCitation>
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,
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<figureCitation id="130B91B9FFAE3207B902FCEFFEAC587A" box="[313,360,866,892]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="4" pageNumber="45">6–7</figureCitation>
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,
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<figureCitation id="130B91B9FFAE3207B94FFCEFFE75587A" box="[372,433,866,892]" captionStart="FIGURES 9 – 10" captionStartId="9.[151,269,1552,1576]" captionTargetBox="[151,1435,418,1526]" captionTargetId="figure@9.[151,1436,416,1528]" captionTargetPageId="9" captionText="FIGURES 9 – 10. SEMs of stridulatory apparatus in T. complanatus; 9) laterosternite 3; 10) interior of elytra showing plectrum. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183537/files/figure.png" pageId="4" pageNumber="45">9–10</figureCitation>
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,
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||
<figureCitation id="130B91B9FFAE3207B986FCEFFE18587A" box="[445,476,866,892]" captionStart="FIGURES 13 – 14" captionStartId="10.[151,269,1627,1651]" captionTargetBox="[151,1435,1278,1602]" captionTargetId="figure@10.[151,1436,1278,1603]" captionTargetPageId="10" captionText="FIGURES 13 – 14. Details of Tritonus spp.; 13) T. complanatus, metafemora, scale bar = 1 mm; 14) T. riambavy, maxillary palpus, scale bar = 0.20 mm." httpUri="https://zenodo.org/record/183539/files/figure.png" pageId="4" pageNumber="45">13</figureCitation>
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,
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<figureCitation id="130B91B9FFAE3207B9D2FCEFFDC3587A" box="[489,519,866,892]" captionStart="FIGURES 15 – 21" captionStartId="11.[151,269,1696,1720]" captionTargetBox="[242,1353,216,1672]" captionTargetId="figure@11.[234,1353,194,1672]" captionTargetPageId="11" captionText="FIGURES 15 – 21. Aedeagus of Tritonus spp., dorsal view; 15) T. riana; 16) T. steineri; 17) T. riambavy; 18) T. cribratus; 19) T. madagascarensis; 20) T. crenulatus; 21) T. complanatus. Scale bar = 0.45 mm." httpUri="https://zenodo.org/record/183540/files/figure.png" pageId="4" pageNumber="45">21</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32ADEB7FFAE3207B8ACFC3FFD9B5EFA" pageId="4" pageNumber="45" type="materials_examined">
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<paragraph id="8B8F8D3CFFAE3207B8ACFC3FFD9B5EFA" blockId="4.[151,1437,946,2012]" pageId="4" pageNumber="45">
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<emphasis id="B944512EFFAE3207B8ACFC3FFE0A58CA" bold="true" box="[151,462,946,972]" pageId="4" pageNumber="45">
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<typeStatus id="548B339EFFAE3207B8ACFC3FFF1658CA" box="[151,210,946,972]" pageId="4" pageNumber="45">Type</typeStatus>
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Material.
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<typeStatus id="548B339EFFAE3207B967FC3FFE0A58CA" box="[348,462,946,972]" pageId="4" pageNumber="45" type="holotype">Holotype</typeStatus>
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</emphasis>
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||
(male): “
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||
<collectingCountry id="F327CDACFFAE3207BA7AFC3FFCD158CA" box="[577,789,946,972]" name="Madagascar" pageId="4" pageNumber="45">MADAGASCAR</collectingCountry>
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(144): Antsiranana/ Andapa: R. Masiaposa (Crossing R.N./ 3b at km 5–6) about & km upstr. Road/ bridge,
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<quantity id="4CC820D9FFAE3207BB2CFC57FCA158F2" box="[791,869,986,1012]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="45" unit="m" value="700.0">700 m</quantity>
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,
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<date id="FF8EABFCFFAE3207BB49FC57FC2A58F2" box="[882,1006,986,1012]" pageId="4" pageNumber="45" value="2001-11-10">10.xi.2001</date>
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/ Gerecke & Goldschmidt” (
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<collectionCode id="ED2115F9FFAE3207BD05FC57FA5458F2" box="[1342,1424,986,1012]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
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).
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<emphasis id="B944512EFFAE3207B8ACFB8FFD855F1A" bold="true" box="[151,577,1026,1052]" pageId="4" pageNumber="45">
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<typeStatus id="548B339EFFAE3207B8ACFB8FFED05F1A" box="[151,276,1026,1052]" pageId="4" pageNumber="45" type="paratype">Paratypes</typeStatus>
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(47):
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<collectingCountry id="F327CDACFFAE3207B95BFB8FFDF85F1A" box="[352,572,1026,1052]" name="Madagascar" pageId="4" pageNumber="45">MADAGASCAR</collectingCountry>
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:
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||
</emphasis>
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||
Antananarivo Province: Manakambahiny,
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||
<date id="FF8EABFCFFAE3207BC05FB8FFB7C5F1A" box="[1086,1208,1026,1052]" pageId="4" pageNumber="45" value="1990-02-15">15.ii.1990</date>
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, W.E. Steiner, leg., “in seepage over exposed granite cliff face” (3,
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<collectionCode id="ED2115F9FFAE3207BAF6FBA7FCEE5F42" box="[717,810,1066,1092]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="45">NMNH</collectionCode>
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,
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||
<collectionCode id="ED2115F9FFAE3207BB0DFBA7FCBF5F42" LSID="urn:lsid:biocol.org:col:33791" box="[822,891,1066,1092]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="45">MCZ</collectionCode>
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||
); Antsiranana Province: Andapa, right affl. R. Ambendrana, downstream of large cascade,
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||
<quantity id="4CC820D9FFAE3207BAA4FBDFFD215F6A" box="[671,741,1106,1132]" metricMagnitude="2" metricUnit="m" metricValue="6.6" pageId="4" pageNumber="45" unit="m" value="660.0">660m</quantity>
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,
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<date id="FF8EABFCFFAE3207BACAFBDFFCB45F6A" box="[753,880,1106,1132]" pageId="4" pageNumber="45" value="2002-11-11">11.xi.2002</date>
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, Gerecke & Goldschmidt, leg., 147 (4,
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<collectionCode id="ED2115F9FFAE3207BD04FBDFFA545F6A" box="[1343,1424,1106,1132]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
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); Andapa, “R. Masiaposa (Crossing R. N. 3b at km 5–6) about & km upstream road bridge”,
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<quantity id="4CC820D9FFAE3207BCFEFBF7FACF5F92" box="[1221,1291,1146,1172]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="45" unit="m" value="700.0">700m</quantity>
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,
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<date id="FF8EABFCFFAE3207BD2CFBF7FA535F92" box="[1303,1431,1146,1172]" pageId="4" pageNumber="45" value="2001-11-10">10.xi.2001</date>
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, Gerecke & Goldschmidt, leg., 144 (14,
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<collectionCode id="ED2115F9FFAE3207BA5FFB2FFD645FBA" box="[612,672,1186,1212]" pageId="4" pageNumber="45">FHC</collectionCode>
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,
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<collectionCode id="ED2115F9FFAE3207BA90FB2FFCC45FBA" box="[683,768,1186,1212]" pageId="4" pageNumber="45">KSEM</collectionCode>
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,
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<collectionCode id="ED2115F9FFAE3207BB30FB2FFC935FBA" box="[779,855,1186,1212]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
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,
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||
<collectionCode id="ED2115F9FFAE3207BB59FB2FFC7F5FBA" box="[866,955,1186,1212]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="45">NMPC</collectionCode>
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); Mt. D’Ambre Forest Reserve, stream at Grand Cascade,
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<quantity id="4CC820D9FFAE3207B96DFB47FE5F5FE2" box="[342,411,1226,1252]" metricMagnitude="2" metricUnit="m" metricValue="6.17" pageId="4" pageNumber="45" unit="m" value="617.0">617m</quantity>
|
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,
|
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<date id="FF8EABFCFFAE3207B99CFB47FDE35FE2" box="[423,551,1226,1252]" pageId="4" pageNumber="45" value="1986-11-13">13.xi.1986</date>
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, J.T. & D.A. Polhemus, CL2278 (1,
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<collectionCode id="ED2115F9FFAE3207BBF4FB47FBC95FE2" LSID="urn:lsid:biocol.org:col:34700" box="[975,1037,1226,1252]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="4" pageNumber="45">CAS</collectionCode>
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); Antsabe,
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<geoCoordinate id="EE04EBFBFFAE3207BCB5FB47FAC95FE2" box="[1166,1293,1226,1252]" direction="south" orientation="latitude" pageId="4" pageNumber="45" precision="5" value="-13.6093">13.6093°S</geoCoordinate>
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,
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<geoCoordinate id="EE04EBFBFFAE3207BD23FB47FA535FE2" box="[1304,1431,1226,1252]" direction="east" orientation="longitude" pageId="4" pageNumber="45" precision="5" value="48.7212">48.7212°E</geoCoordinate>
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,
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<date id="FF8EABFCFFAE3207B8ACFB7FFED35E0A" box="[151,279,1266,1292]" pageId="4" pageNumber="45" value="2004-11-23">23.xi.2004</date>
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, M. Balke, leg. (9,
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<collectionCode id="ED2115F9FFAE3207B9C0FB7FFD8B5E0A" box="[507,591,1266,1292]" pageId="4" pageNumber="45">KSEM</collectionCode>
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,
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<collectionCode id="ED2115F9FFAE3207BA67FB7FFD7C5E0A" box="[604,696,1266,1292]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="45">NHML</collectionCode>
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); Fianarantsoa Province: Andrambouato, large cascade
|
||
<quantity id="4CC820D9FFAE3207BD7DFB7FFABF5E0A" box="[1350,1403,1266,1292]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="45" unit="km" value="3.0">3km</quantity>
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E. of village,
|
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<quantity id="4CC820D9FFAE3207B921FA97FEA45E32" box="[282,352,1306,1332]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="4" pageNumber="45" unit="m" value="820.0">820m</quantity>
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,
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<date id="FF8EABFCFFAE3207B954FA97FE3A5E32" box="[367,510,1306,1332]" pageId="4" pageNumber="45" value="2001-08-20">20.viii.2001</date>
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, Gerecke & Goldschmidt, leg., 0 37 (1,
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<collectionCode id="ED2115F9FFAE3207BBE4FA97FBEB5E32" box="[991,1071,1306,1332]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="45">NMW</collectionCode>
|
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); Abohimahasoa, village hotel Riandrano, alt.
|
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<quantity id="4CC820D9FFAE3207B96BFACFFE615E5A" box="[336,421,1346,1372]" metricMagnitude="3" metricUnit="m" metricValue="1.208" pageId="4" pageNumber="45" unit="m" value="1208.0">1208m</quantity>
|
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,
|
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<geoCoordinate id="EE04EBFBFFAE3207B989FACFFDFC5E5A" box="[434,568,1346,1372]" direction="south" orientation="latitude" pageId="4" pageNumber="45" precision="1" value="-21.103384">21 6.203’S</geoCoordinate>
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,
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<geoCoordinate id="EE04EBFBFFAE3207BA7DFACFFD185E5A" box="[582,732,1346,1372]" direction="east" orientation="longitude" pageId="4" pageNumber="45" precision="1" value="47.2024">47 12.144’E</geoCoordinate>
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,
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<date id="FF8EABFCFFAE3207BAD1FACFFC965E5A" box="[746,850,1346,1372]" pageId="4" pageNumber="45" value="2006-05-06">6.v.2006</date>
|
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, “hygropetric surface next to waterfall”, J. Bergsten, leg., “Loc. P37A” (4,
|
||
<collectionCode id="ED2115F9FFAE3207B9DFFAE7FD845E82" box="[484,576,1386,1412]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="45">NHML</collectionCode>
|
||
); Andringitra National Park,
|
||
<taxonomicName id="4C30F6BFFFAE3207BBA4FAE7FBDC5E82" box="[927,1048,1386,1412]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="54" phylum="Arthropoda" rank="species" species="riambavy">Riambavy</taxonomicName>
|
||
river (Female) cascade,
|
||
<quantity id="4CC820D9FFAE3207BD78FAE7FA5C5E82" box="[1347,1432,1386,1412]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="4" pageNumber="45" unit="m" value="1739.0">1739m</quantity>
|
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, 8.c.2006,
|
||
<geoCoordinate id="EE04EBFBFFAE3207B930FA1FFE4B5EAA" box="[267,399,1426,1452]" direction="south" orientation="latitude" pageId="4" pageNumber="45" precision="1" value="-22.146017">22 8.761’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE04EBFBFFAE3207B9A0FA1FFDEB5EAA" box="[411,559,1426,1452]" direction="east" orientation="longitude" pageId="4" pageNumber="45" precision="1" value="46.891582">46 53.495’E</geoCoordinate>
|
||
, “wet rock surfaces next to high waterfall”, J. Bergsten, leg., “Loc. P39A” (1,
|
||
<collectionCode id="ED2115F9FFAE3207B885FA37FEDE5ED2" box="[190,282,1466,1492]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="45">NHML</collectionCode>
|
||
); Tananarive Province: Tananarive, waterfall and seeps nr. Queen’s palace,
|
||
<quantity id="4CC820D9FFAE3207BCA9FA37FB225ED2" box="[1170,1254,1466,1492]" metricMagnitude="3" metricUnit="m" metricValue="1.31" pageId="4" pageNumber="45" unit="m" value="1310.0">1310m</quantity>
|
||
,
|
||
<date id="FF8EABFCFFAE3207BCC8FA37FAAF5ED2" box="[1267,1387,1466,1492]" pageId="4" pageNumber="45" value="1986-10-24">24.x.1986</date>
|
||
, J.T & D.A. Polhemus, CL2233 (10,
|
||
<collectionCode id="ED2115F9FFAE3207BA2EFA6FFD975EFA" LSID="urn:lsid:biocol.org:col:34700" box="[533,595,1506,1532]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="4" pageNumber="45">CAS</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ADEB7FFAE3207B8FEF987FB685D72" pageId="4" pageNumber="45" type="diagnosis">
|
||
<paragraph id="8B8F8D3CFFAE3207B8FEF987FB685D72" blockId="4.[151,1437,946,2012]" pageId="4" pageNumber="45">
|
||
<emphasis id="B944512EFFAE3207B8FEF987FE815D22" bold="true" box="[197,325,1546,1572]" pageId="4" pageNumber="45">Diagnosis.</emphasis>
|
||
The large size and broadly oval, nearly circular body form serve to easily distinguish this species from all other members of the genus.
|
||
<taxonomicName id="4C30F6BFFFAE3207BAB4F9BEFCB15D4A" box="[655,885,1587,1612]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="crenulatus">
|
||
<emphasis id="B944512EFFAE3207BAB4F9BEFCB15D4A" box="[655,885,1587,1612]" italics="true" pageId="4" pageNumber="45">Tritonus crenulatus</emphasis>
|
||
</taxonomicName>
|
||
is somewhat similar in shape, but is distinctly smaller and possesses strongly crenulate anterior margins of abdominal ventrites 2 and 3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ADEB7FFAE3205B8FEF90FFF245CC3" lastPageId="6" lastPageNumber="47" pageId="4" pageNumber="45" type="description">
|
||
<paragraph id="8B8F8D3CFFAE3205B8FEF90FFCC35D0B" blockId="4.[151,1437,946,2012]" lastBlockId="6.[151,1436,1083,1989]" lastPageId="6" lastPageNumber="47" pageId="4" pageNumber="45">
|
||
<emphasis id="B944512EFFAE3207B8FEF90FFE995D9A" bold="true" box="[197,349,1666,1692]" pageId="4" pageNumber="45">Description.</emphasis>
|
||
Total length 8.2–9.0 mm.
|
||
<emphasis id="B944512EFFAE3207BAA1F90FFD245D9A" bold="true" box="[666,736,1666,1692]" pageId="4" pageNumber="45">Color</emphasis>
|
||
. Dorsum black with general shagreen (
|
||
<figureCitation id="130B91B9FFAE3207BC8FF90FFB3D5D9A" box="[1204,1273,1666,1692]" captionStart="FIGURE 1" captionStartId="5.[151,255,1947,1971]" captionTargetBox="[187,1400,208,1905]" captionTargetId="figure@5.[175,1411,197,1921]" captionTargetPageId="5" captionText="FIGURE 1. Tritonus complanatus paratype, dorsal and lateral habitus." httpUri="https://zenodo.org/record/183533/files/figure.png" pageId="4" pageNumber="45">Fig. 1</figureCitation>
|
||
); most specimens lacking iridescent patches on elytra, with a minority of specimens possess distinct green iridescent spots laterally.
|
||
<emphasis id="B944512EFFAE3207B93FF95FFE8A5DEA" bold="true" box="[260,334,1746,1772]" pageId="4" pageNumber="45">Head.</emphasis>
|
||
Ground punctation fine to moderately coarse; irregular in size (
|
||
<figureCitation id="130B91B9FFAE3207BC79F95FFB4F5DEA" box="[1090,1163,1746,1772]" captionStart="FIGURES 2 – 3" captionStartId="6.[151,269,1008,1032]" captionTargetBox="[175,1411,194,983]" captionTargetId="figure@6.[175,1411,194,984]" captionTargetPageId="6" captionText="FIGURES 2 – 3. SEMs of head, dorsal and ventral views; 2) T. complanatus; 3) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183534/files/figure.png" pageId="4" pageNumber="45">Fig. 2</figureCitation>
|
||
). Systematic punctures distinct; 3–4× as large as largest surrounding ground punctures; on clypeus bearing short decumbent setae; those on frons anterior to each eye relatively long and erect. Antennae with club subequal in length to antennomeres 2–6 combined. Maxillary palpi short, apical palpomere ca. 1.6× length of penultimate palpomere; penultimate palpomere distinctly shorter than segment 2. Mentum with anterior margin slightly convex; anterior half with fine transverse microsculpture; entire mentum with fine to moderately coarse, irregularly scattered punctures. Anterior margin of head very wide, quadrate in shape.
|
||
<emphasis id="B944512EFFAE3207BBE0F84FFBFB5CDA" bold="true" box="[987,1087,1986,2012]" pageId="4" pageNumber="45">Thorax.</emphasis>
|
||
Ground punctation on pronotum and elytra extremely fine, not detectable in some specimens. Pronotal and elytral systematic punctures large and distinct; bearing short setae. Ten rows of fine serial punctures are present in most specimens examined, which appear distinctly larger than ground punctation but less than half the size of the systematic punctures. Prosternum (
|
||
<figureCitation id="130B91B9FFAC3205B988FB3EFE3F5FCB" box="[435,507,1203,1229]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="6" pageNumber="47">Fig. 7</figureCitation>
|
||
) slightly tectiform with a sharp, narrow and glabrous median carina; not distinctly elevated or dilated at anterior margin. Mesoventrite (
|
||
<figureCitation id="130B91B9FFAC3205BB5EFB56FC695FF3" box="[869,941,1243,1269]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="6" pageNumber="47">Fig. 6</figureCitation>
|
||
) elevated medially into a distinct narrow carina; surface of carina narrowly glabrous. Carina slightly depressed, appearing almost concave in lateral view. Metasternum with large elevated glabrous region that is slightly less than half as wide as long. Metafemora with moderately impressed microsculpture over most of ventral surface; proximally in the form of tiny punctures and blending distally to form of fine transverse wavy lines.
|
||
<emphasis id="B944512EFFAC3205BBCCFAF7FBBC5E92" bold="true" box="[1015,1144,1402,1428]" pageId="6" pageNumber="47">Abdomen.</emphasis>
|
||
Posteromedial glabrous region of ventrite 1 reduced, almost appearing absent. Glabrous region of ventrite 2 semi-circular to subtriangular in shape; ca. 0.3 the width of the ventrite. Anterior margins of ventrites 1–3 not crenulate, but with a loosely wrinkled appearance. Aedeagus as in
|
||
<figureCitation id="130B91B9FFAC3205BA96FA7EFCC75D0B" box="[685,771,1523,1549]" captionStart="FIGURES 15 – 21" captionStartId="11.[151,269,1696,1720]" captionTargetBox="[242,1353,216,1672]" captionTargetId="figure@11.[234,1353,194,1672]" captionTargetPageId="11" captionText="FIGURES 15 – 21. Aedeagus of Tritonus spp., dorsal view; 15) T. riana; 16) T. steineri; 17) T. riambavy; 18) T. cribratus; 19) T. madagascarensis; 20) T. crenulatus; 21) T. complanatus. Scale bar = 0.45 mm." httpUri="https://zenodo.org/record/183540/files/figure.png" pageId="6" pageNumber="47">Fig. 21</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF4FDDB4FFAF3206B8ACF816FC4E5CB5" box="[151,906,1947,1971]" httpUri="https://zenodo.org/record/183533/files/figure.png" pageId="5" pageNumber="46" targetBox="[187,1400,208,1905]" targetPageId="5">
|
||
<paragraph id="8B8F8D3CFFAF3206B8ACF816FC4E5CB5" blockId="5.[151,906,1947,1971]" box="[151,906,1947,1971]" pageId="5" pageNumber="46">
|
||
<emphasis id="B944512EFFAF3206B8ACF816FEDD5CB5" bold="true" box="[151,281,1947,1971]" pageId="5" pageNumber="46">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C30F6BFFFAF3206B91BF811FDC35CB5" box="[288,519,1948,1971]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="complanatus">
|
||
<emphasis id="B944512EFFAF3206B91BF811FDC35CB5" box="[288,519,1948,1971]" italics="true" pageId="5" pageNumber="46">Tritonus complanatus</emphasis>
|
||
</taxonomicName>
|
||
paratype, dorsal and lateral habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4FDDB4FFAC3205B8ACFC7DFAFC5F01" box="[151,1336,1008,1032]" httpUri="https://zenodo.org/record/183534/files/figure.png" pageId="6" pageNumber="47" targetBox="[175,1411,194,983]" targetPageId="6">
|
||
<paragraph id="8B8F8D3CFFAC3205B8ACFC7DFAFC5F01" blockId="6.[151,1336,1008,1032]" box="[151,1336,1008,1032]" pageId="6" pageNumber="47">
|
||
<emphasis id="B944512EFFAC3205B8ACFC7DFE875F0E" bold="true" box="[151,323,1008,1032]" pageId="6" pageNumber="47">FIGURES 2–3.</emphasis>
|
||
SEMs of head, dorsal and ventral views; 2)
|
||
<taxonomicName id="4C30F6BFFFAC3205BB27FC7DFC7A5F01" box="[796,958,1008,1031]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="complanatus">
|
||
<emphasis id="B944512EFFAC3205BB27FC7DFC7A5F01" box="[796,958,1008,1031]" italics="true" pageId="6" pageNumber="47">T. complanatus</emphasis>
|
||
</taxonomicName>
|
||
; 3)
|
||
<taxonomicName id="4C30F6BFFFAC3205BBD1FC7DFBA35F01" box="[1002,1127,1008,1031]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="riambavy">
|
||
<emphasis id="B944512EFFAC3205BBD1FC7DFBA35F01" box="[1002,1127,1008,1031]" italics="true" pageId="6" pageNumber="47">T. riambavy</emphasis>
|
||
</taxonomicName>
|
||
. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8F8D3CFFAC3205B8FEF997FF245CC3" blockId="6.[151,1436,1083,1989]" pageId="6" pageNumber="47">
|
||
<emphasis id="B944512EFFAC3205B8FEF997FEFB5D32" bold="true" box="[197,319,1562,1588]" pageId="6" pageNumber="47">Variation.</emphasis>
|
||
Many specimens of
|
||
<taxonomicName id="4C30F6BFFFAC3205BA01F996FD295D32" box="[570,749,1563,1588]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="complanatus">
|
||
<emphasis id="B944512EFFAC3205BA01F996FD295D32" box="[570,749,1563,1588]" italics="true" pageId="6" pageNumber="47">T. complanatus</emphasis>
|
||
</taxonomicName>
|
||
exhibit the normal shiny black and smooth dorsal appearance, which is similar to all other known species in the genus (and most members of the
|
||
<taxonomicName id="4C30F6BFFFAC3205BCEBF9CEFAF95D5A" box="[1232,1341,1603,1628]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B944512EFFAC3205BCEBF9CEFAF95D5A" box="[1232,1341,1603,1628]" italics="true" pageId="6" pageNumber="47">Oocyclus</emphasis>
|
||
</taxonomicName>
|
||
-group). However, there are a number of specimens from several collecting events in which the dorsal surface appears dull and brownish. When viewed under a microscope, the surface of these specimens is rough and coarse; the surface of “normal” specimens is smooth. I originally interpreted this difference to indicate these were two species; however, further examination of other characters, including the genitalia did not reveal any substantive differences. Even more telling, a few specimens also appear to have patches of both
|
||
<typeStatus id="548B339EFFAC3205BC99F886FB245C23" box="[1186,1248,1803,1829]" pageId="6" pageNumber="47">types</typeStatus>
|
||
of dorsal sculpture, leading me to conclude the rough-surfaced specimens represent some aberration, or are possibly caused by physical and/or chemical abrasion. I suspect the latter may be the cause because the entire surface appears to be effaced, such that the ground punctation is no longer visible and the systematic punctures appear shallower.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ADEB7FFAD3204B8FEFF1AFE605BDC" pageId="7" pageNumber="48" type="etymology">
|
||
<paragraph id="8B8F8D3CFFAD3204B8FEFF1AFE605BDC" blockId="7.[151,1436,151,378]" pageId="7" pageNumber="48">
|
||
<emphasis id="B944512EFFAD3204B8FEFF1AFE965BB7" bold="true" box="[197,338,151,177]" pageId="7" pageNumber="48">Etymology.</emphasis>
|
||
Named for the broadly explanate lateral margins of the elytra and pronotum, which are distinctive of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ADEB7FFAD3204B8FEFF6AFDC55A04" box="[197,513,231,258]" pageId="7" pageNumber="48" type="distribution">
|
||
<paragraph id="8B8F8D3CFFAD3204B8FEFF6AFDC55A04" blockId="7.[151,1436,151,378]" box="[197,513,231,258]" pageId="7" pageNumber="48">
|
||
<emphasis id="B944512EFFAD3204B8FEFF6AFEA25A07" bold="true" box="[197,358,231,257]" pageId="7" pageNumber="48">Distribution.</emphasis>
|
||
<collectingCountry id="F327CDACFFAD3204B956FF65FE395A04" box="[365,509,232,258]" name="Madagascar" pageId="7" pageNumber="48">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ADEB7FFAD3204B8FEFE82FD005A7C" pageId="7" pageNumber="48" type="discussion">
|
||
<paragraph id="8B8F8D3CFFAD3204B8FEFE82FD005A7C" blockId="7.[151,1436,151,378]" pageId="7" pageNumber="48">
|
||
<emphasis id="B944512EFFAD3204B8FEFE82FEFA5A2F" bold="true" box="[197,318,271,297]" pageId="7" pageNumber="48">Remarks.</emphasis>
|
||
This species is the most distinctive member of the genus and apparently the most frequently encountered.
|
||
<taxonomicName id="4C30F6BFFFAD3204B90FFEB5FDF75A57" box="[308,563,312,337]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="complanatus">
|
||
<emphasis id="B944512EFFAD3204B90FFEB5FDF75A57" box="[308,563,312,337]" italics="true" pageId="7" pageNumber="48">Tritonus complanatus</emphasis>
|
||
</taxonomicName>
|
||
has been collected in many regions of
|
||
<collectingCountry id="F327CDACFFAD3204BBC7FEB5FB485A54" box="[1020,1164,312,338]" name="Madagascar" pageId="7" pageNumber="48">Madagascar</collectingCountry>
|
||
, and accounts for more than half all the known specimens of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |