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<document id="EDABE9B5D248CFC7CA6648E7F6746D45" ID-DOI="10.5281/zenodo.183532" ID-GBIF-Dataset="90941c20-aece-4da0-b4bf-19e800996840" ID-ISSN="1175-5326" ID-Zenodo-Dep="183532" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920883928" checkinUser="plazi" docAuthor="Short, Andrew Edward Z." docDate="2008" docId="03993C2AFFA83200B8ACFA92FC015E3C" docLanguage="en" docName="zt01855p055.pdf" docOrigin="Zootaxa 1855" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tritonus Mulsant" docType="treatment" docVersion="7" lastPageNumber="44" masterDocId="FFA04452FFAA3203B83BFF8DFFC45B06" masterDocTitle="Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)" masterLastPageNumber="55" masterPageNumber="41" pageNumber="43" updateTime="1698229214647" updateUser="plazi">
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<mods:title id="8ED183FFD2A1DE9BC86F60D14F1D4F4A">Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)</mods:title>
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<mods:namePart id="53E6CCD0F587080FA75AE29AF419CC47">Short, Andrew Edward Z.</mods:namePart>
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<mods:date id="6562FDEE56959F411C208B910AED4666">2008</mods:date>
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<mods:identifier id="9574706CBD4DA8E5CE2B7B1823DBC855" type="DOI">10.5281/zenodo.183532</mods:identifier>
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<treatment id="03993C2AFFA83200B8ACFA92FC015E3C" ID-DOI="http://doi.org/10.5281/zenodo.6232617" ID-GBIF-Taxon="119373674" ID-Zenodo-Dep="6232617" LSID="urn:lsid:plazi:treatment:03993C2AFFA83200B8ACFA92FC015E3C" httpUri="http://treatment.plazi.org/id/03993C2AFFA83200B8ACFA92FC015E3C" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
<subSubSection id="C32ADEB7FFA83201B8ACFA92FEA95E3F" box="[151,365,1311,1337]" pageId="2" pageNumber="43" type="nomenclature">
<paragraph id="8B8F8D3CFFA83201B8ACFA92FEA95E3F" blockId="2.[151,365,1311,1337]" box="[151,365,1311,1337]" pageId="2" pageNumber="43">
<heading id="D0C73A50FFA83201B8ACFA92FEA95E3F" bold="true" box="[151,365,1311,1337]" fontSize="11" level="1" pageId="2" pageNumber="43" reason="1">
<taxonomicName id="4C30F6BFFFA83201B8ACFA92FEA95E3F" ID-CoL="8X7XK" authority="Mulsant" authorityName="Mulsant" box="[151,365,1311,1337]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA83201B8ACFA92FEA95E3F" bold="true" box="[151,365,1311,1337]" pageId="2" pageNumber="43">
<emphasis id="B944512EFFA83201B8ACFA92FF3A5E3F" bold="true" box="[151,254,1311,1337]" italics="true" pageId="2" pageNumber="43">Tritonus</emphasis>
Mulsant
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA83201B8ACFAE7FD905E87" box="[151,596,1386,1409]" pageId="2" pageNumber="43" type="reference_group">
<paragraph id="8B8F8D3CFFA83201B8ACFAE7FD905E87" blockId="2.[151,596,1386,1409]" box="[151,596,1386,1409]" pageId="2" pageNumber="43">
<treatmentCitationGroup id="AB20AA12FFA83201B8ACFAE7FD905E87" box="[151,596,1386,1409]" pageId="2" pageNumber="43">
<taxonomicName id="4C30F6BFFFA83201B8ACFAE7FE0D5E87" ID-CoL="8X7XK" authority="Mulsant, 1844: 378" authorityName="Mulsant" authorityPageNumber="378" authorityYear="1844" box="[151,457,1386,1409]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA83201B8ACFAE7FF345E87" box="[151,240,1386,1409]" italics="true" pageId="2" pageNumber="43">Tritonus</emphasis>
<treatmentCitation id="0A91AB2DFFA83201B8CDFAE7FE0D5E87" author="Mulsant" box="[246,457,1386,1409]" page="378" pageId="2" pageNumber="43" year="1844">
<bibRefCitation id="EFA1F0CDFFA83201B8CDFAE7FE505E87" author="Mulsant" box="[246,404,1386,1409]" pageId="2" pageNumber="55" refString="Mulsant, E. (1844) Description de quelques Palpicornes inedits. Annals de la Societe d'Agriculture de Lyon, 7, 372 - 382." type="journal article" year="1844">Mulsant, 1844</bibRefCitation>
: 378
</treatmentCitation>
</taxonomicName>
. (Masculine)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA83201B8ACFA3FFC5B5ECA" box="[151,927,1458,1484]" pageId="2" pageNumber="43" type="materials_examined">
<paragraph id="8B8F8D3CFFA83201B8ACFA3FFC5B5ECA" blockId="2.[151,1437,1458,2004]" box="[151,927,1458,1484]" pageId="2" pageNumber="43">
<emphasis id="B944512EFFA83201B8ACFA3FFEFA5ECA" bold="true" box="[151,318,1458,1484]" pageId="2" pageNumber="43">
<typeStatus id="548B339EFFA83201B8ACFA3FFF105ECA" box="[151,212,1458,1484]" pageId="2" pageNumber="43">Type</typeStatus>
Species.
</emphasis>
<taxonomicName id="4C30F6BFFFA83201B97EFA3EFD5E5ECA" authority="Mulsant" authorityName="Mulsant" box="[325,666,1458,1484]" class="Insecta" family="Hydrophilidae" genus="Hydrobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="cribratus">
<emphasis id="B944512EFFA83201B97EFA3EFDF75ECA" box="[325,563,1459,1484]" italics="true" pageId="2" pageNumber="43">Hydrobius cribratus</emphasis>
Mulsant
</taxonomicName>
(original designation)
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA83201B8FEFA57FE565D92" pageId="2" pageNumber="43" type="diagnosis">
<paragraph id="8B8F8D3CFFA83201B8FEFA57FE565D92" blockId="2.[151,1437,1458,2004]" pageId="2" pageNumber="43">
<emphasis id="B944512EFFA83201B8FEFA57FE815EF2" bold="true" box="[197,325,1498,1524]" pageId="2" pageNumber="43">Diagnosis.</emphasis>
Body size large: 6.2–9.0 mm. Within the
<taxonomicName id="4C30F6BFFFA83201BB0FFA57FC705EF2" box="[820,948,1498,1524]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="tribe" tribe="Laccobiini">Laccobiini</taxonomicName>
,
<taxonomicName id="4C30F6BFFFA83201BBFBFA56FBE65EF2" box="[960,1058,1499,1524]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA83201BBFBFA56FBE65EF2" box="[960,1058,1499,1524]" italics="true" pageId="2" pageNumber="43">Tritonus</emphasis>
</taxonomicName>
can be characterized by the following characters: Antennae 9-segmented. Abdomen with five ventrites; large glabrous region on ventrites 2– 5. Head, pronotum and elytra each with distinct systematic punctures. Posterolateral margins of the pronotum not spinose. Prosternum, procoxae, and mesoventrite all without spines. The genus contains the largest known members of the tribe.
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA83200B8FEF92FFD275834" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43" type="description">
<paragraph id="8B8F8D3CFFA83200B8FEF92FFD275834" blockId="2.[151,1437,1458,2004]" lastBlockId="3.[151,1437,152,1338]" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
<emphasis id="B944512EFFA83201B8FEF92FFE995DBA" bold="true" box="[197,349,1698,1724]" pageId="2" pageNumber="43">Description.</emphasis>
Form narrowly to very broadly oval.
<emphasis id="B944512EFFA83201BB2CF92FFCA55DBA" bold="true" box="[791,865,1698,1724]" pageId="2" pageNumber="43">Head.</emphasis>
Systematic punctures present on the lateral margins of the clypeus and frons; absent or undetectable on the labrum. Ground punctation fine to coarse; not uniform in size. Labrum small, often retracted beneath or not extending beyond the anterior margin of the clypeus such that it is not visible from above. Mentum quadrate (
<figureCitation id="130B91B9FFA83201BB7BF897FC505C32" box="[832,916,1818,1844]" captionStart="FIGURES 2 – 3" captionStartId="6.[151,269,1008,1032]" captionTargetBox="[175,1411,194,983]" captionTargetId="figure@6.[175,1411,194,984]" captionTargetPageId="6" captionText="FIGURES 2 – 3. SEMs of head, dorsal and ventral views; 2) T. complanatus; 3) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183534/files/figure.png" pageId="2" pageNumber="43">Figs. 2</figureCitation>
b, 3b), slightly depressed at lateral margins; often with fine to coarse microsculpture consisting of fine transverse microreticulation. Maxillary palps very short, always shorter than width of labrum; apical segment longer than penultimate (
<figureCitation id="130B91B9FFA83201BC4BF8E7FB035C82" box="[1136,1223,1898,1924]" captionStart="FIGURES 13 – 14" captionStartId="10.[151,269,1627,1651]" captionTargetBox="[151,1435,1278,1602]" captionTargetId="figure@10.[151,1436,1278,1603]" captionTargetPageId="10" captionText="FIGURES 13 – 14. Details of Tritonus spp.; 13) T. complanatus, metafemora, scale bar = 1 mm; 14) T. riambavy, maxillary palpus, scale bar = 0.20 mm." httpUri="https://zenodo.org/record/183539/files/figure.png" pageId="2" pageNumber="43">Fig. 14</figureCitation>
). Antennae 9-segmented, including a 3-segmented pubescent club. Dorsal surface of the eyes not bulging, in the same plane as the surface of the head.
<emphasis id="B944512EFFA83201B995F837FDD65CD2" bold="true" box="[430,530,1978,2004]" pageId="2" pageNumber="43">Thorax.</emphasis>
Pronotum with systematic punctures forming anterolateral and posterolateral series; each series forming an irregular arc or row; systematic punctures larger than ground punctation. Posterolateral corners of pronotum rounded, never angulate or spinose. Prosternum (
<figureCitation id="130B91B9FFA93200BC04FF4DFB6B5BDC" box="[1087,1199,192,218]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="3" pageNumber="44">Figs. 7–8</figureCitation>
) with a low median carina which may be fine and without a strong ridge (
<taxonomicName id="4C30F6BFFFA93200BB32FF65FCBF5A07" box="[777,891,232,257]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="steineri">
<emphasis id="B944512EFFA93200BB32FF65FCBF5A07" box="[777,891,232,257]" italics="true" pageId="3" pageNumber="44">T. steineri</emphasis>
</taxonomicName>
), or more commonly, strongly ridged (
<taxonomicName id="4C30F6BFFFA93200BD7EFF65FF385A2F" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="B944512EFFA93200BD7EFF65FF385A2F" italics="true" pageId="3" pageNumber="44">T. complanatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C30F6BFFFA93200B930FE9DFE565A2F" box="[267,402,272,297]" class="Insecta" family="Hydrophilidae" genus="Hydrobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="cribratus">
<emphasis id="B944512EFFA93200B930FE9DFE565A2F" box="[267,402,272,297]" italics="true" pageId="3" pageNumber="44">T. cribratus</emphasis>
</taxonomicName>
). Mesoanepiplerual sutures straight, giving the mesoventrite a diamond-shaped appearance (
<figureCitation id="130B91B9FFA93200B8E6FEB5FEE15A54" box="[221,293,312,338]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="3" pageNumber="44">Fig. 6</figureCitation>
). Ten very fine rows of elytral serial punctures are distinct in
<taxonomicName id="4C30F6BFFFA93200BBC0FEB5FB6A5A57" box="[1019,1198,312,337]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="B944512EFFA93200BBC0FEB5FB6A5A57" box="[1019,1198,312,337]" italics="true" pageId="3" pageNumber="44">T. complanatus</emphasis>
</taxonomicName>
, but not discernable (or absent) in the remaining species, although some rows of coarse punctures may represent serial punctures in addition to systematic punctures. Five rows of elytral systematic punctures present although frequently difficult to distinguish from enlarged serial punctures in some species (e.g.
<taxonomicName id="4C30F6BFFFA93200BBC8FE3DFB625ACF" box="[1011,1190,432,457]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="B944512EFFA93200BBC8FE3DFB625ACF" box="[1011,1190,432,457]" italics="true" pageId="3" pageNumber="44">T. complanatus</emphasis>
</taxonomicName>
). Elytra with sutural stria at least in posterior half (
<figureCitation id="130B91B9FFA93200B9CFFE55FDA25AF4" box="[500,614,472,498]" captionStart="FIGURES 4 – 5" captionStartId="7.[151,269,1587,1611]" captionTargetBox="[151,1435,418,1568]" captionTargetId="figure@7.[151,1436,414,1569]" captionTargetPageId="7" captionText="FIGURES 4 – 5. SEMs of elytra; 4) T. complanatus; 5) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183535/files/figure.png" pageId="3" pageNumber="44">Figs. 4–5</figureCitation>
). Interior of elytra with a small roughened region (=plectrum;
<figureCitation id="130B91B9FFA93200BD04FE55FA575AF4" box="[1343,1427,472,498]" captionStart="FIGURES 9 – 10" captionStartId="9.[151,269,1552,1576]" captionTargetBox="[151,1435,418,1526]" captionTargetId="figure@9.[151,1436,416,1528]" captionTargetPageId="9" captionText="FIGURES 9 – 10. SEMs of stridulatory apparatus in T. complanatus; 9) laterosternite 3; 10) interior of elytra showing plectrum. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183537/files/figure.png" pageId="3" pageNumber="44">Fig. 10</figureCitation>
) along the central margin. Procoxae (
<figureCitation id="130B91B9FFA93200BA7BFD8DFD6A591C" box="[576,686,512,538]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="3" pageNumber="44">Figs. 7–8</figureCitation>
) finely pubescent, without thickened setae or spines.
<collectingCountry id="F327CDACFFA93200BD19FD8DFA99591C" box="[1314,1373,512,538]" name="India" pageId="3" pageNumber="44">Hind</collectingCountry>
femora entirely glabrous (
<figureCitation id="130B91B9FFA93200B9A3FDA5FE2B5944" box="[408,495,552,578]" captionStart="FIGURES 13 – 14" captionStartId="10.[151,269,1627,1651]" captionTargetBox="[151,1435,1278,1602]" captionTargetId="figure@10.[151,1436,1278,1603]" captionTargetPageId="10" captionText="FIGURES 13 – 14. Details of Tritonus spp.; 13) T. complanatus, metafemora, scale bar = 1 mm; 14) T. riambavy, maxillary palpus, scale bar = 0.20 mm." httpUri="https://zenodo.org/record/183539/files/figure.png" pageId="3" pageNumber="44">Fig. 13</figureCitation>
). All tarsi with five segments. Meso- and metatarsi with two rows of ventral spicules and a dorsal row of fine setae.
<emphasis id="B944512EFFA93200BA74FDC2FD15596F" bold="true" box="[591,721,591,617]" pageId="3" pageNumber="44">Abdomen.</emphasis>
Abdomen with five exposed ventrites (
<figureCitation id="130B91B9FFA93200BC92FDDDFAC5596C" box="[1193,1281,592,618]" captionStart="FIGURES 11 – 12" captionStartId="10.[151,269,1197,1221]" captionTargetBox="[151,1435,634,1171]" captionTargetId="figure@10.[151,1436,634,1171]" captionTargetPageId="10" captionText="FIGURES 11 – 12. Abdomen of Tritonus spp.; 11) T. riambavy; 12) T. crenulatus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183538/files/figure.png" pageId="3" pageNumber="44">Fig. 11</figureCitation>
). Abdominal ventrites 2 to 5 each with median glabrous region, varying in width and length; ventrite 1 often with a small, posteromedial glabrous margin, but not as extensive as the region of the other ventrites. Anterior margin of ventrites 3 and 4 sometimes slightly to strongly crenulate. Ventrite 5 entire, without apical emargination. Laterosternite 3 with a distinct, ridged stridulatory file (
<figureCitation id="130B91B9FFA93200BB24FD7DFCAF580C" box="[799,875,752,778]" captionStart="FIGURES 9 – 10" captionStartId="9.[151,269,1552,1576]" captionTargetBox="[151,1435,418,1526]" captionTargetId="figure@9.[151,1436,416,1528]" captionTargetPageId="9" captionText="FIGURES 9 – 10. SEMs of stridulatory apparatus in T. complanatus; 9) laterosternite 3; 10) interior of elytra showing plectrum. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183537/files/figure.png" pageId="3" pageNumber="44">Fig. 9</figureCitation>
). Aedeagus of the simple tri-lobed
<typeStatus id="548B339EFFA93200BD19FD7DFA93580C" box="[1314,1367,752,778]" pageId="3" pageNumber="44">type</typeStatus>
, with parameres and median lobe generally unmodified.
</paragraph>
</subSubSection>
<subSubSection id="C32ADEB7FFA93200B8FEFCB2FC015E3C" pageId="3" pageNumber="44" type="discussion">
<paragraph id="8B8F8D3CFFA93200B8FEFCB2FD625F24" blockId="3.[151,1437,152,1338]" pageId="3" pageNumber="44">
<emphasis id="B944512EFFA93200B8FEFCB2FE95585F" bold="true" box="[197,337,831,857]" pageId="3" pageNumber="44">Discussion.</emphasis>
Members of
<taxonomicName id="4C30F6BFFFA93200B9D5FCCDFD94585F" box="[494,592,832,857]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200B9D5FCCDFD94585F" box="[494,592,832,857]" italics="true" pageId="3" pageNumber="44">Tritonus</emphasis>
</taxonomicName>
are rather large water scavenger beetles, and represent the largest members of the tribe
<taxonomicName id="4C30F6BFFFA93200B96EFCE5FE125884" box="[341,470,872,898]" class="Insecta" family="Hydrophilidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="tribe" tribe="Laccobiini">Laccobiini</taxonomicName>
. Indeed, some species (
<taxonomicName id="4C30F6BFFFA93200BAD2FCE5FC5F5887" box="[745,923,872,897]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="complanatus">
<emphasis id="B944512EFFA93200BAD2FCE5FC5F5887" box="[745,923,872,897]" italics="true" pageId="3" pageNumber="44">T. complanatus</emphasis>
</taxonomicName>
) are larger than all but a few lineages of the family as a whole. Although rare in collections (less than
<specimenCount id="9D3646B5FFA93200BB0CFC1DFC2058AC" box="[823,996,912,938]" pageId="3" pageNumber="44" type="generic">100 specimens</specimenCount>
are known to me, more than half representing just one of the seven species), this appears to be an artifact of collector bias rather than actual abundance. Most collectors who have examined hygropetric habitats in
<collectingCountry id="F327CDACFFA93200BB86FC6DFB8F58FC" box="[957,1099,992,1018]" name="Madagascar" pageId="3" pageNumber="44">Madagascar</collectingCountry>
have collected series of the genus, and frequently more than one species.
</paragraph>
<paragraph id="8B8F8D3CFFA93200B8FEFBBDFC015E3C" blockId="3.[151,1437,152,1338]" pageId="3" pageNumber="44">
Within the
<taxonomicName id="4C30F6BFFFA93200B97CFBBDFE705F4F" box="[327,436,1072,1097]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200B97CFBBDFE705F4F" box="[327,436,1072,1097]" italics="true" pageId="3" pageNumber="44">Oocyclus</emphasis>
</taxonomicName>
-group,
<taxonomicName id="4C30F6BFFFA93200BA2BFBBDFDB65F4F" box="[528,626,1072,1097]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200BA2BFBBDFDB65F4F" box="[528,626,1072,1097]" italics="true" pageId="3" pageNumber="44">Tritonus</emphasis>
</taxonomicName>
appears to be most similar to the presently monotypic genus
<taxonomicName id="4C30F6BFFFA93200BD01FBBDFE9B5F74" authority="d'Orchymont" authorityName="d'Orchymont" class="Insecta" family="Hydrophilidae" genus="Scoliopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200BD01FBBDFF725F77" italics="true" pageId="3" pageNumber="44">Scoliopsis</emphasis>
d’Orchymont
</taxonomicName>
, which is known from
<collectingCountry id="F327CDACFFA93200BA49FBD5FD225F74" box="[626,742,1112,1138]" name="Sri Lanka" pageId="3" pageNumber="44">Sri Lanka</collectingCountry>
(and with unpublished records from southern
<collectingCountry id="F327CDACFFA93200BD36FBD5FA8A5F74" box="[1293,1358,1112,1138]" name="India" pageId="3" pageNumber="44">India</collectingCountry>
). Both these genera share broadly glabrous regions on abdominal ventrites, distinct sutural stria, and 9-segmented antennae; conversely,
<taxonomicName id="4C30F6BFFFA93200B9A5FB25FD705FC4" authority="d'Orchymont" authorityName="d'Orchymont" box="[414,692,1192,1218]" class="Insecta" family="Hydrophilidae" genus="Beralitra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200B9A5FB25FDCE5FC7" box="[414,522,1192,1217]" italics="true" pageId="3" pageNumber="44">Beralitra</emphasis>
d’Orchymont
</taxonomicName>
,
<taxonomicName id="4C30F6BFFFA93200BAFAFB25FBC05FC4" authority="Komarek" authorityName="Komarek" box="[705,1028,1192,1218]" class="Insecta" family="Hydrophilidae" genus="Ophthalmocyclus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200BAFAFB25FC4A5FC7" box="[705,910,1192,1217]" italics="true" pageId="3" pageNumber="44">Ophthalmocyclus</emphasis>
Komarek
</taxonomicName>
and
<taxonomicName id="4C30F6BFFFA93200BC7AFB25FB385FC4" authority="Sharp" authorityName="Sharp" box="[1089,1276,1192,1218]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200BC7AFB25FB6A5FC7" box="[1089,1198,1192,1217]" italics="true" pageId="3" pageNumber="44">Oocyclus</emphasis>
Sharp
</taxonomicName>
each possess entirely pubescent abdominal ventrites, lack sutural striae, and have 8-segmented antennae. However, unlike
<taxonomicName id="4C30F6BFFFA93200B8ACFB75FF3D5E17" box="[151,249,1272,1297]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200B8ACFB75FF3D5E17" box="[151,249,1272,1297]" italics="true" pageId="3" pageNumber="44">Tritonus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C30F6BFFFA93200B932FB75FEBB5E17" box="[265,383,1272,1297]" class="Insecta" family="Hydrophilidae" genus="Scoliopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200B932FB75FEBB5E17" box="[265,383,1272,1297]" italics="true" pageId="3" pageNumber="44">Scoliopsis</emphasis>
</taxonomicName>
also possesses spinose posterolateral margins of the pronotum and coarse spines on the prosternum and mesoventrite (features also found in some
<taxonomicName id="4C30F6BFFFA93200BB7CFAADFC705E3F" box="[839,948,1312,1337]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B944512EFFA93200BB7CFAADFC705E3F" box="[839,948,1312,1337]" italics="true" pageId="3" pageNumber="44">Oocyclus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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