141 lines
18 KiB
XML
141 lines
18 KiB
XML
<document id="A2149F7CCA2824FE209CBE3920B1B7C1" ID-DOI="10.5281/zenodo.183532" ID-GBIF-Dataset="90941c20-aece-4da0-b4bf-19e800996840" ID-ISSN="1175-5326" ID-Zenodo-Dep="183532" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920883928" checkinUser="plazi" docAuthor="Short, Andrew Edward Z." docDate="2008" docId="03993C2AFFA6320EB8ACF962FC6058D4" docLanguage="en" docName="zt01855p055.pdf" docOrigin="Zootaxa 1855" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tritonus riambavy Short, 2008, sp.n." docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="FFA04452FFAA3203B83BFF8DFFC45B06" masterDocTitle="Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)" masterLastPageNumber="55" masterPageNumber="41" pageNumber="53" updateTime="1698229214647" updateUser="plazi">
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<mods:mods id="AFDFBE6A367C015BCB6266612FBC1715" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="23460D984B74D1731030C06C5200C748">
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<mods:title id="2F3D6985E919CB0BFF44F1CAC89F8FCB">Revision of the Afrotropical genus Tritonus Mulsant (Coleoptera: Hydrophilidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A424B1AD05E2396934BB14D849810192" type="personal">
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<mods:role id="6F388220A5FB42D223901F0FD6E8CF78">
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<mods:roleTerm id="16D7E92EF4DD5CDE68B12AD3188EC392">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="880656D12AFBCBD4C1F66954D88F8229">Short, Andrew Edward Z.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3FCEA5939CB293E98339C33D69D7CD68">text</mods:typeOfResource>
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<mods:titleInfo id="C28AB32330FF10089DB19090415B7300">
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<mods:title id="2F948D4D96FEB05A0276F843E9103E8A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FC2EA378E23863C1147F375E5AF3E0DA">
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<mods:date id="2440845E39B3108701A1E76EFC39B66F">2008</mods:date>
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<mods:detail id="E3E8AB51D71D792796F0B254801259F0" type="volume">
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<mods:number id="85A5B3E9881A988C9006CAE3841CDDD3">1855</mods:number>
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</mods:detail>
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<mods:extent id="0E24CC9CB72E7847333102CC09B7FFC4" unit="page">
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<mods:start id="33960A298EA113DA6C122AE1DB45B90E">41</mods:start>
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<mods:end id="2F9EE5D3EB055E7A8B5556CF0393F58A">55</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="043AA0D51E520AF62F93D1C33D0B28C6">journal article</mods:classification>
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<mods:identifier id="BD3E7F8CF7B24792A0770DB3A563EF84" type="DOI">10.5281/zenodo.183532</mods:identifier>
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<mods:identifier id="1135CF391F7E3A0312C79B3511E1E9B2" type="GBIF-Dataset">90941c20-aece-4da0-b4bf-19e800996840</mods:identifier>
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<mods:identifier id="85D1715C99FFA9F4E0E84424177D7F76" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E52C38142EE6F2A5C0F1B76C39605204" type="Zenodo-Dep">183532</mods:identifier>
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</mods:mods>
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<treatment id="03993C2AFFA6320EB8ACF962FC6058D4" ID-DOI="http://doi.org/10.5281/zenodo.6232632" ID-GBIF-Taxon="119373680" ID-Zenodo-Dep="6232632" LSID="urn:lsid:plazi:treatment:03993C2AFFA6320EB8ACF962FC6058D4" httpUri="http://treatment.plazi.org/id/03993C2AFFA6320EB8ACF962FC6058D4" lastPageId="13" lastPageNumber="54" pageId="12" pageNumber="53">
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<subSubSection id="C32ADEB7FFA6320FB8ACF962FE735C2A" pageId="12" pageNumber="53" type="nomenclature">
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||
<paragraph id="8B8F8D3CFFA6320FB8ACF962FE7C5C0F" blockId="12.[151,440,1775,1836]" box="[151,440,1775,1801]" pageId="12" pageNumber="53">
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<heading id="D0C73A50FFA6320FB8ACF962FE7C5C0F" bold="true" box="[151,440,1775,1801]" fontSize="11" level="1" pageId="12" pageNumber="53" reason="1">
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||
<emphasis id="B944512EFFA6320FB8ACF962FE7C5C0F" bold="true" box="[151,440,1775,1801]" pageId="12" pageNumber="53">
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<taxonomicName id="4C30F6BFFFA6320FB8ACF962FEB25C0F" ID-CoL="595BX" box="[151,374,1775,1801]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="53" phylum="Arthropoda" rank="species" species="riambavy" status="sp. nov.">
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<emphasis id="B944512EFFA6320FB8ACF962FEB25C0F" bold="true" box="[151,374,1775,1801]" italics="true" pageId="12" pageNumber="53">Tritonus riambavy</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A277EC55FFA6320FB946F962FE7C5C0F" box="[381,440,1775,1801]" pageId="12" pageNumber="53" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8B8F8D3CFFA6320FB8ACF89FFE735C2A" blockId="12.[151,440,1775,1836]" box="[151,439,1810,1836]" pageId="12" pageNumber="53">
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||
(
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<figureCitation id="130B91B9FFA6320FB8A4F89FFF355C2A" box="[159,241,1810,1836]" captionStart="FIGURES 2 – 3" captionStartId="6.[151,269,1008,1032]" captionTargetBox="[175,1411,194,983]" captionTargetId="figure@6.[175,1411,194,984]" captionTargetPageId="6" captionText="FIGURES 2 – 3. SEMs of head, dorsal and ventral views; 2) T. complanatus; 3) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183534/files/figure.png" pageId="12" pageNumber="53">Figs. 3</figureCitation>
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,
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<figureCitation id="130B91B9FFA6320FB8C4F89FFECB5C2A" box="[255,271,1810,1836]" captionStart="FIGURES 4 – 5" captionStartId="7.[151,269,1587,1611]" captionTargetBox="[151,1435,418,1568]" captionTargetId="figure@7.[151,1436,414,1569]" captionTargetPageId="7" captionText="FIGURES 4 – 5. SEMs of elytra; 4) T. complanatus; 5) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183535/files/figure.png" pageId="12" pageNumber="53">5</figureCitation>
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,
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<figureCitation id="130B91B9FFA6320FB927F89FFEE85C2A" box="[284,300,1810,1836]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="12" pageNumber="53">8</figureCitation>
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,
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<figureCitation id="130B91B9FFA6320FB902F89FFE935C2A" box="[313,343,1810,1836]" captionStart="FIGURES 11 – 12" captionStartId="10.[151,269,1197,1221]" captionTargetBox="[151,1435,634,1171]" captionTargetId="figure@10.[151,1436,634,1171]" captionTargetPageId="10" captionText="FIGURES 11 – 12. Abdomen of Tritonus spp.; 11) T. riambavy; 12) T. crenulatus. Scale bars = 1 mm." httpUri="https://zenodo.org/record/183538/files/figure.png" pageId="12" pageNumber="53">11</figureCitation>
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,
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<figureCitation id="130B91B9FFA6320FB95FF89FFE475C2A" box="[356,387,1810,1836]" captionStart="FIGURES 13 – 14" captionStartId="10.[151,269,1627,1651]" captionTargetBox="[151,1435,1278,1602]" captionTargetId="figure@10.[151,1436,1278,1603]" captionTargetPageId="10" captionText="FIGURES 13 – 14. Details of Tritonus spp.; 13) T. complanatus, metafemora, scale bar = 1 mm; 14) T. riambavy, maxillary palpus, scale bar = 0.20 mm." httpUri="https://zenodo.org/record/183539/files/figure.png" pageId="12" pageNumber="53">13</figureCitation>
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,
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<figureCitation id="130B91B9FFA6320FB9ABF89FFE6A5C2A" box="[400,430,1810,1836]" captionStart="FIGURES 15 – 21" captionStartId="11.[151,269,1696,1720]" captionTargetBox="[242,1353,216,1672]" captionTargetId="figure@11.[234,1353,194,1672]" captionTargetPageId="11" captionText="FIGURES 15 – 21. Aedeagus of Tritonus spp., dorsal view; 15) T. riana; 16) T. steineri; 17) T. riambavy; 18) T. cribratus; 19) T. madagascarensis; 20) T. crenulatus; 21) T. complanatus. Scale bar = 0.45 mm." httpUri="https://zenodo.org/record/183540/files/figure.png" pageId="12" pageNumber="53">17</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32ADEB7FFA6320FB8ACF8EFFE305CF2" pageId="12" pageNumber="53" type="materials_examined">
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<paragraph id="8B8F8D3CFFA6320FB8ACF8EFFE305CF2" blockId="12.[151,1437,1890,2036]" pageId="12" pageNumber="53">
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<emphasis id="B944512EFFA6320FB8ACF8EFFE0A5C7A" bold="true" box="[151,462,1890,1916]" pageId="12" pageNumber="53">
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<typeStatus id="548B339EFFA6320FB8ACF8EFFF105C7A" box="[151,212,1890,1916]" pageId="12" pageNumber="53">Type</typeStatus>
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Material.
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<typeStatus id="548B339EFFA6320FB967F8EFFE0A5C7A" box="[348,462,1890,1916]" pageId="12" pageNumber="53" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male): “
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<collectionCode id="ED2115F9FFA6320FBA7BF8EFFD5E5C7A" box="[576,666,1890,1916]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="12" pageNumber="53">NHML</collectionCode>
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742778 MSL 163/
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<collectingCountry id="F327CDACFFA6320FBBB2F8EFFBDF5C7A" box="[905,1051,1890,1916]" name="Madagascar" pageId="12" pageNumber="53">Madagascar</collectingCountry>
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: Fianarantsoa: Andringitra N.P./
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<taxonomicName id="4C30F6BFFFA6320FB8ACF807FED45CA2" box="[151,272,1930,1956]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="54" phylum="Arthropoda" rank="species" species="riambavy">Riambavy</taxonomicName>
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River (female) cascade: alt
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<quantity id="4CC820D9FFA6320FBA60F807FD6F5CA2" box="[603,683,1930,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="12" pageNumber="53" unit="m" value="1739.0">1739m</quantity>
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/ 08:V:2006: Lat –22 8.761 Lon 46 53.495/ Hygropetric habitat, wet rock surfaces next to/ high waterfall: Leg. J. Bergsten Loc P39A” (
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<collectionCode id="ED2115F9FFA6320FBBE1F83FFBF35CCA" box="[986,1079,1970,1996]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="12" pageNumber="53">NHML</collectionCode>
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).
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<emphasis id="B944512EFFA6320FBC71F83FFB385CCA" bold="true" box="[1098,1276,1970,1996]" pageId="12" pageNumber="53">
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<typeStatus id="548B339EFFA6320FBC71F83FFB0C5CCA" box="[1098,1224,1970,1996]" pageId="12" pageNumber="53" type="paratype">Paratypes</typeStatus>
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(3):
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</emphasis>
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Same data as
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<typeStatus id="548B339EFFA6320FB8ACF857FF395CF2" box="[151,253,2010,2036]" pageId="12" pageNumber="53" type="holotype">holotype</typeStatus>
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(3,
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<collectionCode id="ED2115F9FFA6320FB917F857FE415CF2" box="[300,389,2010,2036]" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="12" pageNumber="53">NHML</collectionCode>
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,
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<collectionCode id="ED2115F9FFA6320FB9AAF857FE2C5CF2" box="[401,488,2010,2036]" pageId="12" pageNumber="53">KSEM</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32ADEB7FFA7320EB8FEFF1AFD545BB4" box="[197,656,151,178]" pageId="13" pageNumber="54" type="diagnosis">
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<paragraph id="8B8F8D3CFFA7320EB8FEFF1AFD545BB4" blockId="13.[151,1436,151,978]" box="[197,656,151,178]" pageId="13" pageNumber="54">
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<emphasis id="B944512EFFA7320EB8FEFF1AFE815BB7" bold="true" box="[197,325,151,177]" pageId="13" pageNumber="54">Diagnosis.</emphasis>
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See diagnosis of
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<taxonomicName id="4C30F6BFFFA7320EBA2FFF15FD4D5BB7" box="[532,649,152,177]" class="Insecta" family="Hydrophilidae" genus="Tritonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="steineri">
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<emphasis id="B944512EFFA7320EBA2FFF15FD4D5BB7" box="[532,649,152,177]" italics="true" pageId="13" pageNumber="54">T. steineri</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32ADEB7FFA7320EB8FEFF32FC475884" pageId="13" pageNumber="54" type="description">
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<paragraph id="8B8F8D3CFFA7320EB8FEFF32FC475884" blockId="13.[151,1436,151,978]" pageId="13" pageNumber="54">
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<emphasis id="B944512EFFA7320EB8FEFF32FE995BDF" bold="true" box="[197,349,191,217]" pageId="13" pageNumber="54">Description.</emphasis>
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Total length
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<quantity id="4CC820D9FFA7320EB9C7FF4DFD4B5BDC" box="[508,655,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="6.6" metricValueMin="6.2" pageId="13" pageNumber="54" unit="mm" value="6.4" valueMax="6.6" valueMin="6.2">6.2–6.6 mm</quantity>
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. Color: Dorsum black with general shagreen; with distinct yellow or greenish iridescent patches on elytra.
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<emphasis id="B944512EFFA7320EBA57FF6AFD715A07" bold="true" box="[620,693,231,257]" pageId="13" pageNumber="54">Head.</emphasis>
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Ground punctation coarse; irregular in size (
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<figureCitation id="130B91B9FFA7320EBC84FF65FAC15A04" box="[1215,1285,232,258]" captionStart="FIGURES 2 – 3" captionStartId="6.[151,269,1008,1032]" captionTargetBox="[175,1411,194,983]" captionTargetId="figure@6.[175,1411,194,984]" captionTargetPageId="6" captionText="FIGURES 2 – 3. SEMs of head, dorsal and ventral views; 2) T. complanatus; 3) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183534/files/figure.png" pageId="13" pageNumber="54">Fig. 3</figureCitation>
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). Systematic punctures present but only slightly larger than surrounding ground punctures; on clypeus bearing short decumbent setae; those on frons anterior to each eye relatively long and erect. Antennae with club subequal in length to antennomeres 2–6 combined. Maxillary palpi short, apical segment ca. 1.6× length of penultimate segment; penultimate segment distinctly shorter than segment 2. Mentum with anterior margin slightly convex; anterior half with fine transverse microsculpture; entire mentum with fine to moderately coarse, irregularly scattered punctures. Anterior margin of head evenly rounded.
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<emphasis id="B944512EFFA7320EBB88FE5AFBD35AF7" bold="true" box="[947,1047,471,497]" pageId="13" pageNumber="54">Thorax.</emphasis>
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Ground punctation on pronotum and elytra of very irregular in size, varying from very fine to very coarse. Pronotal and elytral systematic punctures present, but somewhat obscured by coarse ground punctures (
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<figureCitation id="130B91B9FFA7320EBC39FDA5FB885944" box="[1026,1100,552,578]" captionStart="FIGURES 4 – 5" captionStartId="7.[151,269,1587,1611]" captionTargetBox="[151,1435,418,1568]" captionTargetId="figure@7.[151,1436,414,1569]" captionTargetPageId="7" captionText="FIGURES 4 – 5. SEMs of elytra; 4) T. complanatus; 5) T. riambavy. Scale bar = 1 mm." httpUri="https://zenodo.org/record/183535/files/figure.png" pageId="13" pageNumber="54">Fig. 5</figureCitation>
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), which are similar in size; bearing short setae. Prosternum (
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<figureCitation id="130B91B9FFA7320EBA1BFDDDFDA3596C" box="[544,615,592,618]" captionStart="FIGURES 6 – 8" captionStartId="8.[151,269,807,831]" captionTargetBox="[151,1435,178,783]" captionTargetId="figure@8.[151,1436,168,783]" captionTargetPageId="8" captionText="FIGURES 6 – 8. SEMs of thoracic sterna of Tritonus species; 6) meso- and metaventrite of T. complanatus, scale bar = 1 mm; 7) prosternum of T. complanatus, 8) prosternum of T. riambavy. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/183536/files/figure.png" pageId="13" pageNumber="54">Fig. 8</figureCitation>
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) slightly tectiform an indistinct, very narrow and median line that is not distinctly elevated into a carina; not distinctly elevated or dilated at anterior margin. Mesoventrite slightly elevated medially into a carina, with anterior and posterior ends of carina glabrous and more elevated than central, pubescent region. Metasternum with large elevated glabrous region that is three-quarters as wide as long. Metafemora without distinct microsculpture over most of ventral surface.
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<emphasis id="B944512EFFA7320EBC03FD62FB7D580F" bold="true" box="[1080,1209,751,777]" pageId="13" pageNumber="54">Abdomen.</emphasis>
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Posteromedial glabrous region of ventrite 1 distinct, ca. 0.15 the width of the ventrite. Glabrous region of ventrite 2 semi-circular to subtriangular in shape; ca. 0.30 the width of the ventrite. Anterior margins of ventrites 1–3 not crenulate, but with a loosely wrinkled appearance. Aedeagus as in
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<figureCitation id="130B91B9FFA7320EBB12FCE5FCBB5884" box="[809,895,872,898]" captionStart="FIGURES 15 – 21" captionStartId="11.[151,269,1696,1720]" captionTargetBox="[242,1353,216,1672]" captionTargetId="figure@11.[234,1353,194,1672]" captionTargetPageId="11" captionText="FIGURES 15 – 21. Aedeagus of Tritonus spp., dorsal view; 15) T. riana; 16) T. steineri; 17) T. riambavy; 18) T. cribratus; 19) T. madagascarensis; 20) T. crenulatus; 21) T. complanatus. Scale bar = 0.45 mm." httpUri="https://zenodo.org/record/183540/files/figure.png" pageId="13" pageNumber="54">Fig. 17</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32ADEB7FFA7320EB8FEFC02FC5258AC" box="[197,918,911,938]" pageId="13" pageNumber="54" type="etymology">
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<paragraph id="8B8F8D3CFFA7320EB8FEFC02FC5258AC" blockId="13.[151,1436,151,978]" box="[197,918,911,938]" pageId="13" pageNumber="54">
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||
<emphasis id="B944512EFFA7320EB8FEFC02FE9658AF" bold="true" box="[197,338,911,937]" pageId="13" pageNumber="54">Etymology.</emphasis>
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Named for the river from which it was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32ADEB7FFA7320EB8FEFC3AFC6058D4" box="[197,932,951,978]" pageId="13" pageNumber="54" type="distribution">
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||
<paragraph id="8B8F8D3CFFA7320EB8FEFC3AFC6058D4" blockId="13.[151,1436,151,978]" box="[197,932,951,978]" pageId="13" pageNumber="54">
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||
<emphasis id="B944512EFFA7320EB8FEFC3AFEA258D7" bold="true" box="[197,358,951,977]" pageId="13" pageNumber="54">Distribution.</emphasis>
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<collectingCountry id="F327CDACFFA7320EB956FC35FE3958D4" box="[365,509,952,978]" name="Madagascar" pageId="13" pageNumber="54">Madagascar</collectingCountry>
|
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. Known only from the
|
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<typeStatus id="548B339EFFA7320EBB37FC35FCFA58D4" box="[780,830,952,978]" pageId="13" pageNumber="54">type</typeStatus>
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locality.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |