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<document id="737DAC14A3DE74B5753E85340DD1ECFE" ID-CLB-Dataset="38966" ID-DOI="10.11646/zootaxa.4117.2.2" ID-GBIF-Dataset="2a5cc1cb-7377-4778-ba4e-82a8a80946e9" ID-ISSN="1175-5326" ID-Zenodo-Dep="262872" ID-ZooBank="24FDEE28-549D-473B-8742-A2E7B9CFBE7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464675174659" checkinUser="plazi" docAuthor="Yasunaga, Tomohide &amp; Duwal, Ram Keshari" docDate="2016" docId="03990706B347FF929FEA310CBBAD1BC0" docLanguage="en" docName="zootaxa.4117.2.2.pdf.imf" docOrigin="Zootaxa 4117 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pilophorus nakatanii Yasunaga &amp; Duwal, n. sp." docType="treatment" docVersion="10" lastPageNumber="179" masterDocId="FFA07F7EB344FF959F7D363EBB501A6C" masterDocTitle="Three new species of the ant-mimetic plant bug genus Pilophorus from Japan (Hemiptera: Heteroptera: Miridae: Phylinae: Pilophorini)" masterLastPageNumber="182" masterPageNumber="172" pageNumber="175" updateTime="1698662649700" updateUser="ExternalLinkService">
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<mods:title id="F72BCAC2AA5AEA88D3165AF340802867">Three new species of the ant-mimetic plant bug genus Pilophorus from Japan (Hemiptera: Heteroptera: Miridae: Phylinae: Pilophorini)</mods:title>
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<mods:namePart id="40662E194C95ED479197F79B0BCDF9F5">Yasunaga, Tomohide</mods:namePart>
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<mods:namePart id="E1BC0B006D5378B2B24EB55C438EC7CA">Duwal, Ram Keshari</mods:namePart>
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<mods:title id="814B7DC166D75A0CFE4E05111C3DF40B">Zootaxa</mods:title>
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<mods:date id="858C010005582AEF6B5EE25C567CFB9E">2016</mods:date>
<mods:detail id="CE3A652A14F505F127960F0EBD7673C7" type="volume">
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<mods:classification id="004FE2C5297F425E2CE9E4CBB8477FD2">journal article</mods:classification>
<mods:identifier id="EC7EA8F263A229ACAA5D0B8640F72E0A" type="CLB-Dataset">38966</mods:identifier>
<mods:identifier id="DA871D7243A68F1F609A08A81029A44C" type="DOI">10.11646/zootaxa.4117.2.2</mods:identifier>
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<treatment id="03990706B347FF929FEA310CBBAD1BC0" ID-DOI="http://doi.org/10.5281/zenodo.6053576" ID-GBIF-Taxon="120733442" ID-Zenodo-Dep="6053576" LSID="urn:lsid:plazi:treatment:03990706B347FF929FEA310CBBAD1BC0" httpUri="http://treatment.plazi.org/id/03990706B347FF929FEA310CBBAD1BC0" lastPageId="7" lastPageNumber="179" pageId="3" pageNumber="175">
<subSubSection id="C32AE59BB347FF969FEA310CBA321D02" pageId="3" pageNumber="175" type="nomenclature">
<paragraph id="8B8FB610B347FF969FEA310CB9B01D21" blockId="3.[151,736,1842,1902]" box="[151,736,1842,1869]" pageId="3" pageNumber="175">
<heading id="D0C7017CB347FF969FEA310CB9B01D21" bold="true" box="[151,736,1842,1869]" fontSize="11" level="1" pageId="3" pageNumber="175" reason="1">
<emphasis id="B9446A02B347FF969FEA310CB9B01D21" bold="true" box="[151,736,1842,1869]" pageId="3" pageNumber="175">
<taxonomicName id="4C30CD93B347FF969FEA310CB9C11D21" authority="Yasunaga &amp; Duwal" authorityName="Yasunaga &amp; Duwal" box="[151,657,1842,1869]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="175" phylum="Arthropoda" rank="species" species="nakatanii" status="sp. nov.">
<emphasis id="B9446A02B347FF969FEA310CBAC71D20" bold="true" box="[151,407,1842,1868]" italics="true" pageId="3" pageNumber="175">Pilophorus nakatanii</emphasis>
Yasunaga &amp; Duwal
</taxonomicName>
,
<taxonomicNameLabel id="A277D779B347FF969DE3310DB9B01D21" box="[670,736,1843,1869]" pageId="3" pageNumber="175" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8FB610B347FF969FEA316BBA321D02" blockId="3.[151,736,1842,1902]" box="[151,354,1877,1902]" pageId="3" pageNumber="175">
(
<figureCitation id="130BAA95B347FF969FE2316BBBB81D02" box="[159,232,1877,1902]" captionStart="FIGURE 1" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[151,1435,195,1779]" captionTargetId="figure@4.[151,1435,195,1800]" captionTargetPageId="4" captionText="FIGURE 1. Habitus images of new Japanese Pilophorus spp. A. P. nakatanii, holotype male (AMNH _ PBI 00380410). B. Same, dry-preserved condition, right lateral view. C. P. tagoi, male. D. Same, female. E. P. maeharai, holotype male (00380411). F. Same, ventral view. G. Same, female." httpUri="https://zenodo.org/record/262873/files/figure.png" pageId="3" pageNumber="175">Figs. 1</figureCitation>
A–B, 2, 3)
</paragraph>
</subSubSection>
<subSubSection id="C32AE59BB347FF969FEA31A3BECC1DDF" box="[151,1436,1949,1971]" pageId="3" pageNumber="175" type="reference_group">
<paragraph id="8B8FB610B347FF969FEA31A3BECC1DDF" blockId="3.[151,1436,1949,1971]" box="[151,1436,1949,1971]" pageId="3" pageNumber="175">
<treatmentCitationGroup id="AB20913EB347FF969FEA31A3BECC1DDF" box="[151,1436,1949,1971]" pageId="3" pageNumber="175">
<taxonomicName id="4C30CD93B347FF969FEA31A3B9BD1DDF" ID-CoL="77JTR" authority="Miyamoto &amp; Yasunaga, 1989: 163" authorityName="Miyamoto &amp; Yasunaga" authorityPageNumber="163" authorityYear="1989" box="[151,749,1949,1971]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="175" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B9446A02B347FF969FEA31A3BAD31DDF" box="[151,387,1949,1971]" italics="true" pageId="3" pageNumber="175">Pilophorus formosanus</emphasis>
:
<treatmentCitation id="0A919001B347FF969EEC31A3B9BD1DDF" author="Miyamoto" box="[401,749,1949,1971]" page="163" pageId="3" pageNumber="175" year="1989">
<bibRefCitation id="EFA1CBE1B347FF969EEC31A3B9ED1DDF" author="Miyamoto" box="[401,701,1949,1971]" pageId="3" pageNumber="175" refString="Miyamoto, S. &amp; Yasunaga, T. (1989) Hemiptera, Heteroptera. In: Hirashima, Y. (Ed.), A check list of Japanese insects. Entomological Laboratory, Kyushu University, Fukuoka, pp. 151 - 188." type="book chapter" year="1989">Miyamoto &amp; Yasunaga, 1989</bibRefCitation>
: 163
</treatmentCitation>
</taxonomicName>
(list);
<treatmentCitation id="0A919001B347FF969C4D31A0BF551DDF" author="Yasunaga" box="[816,1029,1949,1971]" page="250" pageId="3" pageNumber="175" year="2001">
<bibRefCitation id="EFA1CBE1B347FF969C4D31A0B8841DDF" author="Yasunaga" box="[816,980,1949,1971]" pageId="3" pageNumber="175" refString="Yasunaga, T. (2001) Family Miridae Hahn, plant bugs. In: Yasunaga, T., Takai, M. &amp; Kawasawa, T. (Eds.), A Field Guide to Japanese Bugs. Vol. II. Zenkoku Noson Kyoiku Kyokai, Publishing Co. Ltd., Tokyo, pp. 1 - 96 &amp; 111 - 351. [in Japanese]" type="book chapter" year="2001">Yasunaga, 2001</bibRefCitation>
: 250
</treatmentCitation>
, pl. 91 (diag.) [nec
<treatmentCitation id="0A919001B347FF969BB431A3BEC21DDF" author="Poppius" box="[1225,1426,1949,1971]" page="238" pageId="3" pageNumber="175" year="1914">
<bibRefCitation id="EFA1CBE1B347FF969BB431A3BE0D1DDF" author="Poppius" box="[1225,1373,1949,1971]" pageId="3" pageNumber="175" refString="Poppius, B. (1914) Ubersicht der Pilophorus - Arten nebst Beschreibung verwandter Gattungen (Hem. Het.). Annales de la Societe Entomologique de Belgique, 58, 237 - 254." type="journal article" year="1914">Poppius, 1914</bibRefCitation>
: 238
</treatmentCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF4FE698B340FF919FEA3123B9FE1D1D" httpUri="https://zenodo.org/record/262873/files/figure.png" pageId="4" pageNumber="176" targetBox="[151,1435,195,1779]" targetPageId="4">
<paragraph id="8B8FB610B340FF919FEA3123B9FE1D1D" blockId="4.[151,1436,1821,1905]" pageId="4" pageNumber="176">
<emphasis id="B9446A02B340FF919FEA3123BA461D5F" bold="true" box="[151,278,1821,1843]" pageId="4" pageNumber="176">FIGURE 1.</emphasis>
Habitus images of new Japanese
<taxonomicName id="4C30CD93B340FF919D003123B9BE1D5F" box="[637,750,1821,1843]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="176" phylum="Arthropoda" rank="genus">
<emphasis id="B9446A02B340FF919D003123B9BE1D5F" box="[637,750,1821,1843]" italics="true" pageId="4" pageNumber="176">Pilophorus</emphasis>
</taxonomicName>
spp.
<emphasis id="B9446A02B340FF919C573120B8131D5F" bold="true" box="[810,835,1822,1843]" pageId="4" pageNumber="176">A.</emphasis>
<taxonomicName id="4C30CD93B340FF919C313121B8971D5F" box="[844,967,1821,1844]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="176" phylum="Arthropoda" rank="species" species="nakatanii">
<emphasis id="B9446A02B340FF919C313121B8971D5F" box="[844,967,1821,1844]" italics="true" pageId="4" pageNumber="176">P. nakatanii</emphasis>
</taxonomicName>
, holotype male (AMNH_PBI 00380410).
<emphasis id="B9446A02B340FF919AF93120BECB1D5F" bold="true" box="[1412,1435,1822,1843]" pageId="4" pageNumber="176">B.</emphasis>
Same, dry-preserved condition, right lateral view.
<emphasis id="B9446A02B340FF919DC53102B9801D3E" bold="true" box="[696,720,1852,1874]" pageId="4" pageNumber="176">C.</emphasis>
<taxonomicName id="4C30CD93B340FF919DA03103B8601D3E" box="[733,816,1853,1874]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="176" phylum="Arthropoda" rank="species" species="tagoi">
<emphasis id="B9446A02B340FF919DA03103B8601D3E" box="[733,816,1853,1874]" italics="true" pageId="4" pageNumber="176">P. tagoi</emphasis>
</taxonomicName>
, male.
<emphasis id="B9446A02B340FF919CFA3103B8F01D3E" bold="true" box="[903,928,1853,1874]" pageId="4" pageNumber="176">D.</emphasis>
Same, female.
<emphasis id="B9446A02B340FF919B323103BF361D3E" bold="true" box="[1103,1126,1853,1874]" pageId="4" pageNumber="176">E.</emphasis>
<taxonomicName id="4C30CD93B340FF919B0E3103BFA31D3E" box="[1139,1267,1852,1874]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="176" phylum="Arthropoda" rank="species" species="maeharai">
<emphasis id="B9446A02B340FF919B0E3103BFA31D3E" box="[1139,1267,1852,1874]" italics="true" pageId="4" pageNumber="176">P. maeharai</emphasis>
</taxonomicName>
, holotype male (00380411).
<emphasis id="B9446A02B340FF919E643165BA7A1D1C" bold="true" box="[281,298,1883,1904]" pageId="4" pageNumber="176">F.</emphasis>
Same, ventral view.
<emphasis id="B9446A02B340FF919D7D3165B9441D1D" bold="true" box="[512,532,1883,1905]" pageId="4" pageNumber="176">G.</emphasis>
Same, female.
</paragraph>
</caption>
<caption id="DF4FE698B341FF909FEA3131BE761D49" box="[151,1318,1807,1829]" httpUri="https://zenodo.org/record/262874/files/figure.png" pageId="5" pageNumber="177" targetBox="[156,1425,207,1774]" targetPageId="5">
<paragraph id="8B8FB610B341FF909FEA3131BE761D49" blockId="5.[151,1318,1807,1829]" box="[151,1318,1807,1829]" pageId="5" pageNumber="177">
<emphasis id="B9446A02B341FF909FEA3131BA421D48" bold="true" box="[151,274,1807,1829]" pageId="5" pageNumber="177">FIGURE 2.</emphasis>
Male genitalia (genital segment and parameres) of new Japanese
<taxonomicName id="4C30CD93B341FF909CD03131BF4E1D49" box="[941,1054,1807,1829]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B9446A02B341FF909CD03131BF4E1D49" box="[941,1054,1807,1829]" italics="true" pageId="5" pageNumber="177">Pilophorus</emphasis>
</taxonomicName>
spp. named on the figure.
</paragraph>
</caption>
<subSubSection id="C32AE59BB341FF909FEA3172BE3F1D99" pageId="5" pageNumber="177" type="diagnosis">
<paragraph id="8B8FB610B341FF909FEA3172BE3F1D99" blockId="5.[151,1437,1868,2037]" pageId="5" pageNumber="177">
<emphasis id="B9446A02B341FF909FEA3172BA411D09" bold="true" box="[151,273,1868,1893]" pageId="5" pageNumber="177">Diagnosis.</emphasis>
Recognized by its moderate size; fuscous brown basic coloration; less shining pronotum; very sparse scalelike setae on scutellum; chestnut brown, matt anterior 2/3 of hemelytron; two streaks of scalelike setae on corium (a band on anterior corium not continuing to clavus and the other posterior band confluent); and distinctive shape of endosoma. Most similar to
<taxonomicName id="4C30CD93B341FF909D4C3187B9BC1DBD" authority="Poppius" authorityName="Poppius" box="[561,748,1976,2001]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="177" phylum="Tracheophyta" rank="species" species="niger">
<emphasis id="B9446A02B341FF909D4C3187B9D71DBC" box="[561,647,1977,2000]" italics="true" pageId="5" pageNumber="177">P. niger</emphasis>
Poppius
</taxonomicName>
, based on dorsal color (surface) and vestiture patterns; easily distinguished by apparently smaller size, chestnut anterior corium and clavus, and different form of endosoma.
</paragraph>
</subSubSection>
<subSubSection id="C32AE59BB342FF939FBA36A9B8721931" pageId="6" pageNumber="178" type="description">
<paragraph id="8B8FB610B342FF939FBA36A9B98118E8" blockId="6.[151,1437,151,933]" pageId="6" pageNumber="178">
<emphasis id="B9446A02B342FF939FBA36A9BA091ADC" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="178">Description.</emphasis>
Body generally dark brown, rather small, weakly to moderately constricted (HCR = 0.82–0.88); dorsal surface weakly shining (except for polished posterior 1/3 of hemelytron), with sparsely distributed, simple, brown setae. Head shiny fuscous, triangular. Antenna dark brown, except for wholly creamy white segment III; segment II weakly clavate. Labium shiny reddish brown, reaching apex of metacoxa. Pronotum fuscous, weakly shining, narrow; mesoscutum shiny fuscous; scutellum fuscous, weakly shining, with very sparsely distributed scalelike setae; ostiolar peritreme grayish brown. Hemelytron nearly two-tone colored, with chestnut brown, matte anterior 2/3 and shiny fuscous, posterior 1/3, each demarcated by a confluent band of scalelike setae (
<figureCitation id="130BAA95B342FF939A4F3751BE241BE4" box="[1330,1396,367,392]" captionStart="FIGURE 1" captionStartId="4.[151,250,1821,1843]" captionTargetBox="[151,1435,195,1779]" captionTargetId="figure@4.[151,1435,195,1800]" captionTargetPageId="4" captionText="FIGURE 1. Habitus images of new Japanese Pilophorus spp. A. P. nakatanii, holotype male (AMNH _ PBI 00380410). B. Same, dry-preserved condition, right lateral view. C. P. tagoi, male. D. Same, female. E. P. maeharai, holotype male (00380411). F. Same, ventral view. G. Same, female." httpUri="https://zenodo.org/record/262873/files/figure.png" pageId="6" pageNumber="178">Fig. 1</figureCitation>
A); anterior 1/3 of corium with the other band of scalelike setae that is not continuous to clavus; cuneus with sparsely distributed, scalelike setae along anterior margin; membrane dark grayish brown. Coxa and leg dark brown; median part of procoxa, and apices of shagreened meso- and metacoxa brown; metafemur weakly curved; each tarsus yellowish brown, with darkened apex. Abdomen shiny fuscous; proximal 1/3 of ventral surface with a band of scalelike setae laterally. Male genitalia (
<figureCitation id="130BAA95B342FF939D05341AB9971850" box="[632,711,548,573]" captionStart="FIGURE 2" captionStartId="5.[151,250,1807,1829]" captionTargetBox="[156,1425,207,1774]" captionTargetId="figure@5.[151,1436,193,1786]" captionTargetPageId="5" captionText="FIGURE 2. Male genitalia (genital segment and parameres) of new Japanese Pilophorus spp. named on the figure." httpUri="https://zenodo.org/record/262874/files/figure.png" pageId="6" pageNumber="178">Figs. 2</figureCitation>
,
<figureCitation id="130BAA95B342FF939DA9341AB9B41850" box="[724,740,548,572]" captionStart="FIGURE 3" captionStartId="6.[151,250,1966,1988]" captionTargetBox="[162,1420,996,1937]" captionTargetId="figure@6.[150,1436,985,1945]" captionTargetPageId="6" captionText="FIGURE 3. Male genitalia (endosoma) of new Japanese Pilophorus spp. named on the figure." httpUri="https://zenodo.org/record/262875/files/figure.png" pageId="6" pageNumber="178">3</figureCitation>
): Genital segment (pygophore) rather broad; phallotheca with sharpened apex. Left paramere with a moderate-sized dorsal protuberance. Endosoma C-shaped, medially with a long, straight, saber-like, basally bifurcate process.
</paragraph>
<paragraph id="8B8FB610B342FF939FBA34AEB8721931" blockId="6.[151,1437,151,933]" pageId="6" pageNumber="178">
<emphasis id="B9446A02B342FF939FBA34AEBA2D18C4" bold="true" box="[199,381,656,680]" pageId="6" pageNumber="178">Measurements.</emphasis>
♂/♀: Total body length 3.2−3.3/ 3.6; length from apex of clypeus to cuneal fracture 2.47−2.66/ 2.77; width of head across eyes 0.92−0.95/ 0.95; head height 0.75−0.76/ 0.80; width of vertex 0.42−0.44/ 0.48; lengths of antennal segments I −IV 0.32−0.35, 1.20−1.27, 0.39−0.40, 0.48−0.53/ 0.34, 1.33,?,?; basal width of pronotum 0.31; minimum width across hemelytron 1.14; maximum width across hemelytron 1.33;; and length of metafemur, tibia and tarsus 1.23−1.27, 1.89−1.91, 0.39−0.43/ 1.33, 2.09,? (antennal segments III and IV, and metatarsus missing in available female specimen).
</paragraph>
</subSubSection>
<subSubSection id="C32AE59BB342FF939FBA3556BFA919C9" pageId="6" pageNumber="178" type="etymology">
<paragraph id="8B8FB610B342FF939FBA3556BFA919C9" blockId="6.[151,1437,151,933]" pageId="6" pageNumber="178">
<emphasis id="B9446A02B342FF939FBA3556BA1E19EC" bold="true" box="[199,334,872,896]" pageId="6" pageNumber="178">Etymology.</emphasis>
Named to honor Dr. Yukinobu Nakatani (National Institute of Agro-Environmental Sciences, Tsukuba,
<collectingCountry id="F327F680B342FF939E7E35B2BA1719C8" box="[259,327,908,932]" name="Japan" pageId="6" pageNumber="178">Japan</collectingCountry>
), who is enthusiastically working on the Asian deraeocorine and pilophorine faunas.
</paragraph>
</subSubSection>
<caption id="DF4FE698B342FF939FEA3190BF041DA8" box="[151,1108,1966,1988]" httpUri="https://zenodo.org/record/262875/files/figure.png" pageId="6" pageNumber="178" targetBox="[162,1420,996,1937]" targetPageId="6">
<paragraph id="8B8FB610B342FF939FEA3190BF041DA8" blockId="6.[151,1108,1966,1988]" box="[151,1108,1966,1988]" pageId="6" pageNumber="178">
<emphasis id="B9446A02B342FF939FEA3190BA421DAF" bold="true" box="[151,274,1966,1988]" pageId="6" pageNumber="178">FIGURE 3.</emphasis>
Male genitalia (endosoma) of new Japanese
<taxonomicName id="4C30CD93B342FF939DA73190B81B1DA8" box="[730,843,1966,1988]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="178" phylum="Arthropoda" rank="genus">
<emphasis id="B9446A02B342FF939DA73190B81B1DA8" box="[730,843,1966,1988]" italics="true" pageId="6" pageNumber="178">Pilophorus</emphasis>
</taxonomicName>
spp. named on the figure.
</paragraph>
</caption>
<subSubSection id="C32AE59BB343FF929FBA36A9BB8A1A94" pageId="7" pageNumber="179" type="biology_ecology">
<paragraph id="8B8FB610B343FF929FBA36A9BB8A1A94" blockId="7.[151,1437,151,428]" pageId="7" pageNumber="179">
<emphasis id="B9446A02B343FF929FBA36A9BA771ADC" bold="true" box="[199,295,151,176]" pageId="7" pageNumber="179">Biology.</emphasis>
Although two individuals of this new species were found on Okinawa-pine,
<taxonomicName id="4C30CD93B343FF929BE536A6BA401AB9" authority="Mayr" authorityName="Mayr" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="179" phylum="Tracheophyta" rank="species" species="luchuensis">
<emphasis id="B9446A02B343FF929BE536A6BE061ADC" box="[1176,1366,152,176]" italics="true" pageId="7" pageNumber="179">Pinus luchuensis</emphasis>
Mayr (Pinaceae)
</taxonomicName>
, its breeding host is yet to be confirmed. Collection records suggest
<taxonomicName id="4C30CD93B343FF929B4C3683BFEA1AB8" box="[1073,1210,188,212]" class="Insecta" family="Miridae" genus="Pilophorus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="179" phylum="Arthropoda" rank="species" species="nakatanii">
<emphasis id="B9446A02B343FF929B4C3683BFEA1AB8" box="[1073,1210,188,212]" italics="true" pageId="7" pageNumber="179">P. nakatanii</emphasis>
</taxonomicName>
has a bivoltine life cycle.
</paragraph>
</subSubSection>
<subSubSection id="C32AE59BB343FF929FBA373ABBAD1BC0" pageId="7" pageNumber="179" type="materials_examined">
<paragraph id="8B8FB610B343FF929FBA373AB9C31B2C" blockId="7.[151,1437,151,428]" pageId="7" pageNumber="179">
<emphasis id="B9446A02B343FF929FBA373ABA6D1B70" bold="true" box="[199,317,260,284]" pageId="7" pageNumber="179">
<typeStatus id="548B08B2B343FF929FBA373ABA681B70" box="[199,312,260,284]" pageId="7" pageNumber="179" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂,
<emphasis id="B9446A02B343FF929E14373DBA9C1B70" bold="true" box="[361,460,259,284]" pageId="7" pageNumber="179">
<collectingCountry id="F327F680B343FF929E14373DBA971B70" box="[361,455,259,284]" name="Japan" pageId="7" pageNumber="179">JAPAN</collectingCountry>
:
</emphasis>
Ryukyus: Iriomote Island, Uehara, light trap,
<date id="FF8E90D0B343FF929CAA373ABF071B70" box="[983,1111,260,285]" pageId="7" pageNumber="179" value="2013-06-06">6 Jun 2013</date>
, T. Yasunaga, M. Takai &amp; T. Ishikawa (
<collectionCode id="ED212ED5B343FF929E773716BA311B2C" box="[266,353,296,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="179">AMNH</collectionCode>
_PBI 00380410) (
<collectionCode id="ED212ED5B343FF929D563716B9D71B2C" box="[555,647,296,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="7" pageNumber="179">AMNH</collectionCode>
).
</paragraph>
<paragraph id="8B8FB610B343FF929FBA3772BBAD1BC0" blockId="7.[151,1437,151,428]" pageId="7" pageNumber="179">
<emphasis id="B9446A02B343FF929FBA3772BA161B08" bold="true" box="[199,326,332,356]" pageId="7" pageNumber="179">
<typeStatus id="548B08B2B343FF929FBA3772BA111B08" box="[199,321,332,356]" pageId="7" pageNumber="179" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Ryukyus: Okinawa Island, Kunigami Village, Yona, on
<taxonomicName id="4C30CD93B343FF929CBE3773BF961B09" authority="Mayr" authorityName="Mayr" box="[963,1222,332,357]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="179" phylum="Tracheophyta" rank="species" species="luchuensis">
<emphasis id="B9446A02B343FF929CBE3773BFD01B08" box="[963,1152,332,356]" italics="true" pageId="7" pageNumber="179">Pinus luchuensis</emphasis>
Mayr
</taxonomicName>
,
<date id="FF8E90D0B343FF929BAF3772BEC81B08" box="[1234,1432,332,357]" pageId="7" pageNumber="179" value="1993-05-20" valueMax="1993-05-24" valueMin="1993-05-20">20−24 May 1993</date>
, T. Yasunaga,
<specimenCount id="9D367D99B343FF929E4B374EBA091BE4" box="[310,345,368,392]" pageId="7" pageNumber="179" type="generic">1♂</specimenCount>
(
<collectionCode id="ED212ED5B343FF929E11374EBAEF1BE4" box="[364,447,368,392]" pageId="7" pageNumber="179">TYCN</collectionCode>
); Ishigaki Island, Mt. Bansei (Maese),
<taxonomicName id="4C30CD93B343FF929CF6374FBF711BE4" box="[907,1057,368,392]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="7" pageNumber="179" phylum="Tracheophyta" rank="species" species="luchuensis">
<emphasis id="B9446A02B343FF929CF6374FBF711BE4" box="[907,1057,368,392]" italics="true" pageId="7" pageNumber="179">P. luchuensis</emphasis>
</taxonomicName>
,
<date id="FF8E90D0B343FF929B4E374EBF9A1BE4" box="[1075,1226,368,392]" pageId="7" pageNumber="179" value="2002-09-30">30 Sep 2002</date>
, T. Yasunaga, 1♀ (
<collectionCode id="ED212ED5B343FF929FE237AABBA21BC0" box="[159,242,404,428]" pageId="7" pageNumber="179">TYCN</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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