142 lines
15 KiB
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142 lines
15 KiB
XML
<document id="DABA0D18FCD6C98B6FA8F4A2F68B0175" ID-CLB-Dataset="29643" ID-DOI="http://dx.doi.org/10.3897/dez.65.27033" ID-GBIF-Dataset="8916b689-6eca-4121-9066-01e3ccfdbb56" ID-Pensoft-Pub="1860-1324-2-117" ID-ZooBank="B1A8523CA4634FC4A0C3072C2E78BA02" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1860-1324-2-117" ModsDocOrigin="Deutsche Entomologische Zeitschrift 65 (2)" ModsDocTitle="Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)" checkinTime="1533122252714" checkinUser="pensoft" docAuthor="Salnitska, Maria & Solodovnikov, Alexey" docDate="2018" docId="587BC5818B5E68E2209D0BB05634AAE7" docLanguage="en" docName="DeutEntomolZeit 65(2): 117-159" docOrigin="Deutsche Entomologische Zeitschrift 65 (2)" docSource="http://dx.doi.org/10.3897/dez.65.27033" docTitle="Quedius Stephens 1829" docType="treatment" docVersion="6" lastPageNumber="119" masterDocId="FFF6FFC22708A131FF960567FFF2FFB9" masterDocTitle="Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)" masterLastPageNumber="159" masterPageNumber="117" pageNumber="118" updateTime="1701380021094" updateUser="ExternalLinkService">
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<mods:title id="569CA0D0186982B3453629868D28114F">Revision of the Quedius fauna of Middle Asia (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<mods:namePart id="D6142A2112F71E974DD13B8D93F2A502">Salnitska, Maria</mods:namePart>
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<mods:namePart id="59ADCB60345D82158BD9300DDC8E89D3">Solodovnikov, Alexey</mods:namePart>
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<mods:title id="FDC52703CBF1D7C7E4C2F849B84D3F75">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="F872BB42AF057B16F66283DD7ECE4416">2018</mods:date>
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<mods:number id="F9C98C0940EED85D76952E95FFC58608">65</mods:number>
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<mods:url id="009BAB7980586EB8D0190693963C7C25">http://dx.doi.org/10.3897/dez.65.27033</mods:url>
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<mods:classification id="F9A9A69A3496E98179F5954A7A733475">journal article</mods:classification>
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<mods:identifier id="4CFF97A17440E77952337CFA6C2DE2A6" type="DOI">http://dx.doi.org/10.3897/dez.65.27033</mods:identifier>
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<treatment id="587BC5818B5E68E2209D0BB05634AAE7" ID-GBIF-Taxon="146754224" LSID="urn:lsid:plazi:treatment:587BC5818B5E68E2209D0BB05634AAE7" httpUri="http://treatment.plazi.org/id/587BC5818B5E68E2209D0BB05634AAE7" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118">
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<subSubSection id="CC31AED79CC7BCB3481165FEB3EEDBA8" pageId="1" pageNumber="118" type="nomenclature">
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<paragraph id="D87C1D79C583CFA612C110213ED8378C" pageId="1" pageNumber="118">
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Genus
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<taxonomicName id="06DFA48F38B852701769F362189E7EE9" ID-CoL="8VS2M" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius Stephens, 1829</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="92D24FEC2B298402084B9BFB65E9180B" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118" type="type species">
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<paragraph id="65AF379B72741F17B705D26CAA821310" pageId="1" pageNumber="118">Type species.</paragraph>
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<paragraph id="C0EC3EA7FC9B0E0863450B49518A0572" pageId="1" pageNumber="118">
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<taxonomicName id="D067F9468B41648DD93CD52DA3CE9278" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius levicollis" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="species" species="levicollis">Quedius levicollis</taxonomicName>
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(
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<normalizedToken id="E8DF449DB5BFC76E0A5682B518B50060" originalValue="Brullé">Brulle</normalizedToken>
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, 1832).
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</paragraph>
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<paragraph id="96EA72C2D8982578B34DB27BAA6EC3BE" pageId="1" pageNumber="118">
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According to the latest phylogenetic hypotheses (Solodovnikov, 2006;
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<bibRefCitation id="4562987564E80894BAF1E15209A4DBA4" author="Chatzimanolis, S" journalOrPublisher="Zoologica Scripta" pageId="25" pageNumber="142" pagination="436 - 449" title="Molecular phylogeny of the mega-diverse rove beetle tribe Staphylinini (Insecta, Coleoptera, Staphylinidae)." url="https://doi.org/10.1111/j.1463-6409.2010.00438.x" volume="39" year="2010">Chatzimanolis et al., 2010</bibRefCitation>
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;
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<bibRefCitation id="9F38C73ADCA6E893CBC7D70857DA050B" author="Brunke, AJ" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="25" pageNumber="142" url="https://doi.org/10.1111/cla.12139" year="2016">Brunke et al., 2016</bibRefCitation>
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) the genus
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<taxonomicName id="806C30E55C2D33DC5DB2709F91C1B48D" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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as it stands now in the taxonomic literature (e.g., summaries in Herman, 2001 or
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<normalizedToken id="552044BA0D97E49A8DAF2491BCB6ED3C" originalValue="Schülke">Schuelke</normalizedToken>
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and Smetana, 2015) is a polyphyletic assemblage of species belonging to several different subtribes of
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<taxonomicName id="FDDBBE42E5C464A73E0A1A7DBB7CBB7D" lsidName="" pageId="1" pageNumber="118" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
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. Within the Palaearctic or Middle Asia, all species of
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<taxonomicName id="BC98971D8CF76DC2EE75DC72871AFE23" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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are members of the subtribe
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<taxonomicName id="830F9DA2CD04BAF206BC6BDE66A786AC" lsidName="Quediina" pageId="1" pageNumber="118" rank="subtribe" subtribe="Quediina">Quediina</taxonomicName>
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in the restricted sense of
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<bibRefCitation id="A1B81AF039C44F6DF4B9698D18DBF676" author="Brunke, AJ" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="25" pageNumber="142" url="https://doi.org/10.1111/cla.12139" year="2016">Brunke et al. (2016)</bibRefCitation>
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. Because of the polyphyly,
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<taxonomicName id="76BFD960BAF2C94E9B30484DB9347845" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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in the current composition lacks synapomorphies and clear diagnosis. However, genus descriptions and diagnostic combination of characters that can define any Palaearctic species as a member of the genus
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<taxonomicName id="DF461EC1F2B31B575C72364E9AE14BC4" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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are available in
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<bibRefCitation id="C56EEF409C643D36453F2FA1D9E71D9C" pageId="1" pageNumber="118">Coiffait (1978)</bibRefCitation>
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,
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<bibRefCitation id="BA8418845503334FB24BA288C102A784" author="Smetana, A" journalOrPublisher="Quaestiones Entomologicae" pageId="26" pageNumber="143" pagination="163 - 464" title="Revision of the tribes Quediini and Atanygnathini, Part II. The Himalayan region (Coleoptera: Staphylinidae)." volume="24" year="1988">Smetana (1988)</bibRefCitation>
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,
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<bibRefCitation id="47771BD9EFE3EF4C5990E19EF8F5D800" pageId="1" pageNumber="118">
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Assing and
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<normalizedToken id="48F62E955F696404F9AF2D6B6E780283" originalValue="Schülke">Schuelke</normalizedToken>
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(2012)
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</bibRefCitation>
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and other sources. The diagnosis of the genus
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<taxonomicName id="B6DF0B172D35DB007D62CAB793243029" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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and comparative notes we provide here are tuned for the fauna of Middle Asia.
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</paragraph>
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<paragraph id="01EB8C4F2A7F88EACEE4C37C325E8F19" lastPageId="2" lastPageNumber="119" pageId="1" pageNumber="118">
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Adults and larvae of
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<taxonomicName id="6F9B335EBA09CD90231C8751B55059E1" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="1" pageNumber="118" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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seem to be predators hunting small invertebrates in various, sufficiently hu
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<pageBreakToken id="6EAEB1CEB6114F4A8E8F7C4F41923204" pageId="2" pageNumber="119" start="start">mid</pageBreakToken>
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ground-based debris, mostly in forest leaf litter. In a largely arid region like Middle Asia,
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<taxonomicName id="C0D81EBCD858BC51B50E3876B9E6125C" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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are mainly confined to humid open or forested habitats along creeks or rivers in the lowland or forests, meadows, snowfield margins and talus in the mountains. Some members of the subgenus
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<taxonomicName id="38932ABDB61A100D0CD8B222421ACDC8" genus="Quediina" lsidName="Quediina (Microsaurus)" pageId="2" pageNumber="119" rank="subGenus" subGenus="Microsaurus">Microsaurus</taxonomicName>
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are specialized inhabitants of mammal burrows. Overall, bionomics of the genus in Middle Asia remain largely unstudied.
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</paragraph>
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</subSubSection>
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<subSubSection id="84B6929FED9F34642BD39F7235F61412" pageId="2" pageNumber="119" type="diagnosis">
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<paragraph id="DEB01FF833F5AF766EE6342A81C1D303" pageId="2" pageNumber="119">Diagnosis.</paragraph>
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<paragraph id="DFEEDF7F3F8D9D42F22683CE556206C0" pageId="2" pageNumber="119">
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Medium to large size (body length 3.5-24.0 mm) rove beetles with glossy forebody, infraorbital ridges extended from neck to base of mandibles and pronotal hypomera strongly inflexed under pronotal disk (not visible in lateral view). First segment of antennae at most slightly longer than second and third segments together. Last segment of maxillary palps fusiform, not densely setose. Tarsal formula 5
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<normalizedToken id="B92A36DF4B1CB9E27EA32A18A7C544E3" originalValue="–5–">-5-</normalizedToken>
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5; anterior tarsi widened in both sexes, with pale adhesive setae ventrally, with pair of empodial setae. Males always with distinct apical emargination on abdominal sternite VIII. Aedeagus varies in shape, paramere mostly with sensory peg setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="739F277F88511005F4AE69C29F5293B6" pageId="2" pageNumber="119" type="comparisons">
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<paragraph id="7C85CFEE761DC6E7605CA75086ABC62B" pageId="2" pageNumber="119">Comparison.</paragraph>
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<paragraph id="6DDC0F0B24382A122FE8DDC45CF581F4" pageId="2" pageNumber="119">
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Among other
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<taxonomicName id="32E7FF4639B7C053D42A9657DD5C1838" lsidName="" pageId="2" pageNumber="119" rank="tribe" tribe="Staphylinini">Staphylinini</taxonomicName>
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in Middle Asia
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<taxonomicName id="991E9DFCDE97D6C1CE2ED219BA265005" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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can be sometimes confused with
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<taxonomicName id="035ABAF0AB71E212C0B2323483C0F69C" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
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(subtribe
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<taxonomicName id="ED6E9DC0190CB240676F774178C9B4A8" lsidName="Philonthina" pageId="2" pageNumber="119" rank="subtribe" subtribe="Philonthina">Philonthina</taxonomicName>
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), a genus with somewhat similar habitus and very abundant in the region. Species of
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<taxonomicName id="4D56B21A8AEF1DD4284B8E631AABFD24" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
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, however, do not have long infraorbital ridges, they lack empodial setae and mostly have a pronotal hypomeron well visible in lateral view. Also,
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<taxonomicName id="4EB7840F00F63154FD2105DFB17BF90C" class="Insecta" family="Staphylinidae" genus="Philonthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Philonthus" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Philonthus</taxonomicName>
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mostly possess multiple setiferous punctures in dorsal rows of pronotum (usually at most three in
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<taxonomicName id="F7624D6241FFFFB6667D32AA68DB3413" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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). Smaller species of
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<taxonomicName id="34D493DEFF8793D2E76586F885E18A52" class="Insecta" family="Staphylinidae" genus="Quedius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quedius" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
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may be confused with the genus
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<taxonomicName id="D2BB58A6724B82D68A1299D735F1D6FA" class="Insecta" family="Staphylinidae" genus="Heterothops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heterothops" order="Coleoptera" pageId="2" pageNumber="119" phylum="Arthropoda" rank="genus">Heterothops</taxonomicName>
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(subtribe
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<taxonomicName id="DA765885EFED9FB4EDBB5FB676A5E408" lsidName="Amblyopinina" pageId="2" pageNumber="119" rank="subtribe" subtribe="Amblyopinina">Amblyopinina</taxonomicName>
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), but the latter have very thin acicular apical segments of maxillary palps, and a very different aedeagus without sensory peg setae and reduced median lobe giving the appearance of an absent paramere).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |