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<document id="50BB88CF9780A27F549464680BC8FD0A" ID-CLB-Dataset="121550" ID-DOI="10.1093/zoolinnean/zlab087" ID-GBIF-Dataset="bafaf468-919e-467d-a31a-f8af68920e24" ID-ISSN="0024-4082" ID-Zenodo-Dep="6993173" ID-ZooBank="87E03803-7D69-4393-93CD-24A39D607972" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660323519587" checkinUser="plazi" docAuthor="Gąsiorek, Piotr, Wilamowski, Andrzej, Vončina, Katarzyna &amp; Michalczyk, Łukasz" docDate="2022" docId="586D3B6C0237615CEFF9CE9AFCC0F8E5" docLanguage="en" docName="zlab087.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Barbaria paucigranulata Gąsiorek, Wilamowski, Vončina &amp; Michalczyk, 2022, SP. NOV." docType="treatment" docUuid="A11E6BDF-7E2A-4F0A-BD6A-9BC87FCE147A" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1047" masterDocId="A454431402326157EF21CD2EFFCBFF87" masterDocTitle="Neotropical jewels in the moss: biodiversity, distribution and evolution of the genus Barbaria (Heterotardigrada: Echiniscidae)" masterLastPageNumber="1066" masterPageNumber="1037" pageNumber="1042" updateTime="1699366421497" updateUser="ExternalLinkService">
<mods:mods id="723F876851DC1CEF4DDBBEFE45F29643" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9B3CB872E3110362C4496E3083F3A502">Neotropical jewels in the moss: biodiversity, distribution and evolution of the genus Barbaria (Heterotardigrada: Echiniscidae)</mods:title>
</mods:titleInfo>
<mods:name id="01A6645383E060D71FA1631A3B0BBE1A" type="personal">
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<mods:namePart id="E2A777150DEE2E2EA3CEE0B3D66AA1BE">Gąsiorek, Piotr</mods:namePart>
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<mods:namePart id="1EB1E80E28D3FE095F07DD9C9195219C">Wilamowski, Andrzej</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="397A8A92B90065110305E93A39D5A7DC">Vončina, Katarzyna</mods:namePart>
</mods:name>
<mods:name id="57F0180F4CB3832CF1659C020D5CA5C7" type="personal">
<mods:role id="5DA5779BC0E9EB90A9B1A6BE399A3549">
<mods:roleTerm id="000E594EB0A339565D692F688A602E8D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4DF429FBE2AD889400BFF8489B817FD6">Michalczyk, Łukasz</mods:namePart>
</mods:name>
<mods:typeOfResource id="47FECF9368091883DF7C6B9C26228C42">text</mods:typeOfResource>
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<mods:title id="D75FE423F95BDFFD92AE41E5CE55A086">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="6B67B637D469DF49EF1DA06D505086B6">
<mods:date id="7FBB0E5654C3F89B1F1737E7245122BD">2022</mods:date>
<mods:detail id="BA8922550EC2848DDF37D3E1D5E754B1" type="pubDate">
<mods:number id="C9D164BDA12710BE9B6F968DD9E51D14">2022-01-25</mods:number>
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<mods:detail id="C6C9BC10A4F941E550163AE783AEC363" type="volume">
<mods:number id="6BE50386708642793950041646812711">195</mods:number>
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<mods:start id="13B5D5B0011E07367D69BC187B10A261">1037</mods:start>
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<mods:classification id="D1465770EEF20CCE12D91DAF80F18280">journal article</mods:classification>
<mods:identifier id="1751A14ADA39817E4CF2069A4D70E096" type="CLB-Dataset">121550</mods:identifier>
<mods:identifier id="084D0471868BCC9F6B6C4EAF8A72818C" type="DOI">10.1093/zoolinnean/zlab087</mods:identifier>
<mods:identifier id="7B3D143048AF9DD1A22A197985E45CD6" type="GBIF-Dataset">bafaf468-919e-467d-a31a-f8af68920e24</mods:identifier>
<mods:identifier id="D94A0EE29202498B2DC260F57DD57750" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="4BF34601D87D20D7402AE14CAB4D419C" type="Zenodo-Dep">6993173</mods:identifier>
<mods:identifier id="33343C5CB94B99F8B254A6C883D3C781" type="ZooBank">87E03803-7D69-4393-93CD-24A39D607972</mods:identifier>
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<treatment id="586D3B6C0237615CEFF9CE9AFCC0F8E5" ID-DOI="http://doi.org/10.5281/zenodo.6993207" ID-GBIF-Taxon="199104563" ID-Zenodo-Dep="6993207" LSID="urn:lsid:zoobank.org:act:A11E6BDF-7E2A-4F0A-BD6A-9BC87FCE147A" httpUri="http://treatment.plazi.org/id/586D3B6C0237615CEFF9CE9AFCC0F8E5" lastPageId="11" lastPageNumber="1047" pageId="5" pageNumber="1042">
<subSubSection id="98DED9F102376152EFF9CE9AFD27FC6D" pageId="5" pageNumber="1042" type="nomenclature">
<paragraph id="D07B8A7A02376152EFF9CE9AFD1DFC4C" blockId="5.[194,748,947,1002]" box="[216,726,947,971]" pageId="5" pageNumber="1042">
<heading id="8B333D1602376152EFF9CE9AFD1DFC4C" box="[216,726,947,971]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="1042" reason="3">
<taxonomicName id="17C4F1F902376152EFF9CE9AFD1DFC4C" ID-CoL="dff35314-f814-4f27-8caf-fde71c95393e" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" box="[216,726,947,971]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="1056" phylum="Tardigrada" rank="species" species="paucigranulata" status="SP. NOV.">
<emphasis id="E2B0566802376152EFF9CE9AFD1DFC4C" bold="true" box="[216,726,947,971]" italics="true" pageId="5" pageNumber="1042">BARBARIA PAUCIGRANULATA WILAMOWSKI,</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D07B8A7A02376152EFE3CEFCFD27FC6D" blockId="5.[194,748,947,1002]" box="[194,748,978,1002]" pageId="5" pageNumber="1042">
<heading id="8B333D1602376152EFE3CEFCFD27FC6D" box="[194,748,978,1002]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="1042" reason="3">
<emphasis id="E2B0566802376152EFE3CEFCFD27FC6D" bold="true" box="[194,748,978,1002]" pageId="5" pageNumber="1042">
VONČINA, GĄSIOREK &amp; MICHALCZYK,
<taxonomicNameLabel id="F983EB1302376152EDA8CEFBFD27FC6D" box="[649,748,978,1002]" pageId="5" pageNumber="1042" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="98DED9F102376152EFF0CED7FEA4FB34" pageId="5" pageNumber="1042" type="description">
<paragraph id="D07B8A7A02376152EFF0CED7FD17FB95" blockId="5.[209,732,1017,1042]" box="[209,732,1017,1042]" pageId="5" pageNumber="1042">
(
<figureCitation id="48FF96FF02376152EFFBCED7FE8AFB95" box="[218,321,1017,1042]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="6.[145,224,1739,1761]" captionStartId-1="7.[163,241,1648,1670]" captionStartId-2="8.[143,221,971,993]" captionTargetBox-0="[386,1184,195,1699]" captionTargetBox-1="[291,1315,195,1608]" captionTargetBox-2="[145,1425,195,931]" captionTargetId-0="figure-25@6.[386,1184,195,1699]" captionTargetId-1="figure-103@7.[291,1315,195,1608]" captionTargetId-2="figure-536@8.[145,1425,195,931]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." captionText-1="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." captionText-2="Figure 3. Details of Barbaria paucigranulata (SEM): A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm." figureDoi-0="http://doi.org/10.5281/zenodo.6993175" figureDoi-1="http://doi.org/10.5281/zenodo.6993177" figureDoi-2="http://doi.org/10.5281/zenodo.6993179" httpUri-0="https://zenodo.org/record/6993175/files/figure.png" httpUri-1="https://zenodo.org/record/6993177/files/figure.png" httpUri-2="https://zenodo.org/record/6993179/files/figure.png" pageId="5" pageNumber="1042">FIGS 1–3</figureCitation>
,
<figureCitation id="48FF96FF02376152EE6FCED7FEADFB95" box="[334,358,1017,1042]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="5" pageNumber="1042">7I</figureCitation>
,
<figureCitation id="48FF96FF02376152EE52CED7FE40FB95" box="[371,395,1017,1042]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="5" pageNumber="1042">8I</figureCitation>
,
<figureCitation id="48FF96FF02376152EEB8CED7FE7AFB95" box="[409,433,1017,1042]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="5" pageNumber="1042">9I</figureCitation>
,
<figureCitation id="48FF96FF02376152EE9FCED7FE2EFB95" box="[446,485,1017,1042]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="5" pageNumber="1042">10I</figureCitation>
,
<figureCitation id="48FF96FF02376152EED3CED7FD68FB97" box="[498,675,1017,1042]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="5" pageNumber="1042">11K, L; TABLES</figureCitation>
3, 4)
</paragraph>
<paragraph id="D07B8A7A02376152EF82C90FFCC0FBD1" blockId="5.[163,779,1057,1110]" pageId="5" pageNumber="1042">
<emphasis id="E2B0566802376152EF82C90FFE3BFBB0" box="[163,496,1057,1079]" italics="true" pageId="5" pageNumber="1042">Zoobank registration:</emphasis>
u r n: l s i d: z o o b a n k. org:act:
<uuid id="A462B0AF02376152EFCFC96EFCC0FBD1" box="[238,779,1088,1110]" pageId="5" pageNumber="1042">A11E6BDF-7E2A-4F0A-BD6A-9BC87FCE147A</uuid>
</paragraph>
<paragraph id="D07B8A7A02376152EF82C950FEA4FB34" blockId="5.[163,779,1150,1203]" pageId="5" pageNumber="1042">
<emphasis id="E2B0566802376152EF82C950FE01FB13" box="[163,458,1150,1172]" italics="true" pageId="5" pageNumber="1042">Morphometric data:</emphasis>
w w w. t a r d i g r a d a. n e t / register/0109.htm
</paragraph>
</subSubSection>
<subSubSection id="98DED9F102376152EF82C9F9FE23F986" pageId="5" pageNumber="1042" type="materials_examined">
<paragraph id="D07B8A7A02376152EF82C9F9FE68FAE0" blockId="5.[163,779,1238,1383]" pageId="5" pageNumber="1042">
<materialsCitation id="60AC802702376152EF82C9F9FEABFA8C" ID-GBIF-Occurrence="3888253302" country="Argentina" elevation="2150" latitude="-24.787222" location="vicinity of Rio Rosario" longLatPrecision="21" longitude="-65.725" municipality="Rosario de Lerma Department" pageId="5" pageNumber="1042" specimenCount="1" stateProvince="Salta Province" typeStatus="holotype">
<emphasis id="E2B0566802376152EF82C9F9FEA3FB6B" box="[163,360,1238,1260]" italics="true" pageId="5" pageNumber="1042">Type material:</emphasis>
<typeStatus id="0F7F34D802376152EE58C9F8FE24FB6B" box="[377,495,1238,1260]" pageId="5" pageNumber="1042" type="holotype">Holotype</typeStatus>
(adult female, slide AR.303.02) and
</materialsCitation>
<specimenCount id="C6C241F302376152EE4BC9DBFDC3FA8C" box="[362,520,1269,1291]" pageId="5" pageNumber="1042" type="generic" typeStatus="paratypes">12 paratypes</specimenCount>
(eight adult females on slides AR.302.02, AR.303.01, 3, 5–6 and four juveniles on slides AR.303.03–4). Found together with
<taxonomicName id="17C4F1F902376152EF82C87CFE57FAE1" box="[163,412,1361,1383]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="1042" phylum="Tardigrada" rank="species" species="ollantaytamboensis">
<emphasis id="E2B0566802376152EF82C87CFE57FAE1" box="[163,412,1361,1383]" italics="true" pageId="5" pageNumber="1042">B. ollantaytamboensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D07B8A7A02376152EF82C8BEFE23F986" blockId="5.[163,779,1423,1538]" pageId="5" pageNumber="1042">
<emphasis id="E2B0566802376152EF82C8BEFE98FA22" box="[163,339,1424,1445]" italics="true" pageId="5" pageNumber="1042">Type locality:</emphasis>
<geoCoordinate id="B5F0ECBD02376152EE42C8BEFE3AFA22" box="[355,497,1423,1446]" degrees="24" direction="south" minutes="47" orientation="latitude" pageId="5" pageNumber="1042" precision="15" seconds="14" value="-24.787222">24°47′14″S</geoCoordinate>
,
<geoCoordinate id="B5F0ECBD02376152ED25C8BEFD52FA22" box="[516,665,1423,1446]" degrees="65" direction="west" minutes="43" orientation="longitude" pageId="5" pageNumber="1042" precision="15" seconds="30" value="-65.725">65°43 ′30″W</geoCoordinate>
,
<elevation id="5BE96D4902376152ED8AC8BEFF02FA43" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="1042" unit="m" value="2150.0">
<quantity id="173C279F02376152ED8AC8BEFCC2FA22" box="[683,777,1424,1446]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="1042" unit="m" value="2150.0">2150 m</quantity>
asl
</elevation>
:
<collectingCountry id="A8D3CAEA02376152EFF6C880FE9DFA43" box="[215,342,1454,1476]" name="Argentina" pageId="5" pageNumber="1042">Argentina</collectingCountry>
,
<collectingRegion id="1200449802376152EE44C880FDD6FA43" box="[357,541,1454,1476]" country="Argentina" name="Salta" pageId="5" pageNumber="1042">Salta Province</collectingRegion>
,
<collectingMunicipality id="301F100002376152ED0CC880FEFBFA64" pageId="5" pageNumber="1042">Rosario de Lerma Department</collectingMunicipality>
,
<location id="D51BDCA102376152EE18C8E3FDF8FA64" LSID="urn:lsid:plazi:treatment:586D3B6C0237615CEFF9CE9AFCC0F8E5:D51BDCA102376152EE18C8E3FDF8FA64" box="[313,563,1485,1507]" country="Argentina" latitude="-24.787222" longLatPrecision="21" longitude="-65.725" municipality="Rosario de Lerma Department" name="vicinity of Rio Rosario" pageId="5" pageNumber="1042" stateProvince="Salta Province">vicinity of Río Rosario</location>
; lichen on rock in a shrubland (see also
<tableCitation id="9D46BFC102376152EEA4C8C2FE13F986" box="[389,472,1516,1538]" captionStart="Table 1" captionStartId="2.[105,169,374,396]" captionTargetPageId="2" captionText="Table 1. List of populations used in integrative analyses. Types of analyses: (PCM) imaging and morphometry in PCM, (DNA) DNA sequencing. Number in each analysis indicates how many specimens were utilised in a given method (f, females; m, males; a, adults of unidentified sex; j, juveniles; l, larvae)" httpUri="http://table.plazi.org/id/84BBDAF202306155EF48CC58F98EFE2E" pageId="5" pageNumber="1042" tableUuid="84BBDAF202306155EF48CC58F98EFE2E">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="98DED9F102376152EF82CB0BFE45F910" pageId="5" pageNumber="1042" type="etymology">
<paragraph id="D07B8A7A02376152EF82CB0BFE45F910" blockId="5.[163,780,1573,1687]" pageId="5" pageNumber="1042">
<emphasis id="E2B0566802376152EF82CB0BFEE8F9BD" box="[163,291,1573,1594]" italics="true" pageId="5" pageNumber="1042">Etymology:</emphasis>
From Latin
<emphasis id="E2B0566802376152EE9FCB08FDDBF9BC" box="[446,528,1574,1595]" italics="true" pageId="5" pageNumber="1042">paucus</emphasis>
, few, and
<emphasis id="E2B0566802376152EDA5CB0BFCCFF9BD" box="[644,772,1573,1594]" italics="true" pageId="5" pageNumber="1042">granulatus</emphasis>
, grained, alluding to the scarcity of epicuticular granulation on the dorsal plates. An adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="98DED9F102376150EF82CBEEFACFF8DD" lastPageId="7" lastPageNumber="1044" pageId="5" pageNumber="1042" type="description">
<paragraph id="D07B8A7A02376152EF82CBEEFC09FED2" blockId="5.[163,779,1728,1872]" lastBlockId="5.[827,1444,197,1875]" pageId="5" pageNumber="1042">
<emphasis id="E2B0566802376152EF82CBEEFEE5F952" box="[163,302,1728,1749]" italics="true" pageId="5" pageNumber="1042">Description:</emphasis>
Adult females (i.e. from the third instar onwards, measurements in
<tableCitation id="9D46BFC102376152EEF8CBF0FDE7F973" box="[473,556,1758,1780]" captionStart="Table 3" captionStartId="18.[145,209,196,217]" captionTargetPageId="18" captionText="Table 3. Measurements (in µm) of selected morphological structures of adult females of Barbaria paucigranulata (type series) mounted in Hoyer’s medium. Abbreviations: N, number of specimens/structures measured; RANGE refers to the smallest and the largest structure among all measured specimens; SD, standard deviation; sp, the proportion of the length of a given structure to the length of the scapular plate" httpUri="http://table.plazi.org/id/84BBDAF202206145EFB0CDEAFD0DFEB5" pageId="5" pageNumber="1042" tableUuid="84BBDAF202206145EFB0CDEAFD0DFEB5">Table 3</tableCitation>
). Dark orange body with large red eyes; the pigment and eyes dissolve quickly after mounting in Hoyer’s medium. Body massive with stumpy limbs (
<figureCitation id="48FF96FF02376152EEC7CA14FD93F8D7" box="[486,600,1850,1872]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A, B</figureCitation>
,
<figureCitation id="48FF96FF02376152ED45CA15FDB9F8D7" box="[612,626,1851,1872]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2</figureCitation>
). Cylindrical,
<taxonomicName id="17C4F1F902376152EC1ACDEBFC72FF5D" authorityName="Schultze" authorityYear="1840" box="[827,953,197,218]" class="Heterotardigrada" family="Echiniscidae" genus="Echiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="1042" phylum="Tardigrada" rank="genus">
<emphasis id="E2B0566802376152EC1ACDEBFC72FF5D" box="[827,953,197,218]" italics="true" pageId="5" pageNumber="1042">Echiniscus</emphasis>
</taxonomicName>
-
<typeStatus id="0F7F34D802376152ECE0CDE8FC3FFF5C" box="[961,1012,198,219]" pageId="5" pageNumber="1042">type</typeStatus>
cephalic papillae (secondary clavae) and (primary) clavae; cirri embedded in bulbous cirrophores (
<figureCitation id="48FF96FF02376152ECF1CC2CFBEFFE9F" box="[976,1060,258,280]" captionStart="Figure 3" captionStartId="8.[143,221,971,993]" captionTargetBox="[145,1425,195,931]" captionTargetId="figure-536@8.[145,1425,195,931]" captionTargetPageId="8" captionText="Figure 3. Details of Barbaria paucigranulata (SEM): A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993179" httpUri="https://zenodo.org/record/6993179/files/figure.png" pageId="5" pageNumber="1042">Fig. 3D</figureCitation>
). Cirrus
<emphasis id="E2B0566802376152EBAECC2DFB6BFE9F" box="[1167,1184,259,280]" italics="true" pageId="5" pageNumber="1042">A</emphasis>
is short (&lt;20% of the body length) and with evident, but small cirrophore (
<figureCitation id="48FF96FF02376152EC62CC6EFC78FED2" box="[835,947,320,342]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Fig. 1A, B</figureCitation>
).
</paragraph>
<paragraph id="D07B8A7A02376152EC72CC70FA54F9A7" blockId="5.[827,1444,197,1875]" pageId="5" pageNumber="1042">
Dorsal plate sculpturing of the
<taxonomicName id="17C4F1F902376152EBDECC70FA52FEF4" box="[1279,1433,350,371]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="5" pageNumber="1042" phylum="Tardigrada" rank="species" species="bigranulata">
<emphasis id="E2B0566802376152EBDECC70FA52FEF4" box="[1279,1433,350,371]" italics="true" pageId="5" pageNumber="1042">bigranulata</emphasis>
</taxonomicName>
-
<typeStatus id="0F7F34D802376152EC1ACC50FCB9FE14" box="[827,882,382,403]" pageId="5" pageNumber="1042">type</typeStatus>
, comprising minute, poorly developed pillars (pseudogranulation) and pores (
<figureCitation id="48FF96FF02376152EBEDCCB2FAE0FE36" box="[1228,1323,412,434]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1C</figureCitation>
,
<figureCitation id="48FF96FF02376152EA1ACCB2FA82FE36" box="[1339,1353,412,433]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2</figureCitation>
,
<figureCitation id="48FF96FF02376152EA7BCCB2FA57FE36" box="[1370,1436,412,433]" captionStart="Figure 3" captionStartId="8.[143,221,971,993]" captionTargetBox="[145,1425,195,931]" captionTargetId="figure-536@8.[145,1425,195,931]" captionTargetPageId="8" captionText="Figure 3. Details of Barbaria paucigranulata (SEM): A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993179" httpUri="https://zenodo.org/record/6993179/files/figure.png" pageId="5" pageNumber="1042">3A–C</figureCitation>
,
<figureCitation id="48FF96FF02376152EC1ACC95FC9FFE57" box="[827,852,443,464]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="5" pageNumber="1042">7I</figureCitation>
,
<figureCitation id="48FF96FF02376152EC49CC95FC49FE57" box="[872,898,443,464]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="5" pageNumber="1042">8I</figureCitation>
,
<figureCitation id="48FF96FF02376152ECB7CC95FC64FE57" box="[918,943,443,464]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="5" pageNumber="1042">9I</figureCitation>
,
<figureCitation id="48FF96FF02376152ECE5CC95FC27FE57" box="[964,1004,443,464]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="5" pageNumber="1042">10I</figureCitation>
). Pillars are densely packed and additionally interconnected by thin
<emphasis id="E2B0566802376152EA27CCF4FA87FE68" box="[1286,1356,474,495]" italics="true" pageId="5" pageNumber="1042">striae</emphasis>
in the scapular (
<figureCitation id="48FF96FF02376152EC93CCD6FC34FD8A" box="[946,1023,504,526]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="5" pageNumber="1042">Fig. 7I</figureCitation>
) and caudal (terminal) plates (
<figureCitation id="48FF96FF02376152EA57CCD6FCAAFDAC" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="5" pageNumber="1042">Fig. 10I</figureCitation>
). Pores are slightly larger in the scapular plate, median plate1, posterior portion of median plate 2 and centroposterior portions of paired segmental plates compared with posterolateral portions of paired segmental plates and the caudal plate (
<figureCitation id="48FF96FF02376152EA51CFBFFC82FD42" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1</figureCitation>
,
<figureCitation id="48FF96FF02376152EC78CF9EFCACFD42" box="[857,871,688,709]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2</figureCitation>
); pores of similar diameter are rarely present in the entire dorsum. Pores are always absent in the anterior portion of median plate 2 (
<figureCitation id="48FF96FF02376152EA1FCFC3FA59FC85" box="[1342,1426,749,771]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="5" pageNumber="1042">Fig. 8I</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF02376152EB49CE22FB7FFCA6" box="[1128,1204,780,802]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="5" pageNumber="1042">Fig. 9I</figureCitation>
) and the entirety of median plate 3; and all these areas are covered with epicuticular, multangular granules. Lateralmost portions of the scapular and paired segmental plates can be poreless or with single minute pores (
<figureCitation id="48FF96FF02376152EC65CE8BFC0EFC3D" box="[836,965,933,955]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A, B</figureCitation>
,
<figureCitation id="48FF96FF02376152ECF5CE8BFC29FC3D" box="[980,994,933,954]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2</figureCitation>
). The cephalic plate is large, with a pronounced chalice-shaped anterior incision and with only pillars in the posterior portion of the plate. A broad and strongly sclerotized cervical plate is divided into sculptured anterior portion and smooth posterior portion bordering with the scapular plate (
<figureCitation id="48FF96FF02376152EC62C973FC54FBF5" box="[835,927,1117,1139]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A</figureCitation>
,
<figureCitation id="48FF96FF02376152EC8FC973FC05FBF5" box="[942,974,1117,1138]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2A</figureCitation>
). Lateral sutures in the scapular plate demarcate lateralmost, trapezoidal portions (
<figureCitation id="48FF96FF02376152EA4EC952FCB2FB37" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A, B</figureCitation>
,
<figureCitation id="48FF96FF02376152ECA6C9B4FC6CFB28" box="[903,935,1178,1199]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2B</figureCitation>
). Median plates 1 and 3 are unipartite (the latter with strongly developed granules), median plate 2 is bipartite (
<figureCitation id="48FF96FF02376152EB04C9F6FBBBFB6A" box="[1061,1136,1240,1262]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="5" pageNumber="1042">Fig. 8I</figureCitation>
). Paired segmental plates are without transverse unsculptured bands, as epicuticular granules of anterior portion transition gradually into pillars of the posterior portions (
<figureCitation id="48FF96FF02376152EA57C81AFC98FAE0" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="5" pageNumber="1042">Fig. 9I</figureCitation>
). The caudal plate with short, poorly sclerotized incisions and no signs of faceting (
<figureCitation id="48FF96FF02376152EBC3C85FFAF4FA01" box="[1250,1343,1393,1415]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A</figureCitation>
,
<figureCitation id="48FF96FF02376152EA6FC85FFA97FA01" box="[1358,1372,1393,1414]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2</figureCitation>
,
<figureCitation id="48FF96FF02376152EA4DC85FFA58FA01" box="[1388,1427,1393,1414]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="5" pageNumber="1042">10I</figureCitation>
). Ventral cuticle with minute endocuticular pillars (
<figureCitation id="48FF96FF02376152EC65C880FC63FA44" box="[836,936,1454,1476]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="5" pageNumber="1042">Fig. 11K</figureCitation>
) distributed evenly throughout the entire venter, lacking plates, beside of rarely developed subcephalic plates (
<figureCitation id="48FF96FF02376152EB12C8C5FB45F986" box="[1075,1166,1515,1537]" captionStart="Figure 3" captionStartId="8.[143,221,971,993]" captionTargetBox="[145,1425,195,931]" captionTargetId="figure-536@8.[145,1425,195,931]" captionTargetPageId="8" captionText="Figure 3. Details of Barbaria paucigranulata (SEM): A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993179" httpUri="https://zenodo.org/record/6993179/files/figure.png" pageId="5" pageNumber="1042">Fig. 3D</figureCitation>
). Sexpartite gonopore placed anteriorly to a trilobed anus between legs IV.
</paragraph>
<paragraph id="D07B8A7A02376150EC72CB07FDD4F8DE" blockId="5.[827,1444,197,1875]" lastBlockId="7.[163,779,1737,1882]" lastPageId="7" lastPageNumber="1044" pageId="5" pageNumber="1042">
Pedal plates without pores, their sculpturing consists of poorly developed endocuticular pillars formed as belts in the central portions of the legs (
<figureCitation id="48FF96FF02376152EC65CBABFC02F91D" box="[836,969,1669,1691]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A, B</figureCitation>
,
<figureCitation id="48FF96FF02376152ECFBCBABFBE6F91D" box="[986,1069,1669,1690]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="5" pageNumber="1042">11K, L</figureCitation>
). Thick pulvini on outer side of all legs (
<figureCitation id="48FF96FF02376152ECE9CB8DFB81F93E" box="[968,1098,1699,1721]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1A, B</figureCitation>
,
<figureCitation id="48FF96FF02376152EB7BCB8AFBB1F93E" box="[1114,1146,1700,1721]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2B</figureCitation>
). Dentate collar IV has numerous irregular short teeth (
<figureCitation id="48FF96FF02376152EBE7CBECFAE3F950" box="[1222,1320,1730,1752]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="5" pageNumber="1042">Fig. 11L</figureCitation>
). A small, elongated spine I and a tubby papilla IV (
<figureCitation id="48FF96FF02376152EA61CBCFFA57F971" box="[1344,1436,1761,1783]" captionStart="Figure 1" captionStartId="6.[145,224,1739,1761]" captionTargetBox="[386,1184,195,1699]" captionTargetId="figure-25@6.[386,1184,195,1699]" captionTargetPageId="6" captionText="Figure 1. Habitus of Barbaria paucigranulata (PCM): A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus A; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="5" pageNumber="1042">Figs 1B</figureCitation>
,
<figureCitation id="48FF96FF02376152EC1ACA2EFC90F892" box="[827,859,1792,1813]" captionStart="Figure 2" captionStartId="7.[163,241,1648,1670]" captionTargetBox="[291,1315,195,1608]" captionTargetId="figure-103@7.[291,1315,195,1608]" captionTargetPageId="7" captionText="Figure 2. Habitus of Barbaria paucigranulata (SEM): A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="5" pageNumber="1042">2B</figureCitation>
,
<figureCitation id="48FF96FF02376152EC4ACA2EFC77F892" box="[875,956,1792,1813]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="5" pageNumber="1042">11K, L</figureCitation>
). Claws slightly heteronych with claws IV (
<figureCitation id="48FF96FF02376152EC46CA30FC72F8B4" box="[871,953,1822,1844]" captionStart="Figure 3" captionStartId="8.[143,221,971,993]" captionTargetBox="[145,1425,195,931]" captionTargetId="figure-536@8.[145,1425,195,931]" captionTargetPageId="8" captionText="Figure 3. Details of Barbaria paucigranulata (SEM): A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993179" httpUri="https://zenodo.org/record/6993179/files/figure.png" pageId="5" pageNumber="1042">Fig. 3F</figureCitation>
) higher and more robust than claws I–III (
<figureCitation id="48FF96FF02376152EC65CA13FC51F8D5" box="[836,922,1853,1875]" captionStart="Figure 3" captionStartId="8.[143,221,971,993]" captionTargetBox="[145,1425,195,931]" captionTargetId="figure-536@8.[145,1425,195,931]" captionTargetPageId="8" captionText="Figure 3. Details of Barbaria paucigranulata (SEM): A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm." figureDoi="http://doi.org/10.5281/zenodo.6993179" httpUri="https://zenodo.org/record/6993179/files/figure.png" pageId="5" pageNumber="1042">Fig. 3E</figureCitation>
). Internal claws IV have needle-like spurs more divergent from branches compared to spurs I– III (
<figureCitation id="48FF96FF02356150EFF3CBC6FE9FF97A" box="[210,340,1768,1790]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="7" pageNumber="1044">Fig. 11K, L</figureCitation>
). Cuticular bars below claw bases on the inner side of legs present. Buccal apparatus with a rigid tube and round pharynx containing placoids. Flexible stylet supports present.
</paragraph>
<caption id="84BBDAF202346151EFB0CBE5FE41F8D1" ID-DOI="http://doi.org/10.5281/zenodo.6993175" ID-Zenodo-Dep="6993175" httpUri="https://zenodo.org/record/6993175/files/figure.png" pageId="6" pageNumber="1043" startId="6.[145,224,1739,1761]" targetBox="[386,1184,195,1699]" targetPageId="6">
<paragraph id="D07B8A7A02346151EFB0CBE5FE41F8D1" blockId="6.[145,1425,1739,1878]" pageId="6" pageNumber="1043">
<emphasis id="E2B0566802346151EFB0CBE5FF30F967" bold="true" box="[145,251,1739,1761]" pageId="6" pageNumber="1043">Figure 1.</emphasis>
Habitus of
<taxonomicName id="17C4F1F902346151EE5CCBE5FD1CF966" authority="(PCM)" baseAuthorityName="PCM" box="[381,727,1739,1761]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="6" pageNumber="1043" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B0566802346151EE5CCBE5FD42F966" box="[381,649,1739,1761]" italics="true" pageId="6" pageNumber="1043">Barbaria paucigranulata</emphasis>
(PCM)
</taxonomicName>
: A, holotype (female) in dorsolateral view; B, paratype (female) in lateral view; C, dorsal sculpturing in close-up. Abbreviations: Ia–IIb, paired segmental plates; c, caudal plate; cA, cirrus
<emphasis id="E2B0566802346151EA5ACBC6FA40F979" box="[1403,1419,1768,1790]" italics="true" pageId="6" pageNumber="1043">A</emphasis>
; ce, cirrus externus; ci, cirrus internus; cl, (primary) clava; co, dentate collar IV; cp, cephalic plate; cv, cervical plate; m1–3, median plates; p, cephalic papilla (secondary clava); pIV, papilla IV; pl, pulvinus; pp, pedal plate; sI, spine I; sc, scapular plate. Scale bars in µm.
</paragraph>
</caption>
<caption id="84BBDAF202356150EF82CB5EFE32F924" ID-DOI="http://doi.org/10.5281/zenodo.6993177" ID-Zenodo-Dep="6993177" httpUri="https://zenodo.org/record/6993177/files/figure.png" pageId="7" pageNumber="1044" startId="7.[163,241,1648,1670]" targetBox="[291,1315,195,1608]" targetPageId="7">
<paragraph id="D07B8A7A02356150EF82CB5EFE32F924" blockId="7.[163,1443,1648,1699]" pageId="7" pageNumber="1044">
<emphasis id="E2B0566802356150EF82CB5EFEC1F902" bold="true" box="[163,266,1648,1670]" pageId="7" pageNumber="1044">Figure 2.</emphasis>
Habitus of
<taxonomicName id="17C4F1F902356150EEA9CB5EFD10F901" authority="(SEM)" baseAuthorityName="SEM" box="[392,731,1648,1670]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="7" pageNumber="1044" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B0566802356150EEA9CB5EFD5BF901" box="[392,656,1648,1670]" italics="true" pageId="7" pageNumber="1044">Barbaria paucigranulata</emphasis>
(SEM)
</taxonomicName>
: A, paratype (sex undetermined) in dorsal view; B, paratype (female) in lateral view. Scale bars in µm.
</paragraph>
</caption>
<paragraph id="D07B8A7A02356150EC72CBE7FB51F8BC" blockId="7.[827,1442,1737,1882]" pageId="7" pageNumber="1044">
Juveniles (i.e. the second instar, measurements in
<tableCitation id="9D46BFC102356150EC1ACBC6FC5AF97A" box="[827,913,1768,1790]" captionStart="Table 4" captionStartId="19.[163,227,196,217]" captionTargetPageId="19" captionText="Table 4. Measurements (in µm) of selected morphological structures of juveniles of Barbaria paucigranulata (type series) mounted in Hoyer’s medium. Abbreviations: N, number of specimens/structures measured; RANGE refers to the smallest and the largest structure among all measured specimens; SD, standard deviation; sp, the proportion of the length of a given structure to the length of the scapular plate" httpUri="http://table.plazi.org/id/84BBDAF202216144EF82CDEAFD67FEB5" pageId="7" pageNumber="1044" tableUuid="84BBDAF202216144EF82CDEAFD67FEB5">Table 4</tableCitation>
). Clear morphometric gap between juveniles and adult females. Qualitatively alike adult females, excluding the lack of gonopore.
</paragraph>
<paragraph id="D07B8A7A02356150EC72CA6AFACFF8DD" blockId="7.[827,1442,1737,1882]" box="[851,1284,1860,1882]" pageId="7" pageNumber="1044">Adult males, larvae or eggs not found.</paragraph>
</subSubSection>
<caption id="84BBDAF2023A615FEFAECEE5FE0AFB9B" ID-DOI="http://doi.org/10.5281/zenodo.6993179" ID-Zenodo-Dep="6993179" httpUri="https://zenodo.org/record/6993179/files/figure.png" pageId="8" pageNumber="1045" startId="8.[143,221,971,993]" targetBox="[145,1425,195,931]" targetPageId="8">
<paragraph id="D07B8A7A023A615FEFAECEE5FE0AFB9B" blockId="8.[143,1425,971,1052]" pageId="8" pageNumber="1045">
<emphasis id="E2B05668023A615FEFAECEE5FF3CFC67" bold="true" box="[143,247,971,993]" pageId="8" pageNumber="1045">Figure 3.</emphasis>
Details of
<taxonomicName id="17C4F1F9023A615FEE4CCEE5FD0BFC66" authority="(SEM)" baseAuthorityName="SEM" box="[365,704,971,993]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEE4CCEE5FDBEFC66" box="[365,629,971,993]" italics="true" pageId="8" pageNumber="1045">Barbaria paucigranulata</emphasis>
(SEM)
</taxonomicName>
: A, pores in the scapular plate; B, pores in the posterior portion of the paired segmental plate II; C, pores in the caudal plate; D, cephalic appendages and a pair of subcephalic plates; E, claws I; F, claws IV. Scale bars in µm.
</paragraph>
</caption>
<subSubSection id="98DED9F1023A615CEFB0C97AFCC0F8E5" lastPageId="11" lastPageNumber="1048" pageId="8" pageNumber="1045" type="diagnosis">
<paragraph id="D07B8A7A023A615FEFB0C97AFE81FB2F" blockId="8.[145,760,1108,1192]" pageId="8" pageNumber="1045">
<emphasis id="E2B05668023A615FEFB0C97AFE21FBED" box="[145,490,1108,1130]" italics="true" pageId="8" pageNumber="1045">Differential diagnosis:</emphasis>
The new species
<taxonomicName id="17C4F1F9023A615FEFB0C95AFEABFB0F" box="[145,352,1139,1161]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEFB0C95AFEABFB0F" box="[145,352,1139,1161]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
is distinguished from its congeners (alphabetically):
</paragraph>
<paragraph id="D07B8A7A023A615FEFB0C9EFFD4AF9E3" blockId="8.[145,761,1216,1882]" pageId="8" pageNumber="1045">
•
<taxonomicName id="17C4F1F9023A615FEF90C9EEFE75FB52" box="[177,446,1216,1237]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="bigranulata">
<emphasis id="E2B05668023A615FEF90C9EEFE75FB52" box="[177,446,1216,1237]" italics="true" pageId="8" pageNumber="1045">Barbaria bigranulata</emphasis>
</taxonomicName>
, by the distribution of pores in plates [absent in the anterior portion of median plate 2 (
<figureCitation id="48FF96FF023A615FEE91C9D0FDCFFA94" box="[432,516,1278,1300]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="8" pageNumber="1045">Fig. 8I</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF023A615FEE34C832FEA5FAB6" box="[277,366,1308,1330]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="8" pageNumber="1045">Fig. 9I</figureCitation>
) and the entirety of median plate
<quantity id="173C279F023A615FEFD8C815FEE1FAD6" box="[249,298,1339,1361]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="1045" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="17C4F1F9023A615FEE16C812FDD9FAD7" box="[311,530,1339,1361]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEE16C812FDD9FAD7" box="[311,530,1339,1361]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
vs. present in the anterior portion of m2 (
<figureCitation id="48FF96FF023A615FEEE5C874FDDDFAE8" box="[452,534,1370,1392]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="8" pageNumber="1045">Fig. 8A</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF023A615FEE34C856FEBDFA0A" box="[277,374,1400,1422]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="8" pageNumber="1045">Fig. 9A</figureCitation>
) and the entirety of m
<quantity id="173C279F023A615FEDE0C856FD33FA09" box="[705,760,1400,1422]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="1045" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="17C4F1F9023A615FEF90C8B6FEA9FA2B" box="[177,354,1431,1453]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="bigranulata">
<emphasis id="E2B05668023A615FEF90C8B6FEA9FA2B" box="[177,354,1431,1453]" italics="true" pageId="8" pageNumber="1045">B. bigranulata</emphasis>
</taxonomicName>
], the shape of papilla IV [tubby in
<taxonomicName id="17C4F1F9023A615FEFF2C898FE66FA4C" box="[211,429,1462,1483]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEFF2C898FE66FA4C" box="[211,429,1462,1483]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEEE0C898FDEEFA4C" box="[449,549,1462,1484]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11L</figureCitation>
) vs. elongated in
<taxonomicName id="17C4F1F9023A615FEF90C8FBFE94FA6E" box="[177,351,1492,1514]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="bigranulata">
<emphasis id="E2B05668023A615FEF90C8FBFE94FA6E" box="[177,351,1492,1514]" italics="true" pageId="8" pageNumber="1045">B. bigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEE53C8FAFE1CFA6E" box="[370,471,1492,1514]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11B</figureCitation>
)] and the primary spur morphology [needle-like and adjacent to the claw branch in
<taxonomicName id="17C4F1F9023A615FEE18CB3CFDDCF9A0" box="[313,535,1554,1575]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEE18CB3CFDDCF9A0" box="[313,535,1554,1575]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FED0ECB3CFD0AF9A0" box="[559,705,1554,1576]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11K, L</figureCitation>
) vs. robust, hook-shaped and divergent from the claw branch in
<taxonomicName id="17C4F1F9023A615FEE08CB7EFE1EF9E3" box="[297,469,1615,1637]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="bigranulata">
<emphasis id="E2B05668023A615FEE08CB7EFE1EF9E3" box="[297,469,1615,1637]" italics="true" pageId="8" pageNumber="1045">B. bigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEEC4CB61FDA3F9E3" box="[485,616,1615,1637]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11A, B</figureCitation>
)].
</paragraph>
<paragraph id="D07B8A7A023A615FEFB0CB40FD6DF97A" blockId="8.[145,761,1216,1882]" pageId="8" pageNumber="1045">
•
<taxonomicName id="17C4F1F9023A615FEF90CB40FEB1F904" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" baseAuthorityName="Claps &amp; Rossi" baseAuthorityYear="1997" box="[177,378,1646,1667]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="charrua">
<emphasis id="E2B05668023A615FEF90CB40FEB1F904" box="[177,378,1646,1667]" italics="true" pageId="8" pageNumber="1045">Barbaria charrua</emphasis>
</taxonomicName>
, by the shape of papilla IV [tubby in
<taxonomicName id="17C4F1F9023A615FEFF4CBA3FE61F926" box="[213,426,1676,1698]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEFF4CBA3FE61F926" box="[213,426,1676,1698]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEEE1CBA2FDEFF926" box="[448,548,1676,1698]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11L</figureCitation>
) vs. elongated in
<taxonomicName id="17C4F1F9023A615FEF90CB82FEE5F947" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" baseAuthorityName="Claps &amp; Rossi" baseAuthorityYear="1997" box="[177,302,1707,1729]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="charrua">
<emphasis id="E2B05668023A615FEF90CB82FEE5F947" box="[177,302,1707,1729]" italics="true" pageId="8" pageNumber="1045">B. charrua</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEE63CB85FE6DF947" box="[322,422,1707,1729]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11D</figureCitation>
)] and by the primary spurs [present in
<taxonomicName id="17C4F1F9023A615FEE64CBE4FDD0F958" box="[325,539,1738,1759]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEE64CBE4FDD0F958" box="[325,539,1738,1759]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FED15CBE4FD09F958" box="[564,706,1738,1760]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11K, L</figureCitation>
) vs. typically absent in
<taxonomicName id="17C4F1F9023A615FEEABCBC7FDC8F97A" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" baseAuthorityName="Claps &amp; Rossi" baseAuthorityYear="1997" box="[394,515,1768,1790]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="charrua">
<emphasis id="E2B05668023A615FEEABCBC7FDC8F97A" box="[394,515,1768,1790]" italics="true" pageId="8" pageNumber="1045">B. charrua</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FED33CBC6FD44F979" box="[530,655,1768,1790]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11C, D</figureCitation>
)].
</paragraph>
<paragraph id="D07B8A7A023A615FEFB0CA29FB67FB0F" blockId="8.[145,761,1216,1882]" lastBlockId="8.[809,1426,1108,1897]" pageId="8" pageNumber="1045">
•
<taxonomicName id="17C4F1F9023A615FEF90CA29FEBAF89B" box="[177,369,1799,1820]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="danieli">
<emphasis id="E2B05668023A615FEF90CA29FEBAF89B" box="[177,369,1799,1820]" italics="true" pageId="8" pageNumber="1045">Barbaria danieli</emphasis>
</taxonomicName>
, by the shape of papilla IV [tubby in
<taxonomicName id="17C4F1F9023A615FEFF4CA08FE61F8BC" box="[213,426,1830,1851]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEFF4CA08FE61F8BC" box="[213,426,1830,1851]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEEE1CA08FDEFF8BC" box="[448,548,1830,1852]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11L</figureCitation>
) vs. elongated in
<taxonomicName id="17C4F1F9023A615FEF90CA6BFEECF8DE" box="[177,295,1860,1882]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="danieli">
<emphasis id="E2B05668023A615FEF90CA6BFEECF8DE" box="[177,295,1860,1882]" italics="true" pageId="8" pageNumber="1045">B. danieli</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEE1FCA6AFE5FF8DE" box="[318,404,1860,1882]" captionStart="Figure 12" captionStartId="21.[161,240,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-388@21.[163,1443,195,1071]" captionTargetPageId="21" captionText="Figure 12. Habitus of Barbaria danieli (PCM, female from Alabama). Arrowheads indicate rudimentary papillae on legs II–III. Scale bar in µm." figureDoi="http://doi.org/10.5281/zenodo.6993201" httpUri="https://zenodo.org/record/6993201/files/figure.png" pageId="8" pageNumber="1045">Fig. 12</figureCitation>
)] and by the primary spurs [present in
<taxonomicName id="17C4F1F9023A615FECFCC97BFB78FBEE" box="[989,1203,1108,1130]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FECFCC97BFB78FBEE" box="[989,1203,1108,1130]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEBEDC97AFA91FBED" box="[1228,1370,1108,1130]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11K, L</figureCitation>
) vs. absent in
<taxonomicName id="17C4F1F9023A615FEC98C95AFBEDFB0F" box="[953,1062,1139,1161]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="danieli">
<emphasis id="E2B05668023A615FEC98C95AFBEDFB0F" box="[953,1062,1139,1161]" italics="true" pageId="8" pageNumber="1045">B. danieli</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEB14C95DFB5EFB0F" box="[1077,1173,1139,1161]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11O</figureCitation>
)].
</paragraph>
<paragraph id="D07B8A7A023A615FEC08C9BCFB01F9B1" blockId="8.[809,1426,1108,1897]" pageId="8" pageNumber="1045">
•
<taxonomicName id="17C4F1F9023A615FEC68C9BCFBF6FB20" baseAuthorityName="Michalczyk &amp; Kaczmarek" baseAuthorityYear="2007" box="[841,1085,1170,1191]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="ganczareki">
<emphasis id="E2B05668023A615FEC68C9BCFBF6FB20" box="[841,1085,1170,1191]" italics="true" pageId="8" pageNumber="1045">Barbaria ganczareki</emphasis>
</taxonomicName>
, by the dorsal sculpturing [both pillars and pores easily identifiable in
<taxonomicName id="17C4F1F9023A615FEC68C9FEFBDCFB63" box="[841,1047,1231,1253]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEC68C9FEFBDCFB63" box="[841,1047,1231,1253]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEB07C9E1FBBCFB63" box="[1062,1143,1231,1253]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="8" pageNumber="1045">Figs 7I</figureCitation>
,
<figureCitation id="48FF96FF023A615FEBA5C9E1FB50FB63" box="[1156,1179,1231,1252]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="8" pageNumber="1045">8I</figureCitation>
,
<figureCitation id="48FF96FF023A615FEB88C9E1FB0AFB63" box="[1193,1217,1231,1252]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="8" pageNumber="1045">9I</figureCitation>
,
<figureCitation id="48FF96FF023A615FEBEFC9E1FB3FFB63" box="[1230,1268,1231,1252]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="8" pageNumber="1045">10I</figureCitation>
) vs. pillars so poorly developed that pores become the dominant element of the sculpture in
<taxonomicName id="17C4F1F9023A615FEB88C823FA82FAA6" baseAuthorityName="Michalczyk &amp; Kaczmarek" baseAuthorityYear="2007" box="[1193,1353,1292,1314]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="ganczareki">
<emphasis id="E2B05668023A615FEB88C823FA82FAA6" box="[1193,1353,1292,1314]" italics="true" pageId="8" pageNumber="1045">B. ganczareki</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEA41C822FCACFAC7" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="8" pageNumber="1045">Figs 7C</figureCitation>
,
<figureCitation id="48FF96FF023A615FEC5DC805FC50FAC7" box="[892,923,1323,1344]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="8" pageNumber="1045">8D</figureCitation>
,
<figureCitation id="48FF96FF023A615FEC91C805FC04FAC7" box="[944,975,1323,1344]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="8" pageNumber="1045">9C</figureCitation>
,
<figureCitation id="48FF96FF023A615FECC2C805FBDBFAC7" box="[995,1040,1323,1344]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="8" pageNumber="1045">10C</figureCitation>
)], the pores in the subcephalic region (absent in
<taxonomicName id="17C4F1F9023A615FEB08C864FB35FAD8" box="[1065,1278,1354,1375]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEB08C864FB35FAD8" box="[1065,1278,1354,1375]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
vs. present in
<taxonomicName id="17C4F1F9023A615FEC4CC847FBC0FAFA" baseAuthorityName="Michalczyk &amp; Kaczmarek" baseAuthorityYear="2007" box="[877,1035,1384,1406]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="ganczareki">
<emphasis id="E2B05668023A615FEC4CC847FBC0FAFA" box="[877,1035,1384,1406]" italics="true" pageId="8" pageNumber="1045">B. ganczareki</emphasis>
</taxonomicName>
) and in pedal plates (absent in
<taxonomicName id="17C4F1F9023A615FEC68C8A6FBDDFA1B" box="[841,1046,1415,1437]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEC68C8A6FBDDFA1B" box="[841,1046,1415,1437]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
vs. present in
<taxonomicName id="17C4F1F9023A615FEB9FC8A6FA9CFA1B" baseAuthorityName="Michalczyk &amp; Kaczmarek" baseAuthorityYear="2007" box="[1214,1367,1415,1437]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="ganczareki">
<emphasis id="E2B05668023A615FEB9FC8A6FA9CFA1B" box="[1214,1367,1415,1437]" italics="true" pageId="8" pageNumber="1045">B. ganczareki</emphasis>
</taxonomicName>
) and by the primary spur morphology [needle-like and adjacent to the claw branch in
<taxonomicName id="17C4F1F9023A615FEB9EC8EBFA5BFA5E" box="[1215,1424,1476,1498]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEB9EC8EBFA5BFA5E" box="[1215,1424,1476,1498]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEC70C8CDFC04FA7F" box="[849,975,1507,1529]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11K, L</figureCitation>
) vs. robust, hook-shaped and divergent from the claw branch in
<taxonomicName id="17C4F1F9023A615FEB41CB2CFB33F991" baseAuthorityName="Michalczyk &amp; Kaczmarek" baseAuthorityYear="2007" box="[1120,1272,1537,1559]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="ganczareki">
<emphasis id="E2B05668023A615FEB41CB2CFB33F991" box="[1120,1272,1537,1559]" italics="true" pageId="8" pageNumber="1045">B. ganczareki</emphasis>
</taxonomicName>
, fig.
<quantity id="173C279F023A615FEA0FCB2CFA5AF990" box="[1326,1425,1537,1559]" metricMagnitude="-1" metricUnit="m" metricValue="5.461" metricValueMax="5.588" metricValueMin="5.334" pageId="8" pageNumber="1045" unit="in" value="21.5" valueMax="22.0" valueMin="21.0">21–22 in</quantity>
<bibRefCitation id="B455F78B023A615FEC68CB0EFB70F9B2" author="Michalczyk &amp; Kaczmarek" box="[841,1211,1568,1590]" firstAuthor="Michalczyk" pageId="8" pageNumber="1045" pagination="15 - 25" refId="ref17106" refString="Michalczyk L, Kaczmarek L. 2007. Echiniscus ganczareki, a new species of Tardigrada (Heterotardigrada: Echiniscidae, bigranulatus group) from Costa Rica. Zootaxa 1471: 15 - 25." type="journal article" year="2007">Michalczyk &amp; Kaczmarek (2007)</bibRefCitation>
].
</paragraph>
<paragraph id="D07B8A7A023A615FEC08CB11FA82F8AC" blockId="8.[809,1426,1108,1897]" pageId="8" pageNumber="1045">
•
<taxonomicName id="17C4F1F9023A615FEC68CB11FBD7F9D3" box="[841,1052,1599,1620]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="hannae">
<emphasis id="E2B05668023A615FEC68CB11FBD7F9D3" box="[841,1052,1599,1620]" italics="true" pageId="8" pageNumber="1045">Barbaria hannae</emphasis>
</taxonomicName>
, by the dorsal sculpturing [no smooth plate portions in
<taxonomicName id="17C4F1F9023A615FEB9CCB70FA5AF9F5" box="[1213,1425,1629,1651]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEB9CCB70FA5AF9F5" box="[1213,1425,1629,1651]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEC70CB52FC56F916" box="[849,925,1660,1682]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="8" pageNumber="1045">Figs 8I</figureCitation>
,
<figureCitation id="48FF96FF023A615FEC89CB52FC74F916" box="[936,959,1660,1681]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="8" pageNumber="1045">9I</figureCitation>
,
<figureCitation id="48FF96FF023A615FECEBCB52FC25F916" box="[970,1006,1660,1681]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="8" pageNumber="1045">10I</figureCitation>
) vs. thickened plate portions devoid of sculpturing present in
<taxonomicName id="17C4F1F9023A615FEB60CBB5FB64F937" box="[1089,1199,1691,1712]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="hannae">
<emphasis id="E2B05668023A615FEB60CBB5FB64F937" box="[1089,1199,1691,1712]" italics="true" pageId="8" pageNumber="1045">B. hannae</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEB9ACBB5FAC4F937" box="[1211,1295,1691,1713]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="8" pageNumber="1045">Figs 8E</figureCitation>
,
<figureCitation id="48FF96FF023A615FEA39CBB5FA9FF937" box="[1304,1364,1691,1712]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="8" pageNumber="1045">9D, E</figureCitation>
,
<figureCitation id="48FF96FF023A615FEA7FCBB5FC90F948" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="8" pageNumber="1045">10D, E</figureCitation>
)] and the primary spur morphology [needle-like and adjacent to the claw branch in
<taxonomicName id="17C4F1F9023A615FEBE9CBF7FA5AF96A" box="[1224,1425,1752,1774]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEBE9CBF7FA5AF96A" box="[1224,1425,1752,1774]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEC70CBD9FC1BF88B" box="[849,976,1783,1805]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11K, L</figureCitation>
) vs. robust, hook-shaped and divergent from the claw branch in
<taxonomicName id="17C4F1F9023A615FEB78CA38FB02F8AD" box="[1113,1225,1813,1835]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="hannae">
<emphasis id="E2B05668023A615FEB78CA38FB02F8AD" box="[1113,1225,1813,1835]" italics="true" pageId="8" pageNumber="1045">B. hannae</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023A615FEBF6CA3BFAF9F8AC" box="[1239,1330,1813,1835]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="8" pageNumber="1045">Fig. 11P</figureCitation>
)].
</paragraph>
<paragraph id="D07B8A7A023A615DEC08CA1BFE20F98B" blockId="8.[809,1426,1108,1897]" lastBlockId="10.[138,761,1251,1886]" lastPageId="10" lastPageNumber="1047" pageId="8" pageNumber="1045">
•
<taxonomicName id="17C4F1F9023A615FEC68CA1AFBEBF8CD" box="[841,1056,1844,1866]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="jenningsi">
<emphasis id="E2B05668023A615FEC68CA1AFBEBF8CD" box="[841,1056,1844,1866]" italics="true" pageId="8" pageNumber="1045">Barbaria jenningsi</emphasis>
</taxonomicName>
, by the cirrus
<emphasis id="E2B05668023A615FEBE3CA1AFB18F8CE" box="[1218,1235,1844,1865]" italics="true" pageId="8" pageNumber="1045">A</emphasis>
length (&lt;20% of the body length in
<taxonomicName id="17C4F1F9023A615FEB37CA7DFB29F8EF" box="[1046,1250,1875,1896]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="8" pageNumber="1045" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023A615FEB37CA7DFB29F8EF" box="[1046,1250,1875,1896]" italics="true" pageId="8" pageNumber="1045">B. paucigranulata</emphasis>
</taxonomicName>
vs.&gt; 50% of the body length in
<taxonomicName id="17C4F1F9023B615EEE4FCBD5FE3EF897" box="[366,501,1787,1808]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="9" pageNumber="1046" phylum="Tardigrada" rank="species" species="jenningsi">
<emphasis id="E2B05668023B615EEE4FCBD5FE3EF897" box="[366,501,1787,1808]" italics="true" pageId="9" pageNumber="1046">B. jenningsi</emphasis>
</taxonomicName>
), the
<typeStatus id="0F7F34D8023B615EED16CBD4FDA3F888" box="[567,616,1786,1807]" pageId="9" pageNumber="1046">type</typeStatus>
of perforation in the dorsal plates [pores in
<taxonomicName id="17C4F1F9023B615EED17CA37FCC1F8A9" box="[566,778,1817,1838]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="9" pageNumber="1046" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023B615EED17CA37FCC1F8A9" box="[566,778,1817,1838]" italics="true" pageId="9" pageNumber="1046">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023B615EEFEACA19FED2F8CA" box="[203,281,1847,1869]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="9" pageNumber="1046">Fig. 7I</figureCitation>
) vs. pseudopores in
<taxonomicName id="17C4F1F9023B615EED30CA16FD57F8CA" box="[529,668,1848,1869]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="9" pageNumber="1046" phylum="Tardigrada" rank="species" species="jenningsi">
<emphasis id="E2B05668023B615EED30CA16FD57F8CA" box="[529,668,1848,1869]" italics="true" pageId="9" pageNumber="1046">B. jenningsi</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023B615EED8ECA19FF18F8EC" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="9" pageNumber="1046">Fig. 7E, F</figureCitation>
)] and by the primary spur morphology [slightly heteronych, needle-like and adjacent to the claw branch in
<taxonomicName id="17C4F1F9023B615EECF2CA37FB68F8A9" box="[979,1187,1817,1838]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="9" pageNumber="1046" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B05668023B615EECF2CA37FB68F8A9" box="[979,1187,1817,1838]" italics="true" pageId="9" pageNumber="1046">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023B615EEB95CA37FAFDF8A9" box="[1204,1334,1817,1839]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="9" pageNumber="1046">Fig. 11K, L</figureCitation>
) vs. fully heteronych, robust, hook-shaped and divergent from the claw branch in
<taxonomicName id="17C4F1F9023B615EEB50CA79FB3CF8EB" box="[1137,1271,1879,1900]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="9" pageNumber="1046" phylum="Tardigrada" rank="species" species="jenningsi">
<emphasis id="E2B05668023B615EEB50CA79FB3CF8EB" box="[1137,1271,1879,1900]" italics="true" pageId="9" pageNumber="1046">B. jenningsi</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF023B615EEA27CA78FA4AF8EC" box="[1286,1409,1878,1900]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="9" pageNumber="1046">Fig. 11E, F</figureCitation>
)]. •
<taxonomicName id="17C4F1F90238615DEF90C9CDFE68FB7F" box="[177,419,1251,1272]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="madonnae">
<emphasis id="E2B056680238615DEF90C9CDFE68FB7F" box="[177,419,1251,1272]" italics="true" pageId="10" pageNumber="1047">Barbaria madonnae</emphasis>
</taxonomicName>
, by
<emphasis id="E2B056680238615DEECBC9CAFDE1FB7E" box="[490,554,1252,1273]" italics="true" pageId="10" pageNumber="1047">striae</emphasis>
between pillars in the scapular and the caudal plate [present in
<taxonomicName id="17C4F1F90238615DEFF5C80FFE6CFAB1" box="[212,423,1313,1334]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEFF5C80FFE6CFAB1" box="[212,423,1313,1334]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEE9CC80FFDD9FAB1" box="[445,530,1313,1335]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="10" pageNumber="1047">Figs 7I</figureCitation>
,
<figureCitation id="48FF96FF0238615DED05C80FFD81FAB1" box="[548,586,1313,1334]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="10" pageNumber="1047">10I</figureCitation>
) vs. absent in
<taxonomicName id="17C4F1F90238615DEF90C86EFE8FFAD3" box="[177,324,1343,1365]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="madonnae">
<emphasis id="E2B056680238615DEF90C86EFE8FFAD3" box="[177,324,1343,1365]" italics="true" pageId="10" pageNumber="1047">B. madonnae</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEE70C811FE6CFAD3" box="[337,423,1343,1365]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="10" pageNumber="1047">Figs 7G</figureCitation>
,
<figureCitation id="48FF96FF0238615DEE93C811FE2BFAD3" box="[434,480,1343,1364]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="10" pageNumber="1047">10G</figureCitation>
)], the shape of papilla IV [tubby in
<taxonomicName id="17C4F1F90238615DEE3FC871FE20FAF4" box="[286,491,1374,1396]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEE3FC871FE20FAF4" box="[286,491,1374,1396]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEEDBC870FD9CFAF4" box="[506,599,1374,1396]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11L</figureCitation>
) vs. elongated in
<taxonomicName id="17C4F1F90238615DEFF3C853FEA2FA15" box="[210,361,1405,1426]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="madonnae">
<emphasis id="E2B056680238615DEFF3C853FEA2FA15" box="[210,361,1405,1426]" italics="true" pageId="10" pageNumber="1047">B. madonnae</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEE5AC853FE14FA15" box="[379,479,1405,1427]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11H</figureCitation>
)] and the primary spur morphology [needle-like and adjacent to the claw branch in
<taxonomicName id="17C4F1F90238615DEE03C895FE24FA48" box="[290,495,1466,1488]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEE03C895FE24FA48" box="[290,495,1466,1488]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEEDCC894FDB3FA48" box="[509,632,1466,1488]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11K, L</figureCitation>
) vs. robust, hook-shaped and divergent from the claw branch in
<taxonomicName id="17C4F1F90238615DEF90C8D6FE8EF98B" box="[177,325,1527,1549]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="madonnae">
<emphasis id="E2B056680238615DEF90C8D6FE8EF98B" box="[177,325,1527,1549]" italics="true" pageId="10" pageNumber="1047">B. madonnae</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEE75C8D9FE1EF98A" box="[340,469,1527,1549]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11G, H</figureCitation>
)].
</paragraph>
<caption id="84BBDAF2023B615EEF82CB8EFA6FF953" ID-DOI="http://doi.org/10.5281/zenodo.6993181" ID-Zenodo-Dep="6993181" httpUri="https://zenodo.org/record/6993181/files/figure.png" pageId="9" pageNumber="1046" startId="9.[163,243,1696,1718]" targetBox="[163,1443,195,1656]" targetPageId="9">
<paragraph id="D07B8A7A023B615EEF82CB8EFA6FF953" blockId="9.[163,1444,1696,1748]" pageId="9" pageNumber="1046">
<emphasis id="E2B05668023B615EEF82CB8EFEC6F932" bold="true" box="[163,269,1696,1718]" pageId="9" pageNumber="1046">Figure 4.</emphasis>
Habitus of
<taxonomicName id="17C4F1F9023B615EEEB1CB8EFD75F931" authority="(PCM)" baseAuthorityName="PCM" box="[400,702,1696,1718]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="9" pageNumber="1046" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="E2B05668023B615EEEB1CB8EFDA4F931" box="[400,623,1696,1718]" italics="true" pageId="9" pageNumber="1046">Barbaria weglarskae</emphasis>
(PCM)
</taxonomicName>
: A, holotype (female) in dorsal view; B, dorsal sculpturing in close-up (paratype); C, subcephalic plates (paratype); D, subcephalic plates (holotype); E, gonoporal area (holotype). Scale bars in µm.
</paragraph>
</caption>
<caption id="84BBDAF20238615DEFB0C969FB08FB32" ID-DOI="http://doi.org/10.5281/zenodo.6993185" ID-Zenodo-Dep="6993185" httpUri="https://zenodo.org/record/6993185/files/figure.png" pageId="10" pageNumber="1047" startId="10.[145,224,1095,1117]" targetBox="[244,1320,198,1056]" targetPageId="10">
<paragraph id="D07B8A7A0238615DEFB0C969FB08FB32" blockId="10.[144,1426,1095,1205]" pageId="10" pageNumber="1047">
<emphasis id="E2B056680238615DEFB0C969FF31FBDB" bold="true" box="[145,250,1095,1117]" pageId="10" pageNumber="1047">Figure 5.</emphasis>
Phylogeny of the genus
<taxonomicName id="17C4F1F90238615DED25C969FDAEFBDA" box="[516,613,1095,1117]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="genus">
<emphasis id="E2B056680238615DED25C969FDAEFBDA" box="[516,613,1095,1117]" italics="true" pageId="10" pageNumber="1047">Barbaria</emphasis>
</taxonomicName>
based on five concatenated markers conducted in MrBayes and W-IQ-TREE (values above the nodes signify posterior probabilities, whereas bootstraps are provided below the nodes; maximal supports are denoted with asterisks).
<taxonomicName id="17C4F1F90238615DEE94C9ACFD77FB1F" baseAuthorityName="Richters" baseAuthorityYear="1903" box="[437,700,1154,1176]" class="Heterotardigrada" family="Echiniscidae" genus="Diploechiniscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="oihonnae">
<emphasis id="E2B056680238615DEE94C9ACFD77FB1F" box="[437,700,1154,1176]" italics="true" pageId="10" pageNumber="1047">Diploechiniscus oihonnae</emphasis>
</taxonomicName>
and
<taxonomicName id="17C4F1F90238615DEDCFC9ACFBB4FB1F" box="[750,1151,1154,1176]" class="Heterotardigrada" family="Echiniscidae" genus="Testechiniscus" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="subSpecies" species="spitsbergensis" subSpecies="tropicalis">
<emphasis id="E2B056680238615DEDCFC9ACFBB4FB1F" box="[750,1151,1154,1176]" italics="true" pageId="10" pageNumber="1047">Testechiniscus spitsbergensis tropicalis</emphasis>
</taxonomicName>
were used as an outgroup. The scale bar represents 0.2 substitutions per nucleotide position and refers to the Bayesian inference.
</paragraph>
</caption>
<paragraph id="D07B8A7A0238615DEFB0CB38FE8AF8D9" blockId="10.[138,761,1251,1886]" pageId="10" pageNumber="1047">
•
<taxonomicName id="17C4F1F90238615DEF90CB38FDC8F9AC" box="[177,515,1558,1579]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ollantaytamboensis">
<emphasis id="E2B056680238615DEF90CB38FDC8F9AC" box="[177,515,1558,1579]" italics="true" pageId="10" pageNumber="1047">Barbaria ollantaytamboensis</emphasis>
</taxonomicName>
, by the distribution of pores in plates [absent in the anterior portion of median plate 2 (
<figureCitation id="48FF96FF0238615DEE56CB7DFE0EF9EF" box="[375,453,1619,1641]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="10" pageNumber="1047">Fig. 8I</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF0238615DEF98CB5CFEC1F900" box="[185,266,1650,1672]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="10" pageNumber="1047">Fig. 9I</figureCitation>
) and the entirety of median plate
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<taxonomicName id="17C4F1F90238615DEF90CBBFFEB7F921" box="[177,380,1681,1702]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEF90CBBFFEB7F921" box="[177,380,1681,1702]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
vs. present in the anterior portion of m2 (
<figureCitation id="48FF96FF0238615DEE25CB81FE90F943" box="[260,347,1711,1733]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="10" pageNumber="1047">Fig. 8H</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF0238615DEDBBCB81FD3AF943" box="[666,753,1711,1733]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="10" pageNumber="1047">Fig. 9H</figureCitation>
) and the entirety of m
<quantity id="173C279F0238615DEE96CBE0FE2DF963" box="[439,486,1742,1764]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="1047" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="17C4F1F90238615DEED3CBE1FD3BF964" box="[498,752,1742,1764]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ollantaytamboensis">
<emphasis id="E2B056680238615DEED3CBE1FD3BF964" box="[498,752,1742,1764]" italics="true" pageId="10" pageNumber="1047">B. ollantaytamboensis</emphasis>
</taxonomicName>
] and claw isomorphy [slightly heteronych (heteromorphic) in
<taxonomicName id="17C4F1F90238615DEEAECA22FD95F8A7" box="[399,606,1803,1825]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEEAECA22FD95F8A7" box="[399,606,1803,1825]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DED51CA25FD3BF8A6" box="[624,752,1803,1825]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11K, L</figureCitation>
) vs. isonych (homomorphic) in
<taxonomicName id="17C4F1F90238615DED21CA05FD32F8B8" box="[512,761,1834,1856]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ollantaytamboensis">
<emphasis id="E2B056680238615DED21CA05FD32F8B8" box="[512,761,1834,1856]" italics="true" pageId="10" pageNumber="1047">B. ollantaytamboensis</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEF98CA66FEE0F8D9" box="[185,299,1864,1886]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11I, J</figureCitation>
)].
</paragraph>
<paragraph id="D07B8A7A0238615DEC08C9CAFBFAFA48" blockId="10.[809,1426,1251,1886]" pageId="10" pageNumber="1047">
•
<taxonomicName id="17C4F1F90238615DEC68C9CDFBD6FB7E" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" baseAuthorityName="Pilato" baseAuthorityYear="2007" box="[841,1053,1251,1273]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="quitensis">
<emphasis id="E2B056680238615DEC68C9CDFBD6FB7E" box="[841,1053,1251,1273]" italics="true" pageId="10" pageNumber="1047">Barbaria quitensis</emphasis>
</taxonomicName>
, by the
<typeStatus id="0F7F34D80238615DEB57C9CAFB6CFB7E" box="[1142,1191,1252,1273]" pageId="10" pageNumber="1047">type</typeStatus>
of perforation in the dorsal plates [pores in
<taxonomicName id="17C4F1F90238615DEB62C82DFAC5FA90" box="[1091,1294,1282,1304]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEB62C82DFAC5FA90" box="[1091,1294,1282,1304]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEA3AC82CFAA2FA90" box="[1307,1385,1282,1304]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="10" pageNumber="1047">Figs 7I</figureCitation>
,
<figureCitation id="48FF96FF0238615DEA52C82CFA41FA90" box="[1395,1418,1282,1303]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="10" pageNumber="1047">8I</figureCitation>
,
<figureCitation id="48FF96FF0238615DEC68C80FFCAAFAB1" box="[841,865,1313,1334]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="10" pageNumber="1047">9I</figureCitation>
,
<figureCitation id="48FF96FF0238615DEC4FC80FFC5FFAB1" box="[878,916,1313,1334]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="10" pageNumber="1047">10I</figureCitation>
) vs. pseudopores in
<taxonomicName id="17C4F1F90238615DEB5CC80FFACAFAB1" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" baseAuthorityName="Pilato" baseAuthorityYear="2007" box="[1149,1281,1313,1334]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="quitensis">
<emphasis id="E2B056680238615DEB5CC80FFACAFAB1" box="[1149,1281,1313,1334]" italics="true" pageId="10" pageNumber="1047">B. quitensis</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEA30C80EFAAFFAB1" box="[1297,1380,1312,1334]" captionStart="Figure 7" captionStartId="12.[145,224,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-146@12.[145,1425,195,1487]" captionTargetPageId="12" captionText="Figure 7. Variability of the scapular plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, Barbaria hannae; E, F, Barbaria jenningsi at two focus levels; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993189" httpUri="https://zenodo.org/record/6993189/files/figure.png" pageId="10" pageNumber="1047">Figs 7J</figureCitation>
,
<figureCitation id="48FF96FF0238615DEA50C80FFA41FAB1" box="[1393,1418,1313,1334]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="10" pageNumber="1047">8J</figureCitation>
,
<figureCitation id="48FF96FF0238615DEC68C811FCA8FAD3" box="[841,867,1343,1364]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="10" pageNumber="1047">9J</figureCitation>
,
<figureCitation id="48FF96FF0238615DEC50C811FC51FAD3" box="[881,922,1343,1364]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="10" pageNumber="1047">10J</figureCitation>
)] and the distribution of pores/pseudopores [scarcer on lateralmost portions of the caudal plate (
<figureCitation id="48FF96FF0238615DEC70C852FC60FA15" box="[849,939,1404,1426]" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="10" pageNumber="1047">Fig. 10I</figureCitation>
) in
<taxonomicName id="17C4F1F90238615DECFDC853FB60FA16" box="[988,1195,1404,1426]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DECFDC853FB60FA16" box="[988,1195,1404,1426]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
vs. roughly equally distributed in all portions of the caudal plate (
<figureCitation id="48FF96FF0238615DEA47C8B5FCB9FA48" captionStart="Figure 10" captionStartId="15.[161,239,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-205@15.[163,1443,195,1487]" captionTargetPageId="15" captionText="Figure 10. Variability of the caudal (terminal) plate sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D,E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993195" httpUri="https://zenodo.org/record/6993195/files/figure.png" pageId="10" pageNumber="1047">Fig. 10J</figureCitation>
) in
<taxonomicName id="17C4F1F90238615DECBEC894FBE9FA48" authorityName="Gąsiorek &amp; Wilamowski &amp; Vončina &amp; Michalczyk" authorityYear="2022" baseAuthorityName="Pilato" baseAuthorityYear="2007" box="[927,1058,1466,1487]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="quitensis">
<emphasis id="E2B056680238615DECBEC894FBE9FA48" box="[927,1058,1466,1487]" italics="true" pageId="10" pageNumber="1047">B. quitensis</emphasis>
</taxonomicName>
].
</paragraph>
<paragraph id="D07B8A7A0238615DEC08C8F7FB3BF8A6" blockId="10.[809,1426,1251,1886]" pageId="10" pageNumber="1047">
•
<taxonomicName id="17C4F1F90238615DEC68C8F6FC3CFA69" baseAuthorityName="Ramazzotti" baseAuthorityYear="1964" box="[841,1015,1496,1518]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ranzii">
<emphasis id="E2B056680238615DEC68C8F6FC3CFA69" box="[841,1015,1496,1518]" italics="true" pageId="10" pageNumber="1047">Barbaria ranzii</emphasis>
</taxonomicName>
, by the cirrus
<emphasis id="E2B056680238615DEBB9C8F7FB62FA69" box="[1176,1193,1497,1518]" italics="true" pageId="10" pageNumber="1047">A</emphasis>
length (&lt;20% of the body length in
<taxonomicName id="17C4F1F90238615DECC9C8D6FB65F98B" box="[1000,1198,1527,1549]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DECC9C8D6FB65F98B" box="[1000,1198,1527,1549]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
vs.&gt; 50% of the body length in
<taxonomicName id="17C4F1F90238615DEC93CB38FBDBF9AC" baseAuthorityName="Ramazzotti" baseAuthorityYear="1964" box="[946,1040,1558,1579]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ranzii">
<emphasis id="E2B056680238615DEC93CB38FBDBF9AC" box="[946,1040,1558,1579]" italics="true" pageId="10" pageNumber="1047">B. ranzii</emphasis>
</taxonomicName>
), the distribution of pores in plates [absent in the anterior portion of median plate 2 (
<figureCitation id="48FF96FF0238615DEA46CB1AFCABF9EF" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="10" pageNumber="1047">Fig. 8I</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF0238615DEBABCB7DFB1BF9EF" box="[1162,1232,1619,1641]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="10" pageNumber="1047">Fig. 9I</figureCitation>
) and the entirety of median plate
<quantity id="173C279F0238615DECDDCB5CFBEDF90F" box="[1020,1062,1650,1672]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="1047" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="17C4F1F90238615DEB0CCB5CFB3EF900" box="[1069,1269,1650,1671]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEB0CCB5CFB3EF900" box="[1069,1269,1650,1671]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
vs. present in the anterior portion of m2 (
<figureCitation id="48FF96FF0238615DEB51CBBEFB74F921" box="[1136,1215,1680,1702]" captionStart="Figure 8" captionStartId="13.[163,245,1527,1549]" captionTargetBox="[163,1443,195,1487]" captionTargetId="figure-126@13.[163,1443,195,1487]" captionTargetPageId="13" captionText="Figure 8. Variability of the median plate 2 sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria danieli; D, Barbaria ganczareki; E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993191" httpUri="https://zenodo.org/record/6993191/files/figure.png" pageId="10" pageNumber="1047">Fig. 8K</figureCitation>
), paired segmental plates (
<figureCitation id="48FF96FF0238615DECBDCB81FC24F943" box="[924,1007,1711,1733]" captionStart="Figure 9" captionStartId="14.[145,223,1527,1549]" captionTargetBox="[145,1425,195,1487]" captionTargetId="figure-169@14.[145,1425,195,1487]" captionTargetPageId="14" captionText="Figure 9. Variability of the paired segmental plate II sculpturing (PCM): A, Barbaria bigranulata; B, Barbaria charrua; C, Barbaria ganczareki; D, E, Barbaria hannae; F, Barbaria jenningsi; G, Barbaria madonnae; H, Barbaria ollantaytamboensis; I, Barbaria paucigranulata; J, Barbaria quitensis; K, Barbaria ranzii; L, Barbaria weglarskae. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993193" httpUri="https://zenodo.org/record/6993193/files/figure.png" pageId="10" pageNumber="1047">Fig. 9K</figureCitation>
) and the entirety of m
<quantity id="173C279F0238615DEBD5CB81FAD4F942" box="[1268,1311,1711,1733]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="10" pageNumber="1047" unit="in" value="3.0">3 in</quantity>
<taxonomicName id="17C4F1F90238615DEA09CB9EFA42F942" baseAuthorityName="Ramazzotti" baseAuthorityYear="1964" box="[1320,1417,1712,1733]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ranzii">
<emphasis id="E2B056680238615DEA09CB9EFA42F942" box="[1320,1417,1712,1733]" italics="true" pageId="10" pageNumber="1047">B. ranzii</emphasis>
</taxonomicName>
] and the by secondary spurs directed upwards on external claws IV [absent in
<taxonomicName id="17C4F1F90238615DEBAFCBC3FA9CF886" box="[1166,1367,1772,1794]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEBAFCBC3FA9CF886" box="[1166,1367,1772,1794]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0238615DEA46CBC2FCB8F8A7" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11L</figureCitation>
) vs. present in
<taxonomicName id="17C4F1F90238615DEB3DCA22FBB1F8A6" baseAuthorityName="Ramazzotti" baseAuthorityYear="1964" box="[1052,1146,1804,1825]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="ranzii">
<emphasis id="E2B056680238615DEB3DCA22FBB1F8A6" box="[1052,1146,1804,1825]" italics="true" pageId="10" pageNumber="1047">B. ranzii</emphasis>
</taxonomicName>
,
<figureCitation id="48FF96FF0238615DEBA7CA25FB2AF8A7" box="[1158,1249,1803,1825]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="10" pageNumber="1047">Fig. 11R</figureCitation>
].
</paragraph>
<paragraph id="D07B8A7A0238615CEC08CA04FCC0F8E5" blockId="10.[809,1426,1251,1886]" lastBlockId="11.[195,779,1746,1891]" lastPageId="11" lastPageNumber="1048" pageId="10" pageNumber="1047">
•
<taxonomicName id="17C4F1F90238615DEC68CA04FB88F8B8" box="[841,1091,1834,1855]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="E2B056680238615DEC68CA04FB88F8B8" box="[841,1091,1834,1855]" italics="true" pageId="10" pageNumber="1047">Barbaria weglarskae</emphasis>
</taxonomicName>
, by the cirrus
<emphasis id="E2B056680238615DEA00CA04FAF9F8B8" box="[1313,1330,1834,1855]" italics="true" pageId="10" pageNumber="1047">A</emphasis>
length (&lt;20% of the body length in
<taxonomicName id="17C4F1F90238615DEB9FCA67FA5AF8DA" box="[1214,1425,1864,1886]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="10" pageNumber="1047" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680238615DEB9FCA67FA5AF8DA" box="[1214,1425,1864,1886]" italics="true" pageId="10" pageNumber="1047">B. paucigranulata</emphasis>
</taxonomicName>
vs.&gt; 50% of the body length in
<taxonomicName id="17C4F1F90239615CED7ECBFDFCC9F96F" box="[607,770,1747,1768]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="11" pageNumber="1048" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="E2B056680239615CED7ECBFDFDBAF96F" box="[607,625,1747,1768]" italics="true" pageId="11" pageNumber="1048">B</emphasis>
.
<emphasis id="E2B056680239615CEDA4CBFDFCC9F96F" box="[645,770,1747,1768]" italics="true" pageId="11" pageNumber="1048">weglarskae</emphasis>
</taxonomicName>
) and by the primary spur morphology [needle-like and adjacent to the claw branch in
<taxonomicName id="17C4F1F90239615CED60CA3FFCC0F8A2" box="[577,779,1808,1830]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="11" pageNumber="1048" phylum="Tardigrada" rank="species" species="paucigranulata">
<emphasis id="E2B056680239615CED60CA3FFCC0F8A2" box="[577,779,1808,1830]" italics="true" pageId="11" pageNumber="1048">B. paucigranulata</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0239615CEFEACA01FE82F8C3" box="[203,329,1839,1861]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="11" pageNumber="1048">Fig. 11K, L</figureCitation>
) vs. robust, hook-shaped and divergent from the claw branch in
<taxonomicName id="17C4F1F90239615CEEEFCA60FDACF8E5" box="[462,615,1869,1891]" class="Heterotardigrada" family="Echiniscidae" genus="Barbaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Echiniscoidea" pageId="11" pageNumber="1048" phylum="Tardigrada" rank="species" species="weglarskae">
<emphasis id="E2B056680239615CEEEFCA60FDACF8E5" box="[462,615,1869,1891]" italics="true" pageId="11" pageNumber="1048">B. weglarskae</emphasis>
</taxonomicName>
(
<figureCitation id="48FF96FF0239615CED55CA63FD3EF8E4" box="[628,757,1869,1891]" captionStart="Figure 11" captionStartId="16.[145,223,1525,1547]" captionTargetBox="[146,760,195,1485]" captionTargetId="figure-25@16.[146,760,195,1485]" captionTargetPageId="16" captionText="Figure 11. Claws and leg structures (PCM): A, claws III of Barbaria bigranulata; B, claws IV of Barbaria bigranulata; C, clawsIIofBarbariacharrua;D,clawsIVofBarbariacharrua;E, clawsIIof Barbariajenningsi; F,claws IVofBarbariajenningsi; G, claws III of Barbaria madonnae; H, claws IV of Barbaria madonnae; I, claws II of Barbaria ollantaytamboensis; J,claws IV of Barbaria ollantaytamboensis; K, claws I of Barbaria paucigranulata; L, claws IV of Barbaria paucigranulata; M, claws II of Barbaria weglarskae; N, claws IV of Barbaria weglarskae; O, claws I of Barbaria danieli; P, claws I of Barbaria hannae; Q, claws III of Barbaria quitensis; R, claws IV of Barbaria ranzii. Arrowheads indicate asymmetrically developed primary spurs. Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.6993197" httpUri="https://zenodo.org/record/6993197/files/figure.png" pageId="11" pageNumber="1048">Fig. 11M, N</figureCitation>
)].
</paragraph>
</subSubSection>
</treatment>
</document>
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