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<document id="E5632417EBA41DC564C4EB634691CA27" ID-DOI="10.11646/zootaxa.2360.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5305019" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629659833440" checkinUser="felipe" docAuthor="Baumgardner, D. E. &amp; Mccafferty, W. P." docDate="2010" docId="EC00F4054727FFBCF9B3FE61FB040B45" docLanguage="en" docName="zt02360p033.pdf" docOrigin="Zootaxa 2360 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2360.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptohyphes musseri Allen 1967" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="10398C7D4734FFA8F924FFACFFE20D67" masterDocTitle="Revision of the genus Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) in North and Central America" masterLastPageNumber="33" masterPageNumber="1" pageNumber="20" updateTime="1699091210753" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="24D70DC383E18EAB3E839462C00CF755">Revision of the genus Leptohyphes Eaton (Ephemeroptera: Leptohyphidae) in North and Central America</mods:title>
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<paragraph id="641645134727FFBBF9B3FE61FDC30C80" blockId="19.[151,545,461,487]" box="[151,545,461,487]" pageId="19" pageNumber="20">
<heading id="3F5EF27F4727FFBBF9B3FE61FDC30C80" bold="true" box="[151,545,461,487]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="A3A93E904727FFBBF9B3FE61FDC30C80" ID-CoL="3TQHH" authority="Allen 1967" authorityName="Allen" authorityYear="1967" box="[151,545,461,487]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014727FFBBF9B3FE61FDC30C80" bold="true" box="[151,545,461,487]" pageId="19" pageNumber="20">
<emphasis id="56DD99014727FFBBF9B3FE61FE740C80" bold="true" box="[151,406,461,487]" italics="true" pageId="19" pageNumber="20">Leptohyphes musseri</emphasis>
<bibRefCitation id="003838E24727FFBBF8B9FE61FDC30C80" author="Allen, R. K." box="[413,545,461,487]" pageId="19" pageNumber="20" pagination="350 - 374" refId="ref24721" refString="Allen, R. K. (1967) New species of new world Leptohyphinae (Ephemeroptera: Tricorythidae). Canadian Entomologist, 99, 350 - 374." type="journal article" year="1967">Allen 1967</bibRefCitation>
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<paragraph id="641645134727FFBBF9B3FDBAFC250F4A" blockId="19.[151,967,534,590]" box="[151,967,534,557]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="44B9623D4727FFBBF9B3FDBAFC250F4A" box="[151,967,534,557]" pageId="19" pageNumber="20">
<treatmentCitation id="E50863024727FFBBF9B3FDBAFDCC0F4A" author="Allen, R. K." box="[151,558,534,557]" page="353" pageId="19" pageNumber="20" year="1967">
<taxonomicName id="A3A93E904727FFBBF9B3FDBAFDCC0F4A" ID-CoL="3TQHH" authority="Allen, 1967: 353" authorityName="Allen" authorityPageNumber="353" authorityYear="1967" box="[151,558,534,557]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014727FFBBF9B3FDBAFE950F4A" box="[151,375,534,557]" italics="true" pageId="19" pageNumber="20">Leptohyphes musseri</emphasis>
<bibRefCitation id="003838E24727FFBBF85AFDBAFDCC0F4A" author="Allen, R. K." box="[382,558,534,557]" pageId="19" pageNumber="20" pagination="350 - 374" refId="ref24721" refString="Allen, R. K. (1967) New species of new world Leptohyphinae (Ephemeroptera: Tricorythidae). Canadian Entomologist, 99, 350 - 374." type="journal article" year="1967">Allen, 1967:353</bibRefCitation>
</taxonomicName>
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;
<bibRefCitation id="003838E24727FFBBFB1DFDBAFD5B0F4A" author="Allen, R. K." box="[569,697,534,557]" pageId="19" pageNumber="20" pagination="537 - 558" refId="ref24751" refString="Allen, R. K. (1978) The nymphs of North and Central American Leptohyphes (Ephemeroptera: Tricorythidae). Annals of the Entomological Society of America, 71, 537 - 558." type="journal article" year="1978">Allen, 1978</bibRefCitation>
;
<bibRefCitation id="003838E24727FFBBFBE1FDBAFC200F4A" author="McCafferty, W. P. &amp; Baumgardner, D. E. &amp; Guenther, J. L." box="[709,962,534,557]" pageId="19" pageNumber="20" pagination="201 - 219" refId="ref25182" refString="McCafferty, W. P., Baumgardner, D. E. &amp; Guenther, J. L. (2004) The Ephemeroptera of Central America. Part 1: Guatemala. Transactions of the American Entomological Society, 130, 201 - 219." type="journal article" year="2004">
McCafferty
<emphasis id="56DD99014727FFBBFA63FDBAFC9F0F4A" box="[839,893,534,557]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2004
</bibRefCitation>
.
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</paragraph>
<paragraph id="641645134727FFBBF9B3FD9AFCB80F29" blockId="19.[151,967,534,590]" box="[151,858,566,590]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="44B9623D4727FFBBF9B3FD9AFCB80F29" box="[151,858,566,590]" pageId="19" pageNumber="20">
<treatmentCitation id="E50863024727FFBBF9B3FD9AFD5B0F2A" author="Allen, R. K. &amp; Brusca, R. C." box="[151,697,566,589]" page="353" pageId="19" pageNumber="20" year="1973">
<taxonomicName id="A3A93E904727FFBBF9B3FD9AFD5B0F2A" authority="Allen and Brusca, 1973:353" authorityName="Allen and Brusca" authorityPageNumber="353" authorityYear="1973" box="[151,697,566,589]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014727FFBBF9B3FD9AFE650F2A" box="[151,391,566,589]" italics="true" pageId="19" pageNumber="20">Leptohyphes brunneus</emphasis>
<bibRefCitation id="003838E24727FFBBF8AAFD9AFD5B0F2A" author="Allen, R. K. &amp; Brusca, R. C." box="[398,697,566,589]" pageId="19" pageNumber="20" pagination="83 - 95" refId="ref24788" refString="Allen, R. K. &amp; Brusca, R. C. (1973) New species of Leptohyphinae from Mexico and Central America (Ephemeroptera: Tricorythidae). Canadian Entomologist, 105, 83 - 95." type="journal article" year="1973">Allen and Brusca, 1973:353</bibRefCitation>
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<emphasis id="56DD99014727FFBBFBE4FD9AFCB80F29" bold="true" box="[704,858,566,590]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="4DEE247A4727FFBBFBE4FD9AFCB80F29" box="[704,858,566,590]" pageId="19" pageNumber="20">new synonym</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="2CB316984727FFBBF9B3FDD1FC5C0E55" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="641645134727FFBBF9B3FDD1FC5C0E55" blockId="19.[151,1436,637,1978]" pageId="19" pageNumber="20">
<emphasis id="56DD99014727FFBBF9B3FDD1FEF20FF0" bold="true" box="[151,272,637,663]" pageId="19" pageNumber="20">Diagnosis</emphasis>
: The following combination of characters will distinguish
<taxonomicName id="A3A93E904727FFBBFAE9FDD1FBAB0FF1" authorityName="Allen" authorityYear="1967" box="[973,1097,637,662]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014727FFBBFAE9FDD1FBAB0FF1" box="[973,1097,637,662]" italics="true" pageId="19" pageNumber="20">L. musseri</emphasis>
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larvae from other species in North and Central America: (1) abdominal terga yellow to pale yellowish-brown, with distinct sublateral, submedian and median black macula; (2) frons often with large median brown (usually black) macula between compound eyes; (3) middle and hind femora usually with distinctive black maculation distally, contrasting with the yellow to yellowish-brown coloration of the leg.
</paragraph>
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<subSubSection id="2CB316984727FFBBF9E1FC92FD6A0B77" pageId="19" pageNumber="20" type="description">
<paragraph id="641645134727FFBBF9E1FC92FD6A0B77" blockId="19.[151,1436,637,1978]" pageId="19" pageNumber="20">
<emphasis id="56DD99014727FFBBF9E1FC92FE800E3F" bold="true" box="[197,354,830,856]" pageId="19" pageNumber="20">Description:</emphasis>
<emphasis id="56DD99014727FFBBF84EFC93FE120E3F" box="[362,496,831,856]" italics="true" pageId="19" pageNumber="20">Male Adult</emphasis>
: Unknown.
<emphasis id="56DD99014727FFBBFBA1FC93FD2E0E3F" box="[645,716,831,856]" italics="true" pageId="19" pageNumber="20">Larva</emphasis>
: Length. Body, 5.07.0 mm; caudal filaments, 6.07.0 mm. General coloration yellow to light brown with reddish-brown and black maculation. Head: yellow to light brown; often with large brown or black macula between lateral ocelli (
<figureCitation id="FC9259964727FFBBFAC1FC20FBDD0EC1" box="[997,1087,908,934]" captionStart="FIGURES 4451" captionStartId="17.[151,270,1730,1754]" captionTargetBox="[182,1400,198,1689]" captionTargetId="figure-134@17.[169,1417,194,1706]" captionTargetPageId="17" captionText="FIGURES 4451. Characteristics of larval structures of Leptohyphes murdochi. 44, foreleg (dorsal) [scale bar C]. 45, hindleg (dorsal) [E]. 46, hind claw (ventrolaeral) [B]. FIGURES 4751. Characteristics of larval structures of Leptohyphes musseri. 47, head (anterior) [A]. 48, pro- and mesothorax (dorsal) [A]. 49, foreleg (dorsal) [C]. 50, hindleg [D]. 51, hindclaw (ventrolateral) [F]. Scale bars (mm): A = 0.05; B, F = 0.10; C = 0.40; D, E = 0.50." figureDoi="http://doi.org/10.5281/zenodo.5305035" httpUri="https://zenodo.org/record/5305035/files/figure.png" pageId="19" pageNumber="20">Fig. 47</figureCitation>
), or think black line between compound eyes (
<figureCitation id="FC9259964727FFBBF845FC1EFE590EAB" box="[353,443,946,972]" captionStart="FIGURES 2631" captionStartId="10.[151,271,1929,1953]" captionTargetBox="[194,1396,439,1896]" captionTargetId="figure-168@10.[173,1414,429,1905]" captionTargetPageId="10" captionText="FIGURES 2631. Characteristics of adults and larvae of Leptohyphes ferruginus. 26, forewing, female adult [A]. 27, head and prothorax (dorsal) [scale bar C]. 28, head (dorsal) [B]. 29, foreleg [E]. 30, hindleg [D]. 31, hind claw (ventrolateral) [F]. Scale bars (mm): A = 1.5; B = 0.025; C, F = 0.05; D, E = 0.5." figureDoi="http://doi.org/10.5281/zenodo.5305029" httpUri="https://zenodo.org/record/5305029/files/figure.png" pageId="19" pageNumber="20">Fig. 28</figureCitation>
); antennae pale to yellowish-brown. Thorax: yellow to light brown; pronotum with extensive black maculation laterally and anterolaterally (
<figureCitation id="FC9259964727FFBBFA42FC75FC270E94" box="[870,965,985,1011]" captionStart="FIGURES 4451" captionStartId="17.[151,270,1730,1754]" captionTargetBox="[182,1400,198,1689]" captionTargetId="figure-134@17.[169,1417,194,1706]" captionTargetPageId="17" captionText="FIGURES 4451. Characteristics of larval structures of Leptohyphes murdochi. 44, foreleg (dorsal) [scale bar C]. 45, hindleg (dorsal) [E]. 46, hind claw (ventrolaeral) [B]. FIGURES 4751. Characteristics of larval structures of Leptohyphes musseri. 47, head (anterior) [A]. 48, pro- and mesothorax (dorsal) [A]. 49, foreleg (dorsal) [C]. 50, hindleg [D]. 51, hindclaw (ventrolateral) [F]. Scale bars (mm): A = 0.05; B, F = 0.10; C = 0.40; D, E = 0.50." figureDoi="http://doi.org/10.5281/zenodo.5305035" httpUri="https://zenodo.org/record/5305035/files/figure.png" pageId="19" pageNumber="20">Fig. 48</figureCitation>
); mesonotum with longitudinal black maculation medially, and scattered black maculation laterally; pro- and mesonota without anterolateral projections or median tubercles. Legs. Proleg (
<figureCitation id="FC9259964727FFBBFBE9FB8AFCCB0927" box="[717,809,1062,1088]" captionStart="FIGURES 4451" captionStartId="17.[151,270,1730,1754]" captionTargetBox="[182,1400,198,1689]" captionTargetId="figure-134@17.[169,1417,194,1706]" captionTargetPageId="17" captionText="FIGURES 4451. Characteristics of larval structures of Leptohyphes murdochi. 44, foreleg (dorsal) [scale bar C]. 45, hindleg (dorsal) [E]. 46, hind claw (ventrolaeral) [B]. FIGURES 4751. Characteristics of larval structures of Leptohyphes musseri. 47, head (anterior) [A]. 48, pro- and mesothorax (dorsal) [A]. 49, foreleg (dorsal) [C]. 50, hindleg [D]. 51, hindclaw (ventrolateral) [F]. Scale bars (mm): A = 0.05; B, F = 0.10; C = 0.40; D, E = 0.50." figureDoi="http://doi.org/10.5281/zenodo.5305035" httpUri="https://zenodo.org/record/5305035/files/figure.png" pageId="19" pageNumber="20">Fig. 49</figureCitation>
): femur with transverse row of elongate setae along dorsal surface; large, black macula present distally; anterior margin with scattered robust setae, mostly along basally margin; posterior margin with robust setae along distal margin; tibia with filiform and elongate setae along anterior margin, and row of robust setae along anterioventral surface; posterior margin with few, scattered filiform setae; tarsus with row of elongate setae along anterior margin, and few, scattered filiform setae along posterior margin. Meso- and metalegs (
<figureCitation id="FC9259964727FFBBFBC9FB44FCA70865" box="[749,837,1256,1282]" captionStart="FIGURES 4451" captionStartId="17.[151,270,1730,1754]" captionTargetBox="[182,1400,198,1689]" captionTargetId="figure-134@17.[169,1417,194,1706]" captionTargetPageId="17" captionText="FIGURES 4451. Characteristics of larval structures of Leptohyphes murdochi. 44, foreleg (dorsal) [scale bar C]. 45, hindleg (dorsal) [E]. 46, hind claw (ventrolaeral) [B]. FIGURES 4751. Characteristics of larval structures of Leptohyphes musseri. 47, head (anterior) [A]. 48, pro- and mesothorax (dorsal) [A]. 49, foreleg (dorsal) [C]. 50, hindleg [D]. 51, hindclaw (ventrolateral) [F]. Scale bars (mm): A = 0.05; B, F = 0.10; C = 0.40; D, E = 0.50." figureDoi="http://doi.org/10.5281/zenodo.5305035" httpUri="https://zenodo.org/record/5305035/files/figure.png" pageId="19" pageNumber="20">Fig. 50</figureCitation>
): femora with few robust setae on dorsal surface or none; large, black macula present distally; anterior margin with stout setae along most of margin; posterior margin with fairly regularly spaced elongate setae along distal half of margin; tibiae with few elongate setae along anterior margin; posterior margin with elongate setae along most of margin; tarsus with elongate setae along anterior margins; posterior margin with few, scattered filiform setae, often in pairs. Claws of all legs with three to seven marginal, and a single submarginal denticle (
<figureCitation id="FC9259964727FFBBFA88FA05FBE408A4" box="[940,1030,1449,1475]" captionStart="FIGURES 4451" captionStartId="17.[151,270,1730,1754]" captionTargetBox="[182,1400,198,1689]" captionTargetId="figure-134@17.[169,1417,194,1706]" captionTargetPageId="17" captionText="FIGURES 4451. Characteristics of larval structures of Leptohyphes murdochi. 44, foreleg (dorsal) [scale bar C]. 45, hindleg (dorsal) [E]. 46, hind claw (ventrolaeral) [B]. FIGURES 4751. Characteristics of larval structures of Leptohyphes musseri. 47, head (anterior) [A]. 48, pro- and mesothorax (dorsal) [A]. 49, foreleg (dorsal) [C]. 50, hindleg [D]. 51, hindclaw (ventrolateral) [F]. Scale bars (mm): A = 0.05; B, F = 0.10; C = 0.40; D, E = 0.50." figureDoi="http://doi.org/10.5281/zenodo.5305035" httpUri="https://zenodo.org/record/5305035/files/figure.png" pageId="19" pageNumber="20">Fig. 51</figureCitation>
). Abdomen: terga yellow to light brown, often with black maculation along posterior half of tergite, and on sublateral margins; operculate gill pale to light brown; gill formula 2/5/5/5/1.
</paragraph>
</subSubSection>
<subSubSection id="2CB316984727FFBBF9E1F9B1FF360BE3" pageId="19" pageNumber="20" type="distribution">
<paragraph id="641645134727FFBBF9E1F9B1FF360BE3" blockId="19.[151,1436,637,1978]" pageId="19" pageNumber="20">
<emphasis id="56DD99014727FFBBF9E1F9B1FEBC0B50" bold="true" box="[197,350,1565,1591]" pageId="19" pageNumber="20">Distribution</emphasis>
: This species was previously known from
<collectingCountry id="1CBE05834727FFBBFA75F9B1FC320B50" box="[849,976,1565,1591]" name="Guatemala" pageId="19" pageNumber="20">Guatemala</collectingCountry>
(
<bibRefCitation id="003838E24727FFBBFAFBF9B1FB8C0B50" author="Allen, R. K." box="[991,1134,1565,1591]" pageId="19" pageNumber="20" pagination="537 - 558" refId="ref24751" refString="Allen, R. K. (1978) The nymphs of North and Central American Leptohyphes (Ephemeroptera: Tricorythidae). Annals of the Entomological Society of America, 71, 537 - 558." type="journal article" year="1978">Allen, 1978</bibRefCitation>
;
<bibRefCitation id="003838E24727FFBBFD5DF9B1FA6D0B50" author="McCafferty, W. P. &amp; Baumgardner, D. E. &amp; Guenther, J. L." box="[1145,1423,1565,1591]" pageId="19" pageNumber="20" pagination="201 - 219" refId="ref25182" refString="McCafferty, W. P., Baumgardner, D. E. &amp; Guenther, J. L. (2004) The Ephemeroptera of Central America. Part 1: Guatemala. Transactions of the American Entomological Society, 130, 201 - 219." type="journal article" year="2004">
McCafferty
<emphasis id="56DD99014727FFBBFC2CF9B1FAA00B51" box="[1288,1346,1565,1590]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
, 2004
</bibRefCitation>
),
<collectingCountry id="1CBE05834727FFBBF9B3F9E8FEEB0B39" box="[151,265,1604,1630]" name="Honduras" pageId="19" pageNumber="20">Honduras</collectingCountry>
(
<bibRefCitation id="003838E24727FFBBF83CF9E8FE440B39" author="Allen, R. K." box="[280,422,1604,1630]" pageId="19" pageNumber="20" pagination="537 - 558" refId="ref24751" refString="Allen, R. K. (1978) The nymphs of North and Central American Leptohyphes (Ephemeroptera: Tricorythidae). Annals of the Entomological Society of America, 71, 537 - 558." type="journal article" year="1978">Allen, 1978</bibRefCitation>
), and
<collectingCountry id="1CBE05834727FFBBF8C8F9E8FDA70B39" box="[492,581,1604,1630]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
(
<bibRefCitation id="003838E24727FFBBFB70F9E8FC890B39" author="Allen, R. K. &amp; Brusca, R. C." box="[596,875,1604,1630]" pageId="19" pageNumber="20" pagination="83 - 95" refId="ref24788" refString="Allen, R. K. &amp; Brusca, R. C. (1973) New species of Leptohyphinae from Mexico and Central America (Ephemeroptera: Tricorythidae). Canadian Entomologist, 105, 83 - 95." type="journal article" year="1973">Allen and Brusca, 1973</bibRefCitation>
;
<bibRefCitation id="003838E24727FFBBFA53F9E8FBE60B39" author="Allen, R. K." box="[887,1028,1604,1630]" pageId="19" pageNumber="20" pagination="537 - 558" refId="ref24751" refString="Allen, R. K. (1978) The nymphs of North and Central American Leptohyphes (Ephemeroptera: Tricorythidae). Annals of the Entomological Society of America, 71, 537 - 558." type="journal article" year="1978">Allen, 1978</bibRefCitation>
), and is newly reported from
<collectingCountry id="1CBE05834727FFBBFC7DF9E8FF320BE3" name="Costa Rica" pageId="19" pageNumber="20">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2CB316984727FFBCF9E1F93CFBF60FCF" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="641645134727FFBBF9E1F93CFCDE0ADD" blockId="19.[151,1436,637,1978]" pageId="19" pageNumber="20">
<emphasis id="56DD99014727FFBBF9E1F93CFEB40BCD" bold="true" box="[197,342,1680,1706]" pageId="19" pageNumber="20">Discussion:</emphasis>
<taxonomicName id="A3A93E904727FFBBF844F93DFD930BCD" authorityName="Allen and Brusca" authorityYear="1973" box="[352,625,1681,1706]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014727FFBBF844F93DFD930BCD" box="[352,625,1681,1706]" italics="true" pageId="19" pageNumber="20">Leptohyphes brunneus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="003838E24727FFBBFA70F93DFB630BCC" author="Allen, R. K. &amp; Brusca, R. C." box="[852,1153,1681,1707]" pageId="19" pageNumber="20" pagination="83 - 95" refId="ref24788" refString="Allen, R. K. &amp; Brusca, R. C. (1973) New species of Leptohyphinae from Mexico and Central America (Ephemeroptera: Tricorythidae). Canadian Entomologist, 105, 83 - 95." type="journal article" year="1973">Allen and Brusca (1973)</bibRefCitation>
on the basis of a small series of larvae from several locations in southern
<collectingCountry id="1CBE05834727FFBBFBC9F914FCA90BB5" box="[749,843,1720,1746]" name="Mexico" pageId="19" pageNumber="20">Mexico</collectingCountry>
. The presence of a “large brown to reddish brown macula on the frons between the lateral ocelli” was considered distinctive for this species. It was also described as having abdominal terga that were light brown in color with reddish brown markings, and numerous scattered spines. All specimens of the
<typeStatus id="BB12FBB14727FFBBFBC7F880FCF40A21" box="[739,790,1836,1862]" pageId="19" pageNumber="20">type</typeStatus>
series were examined and found to be badly faded to pale yellow, with none of the specimens having a brown maculation between the lateral ocelli. Other non-type material identified by Allen as
<taxonomicName id="A3A93E904727FFBBFB24F8D5FD6E0AF5" authorityName="Allen and Brusca" authorityYear="1973" box="[512,652,1913,1938]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014727FFBBFB24F8D5FD6E0AF5" box="[512,652,1913,1938]" italics="true" pageId="19" pageNumber="20">L. brunneus</emphasis>
</taxonomicName>
, which is in excellent condition and unfaded, also does not possess the large median brown macula between the lateral ocelli.
</paragraph>
<paragraph id="641645134720FFBCF9E2FF34FBF60FCF" blockId="20.[151,1436,152,1570]" pageId="20" pageNumber="21">
<taxonomicName id="A3A93E904720FFBCF9E2FF34FE5F0DD6" authorityName="Allen" authorityYear="1967" box="[198,445,152,177]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCF9E2FF34FE5F0DD6" box="[198,445,152,177]" italics="true" pageId="20" pageNumber="21">Leptohyphes musseri</emphasis>
</taxonomicName>
was described from
<collectingCountry id="1CBE05834720FFBCFB90FF34FCD10DD5" box="[692,819,152,178]" name="Guatemala" pageId="20" pageNumber="21">Guatemala</collectingCountry>
based upon
<specimenCount id="72AF8E9A4720FFBCFAECFF34FBB00DD5" box="[968,1106,152,178]" pageId="20" pageNumber="21" type="larva">three larvae</specimenCount>
(
<bibRefCitation id="003838E24720FFBCFD47FF34FB110DD5" author="Allen, R. K." box="[1123,1267,152,178]" pageId="20" pageNumber="21" pagination="350 - 374" refId="ref24721" refString="Allen, R. K. (1967) New species of new world Leptohyphinae (Ephemeroptera: Tricorythidae). Canadian Entomologist, 99, 350 - 374." type="journal article" year="1967">Allen, 1967</bibRefCitation>
). In his key to the
<taxonomicName id="A3A93E904720FFBCF9E2FF13FE860DBF" authorityName="Eaton" authorityYear="1882" box="[198,356,191,216]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="56DD99014720FFBCF9E2FF13FE860DBF" box="[198,356,191,216]" italics="true" pageId="20" pageNumber="21">Leptohyphes</emphasis>
</taxonomicName>
larvae of North and Central America,
<bibRefCitation id="003838E24720FFBCFA77FF12FC170DBF" author="Allen, R. K." box="[851,1013,190,216]" pageId="20" pageNumber="21" pagination="537 - 558" refId="ref24751" refString="Allen, R. K. (1978) The nymphs of North and Central American Leptohyphes (Ephemeroptera: Tricorythidae). Annals of the Entomological Society of America, 71, 537 - 558." type="journal article" year="1978">Allen (1978)</bibRefCitation>
distinguished
<taxonomicName id="A3A93E904720FFBCFD94FF13FAD10DBF" authorityName="Allen" authorityYear="1967" box="[1200,1331,191,216]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCFD94FF13FAD10DBF" box="[1200,1331,191,216]" italics="true" pageId="20" pageNumber="21">L. musseri</emphasis>
</taxonomicName>
from
<taxonomicName id="A3A93E904720FFBCFCA7FF13FEE40D99" authorityName="Allen and Brusca" authorityYear="1973" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCFCA7FF13FEE40D99" italics="true" pageId="20" pageNumber="21">L. brunneus</emphasis>
</taxonomicName>
based upon differences in abdominal terga maculations and coloration as well as markings on the legs and head. Specifically,
<taxonomicName id="A3A93E904720FFBCF8D3FEA0FD980C42" authorityName="Allen" authorityYear="1967" box="[503,634,268,293]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCF8D3FEA0FD980C42" box="[503,634,268,293]" italics="true" pageId="20" pageNumber="21">L. musseri</emphasis>
</taxonomicName>
possessed sublateral, submedian, and median black maculae on abdominal terga 6 -9, while
<taxonomicName id="A3A93E904720FFBCF8CDFE9EFD950C2C" authorityName="Allen and Brusca" authorityYear="1973" box="[489,631,306,331]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCF8CDFE9EFD950C2C" box="[489,631,306,331]" italics="true" pageId="20" pageNumber="21">L. brunneus</emphasis>
</taxonomicName>
lacked this maculation and had brown terga instead of yellow. The head of
<taxonomicName id="A3A93E904720FFBCF9D3FEF5FE900C15" authorityName="Allen" authorityYear="1967" box="[247,370,345,370]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCF9D3FEF5FE900C15" box="[247,370,345,370]" italics="true" pageId="20" pageNumber="21">L. musseri</emphasis>
</taxonomicName>
was reported to have a thin black line between the lateral ocelli, and black maculation on the meso- and meta femora, while
<taxonomicName id="A3A93E904720FFBCFB09FE2CFD580CFE" authorityName="Allen and Brusca" authorityYear="1973" box="[557,698,384,409]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCFB09FE2CFD580CFE" box="[557,698,384,409]" italics="true" pageId="20" pageNumber="21">L. brunneus</emphasis>
</taxonomicName>
lacked the thin black line and black maculation on the femora. Recent collections of larvae from
<collectingCountry id="1CBE05834720FFBCFB0EFE0AFD4B0CA7" box="[554,681,422,448]" name="Guatemala" pageId="20" pageNumber="21">Guatemala</collectingCountry>
show numerous individuals (from the same location) with both the black maculation leg characters of
<taxonomicName id="A3A93E904720FFBCFB4EFE61FD0A0C81" authorityName="Allen" authorityYear="1967" box="[618,744,461,486]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCFB4EFE61FD0A0C81" box="[618,744,461,486]" italics="true" pageId="20" pageNumber="21">L. musseri</emphasis>
</taxonomicName>
and the large brown macula between the compound eyes characteristic of
<taxonomicName id="A3A93E904720FFBCF852FE58FDF00F6A" authorityName="Allen and Brusca" authorityYear="1973" box="[374,530,500,525]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCF852FE58FDF00F6A" box="[374,530,500,525]" italics="true" pageId="20" pageNumber="21">L. brunneus</emphasis>
</taxonomicName>
. It appears both these characters, in addition to the abdominal terga characteristics described previously, are highly variable, and often dependent on gender and maturity of individual larvae. In addition, some individual larvae possessed a variety of combinations of these characteristics. Since no other morphological features can be found that distinguish
<taxonomicName id="A3A93E904720FFBCFD80FDC4FAD50FE6" authorityName="Allen and Brusca" authorityYear="1973" box="[1188,1335,616,641]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCFD80FDC4FAD50FE6" box="[1188,1335,616,641]" italics="true" pageId="20" pageNumber="21">L. brunneus</emphasis>
</taxonomicName>
from
<taxonomicName id="A3A93E904720FFBCFCA0FDC4FF100FCF" authorityName="Allen" authorityYear="1967" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCFCA0FDC4FF100FCF" italics="true" pageId="20" pageNumber="21">L. musseri</emphasis>
</taxonomicName>
,
<taxonomicName id="A3A93E904720FFBCF825FD23FE6F0FCF" authorityName="Allen and Brusca" authorityYear="1973" box="[257,397,655,680]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCF825FD23FE6F0FCF" box="[257,397,655,680]" italics="true" pageId="20" pageNumber="21">L. brunneus</emphasis>
</taxonomicName>
is placed as a junior synonym of
<taxonomicName id="A3A93E904720FFBCFA32FD23FBEF0FCF" authorityName="Allen" authorityYear="1967" box="[790,1037,655,680]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCFA32FD23FBEF0FCF" box="[790,1037,655,680]" italics="true" pageId="20" pageNumber="21">Leptohyphes musseri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2CB316984720FFBCF9E1FD19FB040B45" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="641645134720FFBCF9E1FD19FEF10EB9" blockId="20.[151,1436,152,1570]" pageId="20" pageNumber="21">
<emphasis id="56DD99014720FFBCF9E1FD19FDE90FA8" bold="true" box="[197,523,693,719]" pageId="20" pageNumber="21">Type material examined:</emphasis>
<taxonomicName id="A3A93E904720FFBCFB31FD19FC850FA8" authority="Allen" authorityName="Allen" authorityYear="1967" box="[533,871,693,719]" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="musseri">
<emphasis id="56DD99014720FFBCFB31FD19FCF50FA9" box="[533,791,693,718]" italics="true" pageId="20" pageNumber="21">Leptohyphes musseri</emphasis>
<materialsCitation id="D4C14F4E4720FFBCFA04FD19FC850FA8" box="[800,871,693,719]" collectingDate="1962-08-21" collectionCode="FSCA, FAMU, T, CAS, RKA, R" collectorName="G. G. Musser &amp; Soloa &amp; Panajachel &amp; L, G. G. Musser &amp; N. Ayoquezco &amp; Elev. &amp; Same &amp; Tizapan El Alto" country="Guatemala" elevation="1500" location="Tizapan El Alto" municipality="Rio La Pasion" pageId="20" pageNumber="21" specimenCode="E2020.1, CAS #11971" specimenCount="8" specimenCount-larva="8" stateProvince="Oaxaca" typeStatus="holotype">Allen</materialsCitation>
</taxonomicName>
,
<typeStatus id="BB12FBB14720FFBCFA51FD19FBF60FA8" box="[885,1044,693,719]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
(larva):
<collectingCountry id="1CBE05834720FFBCFDA5FD19FAA30FA8" box="[1153,1345,693,719]" name="Guatemala" pageId="20" pageNumber="21">GUATEMALA</collectingCountry>
: Soloa; Panajachel,
<quantity id="A351E8F64720FFBCF803FD70FE9F0F91" box="[295,381,732,758]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">
<elevation id="EF84A2204720FFBCF803FD70FE9F0F91" box="[295,381,732,758]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="101763D34720FFBCF8AEFD70FDFE0F91" box="[394,540,732,758]" pageId="20" pageNumber="21" value="1962-08-21">
<collectingDate id="00539A3B4720FFBCF8AEFD70FDFE0F91" box="[394,540,732,758]" pageId="20" pageNumber="21" value="1962-08-21">21.viii.1962</collectingDate>
</date>
,
<collectorName id="C95C20C54720FFBCFB0DFD70FD590F91" box="[553,699,732,758]" pageId="20" pageNumber="21">G.G. Musser</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFBEEFD70FCF50F91" box="[714,791,732,758]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="20" pageNumber="21" type="Museum">FSCA</collectionCode>
(
<collectionCode id="02B8DDD64720FFBCFA0CFD70FC9D0F91" box="[808,895,732,758]" pageId="20" pageNumber="21">FAMU</collectionCode>
)
<specimenCode id="340FED684720FFBCFAABFD70FC100F91" box="[911,1010,732,758]" collectionCode="FAMU" pageId="20" pageNumber="21">E2020.1</specimenCode>
<collectionCode id="02B8DDD64720FFBCFADFFD70FBF00F91" box="[1019,1042,732,758]" name="Tavera, Department of Geology and Geophysics" pageId="20" pageNumber="21">T</collectionCode>
].
<typeStatus id="BB12FBB14720FFBCFD03FD70FB2D0F91" box="[1063,1231,732,758]" pageId="20" pageNumber="21" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="1CBE05834720FFBCFDFFFD70FA7A0F91" box="[1243,1432,732,758]" name="Guatemala" pageId="20" pageNumber="21">GUATEMALA</collectingCountry>
:
<collectorName id="C95C20C54720FFBCF9B3FCAEFF030E7B" box="[151,225,770,796]" pageId="20" pageNumber="21">Soloa</collectorName>
;
<collectorName id="C95C20C54720FFBCF9CBFCAEFE9A0E7B" box="[239,376,770,796]" pageId="20" pageNumber="21">Panajachel</collectorName>
,
<quantity id="A351E8F64720FFBCF8A2FCAEFE3C0E7B" box="[390,478,770,796]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">
<elevation id="EF84A2204720FFBCF8A2FCAEFE3C0E7B" box="[390,478,770,796]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date id="101763D34720FFBCF8C8FCAEFD670E7B" box="[492,645,770,796]" pageId="20" pageNumber="21" value="1962-08-21">
<collectingDate id="00539A3B4720FFBCF8C8FCAEFD670E7B" box="[492,645,770,796]" pageId="20" pageNumber="21" value="1962-08-21">21.viii.1962</collectingDate>
</date>
, 16
<collectorName id="C95C20C54720FFBCFB97FCAEFC8C0E7B" box="[691,878,770,796]" pageId="20" pageNumber="21">L, G.G. Musser</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFA5AFCAEFC3C0E7B" box="[894,990,770,796]" pageId="20" pageNumber="21">FAMU</collectionCode>
]; same but, 2L [
<collectionCode id="02B8DDD64720FFBCFD83FCAEFB090E7B" box="[1191,1259,770,796]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
].
<taxonomicName id="A3A93E904720FFBCFC24FCAEFEE60E25" authorityName="Allen and Brusca" authorityYear="1973" class="Insecta" family="Leptohyphidae" genus="Leptohyphes" kingdom="Animalia" order="Ephemeroptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="56DD99014720FFBCFC24FCAEFEE60E25" italics="true" pageId="20" pageNumber="21">Leptohyphes brunneus</emphasis>
</taxonomicName>
:
<typeStatus id="BB12FBB14720FFBCF830FC85FE4D0E24" box="[276,431,809,835]" pageId="20" pageNumber="21" type="holotype">HOLOTYPE</typeStatus>
(larva):
<collectingCountry id="1CBE05834720FFBCFB30FC85FD6E0E24" box="[532,652,809,835]" name="Mexico" pageId="20" pageNumber="21">MEXICO</collectingCountry>
:
<collectingRegion id="A66D8BF14720FFBCFBBCFC85FD110E24" box="[664,755,809,835]" country="Mexico" name="Oaxaca" pageId="20" pageNumber="21">Oaxaca</collectingRegion>
; stream
<quantity id="A351E8F64720FFBCFA71FC85FC420E24" box="[853,928,809,835]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="20" pageNumber="21" unit="mi" value="15.0">15 mi.</quantity>
<collectorName id="C95C20C54720FFBCFA8CFC85FBB00E24" box="[936,1106,809,835]" pageId="20" pageNumber="21">N. Ayoquezco</collectorName>
,
<collectorName id="C95C20C54720FFBCFD7AFC85FB7B0E24" box="[1118,1177,809,835]" pageId="20" pageNumber="21">Elev.</collectorName>
6,700', 20.x.68,
<collectionCode id="02B8DDD64720FFBCFC7AFC85FA7E0E24" box="[1374,1436,809,835]" pageId="20" pageNumber="21">RKA</collectionCode>
[
<specimenCode id="340FED684720FFBCF9BAFCFCFEA30E0D" box="[158,321,848,874]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS #11971</specimenCode>
].
<typeStatus id="BB12FBB14720FFBCF870FCFCFE1E0E0D" box="[340,508,848,874]" pageId="20" pageNumber="21" type="paratype">PARATYPES</typeStatus>
:
<collectorName id="C95C20C54720FFBCFB2CFCFCFDA80E0D" box="[520,586,848,874]" pageId="20" pageNumber="21">Same</collectorName>
data as holotype,
<specimenCount id="72AF8E9A4720FFBCFA00FCFCFC610E0D" box="[804,899,848,874]" pageId="20" pageNumber="21" type="larva">8 larvae</specimenCount>
[
<collectionCode id="02B8DDD64720FFBCFAB7FCFCFC370E0D" box="[915,981,848,874]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
];
<collectingRegion id="A66D8BF14720FFBCFACEFCFCFBDC0E0D" box="[1002,1086,848,874]" country="Mexico" name="Jalisco" pageId="20" pageNumber="21">Jalisco</collectingRegion>
;
<collectingMunicipality id="8472DF694720FFBCFD6FFCFCFB160E0D" box="[1099,1268,848,874]" pageId="20" pageNumber="21">Rio La Pasion</collectingMunicipality>
at
<collectorName id="C95C20C54720FFBCFC3EFCFCFF320EF7" pageId="20" pageNumber="21">
<location id="617613C84720FFBCFC3EFCFCFF320EF7" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFC3EFCFCFF320EF7" country="Guatemala" municipality="Rio La Pasion" name="Tizapan El Alto" pageId="20" pageNumber="21" stateProvince="Oaxaca">Tizapan El Alto</location>
</collectorName>
, 16.x.68,
<collectionCode id="02B8DDD64720FFBCF86DFCDAFE820EF7" box="[329,352,886,912]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="21">R</collectionCode>
.
<materialsCitation id="D4C14F4E4720FFBCF841FCDAFEC50ED0" collectionCode="CAS, RKA" collectorName="K. Allen" country="Guatemala" location="Arriaga on Hwy." pageId="20" pageNumber="21" specimenCount="3" specimenCount-larva="3" stateProvince="Chiapas" typeStatus="holotype">
<collectorName id="C95C20C54720FFBCF841FCDAFE330EF7" box="[357,465,886,912]" pageId="20" pageNumber="21">K. Allen</collectorName>
,
<specimenCount id="72AF8E9A4720FFBCF8FBFCDAFDA30EF7" box="[479,577,886,912]" pageId="20" pageNumber="21" type="larva">3 larvae</specimenCount>
[
<collectionCode id="02B8DDD64720FFBCFB75FCDAFD760EF7" box="[593,660,886,912]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
];
<collectingRegion id="A66D8BF14720FFBCFB8EFCDAFCED0EF7" box="[682,783,886,912]" country="Mexico" name="Chiapas" pageId="20" pageNumber="21">Chiapas</collectingRegion>
; stream
<quantity id="A351E8F64720FFBCFA52FCDAFC570EF7" box="[886,949,886,912]" metricMagnitude="4" metricUnit="m" metricValue="1.1265408" pageId="20" pageNumber="21" unit="mi" value="7.0">7 mi.</quantity>
N.
<location id="617613C84720FFBCFAC0FCDAFB530EF7" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFAC0FCDAFB530EF7" box="[996,1201,886,912]" country="Guatemala" name="Arriaga on Hwy." pageId="20" pageNumber="21" stateProvince="Chiapas">Arriaga on Hwy.</location>
190, 23.x.68, elev. 1400',
<collectionCode id="02B8DDD64720FFBCF9C2FC31FEC50ED0" box="[230,295,925,951]" pageId="20" pageNumber="21">RKA</collectionCode>
</materialsCitation>
,
<materialsCitation id="D4C14F4E4720FFBCF817FC31FAC90ED0" box="[307,1323,925,951]" collectionCode="CAS, R" collectorName="Huajintlan &amp; Elev" country="Guatemala" location="Amacuzuc" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Morelos" typeStatus="holotype">
1L [
<collectionCode id="02B8DDD64720FFBCF840FC31FE470ED0" box="[356,421,925,951]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
];
<collectingRegion id="A66D8BF14720FFBCF89DFC31FDFC0ED0" box="[441,542,925,951]" country="Mexico" name="Morelos" pageId="20" pageNumber="21">Morelos</collectingRegion>
;
<location id="617613C84720FFBCFB0FFC31FD4A0ED0" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFB0FFC31FD4A0ED0" box="[555,680,925,951]" country="Guatemala" name="Amacuzuc" pageId="20" pageNumber="21" stateProvince="Morelos">Amacuzuc</location>
at
<collectorName id="C95C20C54720FFBCFBE9FC31FCAA0ED0" box="[717,840,925,951]" pageId="20" pageNumber="21">Huajintlan</collectorName>
on hwy. 95,
<collectorName id="C95C20C54720FFBCFAC7FC31FBFB0ED0" box="[995,1049,925,951]" pageId="20" pageNumber="21">Elev</collectorName>
3,200', 29/30.vii.66,
<collectionCode id="02B8DDD64720FFBCFC30FC31FAC90ED0" box="[1300,1323,925,951]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="21">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFC0BFC31FEEC0EB9" collectionCode="CAS" collectorName="Allen" country="Guatemala" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Morelos" typeStatus="holotype">
K.
<collectorName id="C95C20C54720FFBCFC70FC31FA7A0ED0" box="[1364,1432,925,951]" pageId="20" pageNumber="21">Allen</collectorName>
, 6L [
<collectionCode id="02B8DDD64720FFBCF9E2FC68FEE50EB9" box="[198,263,964,990]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="641645134720FFBCF9E2FC46FB040B45" blockId="20.[151,1436,152,1570]" pageId="20" pageNumber="21">
<emphasis id="56DD99014720FFBCF9E2FC46FDF10963" bold="true" box="[198,531,1002,1028]" pageId="20" pageNumber="21">Other material examined:</emphasis>
<materialsCitation id="D4C14F4E4720FFBCFB3AFC46FC27094C" collectingDate="2000-06-17" collectionCode="WDS, TAMU" country="Costa Rica" county="Km" latitude="8.602778" location="Rio Coloradito" longLatPrecision="21" longitude="-82.90195" municipality="Coloradito Norte" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Puntarenas">
<emphasis id="56DD99014720FFBCFB3AFC46FD350963" bold="true" box="[542,727,1002,1028]" pageId="20" pageNumber="21">
<collectingCountry id="1CBE05834720FFBCFB3AFC46FD350963" box="[542,727,1002,1028]" name="Costa Rica" pageId="20" pageNumber="21">COSTA RICA</collectingCountry>
</emphasis>
:
<collectingRegion id="A66D8BF14720FFBCFBCFFC46FC8D0963" box="[747,879,1002,1028]" country="Costa Rica" name="Puntarenas" pageId="20" pageNumber="21">Puntarenas</collectingRegion>
; 1
<collectingCounty id="8D773D9F4720FFBCFABCFC46FC260963" box="[920,964,1002,1028]" pageId="20" pageNumber="21">Km</collectingCounty>
S
<collectingMunicipality id="8472DF694720FFBCFACFFC46FB580963" box="[1003,1210,1002,1028]" pageId="20" pageNumber="21">Coloradito Norte</collectingMunicipality>
,
<location id="617613C84720FFBCFDEDFC46FA990963" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFDEDFC46FA990963" box="[1225,1403,1002,1028]" country="Costa Rica" county="Km" latitude="8.602778" longLatPrecision="21" longitude="-82.90195" municipality="Coloradito Norte" name="Rio Coloradito" pageId="20" pageNumber="21" stateProvince="Puntarenas">Rio Coloradito</location>
at
<location id="617613C84720FFBCF9B3FBBDFF34094C" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF9B3FBBDFF34094C" box="[151,214,1041,1067]" country="Costa Rica" county="Km" latitude="8.602778" longLatPrecision="21" longitude="-82.90195" municipality="Coloradito Norte" name="Hwy." pageId="20" pageNumber="21" stateProvince="Puntarenas">Hwy.</location>
2, (
<geoCoordinate id="019D23D44720FFBCF821FBBDFE6D094C" box="[261,399,1041,1067]" degrees="08" direction="north" minutes="36" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="10" value="8.602778">N08°3610</geoCoordinate>
:;
<geoCoordinate id="019D23D44720FFBCF885FBBDFDD9094C" box="[417,571,1041,1067]" degrees="82" direction="west" minutes="54" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="07" value="-82.90195">W82°5407”</geoCoordinate>
),
<date id="101763D34720FFBCFB75FBBDFD30094C" box="[593,722,1041,1067]" pageId="20" pageNumber="21" value="2000-06-17">
<collectingDate id="00539A3B4720FFBCFB75FBBDFD30094C" box="[593,722,1041,1067]" pageId="20" pageNumber="21" value="2000-06-17">17.vi.2000</collectingDate>
</date>
, 4L,
<collectionCode id="02B8DDD64720FFBCFA2BFBBDFCB2094C" box="[783,848,1041,1067]" pageId="20" pageNumber="21">WDS</collectionCode>
[
<collectionCode id="02B8DDD64720FFBCFA44FBBDFC5C094C" box="[864,958,1041,1067]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFAF4FBBCFEE009F8" collectingDate="2000-06-19" collectingDateMax="2001-06-22" collectingDateMin="2000-06-19" collectionCode="DEB, TAMU, WDS" collectorName="NE Dominical" country="Costa Rica" county="Rio Pedregoso" elevation="610" latitude="9.28" location="Km" longLatPrecision="21" longitude="-83.82278" municipality="Hwy." pageId="20" pageNumber="21" specimenCount="1" stateProvince="San Jose">
<emphasis id="56DD99014720FFBCFAF4FBBCFBDC094D" bold="true" box="[976,1086,1040,1066]" pageId="20" pageNumber="21">
<collectingRegion id="A66D8BF14720FFBCFAF4FBBCFBDC094D" box="[976,1086,1040,1066]" country="Costa Rica" name="San Jose" pageId="20" pageNumber="21">San Jose</collectingRegion>
</emphasis>
;
<collectingCounty id="8D773D9F4720FFBCFD6AFBBDFB19094C" box="[1102,1275,1041,1067]" pageId="20" pageNumber="21">Río Pedregoso</collectingCounty>
at
<collectingMunicipality id="8472DF694720FFBCFC05FBBDFA83094C" box="[1313,1377,1041,1067]" pageId="20" pageNumber="21">Hwy.</collectingMunicipality>
243, ca. 4
<location id="617613C84720FFBCF9F8FB94FEEB0935" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF9F8FB94FEEB0935" box="[220,265,1080,1106]" country="Costa Rica" county="Rio Pedregoso" latitude="9.28" longLatPrecision="21" longitude="-83.82278" municipality="Hwy." name="Km" pageId="20" pageNumber="21" stateProvince="San Jose">Km</location>
S.
<location id="617613C84720FFBCF811FB94FD8E0935" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF811FB94FD8E0935" box="[309,620,1080,1106]" country="Costa Rica" county="Rio Pedregoso" latitude="9.28" longLatPrecision="21" longitude="-83.82278" municipality="Hwy." name="San Isidro de El General" pageId="20" pageNumber="21" stateProvince="San Jose">San Isidro de El General</location>
(
<geoCoordinate id="019D23D44720FFBCFBA4FB94FCF90935" box="[640,795,1080,1106]" degrees="09" direction="north" minutes="21" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="15" value="9.354167">N09°2115”</geoCoordinate>
;
<geoCoordinate id="019D23D44720FFBCFA0FFB94FC2E0935" box="[811,972,1080,1106]" degrees="83" direction="west" minutes="43" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="35" value="-83.72639">W83°4335”</geoCoordinate>
, elev.
<quantity id="A351E8F64720FFBCFD3AFB94FB9E0935" box="[1054,1148,1080,1106]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="20" pageNumber="21" unit="ft" value="2000.0">
<elevation id="EF84A2204720FFBCFD3AFB94FB9E0935" box="[1054,1148,1080,1106]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="20" pageNumber="21" unit="ft" value="2000.0">2000 ft</elevation>
</quantity>
),
<date id="101763D34720FFBCFDB1FB94FAFF0935" box="[1173,1309,1080,1106]" pageId="20" pageNumber="21" value="2001-06-22">
<collectingDate id="00539A3B4720FFBCFDB1FB94FAFF0935" box="[1173,1309,1080,1106]" pageId="20" pageNumber="21" value="2001-06-22">22.vi.2001</collectingDate>
</date>
, 6L,
<collectionCode id="02B8DDD64720FFBCFC44FB94FA7E0935" box="[1376,1436,1080,1106]" pageId="20" pageNumber="21">DEB</collectionCode>
[
<collectionCode id="02B8DDD64720FFBCF9BAFBF2FF1E091F" box="[158,252,1118,1144]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
];
<collectorName id="C95C20C54720FFBCF82BFBF2FE5D091F" box="[271,447,1118,1144]" pageId="20" pageNumber="21">NE Dominical</collectorName>
, unnamed stream (
<geoCoordinate id="019D23D44720FFBCFBBBFBF2FCD6091F" box="[671,820,1118,1144]" degrees="09" direction="north" minutes="16" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="48" value="9.28">N09°1648”</geoCoordinate>
;
<geoCoordinate id="019D23D44720FFBCFA66FBF2FC3E091F" box="[834,988,1118,1144]" degrees="83" direction="west" minutes="49" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="22" value="-83.82278">W83°4922”</geoCoordinate>
, elev.
<quantity id="A351E8F64720FFBCFD0CFBF2FB94091F" box="[1064,1142,1118,1144]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="m" value="200.0">
<elevation id="EF84A2204720FFBCFD0CFBF2FB94091F" box="[1064,1142,1118,1144]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="21" unit="m" value="200.0">200 m</elevation>
</quantity>
),
<date id="101763D34720FFBCFDA9FBF2FAEC091F" box="[1165,1294,1118,1144]" pageId="20" pageNumber="21" value="2000-06-19">
<collectingDate id="00539A3B4720FFBCFDA9FBF2FAEC091F" box="[1165,1294,1118,1144]" pageId="20" pageNumber="21" value="2000-06-19">19.vi.2000</collectingDate>
</date>
, 15L,
<collectionCode id="02B8DDD64720FFBCFC7EFBF2FA79091F" box="[1370,1435,1118,1144]" pageId="20" pageNumber="21">WDS</collectionCode>
[
<collectionCode id="02B8DDD64720FFBCF9BAFB29FF1E09F8" box="[158,252,1157,1183]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCF82AFB29FA7A09F8" box="[270,1432,1157,1183]" collectingDate="1991-02-20" collectionCode="TAMU, KM" collectorName="Esther Gonzalez" country="Guatemala" location="Rio Latoma" municipality="Puerta Parada" pageId="20" pageNumber="21" specimenCount="1">
<emphasis id="56DD99014720FFBCF82AFB29FE2C09F8" bold="true" box="[270,462,1157,1183]" pageId="20" pageNumber="21">
<collectingCountry id="1CBE05834720FFBCF82AFB29FE2C09F8" box="[270,462,1157,1183]" name="Guatemala" pageId="20" pageNumber="21">GUATEMALA</collectingCountry>
</emphasis>
:
<collectingMunicipality id="8472DF694720FFBCF8FAFB29FD6609F8" box="[478,644,1157,1183]" pageId="20" pageNumber="21">Puerta Parada</collectingMunicipality>
,
<date id="101763D34720FFBCFBB5FB29FCE809F8" box="[657,778,1157,1183]" pageId="20" pageNumber="21" value="1991-02-20">
<collectingDate id="00539A3B4720FFBCFBB5FB29FCE809F8" box="[657,778,1157,1183]" pageId="20" pageNumber="21" value="1991-02-20">20.ii.1991</collectingDate>
</date>
, 1L, M.
<collectorName id="C95C20C54720FFBCFA56FB29FBD609F8" box="[882,1076,1157,1183]" pageId="20" pageNumber="21">Esther Gonzalez</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFD66FB29FB4209F8" box="[1090,1184,1157,1183]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
];
<location id="617613C84720FFBCFD90FB29FADD09F8" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFD90FB29FADD09F8" box="[1204,1343,1157,1183]" country="Guatemala" municipality="Puerta Parada" name="Rio Latoma" pageId="20" pageNumber="21">Río Latoma</location>
at
<collectionCode id="02B8DDD64720FFBCFC41FB29FA7A09F8" box="[1381,1432,1157,1183]" country="Russia" name="Kotel'nich Museum" pageId="20" pageNumber="21">KM</collectionCode>
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCF9B3FB00FD1A09A1" box="[151,760,1196,1222]" collectionCode="R" country="Guatemala" county="Hwy." location="Temp" municipality="Elev." pageId="20" pageNumber="21" specimenCount="1">
182 on
<collectingCounty id="8D773D9F4720FFBCF9CBFB00FECF09A1" box="[239,301,1196,1222]" pageId="20" pageNumber="21">Hwy.</collectingCounty>
#2,
<collectingMunicipality id="8472DF694720FFBCF845FB00FE7E09A1" box="[353,412,1196,1222]" pageId="20" pageNumber="21">Elev.</collectingMunicipality>
2300',
<location id="617613C84720FFBCF8D6FB00FDD609A1" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF8D6FB00FDD609A1" box="[498,564,1196,1222]" country="Guatemala" county="Hwy." municipality="Elev." name="Temp" pageId="20" pageNumber="21">Temp</location>
82F, 24-x-68,
<collectionCode id="02B8DDD64720FFBCFBC5FB00FD1A09A1" box="[737,760,1196,1222]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="21">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFBD8FB00FC0909A1" box="[764,1003,1196,1222]" collectionCode="CAS" collectorName="R.K. Allen" country="Guatemala" pageId="20" pageNumber="21" specimenCount="1" specimenCount-larva="1">
<collectorName id="C95C20C54720FFBCFBD8FB00FCD509A1" box="[764,823,1196,1222]" name="R.K. Allen" pageId="20" pageNumber="21">K.A.</collectorName>
,
<specimenCount id="72AF8E9A4720FFBCFA67FB00FC7109A1" box="[835,915,1196,1222]" pageId="20" pageNumber="21" type="larva">1 larva</specimenCount>
(
<collectionCode id="02B8DDD64720FFBCFA80FB00FC0009A1" box="[932,994,1196,1222]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="D4C14F4E4720FFBCFAD3FB00FCC50874" collectingDate="1995-04-05" collectingDateMax="2001-07-14" collectingDateMin="1995-04-05" collectionCode="DEB, TAMU" collectorName="Km W. Tucuru &amp; Bryan Yates" country="Guatemala" county="Rio Cahabon" elevation="1500" latitude="15.319166" location="San Julian" longLatPrecision="21" longitude="-90.31833" municipality="Hwy." pageId="20" pageNumber="21" specimenCount="1" stateProvince="Alta Verapaz">
<collectingRegion id="A66D8BF14720FFBCFAD3FB00FB7109A1" box="[1015,1171,1196,1222]" country="Guatemala" name="Alta Verapaz" pageId="20" pageNumber="21">Alta Verapaz</collectingRegion>
;
<collectingCounty id="8D773D9F4720FFBCFDBBFB00FADB09A1" box="[1183,1337,1196,1222]" pageId="20" pageNumber="21">Río Cahabon</collectingCounty>
at
<collectingMunicipality id="8472DF694720FFBCFC79FB00FA7E09A1" box="[1373,1436,1196,1222]" pageId="20" pageNumber="21">Hwy.</collectingMunicipality>
7E,
<location id="617613C84720FFBCF9E3FB7EFEA1098B" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF9E3FB7EFEA1098B" box="[199,323,1234,1260]" country="Guatemala" county="Rio Cahabon" latitude="15.319166" longLatPrecision="21" longitude="-90.31833" municipality="Hwy." name="San Julian" pageId="20" pageNumber="21" stateProvince="Alta Verapaz">San Julian</location>
(
<geoCoordinate id="019D23D44720FFBCF870FB7EFE0B098B" box="[340,489,1234,1260]" degrees="15" direction="north" minutes="19" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="09" value="15.319166">N15°1909”</geoCoordinate>
;
<geoCoordinate id="019D23D44720FFBCF8D2FB7EFD72098B" box="[502,656,1234,1260]" degrees="90" direction="west" minutes="19" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="06" value="-90.31833">W90°1906”</geoCoordinate>
, elev.
<quantity id="A351E8F64720FFBCFBF9FB7EFCD8098B" box="[733,826,1234,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">
<elevation id="EF84A2204720FFBCFBF9FB7EFCD8098B" box="[733,826,1234,1260]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="21" unit="m" value="1500.0">1500 m</elevation>
</quantity>
),
<date id="101763D34720FFBCFA75FB7EFC38098B" box="[849,986,1234,1260]" pageId="20" pageNumber="21" value="2001-07-14">
<collectingDate id="00539A3B4720FFBCFA75FB7EFC38098B" box="[849,986,1234,1260]" pageId="20" pageNumber="21" value="2001-07-14">14.vii.2001</collectingDate>
</date>
, 30L,
<collectionCode id="02B8DDD64720FFBCFD03FB7EFB80098B" box="[1063,1122,1234,1260]" pageId="20" pageNumber="21">DEB</collectionCode>
[
<collectionCode id="02B8DDD64720FFBCFD55FB7EFB2D098B" box="[1137,1231,1234,1260]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
];
<location id="617613C84720FFBCFDC7FB7EFA63098B" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFDC7FB7EFA63098B" box="[1251,1409,1234,1260]" country="Guatemala" county="Rio Cahabon" latitude="15.319166" longLatPrecision="21" longitude="-90.31833" municipality="Hwy." name="Rio Polochic" pageId="20" pageNumber="21" stateProvince="Alta Verapaz">Río Polochic</location>
, 2
<collectorName id="C95C20C54720FFBCF9B3FB55FEB00874" box="[151,338,1273,1299]" pageId="20" pageNumber="21">Km W. Tucuru</collectorName>
,
<date id="101763D34720FFBCF844FB55FE340874" box="[352,470,1273,1299]" pageId="20" pageNumber="21" value="1995-04-05">
<collectingDate id="00539A3B4720FFBCF844FB55FE340874" box="[352,470,1273,1299]" pageId="20" pageNumber="21" value="1995-04-05">5.iv.1995</collectingDate>
</date>
, 4L,
<collectorName id="C95C20C54720FFBCFB33FB55FD4F0874" box="[535,685,1273,1299]" pageId="20" pageNumber="21">Bryan Yates</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFB9BFB55FCC20874" box="[703,800,1273,1299]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="D4C14F4E4720FFBCFA11FB55FBE0085D" collectingDate="2001-07-13" collectionCode="DEB, TAMU" country="Guatemala" elevation="1050" latitude="15.064445" location="San Jeronimo" longLatPrecision="21" longitude="-90.23416" municipality="Rio San Jeronimo" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Baja Verapaz">
<collectingRegion id="A66D8BF14720FFBCFA11FB55FC3F0874" box="[821,989,1273,1299]" country="Guatemala" name="Baja Verapaz" pageId="20" pageNumber="21">Baja Verapaz</collectingRegion>
;
<collectingMunicipality id="8472DF694720FFBCFACFFB55FB280874" box="[1003,1226,1273,1299]" pageId="20" pageNumber="21">Río San Jeronimo</collectingMunicipality>
at
<location id="617613C84720FFBCFDD7FB55FA7E0874" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFDD7FB55FA7E0874" box="[1267,1436,1273,1299]" country="Guatemala" latitude="15.064445" longLatPrecision="21" longitude="-90.23416" municipality="Rio San Jeronimo" name="San Jeronimo" pageId="20" pageNumber="21" stateProvince="Baja Verapaz">San Jeronimo</location>
(
<geoCoordinate id="019D23D44720FFBCF984FA8CFED0085D" box="[160,306,1312,1338]" degrees="15" direction="north" minutes="03" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="52" value="15.064445">N15°0352”</geoCoordinate>
;
<geoCoordinate id="019D23D44720FFBCF81BFA8CFE35085D" box="[319,471,1312,1338]" degrees="90" direction="west" minutes="14" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="03" value="-90.23416">W90°1403”</geoCoordinate>
, elev.
<quantity id="A351E8F64720FFBCFB05FA8CFD9F085D" box="[545,637,1312,1338]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="21" unit="m" value="1050.0">
<elevation id="EF84A2204720FFBCFB05FA8CFD9F085D" box="[545,637,1312,1338]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="20" pageNumber="21" unit="m" value="1050.0">1050 m</elevation>
</quantity>
),
<date id="101763D34720FFBCFBB6FA8CFCF8085D" box="[658,794,1312,1338]" pageId="20" pageNumber="21" value="2001-07-13">
<collectingDate id="00539A3B4720FFBCFBB6FA8CFCF8085D" box="[658,794,1312,1338]" pageId="20" pageNumber="21" value="2001-07-13">13.vii.2001</collectingDate>
</date>
, 4L,
<collectionCode id="02B8DDD64720FFBCFA71FA8CFC72085D" box="[853,912,1312,1338]" pageId="20" pageNumber="21">DEB</collectionCode>
[
<collectionCode id="02B8DDD64720FFBCFAB9FA8CFC19085D" box="[925,1019,1312,1338]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFD29FAB3FF5C0807" country="Honduras" location="5 mi. So." pageId="20" pageNumber="21" specimenCount="1" stateProvince="Comayagua">
<emphasis id="56DD99014720FFBCFD29FAB3FB5C085E" bold="true" box="[1037,1214,1311,1337]" pageId="20" pageNumber="21">
<collectingCountry id="1CBE05834720FFBCFD29FAB3FB5B085E" box="[1037,1209,1311,1337]" name="Honduras" pageId="20" pageNumber="21">HONDURAS</collectingCountry>
:
</emphasis>
<collectingRegion id="A66D8BF14720FFBCFDE1FA8CFAB6085D" box="[1221,1364,1312,1338]" country="Honduras" name="Comayagua" pageId="20" pageNumber="21">Comayagua</collectingRegion>
,
<location id="617613C84720FFBCFC44FA8CFF5C0807" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFC44FA8CFF5C0807" country="Honduras" name="5 mi. So." pageId="20" pageNumber="21" stateProvince="Comayagua">
<quantity id="A351E8F64720FFBCFC44FA8CFA7E085D" box="[1376,1436,1312,1338]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="20" pageNumber="21" unit="mi" value="5.0">5 mi.</quantity>
So.
</location>
</materialsCitation>
<materialsCitation id="D4C14F4E4720FFBCF9E3FAEAFB9D0807" box="[199,1151,1350,1376]" collectingDate="1964-10-17" collectionCode="CAS" collectorName="L, J. S. Packer" country="Honduras" location="Hwy." pageId="20" pageNumber="21" specimenCount="1" stateProvince="Comayagua">
<collectingRegion id="A66D8BF14720FFBCF9E3FAEAFEB50807" box="[199,343,1350,1376]" country="Honduras" name="Comayagua" pageId="20" pageNumber="21">Comayagua</collectingRegion>
on
<location id="617613C84720FFBCF8ACFAEAFE2A0807" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF8ACFAEAFE2A0807" box="[392,456,1350,1376]" country="Honduras" name="Hwy." pageId="20" pageNumber="21" stateProvince="Comayagua">Hwy.</location>
#1 at bridge, stream.,
<date id="101763D34720FFBCFBFAFAEAFCB80807" box="[734,858,1350,1376]" pageId="20" pageNumber="21" value="1964-10-17">
<collectingDate id="00539A3B4720FFBCFBFAFAEAFCB80807" box="[734,858,1350,1376]" pageId="20" pageNumber="21" value="1964-10-17">17.x.1964</collectingDate>
</date>
, 3
<collectorName id="C95C20C54720FFBCFA53FAEAFBC00807" box="[887,1058,1350,1376]" pageId="20" pageNumber="21">L, J.S. Packer</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFD17FAEAFB940807" box="[1075,1142,1350,1376]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFDAFFAEAFC9E08C9" collectingDate="1964-10-31" collectingDateMax="1964-11-06" collectingDateMin="1964-10-31" collectionCode="CAS, FAMU" collectorName="E. Zamorano &amp; L, J. S. Packer &amp; E. Guaimaca" country="Honduras" location="Guaimaca on Hwy." municipality="Zamorano on Hwy." pageId="20" pageNumber="21" specimenCount="1" stateProvince="El Paraiso">
<emphasis id="56DD99014720FFBCFDAFFAEAFAF00807" bold="true" box="[1163,1298,1350,1376]" pageId="20" pageNumber="21">
<collectingRegion id="A66D8BF14720FFBCFDAFFAEAFAF00807" box="[1163,1298,1350,1376]" country="Honduras" name="El Paraiso" pageId="20" pageNumber="21">El Paraiso</collectingRegion>
</emphasis>
;
<quantity id="A351E8F64720FFBCFC00FAEAFA970807" box="[1316,1397,1350,1376]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="20" pageNumber="21" unit="km" value="38.0">38 km</quantity>
.
<collectorName id="C95C20C54720FFBCFCA6FAEAFEED08E0" pageId="20" pageNumber="21">
E.
<collectingMunicipality id="8472DF694720FFBCF9B3FAC1FEED08E0" box="[151,271,1389,1415]" pageId="20" pageNumber="21">Zamorano</collectingMunicipality>
</collectorName>
on Hwy. #4, stream.,
<date id="101763D34720FFBCFB30FAC1FD6E08E0" box="[532,652,1389,1415]" pageId="20" pageNumber="21" value="1964-10-31">
<collectingDate id="00539A3B4720FFBCFB30FAC1FD6E08E0" box="[532,652,1389,1415]" pageId="20" pageNumber="21" value="1964-10-31">31.x.1964</collectingDate>
</date>
, 7
<collectorName id="C95C20C54720FFBCFB8CFAC1FCA808E0" box="[680,842,1389,1415]" pageId="20" pageNumber="21">L, J.S. Packer</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFA7EFAC1FC7908E0" box="[858,923,1389,1415]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
].
<emphasis id="56DD99014720FFBCFA8BFAC1FBCB08E0" bold="true" box="[943,1065,1389,1415]" pageId="20" pageNumber="21">Francisco</emphasis>
; Morazan,
<quantity id="A351E8F64720FFBCFD94FAC1FB1E08E0" box="[1200,1276,1389,1415]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="20" pageNumber="21" unit="mi" value="10.0">10 mi.</quantity>
<collectorName id="C95C20C54720FFBCFC20FAC1FA7E08E0" box="[1284,1436,1389,1415]" pageId="20" pageNumber="21">
E.
<location id="617613C84720FFBCFC01FAC1FA7E08E0" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFC01FAC1FA7E08E0" box="[1317,1436,1389,1415]" country="Honduras" municipality="Zamorano on Hwy." name="Guaimaca on Hwy." pageId="20" pageNumber="21" stateProvince="El Paraiso">Guaimaca</location>
</collectorName>
on Hwy. #3, Small stream,
<date id="101763D34720FFBCF8FEFA38FDA908C9" box="[474,587,1428,1454]" pageId="20" pageNumber="21" value="1964-11-06">
<collectingDate id="00539A3B4720FFBCF8FEFA38FDA908C9" box="[474,587,1428,1454]" pageId="20" pageNumber="21" value="1964-11-06">6.xi.1964</collectingDate>
</date>
, 4
<collectorName id="C95C20C54720FFBCFB43FA38FCE808C9" box="[615,778,1428,1454]" pageId="20" pageNumber="21">L, J.S. Packer</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFA3DFA38FC9708C9" box="[793,885,1428,1454]" pageId="20" pageNumber="21">FAMU</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFAACFA3FFA7A08B3" collectingDate="1964-11-06" collectionCode="CAS" collectorName="L, J. S. Packer" country="Honduras" county="Juticalpa on Hwy." location="Juticalpa" municipality="Rio Telica" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Olancho">
<emphasis id="56DD99014720FFBCFAACFA3FFC1008CA" bold="true" box="[904,1010,1427,1453]" pageId="20" pageNumber="21">
<collectingRegion id="A66D8BF14720FFBCFAACFA3FFC1008CA" box="[904,1010,1427,1453]" country="Honduras" name="Olancho" pageId="20" pageNumber="21">Olancho</collectingRegion>
</emphasis>
;
<quantity id="A351E8F64720FFBCFD26FA38FBDD08C9" box="[1026,1087,1428,1454]" metricMagnitude="3" metricUnit="m" metricValue="9.656064" pageId="20" pageNumber="21" unit="mi" value="6.0">6 mi.</quantity>
E.
<collectingCounty id="8D773D9F4720FFBCFD4CFA38FADF08C9" box="[1128,1341,1428,1454]" pageId="20" pageNumber="21">Juticalpa on Hwy.</collectingCounty>
#3,
<collectingMunicipality id="8472DF694720FFBCFC56FA38FF0308B3" pageId="20" pageNumber="21">Río Telica</collectingMunicipality>
,
<date id="101763D34720FFBCF9C8FA16FEBF08B3" box="[236,349,1466,1492]" pageId="20" pageNumber="21" value="1964-11-06">
<collectingDate id="00539A3B4720FFBCF9C8FA16FEBF08B3" box="[236,349,1466,1492]" pageId="20" pageNumber="21" value="1964-11-06">6.xi.1964</collectingDate>
</date>
, 1
<collectorName id="C95C20C54720FFBCF850FA16FDFA08B3" box="[372,536,1466,1492]" pageId="20" pageNumber="21">L, J.S. Packer</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFB02FA16FD8508B3" box="[550,615,1466,1492]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
];
<quantity id="A351E8F64720FFBCFB5EFA16FD2608B3" box="[634,708,1466,1492]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="20" pageNumber="21" unit="mi" value="10.0">10 mi.</quantity>
W.
<location id="617613C84720FFBCFBD6FA16FCBF08B3" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFBD6FA16FCBF08B3" box="[754,861,1466,1492]" country="Honduras" county="Juticalpa on Hwy." municipality="Rio Telica" name="Juticalpa" pageId="20" pageNumber="21" stateProvince="Olancho">Juticalpa</location>
,
<location id="617613C84720FFBCFA4CFA16FBE908B3" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFA4CFA16FBE908B3" box="[872,1035,1466,1492]" country="Honduras" county="Juticalpa on Hwy." municipality="Rio Telica" name="Rio Juticalpa." pageId="20" pageNumber="21" stateProvince="Olancho">Río Juticalpa.</location>
,
<date id="101763D34720FFBCFD32FA16FB6508B3" box="[1046,1159,1466,1492]" pageId="20" pageNumber="21" value="1964-11-06">
<collectingDate id="00539A3B4720FFBCFD32FA16FB6508B3" box="[1046,1159,1466,1492]" pageId="20" pageNumber="21" value="1964-11-06">6.xi.1964</collectingDate>
</date>
, 1
<collectorName id="C95C20C54720FFBCFDBAFA16FAA008B3" box="[1182,1346,1466,1492]" pageId="20" pageNumber="21">L, J.S. Packer</collectorName>
[
<collectionCode id="02B8DDD64720FFBCFC74FA16FA7308B3" box="[1360,1425,1466,1492]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="20" pageNumber="21" type="Museum">CAS</collectionCode>
]
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCF9B3FA4CFDF2089C" box="[151,528,1504,1531]" collectionCode="MP" country="Mexico" location="Queretaro" pageId="20" pageNumber="21" specimenCode="MP10" specimenCount="1" stateProvince="Queretaro de Arteaga">
<emphasis id="56DD99014720FFBCF9B3FA4CFE4D089D" bold="true" box="[151,431,1504,1530]" pageId="20" pageNumber="21">
<collectingCountry id="1CBE05834720FFBCF9B3FA4CFEF9089D" box="[151,283,1504,1530]" name="Mexico" pageId="20" pageNumber="21">MEXICO</collectingCountry>
:
<collectingRegion id="A66D8BF14720FFBCF80DFA4CFE4D089D" box="[297,431,1504,1530]" country="Mexico" name="Queretaro de Arteaga" pageId="20" pageNumber="21">Queretaro</collectingRegion>
</emphasis>
;
<specimenCode id="340FED684720FFBCF8E6FA4DFDF2089C" box="[450,528,1505,1531]" collectionCode="MP" country="0" name="Mohonk Preserve, Inc." pageId="20" pageNumber="21">MP10</specimenCode>
</materialsCitation>
:
<materialsCitation id="D4C14F4E4720FFBCFB39FA4DFAA9089C" box="[541,1355,1505,1531]" collectingDate="1997-08-28" collectionCode="R" country="Mexico" location="Agua Fria" municipality="Pinal de Amoles" pageId="20" pageNumber="21" specimenCount="1">
<collectingMunicipality id="8472DF694720FFBCFB39FA4DFD0F089C" box="[541,749,1505,1531]" pageId="20" pageNumber="21">Pinal de Amoles</collectingMunicipality>
,
<location id="617613C84720FFBCFBDEFA4DFC94089C" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCFBDEFA4DFC94089C" box="[762,886,1505,1531]" country="Mexico" municipality="Pinal de Amoles" name="Agua Fria" pageId="20" pageNumber="21">Agua Fria</location>
(UTM 2338413),
<date id="101763D34720FFBCFD7FFA4DFB11089C" box="[1115,1267,1505,1531]" pageId="20" pageNumber="21" value="1997-08-28">
<collectingDate id="00539A3B4720FFBCFD7FFA4DFB11089C" box="[1115,1267,1505,1531]" pageId="20" pageNumber="21" value="1997-08-28">28.viii.1997</collectingDate>
</date>
, 3L,
<collectionCode id="02B8DDD64720FFBCFC10FA4DFAA9089C" box="[1332,1355,1505,1531]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="21">R</collectionCode>
</materialsCitation>
.
<materialsCitation id="D4C14F4E4720FFBCFC7CFA4DFB000B45" collectingDate="2000-07-13" collectionCode="TAMU, WDS" country="Mexico" county="Jones" latitude="21.265556" location="Rio Verdito" longLatPrecision="21" longitude="-99.056946" municipality="Neblinas" pageId="20" pageNumber="21" specimenCount="1">
<collectingCounty id="8D773D9F4720FFBCFC7CFA4DFA79089C" box="[1368,1435,1505,1531]" pageId="20" pageNumber="21">Jones</collectingCounty>
[
<collectionCode id="02B8DDD64720FFBCF9B9F9A4FF180B45" box="[157,250,1544,1570]" country="USA" lsid="urn:lsid:biocol.org:col:14193" name="Texas A&amp;amp;M University" pageId="20" pageNumber="21" type="Herbarium">TAMU</collectionCode>
];
<collectingMunicipality id="8472DF694720FFBCF82AF9A4FE9B0B45" box="[270,377,1544,1570]" pageId="20" pageNumber="21">Neblinas</collectingMunicipality>
,
<location id="617613C84720FFBCF8A0F9A4FDEE0B45" LSID="urn:lsid:plazi:treatment:EC00F4054727FFBCF9B3FE61FB040B45:617613C84720FFBCF8A0F9A4FDEE0B45" box="[388,524,1544,1570]" country="Mexico" county="Jones" latitude="21.265556" longLatPrecision="21" longitude="-99.056946" municipality="Neblinas" name="Rio Verdito" pageId="20" pageNumber="21">Río Verdito</location>
(
<geoCoordinate id="019D23D44720FFBCFB39F9A4FD520B45" box="[541,688,1544,1570]" degrees="21" direction="north" minutes="15" orientation="latitude" pageId="20" pageNumber="21" precision="15" seconds="56" value="21.265556">N21°1556”</geoCoordinate>
;
<geoCoordinate id="019D23D44720FFBCFB98F9A4FCB70B45" box="[700,853,1544,1570]" degrees="99" direction="west" minutes="03" orientation="longitude" pageId="20" pageNumber="21" precision="15" seconds="25" value="-99.056946">W99°0325”</geoCoordinate>
),
<date id="101763D34720FFBCFA4EF9A4FC100B45" box="[874,1010,1544,1570]" pageId="20" pageNumber="21" value="2000-07-13">
<collectingDate id="00539A3B4720FFBCFA4EF9A4FC100B45" box="[874,1010,1544,1570]" pageId="20" pageNumber="21" value="2000-07-13">13.vii.2000</collectingDate>
</date>
, 1L,
<collectionCode id="02B8DDD64720FFBCFD0AF9A4FB8D0B45" box="[1070,1135,1544,1570]" pageId="20" pageNumber="21">WDS</collectionCode>
[
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.
</paragraph>
</subSubSection>
</treatment>
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