437 lines
53 KiB
XML
437 lines
53 KiB
XML
<document id="5D27F41D650B4B34181A3728B0416D3A" ID-CLB-Dataset="39225" ID-DOI="10.11646/zootaxa.4059.2.2" ID-GBIF-Dataset="65f3dd28-09dd-4739-99cc-0031cbd72b0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="253812" ID-ZooBank="5D7E8244-3A68-40A7-ABCF-8932C6492948" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461361456702" checkinUser="plazi" docAuthor="Steinhoff, Philip O. M. & Uhl, Gabriele" docDate="2015" docId="D04487FBA679FFBEFF78F973FCB9151E" docLanguage="en" docName="zt04059p276.pdf" docOrigin="Zootaxa 4059 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia scutellum Laidlaw 1932" docType="treatment" docVersion="8" lastPageNumber="262" masterDocId="2C7DFF83A678FFBBFFEFFF94FFB81013" masterDocTitle="Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis" masterLastPageNumber="276" masterPageNumber="257" pageNumber="258" updateTime="1698661837837" updateUser="ExternalLinkService">
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<mods:titleInfo id="34205B5817E69611F6C7E7648689B617">
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<mods:title id="7337889F9784D0C8684541541538194C">Taxonomy and nomenclature of some mainland SE-Asian Coeliccia species (Odonata, Platycnemididae) using micro-CT analysis</mods:title>
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<mods:namePart id="1FE23628E8A8B2D9FDDE216FA94FD974">Steinhoff, Philip O. M.</mods:namePart>
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<mods:namePart id="2655C703823DE9C863578B84FD953290">Uhl, Gabriele</mods:namePart>
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<mods:typeOfResource id="ACEC039384BE9AF46354C93977AE64A5">text</mods:typeOfResource>
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<mods:title id="CC3E4B580C0AB85168FE7B9E9F792631">Zootaxa</mods:title>
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<mods:part id="85038A1483388F73B01FFAF8B1BB8B66">
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<mods:date id="28FA70044404CB4DE5671715BD3E40B5">2015</mods:date>
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<mods:detail id="321D4D9B494E68E3B5936AD75BF8C724" type="volume">
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<mods:number id="236781EB4B617969F8F58A4F95E9B35A">4059</mods:number>
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<mods:number id="DBB27CA447AA7F03EFC13219C4768B7F">2</mods:number>
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<mods:identifier id="44034F1E7D0EBC5E304FE2739EB2246F" type="DOI">10.11646/zootaxa.4059.2.2</mods:identifier>
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<treatment id="D04487FBA679FFBEFF78F973FCB9151E" ID-DOI="http://doi.org/10.5281/zenodo.6100439" ID-GBIF-Taxon="119648603" ID-Zenodo-Dep="6100439" LSID="urn:lsid:plazi:treatment:D04487FBA679FFBEFF78F973FCB9151E" httpUri="http://treatment.plazi.org/id/D04487FBA679FFBEFF78F973FCB9151E" lastPageId="5" lastPageNumber="262" pageId="1" pageNumber="258">
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<subSubSection id="10F76566A679FFBAFF78F973FD821711" box="[151,570,1767,1794]" pageId="1" pageNumber="258" type="nomenclature">
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<paragraph id="585236EDA679FFBAFF78F973FD821711" blockId="1.[151,570,1767,1794]" box="[151,570,1767,1794]" pageId="1" pageNumber="258">
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<heading id="031A8181A679FFBAFF78F973FD821711" bold="true" box="[151,570,1767,1794]" fontSize="11" level="1" pageId="1" pageNumber="258" reason="1">
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<taxonomicName id="9FED4D6EA679FFBAFF78F973FD821711" ID-CoL="WQDD" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[151,570,1767,1794]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="scutellum">
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<emphasis id="6A99EAFFA679FFBAFF78F973FD821711" bold="true" box="[151,570,1767,1794]" pageId="1" pageNumber="258">
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<emphasis id="6A99EAFFA679FFBAFF78F973FE3C1712" bold="true" box="[151,388,1767,1793]" italics="true" pageId="1" pageNumber="258">Coeliccia scutellum</emphasis>
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<bibRefCitation id="3C7C4B1CA679FFBAFE64F97CFD821711" author="Laidlaw" box="[395,570,1768,1794]" pageId="1" pageNumber="258" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="10F76566A679FFB9FF78F8BAFE6E110F" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258" type="reference_group">
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<paragraph id="585236EDA679FFBAFF78F8BAFA2417C4" blockId="1.[151,1436,1838,2007]" pageId="1" pageNumber="258">
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<taxonomicName id="9FED4D6EA679FFBAFF78F8BAFD9D1754" ID-CoL="WQDK" authority="Asahina, 1997" authorityName="Asahina" authorityYear="1997" box="[151,549,1838,1863]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="6A99EAFFA679FFBAFF78F8BAFEC21755" box="[151,378,1838,1862]" italics="true" pageId="1" pageNumber="258">Coeliccia tomokunii</emphasis>
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<bibRefCitation id="3C7C4B1CA679FFBAFE6DF8BAFD9D1754" author="Asahina" box="[386,549,1838,1863]" pageId="1" pageNumber="258" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina, 1997</bibRefCitation>
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</taxonomicName>
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has been considered a junior synonym of
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<taxonomicName id="9FED4D6EA679FFBAFBEAF8BBFEF01778" ID-CoL="WQDD" authority="Laidlaw, 1932 (Wilson & Reels 2001)" authorityName="Laidlaw, 1932 (Wilson & Reels" authorityYear="2001" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="scutellum">
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<emphasis id="6A99EAFFA679FFBAFBEAF8BBFB2A1755" box="[1029,1170,1838,1862]" italics="true" pageId="1" pageNumber="258">C. scutellum</emphasis>
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<bibRefCitation id="3C7C4B1CA679FFBAFB75F8BAFA831754" author="Laidlaw" box="[1178,1339,1838,1863]" pageId="1" pageNumber="258" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
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(
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<bibRefCitation id="3C7C4B1CA679FFBAFAA5F8BAFEF81778" author="Wilson" pageId="1" pageNumber="258" refString="Wilson, K. D. P. & Reels, G. T. (2001) Odonata of Hainan, China. Odonatologica, 30 (2), 145 - 208." type="journal article" year="2001">Wilson & Reels 2001</bibRefCitation>
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)
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</taxonomicName>
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.
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<bibRefCitation id="3C7C4B1CA679FFBAFEB6F8C6FD841778" author="Asahina" box="[345,572,1874,1899]" pageId="1" pageNumber="258" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina (1997: 24)</bibRefCitation>
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indicated this possibility in his original description of
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<taxonomicName id="9FED4D6EA679FFBAFB3EF8C7FADF1778" ID-CoL="WQDK" box="[1233,1383,1875,1899]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="6A99EAFFA679FFBAFB3EF8C7FADF1778" box="[1233,1383,1875,1899]" italics="true" pageId="1" pageNumber="258">C. tomokunii</emphasis>
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</taxonomicName>
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and suggested that “...a careful comparison of the
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<typeStatus id="8756884FA679FFBAFD4BF8E3FD6C179C" box="[676,724,1911,1935]" pageId="1" pageNumber="258">type</typeStatus>
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specimen is needed between the two species.” In order to test this possible synonymy we compared the
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<typeStatus id="8756884FA679FFBAFD94F80EFD5A17A0" box="[635,738,1946,1971]" pageId="1" pageNumber="258" type="lectotype">lectotype</typeStatus>
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of
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<taxonomicName id="9FED4D6EA679FFBAFCE3F80FFC2217A0" ID-CoL="WQDD" box="[780,922,1947,1971]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="scutellum">
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<emphasis id="6A99EAFFA679FFBAFCE3F80FFC2217A0" box="[780,922,1947,1971]" italics="true" pageId="1" pageNumber="258">C. scutellum</emphasis>
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</taxonomicName>
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with specimens that were collected near the
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<typeStatus id="8756884FA679FFBAFF78F82BFF7F17C4" box="[151,199,1983,2007]" pageId="1" pageNumber="258">type</typeStatus>
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locality of
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<emphasis id="6A99EAFFA679FFBAFEB8F82BFE4C17C5" box="[343,500,1982,2006]" italics="true" pageId="1" pageNumber="258">
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<taxonomicName id="9FED4D6EA679FFBAFEB8F82BFE4817C5" ID-CoL="WQDK" box="[343,496,1982,2006]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="tomokunii">C. tomokunii</taxonomicName>
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,
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</emphasis>
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since the
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<typeStatus id="8756884FA679FFBAFD9AF82BFD1D17C4" box="[629,677,1983,2007]" pageId="1" pageNumber="258">type</typeStatus>
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specimens of
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<taxonomicName id="9FED4D6EA679FFBAFCB9F82BFC5417C5" ID-CoL="WQDK" box="[854,1004,1982,2006]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="258" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="6A99EAFFA679FFBAFCB9F82BFC5417C5" box="[854,1004,1982,2006]" italics="true" pageId="1" pageNumber="258">C. tomokunii</emphasis>
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</taxonomicName>
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are not accessible and may be lost.
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</paragraph>
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<paragraph id="585236EDA67AFFB9FF78FF03FE6E110F" blockId="2.[151,1436,151,285]" pageId="2" pageNumber="259">
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Descriptions and illustrations of specimens are provided and compared with the original descriptions by
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<bibRefCitation id="3C7C4B1CA67AFFB9FAAFFF03FF5910C7" author="Laidlaw" pageId="2" pageNumber="259" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw (1932)</bibRefCitation>
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and
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<bibRefCitation id="3C7C4B1CA67AFFB9FECEFF28FE6E10C7" author="Asahina" box="[289,470,188,213]" pageId="2" pageNumber="259" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina (1997)</bibRefCitation>
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.
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<bibRefCitation id="3C7C4B1CA67AFFB9FE09FF28FD2010C7" author="Laidlaw" box="[486,664,188,213]" pageId="2" pageNumber="259" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw (1932)</bibRefCitation>
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gave only a short description and a rough sketch of the thoracic markings of
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<taxonomicName id="9FED4D6EA67AFFB9FEC9FF74FE0B10EB" ID-CoL="WQDD" box="[294,435,224,248]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="scutellum">
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<emphasis id="6A99EAFFA67AFFB9FEC9FF74FE0B10EB" box="[294,435,224,248]" italics="true" pageId="2" pageNumber="259">C. scutellum</emphasis>
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</taxonomicName>
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, whereas Asahina's (1997) description includes illustrations of the head, thorax and anal appendages of
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<taxonomicName id="9FED4D6EA67AFFB9FED0FE90FE77110F" ID-CoL="WQDK" box="[319,463,260,284]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="tomokunii">
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<emphasis id="6A99EAFFA67AFFB9FED0FE90FE77110F" box="[319,463,260,284]" italics="true" pageId="2" pageNumber="259">C. tomokunii</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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||
<caption id="0C926665A67AFFB9FF78F91FFCE416CD" httpUri="https://zenodo.org/record/253813/files/figure.png" pageId="2" pageNumber="259" targetBox="[317,1269,311,1658]" targetPageId="2">
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<paragraph id="585236EDA67AFFB9FF78F91FFCE416CD" blockId="2.[151,1436,1675,1758]" pageId="2" pageNumber="259">
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<emphasis id="6A99EAFFA67AFFB9FF78F91FFEAC16B3" bold="true" box="[151,276,1675,1697]" pageId="2" pageNumber="259">FIGURE 1.</emphasis>
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Mounting methods and detail of genital ligula.
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<emphasis id="6A99EAFFA67AFFB9FCEEF91FFCAB16B3" bold="true" box="[769,787,1675,1696]" pageId="2" pageNumber="259">A</emphasis>
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Pinned specimen (
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<taxonomicName id="9FED4D6EA67AFFB9FC36F91FFB1B16B2" box="[985,1187,1675,1697]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="scutellum">
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<emphasis id="6A99EAFFA67AFFB9FC36F91FFB1B16B2" box="[985,1187,1675,1697]" italics="true" pageId="2" pageNumber="259">Coeliccia scutellum</emphasis>
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||
</taxonomicName>
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||
) mounted for micro-CT scan,
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<emphasis id="6A99EAFFA67AFFB9FF3EF93EFF5A16AC" bold="true" box="[209,226,1706,1727]" pageId="2" pageNumber="259">B</emphasis>
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papered specimen (
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<taxonomicName id="9FED4D6EA67AFFB9FE5FF93DFDC116AC" box="[432,633,1705,1727]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="scutellum">
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<emphasis id="6A99EAFFA67AFFB9FE5FF93DFDC116AC" box="[432,633,1705,1727]" italics="true" pageId="2" pageNumber="259">Coeliccia scutellum</emphasis>
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||
</taxonomicName>
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) mounted for micro-CT scan,
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<emphasis id="6A99EAFFA67AFFB9FC5CF93DFC7D16AC" bold="true" box="[947,965,1705,1727]" pageId="2" pageNumber="259">C</emphasis>
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macro-photograph (Zeiss MCr Camera) of
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<taxonomicName id="9FED4D6EA67AFFB9FA6BF93EFF4016CD" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="scutellum">
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||
<emphasis id="6A99EAFFA67AFFB9FA6BF93EFF4016CD" italics="true" pageId="2" pageNumber="259">C. scutellum</emphasis>
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||
</taxonomicName>
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||
genital ligula, showing thinness of flaps.
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||
<emphasis id="6A99EAFFA67AFFB9FD71F95CFCBF16CD" box="[670,775,1736,1758]" italics="true" pageId="2" pageNumber="259">Scale bar:</emphasis>
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||
250Μm.
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</paragraph>
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||
</caption>
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||
<subSubSection id="10F76566A67AFFB9FF28F891FEBD17E6" pageId="2" pageNumber="259" type="materials_examined">
|
||
<paragraph id="585236EDA67AFFB9FF28F891FDBA1751" blockId="2.[151,1436,1797,2037]" pageId="2" pageNumber="259">
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||
<emphasis id="6A99EAFFA67AFFB9FF28F891FE0B170D" bold="true" box="[199,435,1797,1822]" pageId="2" pageNumber="259">Material examined.</emphasis>
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||
<typeStatus id="8756884FA67AFFB9FE52F892FE4D170D" box="[445,501,1798,1822]" pageId="2" pageNumber="259">Type</typeStatus>
|
||
material:
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||
<specimenCount id="4EEBFD64A67AFFB9FD83F892FD37170D" box="[620,655,1798,1822]" pageId="2" pageNumber="259" type="generic">1♂</specimenCount>
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||
<typeStatus id="8756884FA67AFFB9FD76F892FCB6170D" box="[665,782,1798,1822]" pageId="2" pageNumber="259" type="lectotype">Lectotype</typeStatus>
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||
: Bao Ha,
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||
<collectingCountry id="20FA767DA67AFFB9FC6CF891FC6A170D" box="[899,978,1797,1822]" name="Vietnam" pageId="2" pageNumber="259">Tonkin</collectingCountry>
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||
(
|
||
<collectingCountry id="20FA767DA67AFFB9FC0BF891FBF1170D" box="[996,1097,1797,1822]" name="Vietnam" pageId="2" pageNumber="259">Vietnam</collectingCountry>
|
||
),
|
||
<date id="2C53102DA67AFFB9FBB1F892FB67170D" box="[1118,1247,1798,1822]" pageId="2" pageNumber="259" value="1924-04-20">20.04.1924</date>
|
||
, leg. H. Stevens [
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||
<collectionCode id="3EFCAE28A67AFFB9FF72F8BEFF4F1752" box="[157,247,1834,1857]" pageId="2" pageNumber="259">BMNH</collectionCode>
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||
,
|
||
<collectionCode id="3EFCAE28A67AFFB9FEEDF8BEFED41752" LSID="urn:lsid:biocol.org:col:34665" box="[258,364,1834,1857]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="259">NHMUK</collectionCode>
|
||
010266912].
|
||
</paragraph>
|
||
<paragraph id="585236EDA67AFFB9FF28F8D9FEBD17E6" blockId="2.[151,1436,1797,2037]" pageId="2" pageNumber="259">
|
||
Other material: Total
|
||
<specimenCount id="4EEBFD64A67AFFB9FE54F8D9FE591775" box="[443,481,1869,1894]" pageId="2" pageNumber="259" type="generic">6♂</specimenCount>
|
||
:
|
||
<specimenCount id="4EEBFD64A67AFFB9FE02F8DAFDA81775" box="[493,528,1870,1894]" pageId="2" pageNumber="259" type="generic">1♂</specimenCount>
|
||
Mt. Tam
|
||
<collectingCountry id="20FA767DA67AFFB9FD6FF8DAFD091776" box="[640,689,1870,1893]" name="Iceland" pageId="2" pageNumber="259">Dao</collectingCountry>
|
||
, Vinh Phu Province, northern
|
||
<collectingCountry id="20FA767DA67AFFB9FBE7F8D9FBD21775" box="[1032,1130,1869,1894]" name="Vietnam" pageId="2" pageNumber="259">Vietnam</collectingCountry>
|
||
, 1994, leg. M. Hämäläinen [
|
||
<collectionCode id="3EFCAE28A67AFFB9FF72F8E6FF42179A" LSID="urn:lsid:biocol.org:col:34992" box="[157,250,1906,1929]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="259">RMNH</collectionCode>
|
||
].
|
||
<specimenCount id="4EEBFD64A67AFFB9FEE0F8E6FE8A1799" box="[271,306,1906,1930]" pageId="2" pageNumber="259" type="generic">1♂</specimenCount>
|
||
“Stream below Thac Bac” Tam
|
||
<collectingCountry id="20FA767DA67AFFB9FD5CF8E6FD5C179A" box="[691,740,1906,1929]" name="Iceland" pageId="2" pageNumber="259">Dao</collectingCountry>
|
||
, Vinh Phu Province, northern
|
||
<collectingCountry id="20FA767DA67AFFB9FBA7F8E5FB121799" box="[1096,1194,1905,1930]" name="Vietnam" pageId="2" pageNumber="259">Vietnam</collectingCountry>
|
||
,
|
||
<date id="2C53102DA67AFFB9FB56F8E5FA821799" box="[1209,1338,1905,1930]" pageId="2" pageNumber="259" value="2008-06-26">26.06.2008</date>
|
||
, leg. M. Hämäläinen [
|
||
<collectionCode id="3EFCAE28A67AFFB9FEC2F802FE3217BE" LSID="urn:lsid:biocol.org:col:34992" box="[301,394,1942,1965]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="259">RMNH</collectionCode>
|
||
].
|
||
<specimenCount id="4EEBFD64A67AFFB9FE72F802FE7817BD" box="[413,448,1942,1966]" pageId="2" pageNumber="259" type="generic">1♂</specimenCount>
|
||
same locality,
|
||
<date id="2C53102DA67AFFB9FD82F801FD5517BD" box="[621,749,1941,1966]" pageId="2" pageNumber="259" value="2008-06-28">28.06.2008</date>
|
||
, leg. M. Hämäläinen [
|
||
<collectionCode id="3EFCAE28A67AFFB9FC03F802FBF117BE" LSID="urn:lsid:biocol.org:col:34992" box="[1004,1097,1942,1965]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="259">RMNH</collectionCode>
|
||
].
|
||
<specimenCount id="4EEBFD64A67AFFB9FBB4F802FBC717BD" box="[1115,1151,1942,1966]" pageId="2" pageNumber="259" type="generic">3♂</specimenCount>
|
||
“Rhododendron & Fivelakes trail” Bach Ma National Park, Thua Tien Hue Province, central
|
||
<collectingCountry id="20FA767DA67AFFB9FC72F82DFC4717C1" box="[925,1023,1977,2002]" name="Vietnam" pageId="2" pageNumber="259">Vietnam</collectingCountry>
|
||
, 15–
|
||
<date id="2C53102DA67AFFB9FBDEF82DFB0C17C1" box="[1073,1204,1977,2002]" pageId="2" pageNumber="259" value="2008-06-18">18.06.2008</date>
|
||
, leg. M. Hämäläinen [
|
||
<collectionCode id="3EFCAE28A67AFFB9FF72F84AFF4217E6" LSID="urn:lsid:biocol.org:col:34992" box="[157,250,2014,2037]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="259">RMNH</collectionCode>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="0C926665A67BFFB8FF78F98DFA25165D" httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="3" pageNumber="260" targetBox="[317,1270,200,1525]" targetPageId="3">
|
||
<paragraph id="585236EDA67BFFB8FF78F98DFA25165D" blockId="3.[151,1437,1561,1614]" pageId="3" pageNumber="260">
|
||
<emphasis id="6A99EAFFA67BFFB8FF78F98DFEAA163D" bold="true" box="[151,274,1561,1583]" pageId="3" pageNumber="260">FIGURE 2.</emphasis>
|
||
<taxonomicName id="9FED4D6EA67BFFB8FEF7F98DFE58163C" box="[280,480,1561,1583]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="scutellum">
|
||
<emphasis id="6A99EAFFA67BFFB8FEF7F98DFE58163C" box="[280,480,1561,1583]" italics="true" pageId="3" pageNumber="260">Coeliccia scutellum</emphasis>
|
||
</taxonomicName>
|
||
Lectotype.
|
||
<emphasis id="6A99EAFFA67BFFB8FDB7F98DFDD2163D" bold="true" box="[600,618,1561,1582]" pageId="3" pageNumber="260">A</emphasis>
|
||
head dorsolateral,
|
||
<emphasis id="6A99EAFFA67BFFB8FCC5F98EFC83163C" bold="true" box="[810,827,1562,1583]" pageId="3" pageNumber="260">B</emphasis>
|
||
prothorax and thorax dorsal,
|
||
<emphasis id="6A99EAFFA67BFFB8FB8BF98DFBCE163C" bold="true" box="[1124,1142,1561,1583]" pageId="3" pageNumber="260">C</emphasis>
|
||
prothorax and thorax lateral,
|
||
<emphasis id="6A99EAFFA67BFFB8FF78F9ACFF11165E" bold="true" box="[151,169,1592,1613]" pageId="3" pageNumber="260">D</emphasis>
|
||
anal appendages lateral,
|
||
<emphasis id="6A99EAFFA67BFFB8FE48F9ACFE00165E" bold="true" box="[423,440,1592,1613]" pageId="3" pageNumber="260">E</emphasis>
|
||
anal appendages dorsolateral,
|
||
<emphasis id="6A99EAFFA67BFFB8FD02F9ACFD44165E" bold="true" box="[749,764,1592,1613]" pageId="3" pageNumber="260">F</emphasis>
|
||
anal appendages dorsal,
|
||
<emphasis id="6A99EAFFA67BFFB8FC16F9ACFBB5165D" bold="true" box="[1017,1037,1592,1614]" pageId="3" pageNumber="260">G</emphasis>
|
||
left Fw,
|
||
<emphasis id="6A99EAFFA67BFFB8FB8AF9ACFBC1165E" bold="true" box="[1125,1145,1592,1613]" pageId="3" pageNumber="260">H</emphasis>
|
||
left Hw.
|
||
<emphasis id="6A99EAFFA67BFFB8FB39F9ACFAF0165D" box="[1238,1352,1592,1614]" italics="true" pageId="3" pageNumber="260">Scale bars:</emphasis>
|
||
500Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="10F76566A67BFFBEFF28F9E1FDA01296" lastPageId="5" lastPageNumber="262" pageId="3" pageNumber="260" type="description">
|
||
<paragraph id="585236EDA67BFFB8FF28F9E1FCDE16C5" blockId="3.[151,1437,1653,2038]" pageId="3" pageNumber="260">
|
||
<emphasis id="6A99EAFFA67BFFB8FF28F9E1FDA1169D" bold="true" box="[199,537,1653,1678]" pageId="3" pageNumber="260">
|
||
Description of the
|
||
<typeStatus id="8756884FA67BFFB8FE49F9E1FDAB169D" box="[422,531,1653,1678]" pageId="3" pageNumber="260" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Head (
|
||
<figureCitation id="C0D62A68A67BFFB8FD83F9E1FD08169D" box="[620,688,1653,1678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="3" pageNumber="260">Fig. 2</figureCitation>
|
||
A): Completely black except for diffuse greyish patches reaching out apically from the ocelli; typical postocular spots only just visible, the coloration faded away almost entirely. Prothorax (
|
||
<figureCitation id="C0D62A68A67BFFB8FEFBF929FEEF16C5" box="[276,343,1725,1750]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="3" pageNumber="260">Fig. 2</figureCitation>
|
||
B–C): All black, posterior lobe slightly raised.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67BFFB8FF28F975FE341799" blockId="3.[151,1437,1653,2038]" pageId="3" pageNumber="260">
|
||
Synthorax (
|
||
<figureCitation id="C0D62A68A67BFFB8FEA3F975FE2916E9" box="[332,401,1761,1786]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="3" pageNumber="260">Fig. 2</figureCitation>
|
||
B–C): Black dorsally, with two big yellow markings (
|
||
<figureCitation id="C0D62A68A67BFFB8FBE4F975FBEA16E9" box="[1035,1106,1761,1786]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="3" pageNumber="260">Fig. 2</figureCitation>
|
||
B). Synthorax laterally black with two big yellow markings, the lower one covering most of the metepimeron, the upper one covering the posterior part of the metepisternum up to the metathoracic spiracle, dorsally reaching slightly onto the mesepisternum. A small but distinct black spot near the upper margin of the metepisternal yellow marking; mesepisternum black.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67BFFB8FF28F801FCAE17E5" blockId="3.[151,1437,1653,2038]" pageId="3" pageNumber="260">
|
||
Wings (
|
||
<figureCitation id="C0D62A68A67BFFB8FEF2F801FEE717BD" box="[285,351,1941,1966]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="3" pageNumber="260">Fig. 2</figureCitation>
|
||
G–H): Hyaline with black Pt, covering nearly two cells in Fw and Hw. 21 Px in Fw, 20 Px in Hw.
|
||
<collectingCountry id="20FA767DA67BFFB8FF78F82EFF0217C1" box="[151,186,1978,2002]" name="Poland" pageId="3" pageNumber="260">RP</collectingCountry>
|
||
2 arising slightly proximal to Px
|
||
<quantity id="9F159B08A67BFFB8FDC7F82EFDD917C1" box="[552,609,1977,2002]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="260" unit="in" value="10.0">10 in</quantity>
|
||
Fw, between Px 7 and Px
|
||
<quantity id="9F159B08A67BFFB8FC67F82EFC0A17C1" box="[904,946,1977,2002]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="3" pageNumber="260" unit="in" value="8.0">8 in</quantity>
|
||
Hw. IR1 arises at Px
|
||
<quantity id="9F159B08A67BFFB8FB4BF82EFB6517C1" box="[1188,1245,1977,2002]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="3" pageNumber="260" unit="in" value="13.0">13 in</quantity>
|
||
Fw, and at Px
|
||
<quantity id="9F159B08A67BFFB8FA6EF82EFF1517E5" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="3" pageNumber="260" unit="in" value="11.0">11 in</quantity>
|
||
Hw. IR2 arises slightly distal to subnodus in all wings.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67CFFBFFF28FF03FCD610A3" blockId="4.[151,1437,151,645]" box="[199,878,151,176]" pageId="4" pageNumber="261">Abdomen: Slender, black with whitish lateral spots on S3–6.</paragraph>
|
||
<paragraph id="585236EDA67CFFBFFF28FF28FD9C1153" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="261">
|
||
Anal appendages (
|
||
<figureCitation id="C0D62A68A67CFFBFFE71FF28FE5C10C7" box="[414,484,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="4" pageNumber="261">Fig. 2</figureCitation>
|
||
D–F): Paraprocts slightly overtopping cerci in length. Cerci with small, medially directed spine on upper inner border; followed dorsoapically by a rounded elevation, and with distinct ventromedial tooth arising subapically. This tooth is difficult to see in the
|
||
<typeStatus id="8756884FA67CFFBFFC33FE90FBFD110E" box="[988,1093,260,285]" pageId="4" pageNumber="261" type="lectotype">lectotype</typeStatus>
|
||
, as the cerci are turned so that the tooth is rather directed inwards.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67CFFBFFF28FED8FBAE11BE" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="261">
|
||
Genital ligula (
|
||
<figureCitation id="C0D62A68A67CFFBFFE99FED8FE041176" box="[374,444,332,357]" captionStart="FIGURE 5" captionStartId="7.[151,250,1111,1133]" captionTargetBox="[201,1380,199,1085]" captionTargetId="figure@7.[198,1388,193,1090]" captionTargetPageId="7" captionText="FIGURE 5. Genital ligulae as reconstructed from micro-CT (orientation bars: dl dorsolateral, l lateral, p posterior, v ventral, vl ventrolateral). A – B Genital ligula of C. scutellum lectotype C – D genital ligula of C. scutellum from Tam Đ ảo, Vietnam E – F genital ligula of C. hainanense lectotype." httpUri="https://zenodo.org/record/253817/files/figure.png" pageId="4" pageNumber="261">Fig. 5</figureCitation>
|
||
A–B, Supplementary
|
||
<figureCitation id="C0D62A68A67CFFBFFD2FFED8FCBD1177" box="[704,773,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1675,1697]" captionTargetBox="[317,1269,311,1658]" captionTargetId="figure@2.[317,1270,311,1658]" captionTargetPageId="2" captionText="FIGURE 1. Mounting methods and detail of genital ligula. A Pinned specimen (Coeliccia scutellum) mounted for micro-CT scan, B papered specimen (Coeliccia scutellum) mounted for micro-CT scan, C macro-photograph (Zeiss MCr Camera) of C. scutellum genital ligula, showing thinness of flaps. Scale bar: 250 Μm." httpUri="https://zenodo.org/record/253813/files/figure.png" pageId="4" pageNumber="261">Fig. 1</figureCitation>
|
||
): First segment simple with distal bend; second segment curved with well-developed inner fold, without terminal fold; third segment flat, median part distinctly raised, with two recurved flagella arising medially from raised part, bordering first segment.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67CFFBFFF28FE2DFB9611C3" blockId="4.[151,1437,151,645]" box="[199,1070,439,464]" pageId="4" pageNumber="261">
|
||
<emphasis id="6A99EAFFA67CFFBFFF28FE2DFE0D11DC" box="[199,437,440,464]" italics="true" pageId="4" pageNumber="261">Measurements (mm):</emphasis>
|
||
<typeStatus id="8756884FA67CFFBFFE53FE2CFD8811C3" box="[444,560,440,464]" pageId="4" pageNumber="261" type="lectotype">Lectotype</typeStatus>
|
||
: Total length: 49, Abd.: 42, Fw/Hw (left): 21.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67CFFBFFF28FE49FDCC120B" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="261">
|
||
Measurement ranges of the six studied specimen collected near the
|
||
<typeStatus id="8756884FA67CFFBFFC3FFE49FBB811E6" box="[976,1024,477,501]" pageId="4" pageNumber="261">type</typeStatus>
|
||
locality: Total length: 55–61, Abd.: 47–52, Fw (left): 29–34, Hw (left): 29–34.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67CFFBFFF28FDB0FEE91296" blockId="4.[151,1437,151,645]" pageId="4" pageNumber="261">
|
||
<emphasis id="6A99EAFFA67CFFBFFF28FDB0FE83122E" bold="true" box="[199,315,548,573]" pageId="4" pageNumber="261">Variation.</emphasis>
|
||
Head (
|
||
<figureCitation id="C0D62A68A67CFFBFFE7AFDB0FE62122F" box="[405,474,548,573]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="4" pageNumber="261">Fig. 3</figureCitation>
|
||
A): Specimens from near the
|
||
<typeStatus id="8756884FA67CFFBFFCD3FDB1FCD4122E" box="[828,876,549,573]" pageId="4" pageNumber="261">type</typeStatus>
|
||
locality show more extensive whitish coloration reaching out from the lateral ocelli towards the antennal bases, which have a whitish base; postocular spots distinct, yellowish white.
|
||
</paragraph>
|
||
<caption id="0C926665A67CFFBFFF78F8EFFBB817A3" httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="4" pageNumber="261" targetBox="[319,1273,697,1878]" targetPageId="4">
|
||
<paragraph id="585236EDA67CFFBFFF78F8EFFBB817A3" blockId="4.[151,1436,1915,1968]" pageId="4" pageNumber="261">
|
||
<emphasis id="6A99EAFFA67CFFBFFF78F8EFFEAA1783" bold="true" box="[151,274,1915,1937]" pageId="4" pageNumber="261">FIGURE 3.</emphasis>
|
||
<taxonomicName id="9FED4D6EA67CFFBFFEF6F8EFFD961782" authority="Tam D" authorityName="Tam D" box="[281,558,1915,1937]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="scutellum">
|
||
<emphasis id="6A99EAFFA67CFFBFFEF6F8EFFE5A1782" box="[281,482,1915,1937]" italics="true" pageId="4" pageNumber="261">Coeliccia scutellum</emphasis>
|
||
Tam Đ
|
||
</taxonomicName>
|
||
ảo, Vietnam.
|
||
<emphasis id="6A99EAFFA67CFFBFFD5BF8EFFD7E1783" bold="true" box="[692,710,1915,1936]" pageId="4" pageNumber="261">A</emphasis>
|
||
head dorsolateral,
|
||
<emphasis id="6A99EAFFA67CFFBFFC67F8EFFC211783" bold="true" box="[904,921,1915,1936]" pageId="4" pageNumber="261">B</emphasis>
|
||
thorax dorsal,
|
||
<emphasis id="6A99EAFFA67CFFBFFBDEF8EFFBFB1782" bold="true" box="[1073,1091,1915,1937]" pageId="4" pageNumber="261">C</emphasis>
|
||
thorax lateral,
|
||
<emphasis id="6A99EAFFA67CFFBFFB33F8EFFB561783" bold="true" box="[1244,1262,1915,1936]" pageId="4" pageNumber="261">D</emphasis>
|
||
anal appendages dorsal,
|
||
<emphasis id="6A99EAFFA67CFFBFFF0EF80EFF4A17BC" bold="true" box="[225,242,1946,1967]" pageId="4" pageNumber="261">E</emphasis>
|
||
anal appendages lateral,
|
||
<emphasis id="6A99EAFFA67CFFBFFE1EF80EFDB817BC" bold="true" box="[497,512,1946,1967]" pageId="4" pageNumber="261">F</emphasis>
|
||
anal appendages dorsolateral.
|
||
<emphasis id="6A99EAFFA67CFFBFFCD7F80EFC1317A3" box="[824,939,1946,1968]" italics="true" pageId="4" pageNumber="261">Scale bars:</emphasis>
|
||
500Μm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="585236EDA67DFFBEFF28FF03FA391177" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="262">
|
||
Synthorax (
|
||
<figureCitation id="C0D62A68A67DFFBEFEA5FF03FE3410A3" box="[330,396,151,176]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="5" pageNumber="262">Fig. 3</figureCitation>
|
||
B–C): Lateral yellow markings of the specimens from near the
|
||
<typeStatus id="8756884FA67DFFBEFBB3FF0CFB3410A3" box="[1116,1164,152,176]" pageId="5" pageNumber="262">type</typeStatus>
|
||
locality agree well with the
|
||
<typeStatus id="8756884FA67DFFBEFF29FF28FE8810C6" box="[198,304,188,213]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
, but there is great variation in the extent of the two dorsal, escutcheon-shaped spots. In some specimens they cover about one third of the dorsal part of the synthorax as in the
|
||
<typeStatus id="8756884FA67DFFBEFBC5FF4BFB2C10EB" box="[1066,1172,223,248]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
; in other specimens the spots cover up to two thirds of the dorsal synthorax (
|
||
<figureCitation id="C0D62A68A67DFFBEFD08FE90FC91110F" box="[743,809,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="5" pageNumber="262">Fig. 3</figureCitation>
|
||
B–C). The coloration, described by
|
||
<bibRefCitation id="3C7C4B1CA67DFFBEFB2CFE90FA24110F" author="Laidlaw" box="[1219,1436,260,285]" pageId="5" pageNumber="262" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw (1932: 22)</bibRefCitation>
|
||
as “…citron or sulphur-yellow…” also varies in the specimen from the
|
||
<typeStatus id="8756884FA67DFFBEFC25FEBCFC421153" box="[970,1018,296,320]" pageId="5" pageNumber="262">type</typeStatus>
|
||
locality; some specimen have citron markings as in the
|
||
<typeStatus id="8756884FA67DFFBEFE85FED8FE6C1176" box="[362,468,332,357]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
, while in others the markings are of a rather deep golden yellow color (
|
||
<figureCitation id="C0D62A68A67DFFBEFB1CFED8FA8E1177" box="[1267,1334,332,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="5" pageNumber="262">Fig. 3</figureCitation>
|
||
B–C).
|
||
</paragraph>
|
||
<paragraph id="585236EDA67DFFBEFF28FEFBFB74120B" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="262">
|
||
Anal appendages (
|
||
<figureCitation id="C0D62A68A67DFFBEFE79FEFBFE6F119B" box="[406,471,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="5" pageNumber="262">Fig. 3</figureCitation>
|
||
D–F): In some of the other specimens examined, the anal appendages dried in another position than in the
|
||
<typeStatus id="8756884FA67DFFBEFE94FE00FE5D11BE" box="[379,485,404,429]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
, so that the ventromedial tooth is directed straight downward and the small spine is directed rather obliquely inward (
|
||
<figureCitation id="C0D62A68A67DFFBEFDF3FE23FDD911C3" box="[540,609,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="5" pageNumber="262">Fig. 3</figureCitation>
|
||
D–F).
|
||
<bibRefCitation id="3C7C4B1CA67DFFBEFD57FE23FC2F11C3" author="Asahina" box="[696,919,439,464]" pageId="5" pageNumber="262" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina (1997: 23)</bibRefCitation>
|
||
did not describe the anal appendages for
|
||
<taxonomicName id="9FED4D6EA67DFFBEFA6DFE2CFEBF11E7" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="tomokunii">
|
||
<emphasis id="6A99EAFFA67DFFBEFA6DFE2CFEBF11E7" italics="true" pageId="5" pageNumber="262">C. tomokunii</emphasis>
|
||
</taxonomicName>
|
||
but mentioned that the superior appendages are “provided with inflated head…”. Depending on the position of these appendages, this impression may arise from the rounded dorsoapical elevation.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67DFFBEFF28FDB0FDA01296" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="262">
|
||
Genital ligula (
|
||
<figureCitation id="C0D62A68A67DFFBEFE9DFDB0FE0E122E" box="[370,438,548,573]" captionStart="FIGURE 5" captionStartId="7.[151,250,1111,1133]" captionTargetBox="[201,1380,199,1085]" captionTargetId="figure@7.[198,1388,193,1090]" captionTargetPageId="7" captionText="FIGURE 5. Genital ligulae as reconstructed from micro-CT (orientation bars: dl dorsolateral, l lateral, p posterior, v ventral, vl ventrolateral). A – B Genital ligula of C. scutellum lectotype C – D genital ligula of C. scutellum from Tam Đ ảo, Vietnam E – F genital ligula of C. hainanense lectotype." httpUri="https://zenodo.org/record/253817/files/figure.png" pageId="5" pageNumber="262">Fig. 5</figureCitation>
|
||
C–D, Supplementary
|
||
<figureCitation id="C0D62A68A67DFFBEFD5AFDB0FD43122F" box="[693,763,548,573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1561,1583]" captionTargetBox="[317,1270,200,1525]" captionTargetId="figure@3.[317,1270,193,1540]" captionTargetPageId="3" captionText="FIGURE 2. Coeliccia scutellum Lectotype. A head dorsolateral, B prothorax and thorax dorsal, C prothorax and thorax lateral, D anal appendages lateral, E anal appendages dorsolateral, F anal appendages dorsal, G left Fw, H left Hw. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253814/files/figure.png" pageId="5" pageNumber="262">Fig. 2</figureCitation>
|
||
): All specimens from near the
|
||
<typeStatus id="8756884FA67DFFBEFBB2FDB1FB35122E" box="[1117,1165,549,573]" pageId="5" pageNumber="262">type</typeStatus>
|
||
locality with two small thin flaps arising laterally from raised median part of segment three, posterior of origin of flagella (
|
||
<figureCitation id="C0D62A68A67DFFBEFB16FDD3FA841273" box="[1273,1340,583,608]" captionStart="FIGURE 1" captionStartId="2.[151,250,1675,1697]" captionTargetBox="[317,1269,311,1658]" captionTargetId="figure@2.[317,1270,311,1658]" captionTargetPageId="2" captionText="FIGURE 1. Mounting methods and detail of genital ligula. A Pinned specimen (Coeliccia scutellum) mounted for micro-CT scan, B papered specimen (Coeliccia scutellum) mounted for micro-CT scan, C macro-photograph (Zeiss MCr Camera) of C. scutellum genital ligula, showing thinness of flaps. Scale bar: 250 Μm." httpUri="https://zenodo.org/record/253813/files/figure.png" pageId="5" pageNumber="262">Fig. 1</figureCitation>
|
||
C, 5 C– D). Flaps missing in the
|
||
<typeStatus id="8756884FA67DFFBEFE45FDF8FDAC1296" box="[426,532,620,645]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="10F76566A67DFFBEFF28FD1BFB9914B2" pageId="5" pageNumber="262" type="discussion">
|
||
<paragraph id="585236EDA67DFFBEFF28FD1BFEA81403" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="262">
|
||
<emphasis id="6A99EAFFA67DFFBEFF28FD1BFE8312BB" bold="true" box="[199,315,655,680]" pageId="5" pageNumber="262">Remarks.</emphasis>
|
||
The specimens of
|
||
<taxonomicName id="9FED4D6EA67DFFBEFDFAFD04FD4D12BB" box="[533,757,656,680]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="scutellum">
|
||
<emphasis id="6A99EAFFA67DFFBEFDFAFD04FD4D12BB" box="[533,757,656,680]" italics="true" pageId="5" pageNumber="262">Coeliccia scutellum</emphasis>
|
||
</taxonomicName>
|
||
studied show rather strong variation in coloration, size and shape of dorsal markings. In order to visualize this variability, we illustrate a specimen that is most different from the
|
||
<typeStatus id="8756884FA67DFFBEFF2DFD43FE9112E3" box="[194,297,727,752]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
in all mentioned characters (
|
||
<figureCitation id="C0D62A68A67DFFBEFD97FD43FD0612E3" box="[632,702,727,752]" captionStart="FIGURE 3" captionStartId="4.[151,250,1915,1937]" captionTargetBox="[319,1273,697,1878]" captionTargetId="figure@4.[313,1274,697,1894]" captionTargetPageId="4" captionText="FIGURE 3. Coeliccia scutellum Tam Đ ảo, Vietnam. A head dorsolateral, B thorax dorsal, C thorax lateral, D anal appendages dorsal, E anal appendages lateral, F anal appendages dorsolateral. Scale bars: 500 Μm." httpUri="https://zenodo.org/record/253815/files/figure.png" pageId="5" pageNumber="262">Fig. 3</figureCitation>
|
||
). The genital ligula of all examined specimens, differs from the genital ligula of the
|
||
<typeStatus id="8756884FA67DFFBEFE94FD68FE591306" box="[379,481,764,789]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
in having two thin lateral flaps on segment three (
|
||
<figureCitation id="C0D62A68A67DFFBEFBF5FD68FBE51307" box="[1050,1117,764,789]" captionStart="FIGURE 1" captionStartId="2.[151,250,1675,1697]" captionTargetBox="[317,1269,311,1658]" captionTargetId="figure@2.[317,1270,311,1658]" captionTargetPageId="2" captionText="FIGURE 1. Mounting methods and detail of genital ligula. A Pinned specimen (Coeliccia scutellum) mounted for micro-CT scan, B papered specimen (Coeliccia scutellum) mounted for micro-CT scan, C macro-photograph (Zeiss MCr Camera) of C. scutellum genital ligula, showing thinness of flaps. Scale bar: 250 Μm." httpUri="https://zenodo.org/record/253813/files/figure.png" pageId="5" pageNumber="262">Fig. 1</figureCitation>
|
||
C, 5 C–D). The
|
||
<typeStatus id="8756884FA67DFFBEFAF4FD68FA3A1306" box="[1307,1410,764,789]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
is the only specimen without flaps on the genital ligula, although the shape of the ligula is otherwise consistent with that found in the other specimens. Since there are no other structural differences between the
|
||
<typeStatus id="8756884FA67DFFBEFB38FCD0FA86134E" box="[1239,1342,836,861]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
and the other examined specimens, it seems possible that the flaps have eroded in the
|
||
<typeStatus id="8756884FA67DFFBEFC18FCF3FBD91393" box="[1015,1121,871,896]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
. Alternatively, within the 70 years that passed between the collection of the
|
||
<typeStatus id="8756884FA67DFFBEFD5AFC18FCA413B6" box="[693,796,908,933]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
and the collection of the other specimens examined the genital ligula might have changed due to sexual selection or genetic drift. It seems also possible that there is variation in the shape of the genital ligula, with the
|
||
<typeStatus id="8756884FA67DFFBEFD1CFC40FCE213FE" box="[755,858,980,1005]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
without flaps representing the extreme end of the variability.
|
||
</paragraph>
|
||
<paragraph id="585236EDA67DFFBEFF28FB88FB9914B2" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="262">
|
||
<collectingCountry id="20FA767DA67DFFBEFF28FB88FF5E1427" box="[199,230,1052,1076]" name="American Samoa" pageId="5" pageNumber="262">As</collectingCountry>
|
||
the somatic and genital variability is strong between
|
||
<typeStatus id="8756884FA67DFFBEFC87FB88FC761426" box="[872,974,1052,1077]" pageId="5" pageNumber="262" type="lectotype">lectotype</typeStatus>
|
||
and all other examined specimens, we suggest that
|
||
<taxonomicName id="9FED4D6EA67DFFBEFEC5FBD4FE06144B" box="[298,446,1088,1112]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="tomokunii">
|
||
<emphasis id="6A99EAFFA67DFFBEFEC5FBD4FE06144B" box="[298,446,1088,1112]" italics="true" pageId="5" pageNumber="262">C. tomokunii</emphasis>
|
||
</taxonomicName>
|
||
is synonymous with
|
||
<taxonomicName id="9FED4D6EA67DFFBEFD55FBD4FCF1144B" box="[698,841,1088,1112]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="scutellum">
|
||
<emphasis id="6A99EAFFA67DFFBEFD55FBD4FCF1144B" box="[698,841,1088,1112]" italics="true" pageId="5" pageNumber="262">C. scutellum</emphasis>
|
||
</taxonomicName>
|
||
. Investigating the variability in a long series of
|
||
<taxonomicName id="9FED4D6EA67DFFBEFA6DFBD4FEBA146F" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="scutellum">
|
||
<emphasis id="6A99EAFFA67DFFBEFA6DFBD4FEBA146F" italics="true" pageId="5" pageNumber="262">C. scutellum</emphasis>
|
||
</taxonomicName>
|
||
, in particular the assessment of whether there is continuous variation in flap size or bimodal distribution, would help to clarify the taxonomic status. We propose the following synonymy:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="10F76566A67DFFBEFF78FB44FCB9151E" pageId="5" pageNumber="262" type="reference_group">
|
||
<paragraph id="585236EDA67DFFBEFF78FB44FDA314FB" blockId="5.[151,769,1231,1293]" box="[151,539,1231,1256]" pageId="5" pageNumber="262">
|
||
<taxonomicName id="9FED4D6EA67DFFBEFF78FB44FDA314FB" ID-CoL="WQDD" authority="Laidlaw, 1932" authorityName="Laidlaw" authorityYear="1932" box="[151,539,1231,1256]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="scutellum">
|
||
<emphasis id="6A99EAFFA67DFFBEFF78FB44FECD14FB" box="[151,373,1232,1256]" italics="true" pageId="5" pageNumber="262">Coeliccia scutellum</emphasis>
|
||
<bibRefCitation id="3C7C4B1CA67DFFBEFE93FB5BFDA314FB" author="Laidlaw" box="[380,539,1231,1256]" pageId="5" pageNumber="262" refString="Laidlaw, F. F. (1932) A revision of the genus Coeliccia (Order Odonata). Records of the Indian Museum, 34, 7 - 42, 3 pls." type="journal article" year="1932">Laidlaw, 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="585236EDA67DFFBEFF28FB60FCB9151E" blockId="5.[151,769,1231,1293]" box="[199,769,1268,1293]" pageId="5" pageNumber="262">
|
||
Syn.
|
||
<taxonomicName id="9FED4D6EA67DFFBEFEEFFB60FD36151F" ID-CoL="WQDK" authority="Asahina, 1997" authorityName="Asahina" authorityYear="1997" box="[256,654,1268,1293]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="tomokunii">
|
||
<emphasis id="6A99EAFFA67DFFBEFEEFFB60FE5B151F" box="[256,483,1268,1292]" italics="true" pageId="5" pageNumber="262">Coeliccia tomokunii</emphasis>
|
||
<bibRefCitation id="3C7C4B1CA67DFFBEFE05FB60FD36151F" author="Asahina" box="[490,654,1268,1293]" pageId="5" pageNumber="262" refString="Asahina, S. (1997) Records of the northernVietnamese Odonata taken by the expedition members from the National Science Museum, Tokyo, 5: Coenagrionidae, Protoneuridae, Platycnemididae. Bulletin of the National Science Museum, Tokyo (A), 23, 17 - 34." type="journal article" year="1997">Asahina, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="71AA5784A67DFFBEFD76FB60FCB9151E" box="[665,769,1268,1293]" pageId="5" pageNumber="262">
|
||
<emphasis id="6A99EAFFA67DFFBEFD76FB60FCB9151E" bold="true" box="[665,769,1268,1293]" pageId="5" pageNumber="262">syn. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |