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<document id="8E3448CF1A52F8E3D4B6A5E59C9CD806" ID-DOI="10.11646/zootaxa.4092.1.7" ID-GBIF-Dataset="fe10fcea-0b20-49d9-969d-44072b9a2e50" ID-ISSN="1175-5326" ID-Zenodo-Dep="256045" ID-ZooBank="5F6A20FB-CE78-4F9D-9777-02BD773B3924" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458037658074" checkinUser="plazi" docAuthor="Hodgson, Chris J., Gunawardana, D. N. &amp; Richmond, J. E." docDate="2016" docId="C918B769FFA1FFB8FF3C6158FE007063" docLanguage="en" docName="15620-5145-1-PB.pdf.imf" docOrigin="Zootaxa 4092 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kalasiris martini Hodgson &amp; Richmond, spec. n." docType="treatment" docUuid="41D7CF0C-1BB9-46E6-908E-4E95C111EABB" docUuidSource="ZooBank" docVersion="11" lastPageNumber="127" masterDocId="3521CF11FFA0FFBDFFAB622EFFF7751B" masterDocTitle="A new species in the indigenous New Zealand soft scale insect genus Kalasiris Henderson &amp; Hodgson (Hemiptera: Sternorrhyncha: Coccomorpha: Coccidae) on Gahnia setifolia (Cyperaceae)" masterLastPageNumber="128" masterPageNumber="122" pageNumber="123" updateTime="1698184572017" updateUser="plazi">
<mods:mods id="B1896054BCC3E38C76BD7A81F804757E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1B0855B50B1C98FFE622D48E50E0C6CD">
<mods:title id="6DD4723DD1735CD0C265216A781A8D0A">A new species in the indigenous New Zealand soft scale insect genus Kalasiris Henderson &amp; Hodgson (Hemiptera: Sternorrhyncha: Coccomorpha: Coccidae) on Gahnia setifolia (Cyperaceae)</mods:title>
</mods:titleInfo>
<mods:name id="0962941A40192922593FAF9FD921F2DE" type="personal">
<mods:role id="D0A84BE3AC7DB9BBD62F2132A4C3308B">
<mods:roleTerm id="7259A2ED2DCDD5D16B34EC7D90EDB4DA">Author</mods:roleTerm>
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<mods:namePart id="5FDC50FDCD62FE76A00C494FFE0C34ED">Hodgson, Chris J.</mods:namePart>
</mods:name>
<mods:name id="F9C5A82A1E7CC1C5A25B2C39E2288339" type="personal">
<mods:role id="B9FCC60653E31529867D7E4210EA142F">
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<mods:namePart id="9BEFA3FEEA5E2D942DF9FED0681991A5">Gunawardana, D. N.</mods:namePart>
</mods:name>
<mods:name id="C62140CBC98CD7073CB14716C7E6FCFE" type="personal">
<mods:role id="E2771D6C4223798EA367AAC0BA37254F">
<mods:roleTerm id="33B368AC3CC7A59AFE1D1772B5E3D808">Author</mods:roleTerm>
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<mods:namePart id="247EADA786B3476B58AAB41E0A0B3F67">Richmond, J. E.</mods:namePart>
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<mods:typeOfResource id="FCE154C1B47AC3018590E1E62E16FA01">text</mods:typeOfResource>
<mods:relatedItem id="C33FCB666D00BB3EA04763FCAE05B501" type="host">
<mods:titleInfo id="241A93FD989D82D9B20072E2C3904FEC">
<mods:title id="8D3D9789C394A8099517DBC173EEE6DC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E232F2D4FE448607C0A9F0D1793AFE7C">
<mods:date id="510F2137A8B949C99B11FBF455D2734F">2016</mods:date>
<mods:detail id="D7F9D750E29C619A4F411906AF33BBEF" type="volume">
<mods:number id="FD011831CB89E16BE62BD98C7517845F">4092</mods:number>
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<mods:detail id="13220FEA5B5A000D92C4D7B068E8A69B" type="issue">
<mods:number id="288A066C5CF773A1B4486AC950CD0AF8">1</mods:number>
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<mods:classification id="EF9C4827F6B422B8EBFB0131DA5CC636">journal article</mods:classification>
<mods:identifier id="A7D5AB2B4ED9D894DC1D828D07741D69" type="DOI">10.11646/zootaxa.4092.1.7</mods:identifier>
<mods:identifier id="AA9A12E4A6DF8DE41B9969F8643EE01C" type="GBIF-Dataset">fe10fcea-0b20-49d9-969d-44072b9a2e50</mods:identifier>
<mods:identifier id="EDF58493D17C3E501809213A9465BA1C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6BCFF9759723922503BB3BE63DAABC79" type="Zenodo-Dep">256045</mods:identifier>
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<treatment id="C918B769FFA1FFB8FF3C6158FE007063" ID-DOI="http://doi.org/10.5281/zenodo.6081482" ID-GBIF-Taxon="117912021" ID-Zenodo-Dep="6081482" LSID="urn:lsid:zoobank.org:act:41D7CF0C-1BB9-46E6-908E-4E95C111EABB" httpUri="http://treatment.plazi.org/id/C918B769FFA1FFB8FF3C6158FE007063" lastPageId="5" lastPageNumber="127" pageId="1" pageNumber="123">
<subSubSection id="09AB55F4FFA1FFBCFF3C6158FBF17059" pageId="1" pageNumber="123" type="nomenclature">
<paragraph id="410E067FFFA1FFBCFF3C6158FD1A768B" blockId="1.[151,749,886,946]" box="[151,749,886,912]" pageId="1" pageNumber="123">
<heading id="1A46B113FFA1FFBCFF3C6158FD1A768B" bold="true" box="[151,749,886,912]" fontSize="11" level="1" pageId="1" pageNumber="123" reason="1">
<emphasis id="73C5DA6DFFA1FFBCFF3C6158FD1A768B" bold="true" box="[151,749,886,912]" pageId="1" pageNumber="123">
<taxonomicName id="86B17DFCFFA1FFBCFF3C6158FD73768B" authority="Hodgson &amp; Richmond" authorityName="Hodgson &amp; Richmond" box="[151,644,886,912]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="123" phylum="Arthropoda" rank="species" species="martini" status="sp. nov.">
<emphasis id="73C5DA6DFFA1FFBCFF3C6158FE93768B" bold="true" box="[151,356,886,912]" italics="true" pageId="1" pageNumber="123">Kalasiris martini</emphasis>
Hodgson &amp; Richmond
</taxonomicName>
,
<taxonomicNameLabel id="68F66716FFA1FFBCFD3A6158FD1A768B" box="[657,749,886,912]" pageId="1" pageNumber="123" rank="species">spec. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="410E067FFFA1FFBCFF3C61B7FED076A9" blockId="1.[151,749,886,946]" box="[151,295,921,946]" pageId="1" pageNumber="123">
(
<figureCitation id="D98A1AFAFFA1FFBCFF3461B7FF1276A9" box="[159,229,921,946]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1964]" captionTargetBox="[374,1215,223,1915]" captionTargetId="figure@2.[347,1240,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of Kalasiris martini Hodgson &amp; Richmond, spec. n. 1 a: adult female, showing waxy test and reticulation plates; 1 b: an old female with body showing dark at anterior end and remainder of test filled with eggs; 1 c: test of second-instar male showing arrangement of reticulation plates." httpUri="https://zenodo.org/record/256046/files/figure.png" pageId="1" pageNumber="123">Figs 1</figureCitation>
&amp;
<figureCitation id="D98A1AFAFFA1FFBCFEA261B7FEEE76A9" box="[265,281,921,946]" captionStart="FIGURE 2" captionStartId="3.[151,250,2009,2030]" captionTargetBox="[188,1377,283,1870]" captionTargetId="figure@3.[185,1384,283,1872]" captionTargetPageId="3" captionText="FIGURE 2. Adult female Kalasiris martini Hodgson &amp; Richmond, spec. n." httpUri="https://zenodo.org/record/256047/files/figure.png" pageId="1" pageNumber="123">2</figureCitation>
).
</paragraph>
<paragraph id="410E067FFFA1FFBCFF3C61F2FD247180" blockId="1.[151,1437,988,1979]" pageId="1" pageNumber="123">
Material examined.
<typeStatus id="9E0AB8DDFFA1FFBCFE2061F2FE0F76EE" box="[395,504,988,1013]" pageId="1" pageNumber="123" type="holotype">Holotype</typeStatus>
and
<typeStatus id="9E0AB8DDFFA1FFBCFD9D61F3FD5576EF" box="[566,674,989,1012]" pageId="1" pageNumber="123" type="paratype">paratype</typeStatus>
adult females:
<collectingCountry id="39A646EFFFA1FFBCFCF761F3FBDF76EF" box="[860,1064,988,1012]" name="New Zealand" pageId="1" pageNumber="123">NEW ZEALAND</collectingCountry>
(AK), Avondale South Domain; Blockhouse Bay, on
<taxonomicName id="86B17DFCFFA1FFBCFE2861D3FD2E710D" box="[387,729,1021,1046]" class="Liliopsida" family="Cyperaceae" genus="Gahnia" kingdom="Plantae" order="Poales" pageId="1" pageNumber="123" phylum="Tracheophyta" rank="species" species="setifolia">
<emphasis id="73C5DA6DFFA1FFBCFE2861D3FDCE710D" box="[387,569,1021,1046]" italics="true" pageId="1" pageNumber="123">Gahnia setifolia</emphasis>
(Cyperaceae)
</taxonomicName>
,
<date id="350F20BFFFA1FFBCFD4D61D3FC96710D" box="[742,865,1021,1046]" pageId="1" pageNumber="123" value="2014-09-15">15.ix.2014</date>
, N.A. Martin (
<collectingCountry id="39A646EFFFA1FFBCFBA261D3FB55710D" box="[1033,1186,1021,1046]" name="New Zealand" pageId="1" pageNumber="123">New Zealand</collectingCountry>
Arthropod Collection (NZAC)): one slide with two specimens in good condition,
<typeStatus id="9E0AB8DDFFA1FFBCFC9E6630FC60712C" box="[821,919,1054,1079]" pageId="1" pageNumber="123" type="holotype">holotype</typeStatus>
specimen clearly marked near center of slide. Also
<typeStatus id="9E0AB8DDFFA1FFBCFF7F666EFEC8714C" box="[212,319,1088,1111]" pageId="1" pageNumber="123" type="paratype">paratype</typeStatus>
adult females: as previous but dated
<date id="350F20BFFFA1FFBCFD456611FC927143" box="[750,869,1087,1112]" pageId="1" pageNumber="123" value="2014-09-16">16.ix.2014</date>
(NZAC): two slides each with a single specimen (fair-good) and one slide with a pharate pupa. [Note that Avondale South Domain has recently been renamed Gittos Domain and is also known as Sandy Bay Reserve].
</paragraph>
<paragraph id="410E067FFFA1FFBCFF6C668BFCDE71A6" blockId="1.[151,1437,988,1979]" box="[199,809,1188,1213]" pageId="1" pageNumber="123">Note. Described from all four adult female specimens.</paragraph>
<paragraph id="410E067FFFA1FFBCFF6C66EBFBF17059" blockId="1.[151,1437,988,1979]" pageId="1" pageNumber="123">
Adult female.
<emphasis id="73C5DA6DFFA1FFBCFEDD66EBFD9571C5" box="[374,610,1221,1246]" italics="true" pageId="1" pageNumber="123">Unmounted material</emphasis>
. Fresh specimens greenish yellow, covered in a transparent glassy test, with 29 triangular waxy extensions around margin (
<figureCitation id="D98A1AFAFFA1FFBCFD5F66C8FCCF71E4" box="[756,824,1254,1279]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1964]" captionTargetBox="[374,1215,223,1915]" captionTargetId="figure@2.[347,1240,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of Kalasiris martini Hodgson &amp; Richmond, spec. n. 1 a: adult female, showing waxy test and reticulation plates; 1 b: an old female with body showing dark at anterior end and remainder of test filled with eggs; 1 c: test of second-instar male showing arrangement of reticulation plates." httpUri="https://zenodo.org/record/256046/files/figure.png" pageId="1" pageNumber="123">Fig. 1</figureCitation>
), forming a ledge above a short vertical side. Older specimens with post-reproductive adult crumpled up at anterior end, appearing dark through test (
<figureCitation id="D98A1AFAFFA1FFBCFA9F6729FA8E703B" box="[1332,1401,1287,1312]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1964]" captionTargetBox="[374,1215,223,1915]" captionTargetId="figure@2.[347,1240,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of Kalasiris martini Hodgson &amp; Richmond, spec. n. 1 a: adult female, showing waxy test and reticulation plates; 1 b: an old female with body showing dark at anterior end and remainder of test filled with eggs; 1 c: test of second-instar male showing arrangement of reticulation plates." httpUri="https://zenodo.org/record/256046/files/figure.png" pageId="1" pageNumber="123">Fig. 1</figureCitation>
b); remainder of test full of eggs. Young specimens in alcohol pale to dark brown.
</paragraph>
</subSubSection>
<subSubSection id="09AB55F4FFA1FFBCFF6C6764FF1C72A1" pageId="1" pageNumber="123" type="materials_examined">
<paragraph id="410E067FFFA1FFBCFF6C6764FE1A709E" blockId="1.[151,1437,988,1979]" pageId="1" pageNumber="123">
<emphasis id="73C5DA6DFFA1FFBCFF6C6764FE667078" box="[199,401,1354,1379]" italics="true" pageId="1" pageNumber="123">Mounted material</emphasis>
. Body elongate oval; length
<quantity id="8649AB9AFFA1FFBCFD7C6765FC957079" box="[727,866,1355,1379]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="7.9" metricValueMin="3.7" pageId="1" pageNumber="123" unit="mm" value="5.8" valueMax="7.9" valueMin="3.7">3.77.9 mm</quantity>
; breadth
<quantity id="8649AB9AFFA1FFBCFC626765FB957079" box="[969,1122,1355,1379]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="2.25" metricValueMin="1.1" pageId="1" pageNumber="123" unit="mm" value="1.675" valueMax="2.25" valueMin="1.1">1.12.25 mm</quantity>
; anal cleft about 1/9th body length; stigmatic clefts absent.
</paragraph>
<paragraph id="410E067FFFA1FFBCFF6C67A0FF1C72A1" blockId="1.[151,1437,988,1979]" pageId="1" pageNumber="123">
<emphasis id="73C5DA6DFFA1FFBCFF6C67A0FEDE70BE" box="[199,297,1422,1445]" italics="true" pageId="1" pageNumber="123">Dorsum.</emphasis>
Derm membranous. Dorsal setae absent. Dorsal pores in a reticulate pattern, with reticulation areas in probably 9 longitudinal rows, with 8 (9?) reticulation areas medially between anal plates and anterior margin and with 29 areas around margin (but see under Discussion below). Dorsal pores of possibly five
<typeStatus id="9E0AB8DDFFA1FFBCFA8467FEFA9870FC" box="[1327,1391,1488,1511]" pageId="1" pageNumber="123">types</typeStatus>
: (i) microductules, each about 0.5 µm wide, with a long balloon-like inner ductule, main pore marking position of reticulation lines, possibly absent elsewhere; (ii) simple pores, each small and probably flat, about 1.0 µm wide, sparse throughout dorsum, including between reticulation lines; (iii) a slightly larger simple pore, each about 2.0 2.5 µm wide with a dark centre, sparse within reticulation lines and possibly forming a sparse line along margin; (iv) slightly larger, sclerotised macropores, each probably convex and 3.03.5 µm wide, delimiting submedial reticulation lines, at least on abdomen, with a few extending onto thorax and also in more medial radial lines, and (v) a similar-sized simple pore, each rather variable in size, 2.04.0 µm wide, probably flat, in a sparse band or line on each side of anal plates. Preopercular pores, dorsal tubercles and dorsal tubular ducts absent. Anal plates together widest at anterior 1/5th to 1/4th, tapering to a narrow apex, each 180200 µm long, combined widths 130 165 µm; with 02 minute pores on dorsal surface of each plate; length of setae: anterior inner margin setae 1325 µm; posterior inner margin 1620 µm; apical 3540 um and outer margin setae 1620 µm (actually on dorsal surface near apex); each anal plate with a narrow supporting bar. Anogenital fold with 2 pairs of setae along anterior margin and 1 or 2 setae on each lateral margin; longest 5055 µm long. Anal tube quite long; anal ring with 6 setae.
</paragraph>
</subSubSection>
<caption id="15CE56F7FFA2FFBFFF3C65B9FCE372F1" httpUri="https://zenodo.org/record/256046/files/figure.png" pageId="2" pageNumber="124" targetBox="[374,1215,223,1915]" targetPageId="2">
<paragraph id="410E067FFFA2FFBFFF3C65B9FCE372F1" blockId="2.[151,1436,1942,2026]" pageId="2" pageNumber="124">
<emphasis id="73C5DA6DFFA2FFBFFF3C65B9FEE072B7" bold="true" box="[151,279,1943,1964]" pageId="2" pageNumber="124">FIGURE 1.</emphasis>
Photographs of
<taxonomicName id="86B17DFCFFA2FFBFFE6765B8FC8C72B7" authority="Hodgson &amp; Richmond" authorityName="Hodgson &amp; Richmond" box="[460,891,1942,1964]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="124" phylum="Arthropoda" rank="species" species="martini" status="sp. nov.">
<emphasis id="73C5DA6DFFA2FFBFFE6765B8FD8872B7" box="[460,639,1942,1964]" italics="true" pageId="2" pageNumber="124">Kalasiris martini</emphasis>
Hodgson &amp; Richmond
</taxonomicName>
,
<taxonomicNameLabel id="68F66716FFA2FFBFFC2265B9FC2A72B7" box="[905,989,1943,1964]" pageId="2" pageNumber="124" rank="species">
<emphasis id="73C5DA6DFFA2FFBFFC2265B9FC2A72B7" bold="true" box="[905,989,1943,1964]" pageId="2" pageNumber="124">spec. n.</emphasis>
</taxonomicNameLabel>
1a: adult female, showing waxy test and reticulation plates; 1b: an old female with body showing dark at anterior end and remainder of test filled with eggs; 1c: test of second-instar male showing arrangement of reticulation plates.
</paragraph>
</caption>
<caption id="15CE56F7FFA3FFBEFF3C65F7FC6E72F4" box="[151,921,2009,2031]" httpUri="https://zenodo.org/record/256047/files/figure.png" pageId="3" pageNumber="125" targetBox="[188,1377,283,1870]" targetPageId="3">
<paragraph id="410E067FFFA3FFBEFF3C65F7FC6E72F4" blockId="3.[151,921,2009,2031]" box="[151,921,2009,2031]" pageId="3" pageNumber="125">
<emphasis id="73C5DA6DFFA3FFBEFF3C65F7FEE572F5" bold="true" box="[151,274,2009,2030]" pageId="3" pageNumber="125">FIGURE 2.</emphasis>
Adult female
<taxonomicName id="86B17DFCFFA3FFBEFE0F65F7FCB772F4" authority="Hodgson &amp; Richmond" authorityName="Hodgson &amp; Richmond" box="[420,832,2009,2031]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="125" phylum="Arthropoda" rank="species" species="martini" status="sp. nov.">
<emphasis id="73C5DA6DFFA3FFBEFE0F65F7FDA672F4" box="[420,593,2009,2031]" italics="true" pageId="3" pageNumber="125">Kalasiris martini</emphasis>
Hodgson &amp; Richmond
</taxonomicName>
,
<taxonomicNameLabel id="68F66716FFA3FFBEFCE165F4FC6E72F4" box="[842,921,2010,2031]" pageId="3" pageNumber="125" rank="species">spec. n.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="09AB55F4FFA4FFB9FF6C62B6FCDB734B" pageId="4" pageNumber="126" type="description">
<paragraph id="410E067FFFA4FFB9FF6C62B6FB7F7493" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C62B6FED575B4" box="[199,290,152,175]" italics="true" pageId="4" pageNumber="126">Margin.</emphasis>
Marginal setae in a single line, of two sizes, both lanceolate and spinose, with rather blunt apices and narrow basal sockets: (i) shorter 1620 µm long, with 5163 on each side between stigmatic clefts, and (ii) larger setae, similar in shape to (i) but each 3045 µm long, located at ends of each reticulation line, with 8 between stigmatic areas anteriorly; three on each side between stigmatic areas and seven on each abdominal margin; marginal setae extending only a short distance up anal cleft margin. Stigmatic areas without stigmatic sclerotisations and without a cleft, each area with a single rather blunt spine, generally slightly curved, with broad basal sockets; each 5060 µm long. Eyespot present on margin, each about 810 µm long.
</paragraph>
<paragraph id="410E067FFFA4FFB9FF6C63BBFC1E7167" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C63BBFEFB74B7" box="[199,268,405,428]" italics="true" pageId="4" pageNumber="126">Venter</emphasis>
. Derm membranous. Pregenital disc-pores each about 8 µm wide with mainly 10 loculi; in bands across all abdominal segments but most abundant posteriorly; each band with a mediolateral group; with none laterad to metacoxae. Spiracular disc-pores each about 45 µm wide with mainly 5 loculi, in bands 14 pores wide; with
<quantity id="8649AB9AFFA4FFB9FAD963F2FF387703" metricMagnitude="-1" metricUnit="m" metricValue="8.509" metricValueMax="10.668" metricValueMin="6.35" pageId="4" pageNumber="126" unit="in" value="33.5" valueMax="42.0" valueMin="25.0">25 42 in</quantity>
each band; with a few extending medially over each peritreme. Ventral microducts of one size, each about 1.5 µm wide, in a sparse submarginal band and in a dense group posterior to mouthparts. With 02 preantennal pores just anterior to each antenna. Ventral tubular ducts of two
<typeStatus id="9E0AB8DDFFA4FFB9FCB66066FCAF7744" box="[797,856,584,607]" pageId="4" pageNumber="126">types</typeStatus>
as for genus: (i) shorter duct with outer ductule 15 17 µm long and with a fine inner ductule 1114 µm long, frequent on either side of anal cleft and across all abdominal segments in a line or band anterior to multilocular disc-pores, but very sparse or absent medially, and (ii) longer ducts, each with outer ductule about 25 µm long and inner ductule 8 µm long, in a broad submarginal band, extending medially to spiracles and also a few present between clypeolabral shield and antennae. Ventral setae: ventral anal lobe setae 1525 µm long; with one pair of anterior anal cleft setae; with long pregenital setae restricted to pairs medially on abdominal segments VI and VII; short setae medially on each abdominal segment few; with 2 or 3 minute setae near each coxa but with one longer seta near each procoxa; with 3 pairs of interantennal setae, posteriormost setae unusually long; with 13 submarginal setae on each side between stigmatic areas. Antennae 8-segmented, with distinct segmentation between segments III and IV, and between IV and V; total length 500550 µm; length of apical setae 8090 µm; fleshy seta on each antenna particularly long. Length of clypeolabral shield 165170 µm. Width of spiracular peritremes: anterior 5060 µm, posterior 5870 µm. Legs elongate, with a separate tibia and tarsus, but no tibio-tarsal articulatory sclerosis; distribution of leg setae as for family; claws with a minute denticle; claw digitules similar and broad; tarsal digitules dissimilar, one shorter and narrower than other; tarsal campaniform pores absent; lengths (metathoracic): coxa 160175 µm, trochanter + femur 225240 µm, tibia 165175 µm, tarsus 100110 µm, claw 2023 µm.
</paragraph>
<paragraph id="410E067FFFA4FFB9FF6C66A7FEFC702B" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C66A7FEF071BB" box="[199,263,1161,1184]" italics="true" pageId="4" pageNumber="126">Notes</emphasis>
. Although the exact layout of the reticulation lines on the mounted specimens was hard to determine, the presence of 14 larger marginal setae on each side clearly suggests that there are a total of 29 marginal waxy plates. The small size of the pores and their sparseness made delineation of the other rows of reticulation plates difficult but the photographs strongly suggest that there are three pairs in addition to the central line. These are not shown in the figure.
</paragraph>
<paragraph id="410E067FFFA4FFB9FF6C6712FE497337" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
Hodgson &amp; Henderson (2000) suggested that the following adult female characteristics diagnosed
<taxonomicName id="86B17DFCFFA4FFB9FA866712FA63704E" box="[1325,1428,1340,1365]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="126" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA4FFB9FA866712FA63704E" box="[1325,1428,1340,1365]" italics="true" pageId="4" pageNumber="126">Kalasiris</emphasis>
</taxonomicName>
: (i) marginal setae particularly abundant, with some extending up margins of anal cleft; (ii) at least a few larger simple pores dorsally on either side of anal plates; (iii) two
<typeStatus id="9E0AB8DDFFA4FFB9FC2967ABFC4A7087" box="[898,957,1413,1436]" pageId="4" pageNumber="126">types</typeStatus>
of ventral tubular ducts, a longer
<typeStatus id="9E0AB8DDFFA4FFB9FAC767ABFA6B7087" box="[1388,1436,1413,1436]" pageId="4" pageNumber="126">type</typeStatus>
submarginally and a shorter
<typeStatus id="9E0AB8DDFFA4FFB9FE746787FDF870DB" box="[479,527,1449,1472]" pageId="4" pageNumber="126">type</typeStatus>
medially and mediolaterally on abdominal segments; (iv) ventral tubular ducts present in a submarginal band near to or on margin, the ducts often lying radially, and (v) ventral tubular ducts absent medially on thorax and restricted to between clypeolabral shield and antennae on head. Clearly
<taxonomicName id="86B17DFCFFA4FFB9FA8C67DFFA6B7313" box="[1319,1436,1521,1544]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="126" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="73C5DA6DFFA4FFB9FA8C67DFFA6B7313" box="[1319,1436,1521,1544]" italics="true" pageId="4" pageNumber="126">K. martini</emphasis>
</taxonomicName>
agrees with this diagnosis.
</paragraph>
<paragraph id="410E067FFFA4FFB9FF6C6419FCDB734B" blockId="4.[151,1437,151,2013]" box="[199,812,1591,1616]" pageId="4" pageNumber="126">Male test and pupa. Described from single specimen.</paragraph>
</subSubSection>
<subSubSection id="09AB55F4FFA4FFB9FF6C6472FD437233" pageId="4" pageNumber="126" type="materials_examined">
<paragraph id="410E067FFFA4FFB9FF6C6472FA7473A6" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C6472FE63736E" box="[199,404,1628,1653]" italics="true" pageId="4" pageNumber="126">Mounted material</emphasis>
.
<emphasis id="73C5DA6DFFA4FFB9FE0D6472FDE5736F" box="[422,530,1628,1653]" italics="true" pageId="4" pageNumber="126">Male test</emphasis>
(
<figureCitation id="D98A1AFAFFA4FFB9FD8E6472FD9A736F" box="[549,621,1628,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1943,1964]" captionTargetBox="[374,1215,223,1915]" captionTargetId="figure@2.[347,1240,195,1934]" captionTargetPageId="2" captionText="FIGURE 1. Photographs of Kalasiris martini Hodgson &amp; Richmond, spec. n. 1 a: adult female, showing waxy test and reticulation plates; 1 b: an old female with body showing dark at anterior end and remainder of test filled with eggs; 1 c: test of second-instar male showing arrangement of reticulation plates." httpUri="https://zenodo.org/record/256046/files/figure.png" pageId="4" pageNumber="126">Figs 1</figureCitation>
c &amp; 3): Glassy with a reticulate pattern of polygonal plates, in seven longitudinal rows. Length of test at least
<quantity id="8649AB9AFFA4FFB9FDCD64AEFD5C7383" box="[614,683,1664,1688]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="126" unit="mm" value="2.0">2 mm</quantity>
. Test with 8 medial plates anterior to anal plates, and with more or less 3 lateral lines of plates on each side, each plate of rather variable shape; with 29 plates around outer margin.
</paragraph>
<paragraph id="410E067FFFA4FFB9FF6C64E7FD437233" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C64E7FEF573FB" box="[199,258,1737,1760]" italics="true" pageId="4" pageNumber="126">Pupa</emphasis>
(
<figureCitation id="D98A1AFAFFA4FFB9FEBF64E9FEAB73FB" box="[276,348,1735,1760]" captionStart="FIGURE 4" captionStartId="6.[151,250,884,905]" captionTargetBox="[163,1424,212,848]" captionTargetId="figure@6.[151,1435,193,863]" captionTargetPageId="6" captionText="FIGURE 4. Pupal. Kalasiris martini Hodgson &amp; Richmond, spec. n." httpUri="https://zenodo.org/record/256049/files/figure.png" pageId="4" pageNumber="126">Fig. 4</figureCitation>
): Division into head, thorax and abdomen clear, although segmentation obscure apart from on abdomen. Derm membranous, with small dermal spinules. All ducts and pores (bar spiracular disc-pores) absent and setae few. Of moderate size: length
<quantity id="8649AB9AFFA4FFB9FDFC653EFD587233" box="[599,687,1808,1832]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="4" pageNumber="126" unit="mm" value="1.8">1.8 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="09AB55F4FFA4FFB8FF6C651AFE007063" lastPageId="5" lastPageNumber="127" pageId="4" pageNumber="126" type="description">
<paragraph id="410E067FFFA4FFB9FF6C651AFCAE726B" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C651AFEFB7256" box="[199,268,1844,1869]" italics="true" pageId="4" pageNumber="126">Head.</emphasis>
Without mouthparts or eyes. Setae: two pairs of setae present posteriorly on dorsum. Antennae ten segmented, total length 900 µm (ratio to total body length 1:2).
</paragraph>
<paragraph id="410E067FFFA4FFB8FF6C6552FBFA75E3" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,1400]" lastPageId="5" lastPageNumber="127" pageId="4" pageNumber="126">
<emphasis id="73C5DA6DFFA4FFB9FF6C6552FED7728E" box="[199,288,1916,1941]" italics="true" pageId="4" pageNumber="126">Thorax.</emphasis>
No setae located. With 1215 spiracular disc-pores associated with each anterior spiracle, distributed anteriorly and laterad to peritreme, extending medially full length of spiracular apodeme; with 1 disc-pore just anterior to each posterior spiracle; shape and number of loculi in each disc-pore highly variable. Spiracles: width of peritremes 25 µm. Legs each with a small triangular finger on apex, probably an incipient claw; length of metathoracic legs 950 µm. With a pair of long wing-buds on each side, extending to about abdominal segment III, mildly sclerotised: length 725 µm, width 225 µm (ratio length to width 1:0.32).
</paragraph>
<paragraph id="410E067FFFA5FFB8FF6C632AFE047727" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="127">
<emphasis id="73C5DA6DFFA5FFB8FF6C632AFECE7406" box="[199,313,260,285]" italics="true" pageId="5" pageNumber="127">Abdomen.</emphasis>
Segmentation fairly distinct, anteriormost segment considered to represent segment II (see Hodgson &amp; Henderson, 2000, p. 54), so that there are seven visible segments (segments II to VIII) anterior to penial sheath. With 2 ante-anal setae dorsally on segment VIII; with 1 pair of small ventral setae on segments IIVII; dorsopleural setae: 1 pair on segments VVIII; 0 or 1 small ventropleural seta on segments VVII. Lateral lobe of VII rather pointed, each about half length of penial sheath, each with a long apical seta (45 µm long) and two shorter setae laterally. Segment VIII with a pair of prominent, mildly sclerotised lobes dorsally on either side of penial sheath, each with 2 stout setae. Penial sheath longer than lateral lobes of segment VII and a little longer than wide (165 µm long, 125 µm wide at base; ratio length to width 1:0.75); with a pair of minute setae on dorsal surface and numerous small pores on apex.
</paragraph>
<paragraph id="410E067FFFA5FFB8FF6C6066FF217647" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="127">
<emphasis id="73C5DA6DFFA5FFB8FF6C6066FEE77744" box="[199,272,584,607]" italics="true" pageId="5" pageNumber="127">Notes:</emphasis>
The pupae of
<taxonomicName id="86B17DFCFFA5FFB8FE186069FDED777B" box="[435,538,583,608]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FE186069FDED777B" box="[435,538,583,608]" italics="true" pageId="5" pageNumber="127">Kalasiris</emphasis>
</taxonomicName>
species are rather similar to those of
<taxonomicName id="86B17DFCFFA5FFB8FC156069FBAD777B" box="[958,1114,583,608]" class="Insecta" family="Coccidae" genus="Aphenochiton" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FC156069FBAD777B" box="[958,1114,583,608]" italics="true" pageId="5" pageNumber="127">Aphenochiton</emphasis>
</taxonomicName>
,
<taxonomicName id="86B17DFCFFA5FFB8FBC36069FAF2777B" box="[1128,1285,583,608]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FBC36069FAF2777B" box="[1128,1285,583,608]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
</taxonomicName>
(
<emphasis id="73C5DA6DFFA5FFB8FABE6069FAB6777B" box="[1301,1345,583,608]" italics="true" pageId="5" pageNumber="127">fagi</emphasis>
-group),
<taxonomicName id="86B17DFCFFA5FFB8FF3C6042FED7779E" box="[151,288,620,645]" class="Insecta" family="Coccidae" genus="Ctenochiton" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FF3C6042FED7779E" box="[151,288,620,645]" italics="true" pageId="5" pageNumber="127">Ctenochiton</emphasis>
</taxonomicName>
and
<taxonomicName id="86B17DFCFFA5FFB8FECB6042FDF7779E" box="[352,512,620,645]" class="Insecta" family="Coccidae" genus="Umbonichiton" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FECB6042FDF7779E" box="[352,512,620,645]" italics="true" pageId="5" pageNumber="127">Umbonichiton</emphasis>
</taxonomicName>
from which it is difficult to separate them. The pupae of
<taxonomicName id="86B17DFCFFA5FFB8FB6B6042FAD0779E" box="[1216,1319,620,645]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FB6B6042FAD0779E" box="[1216,1319,620,645]" italics="true" pageId="5" pageNumber="127">Kalasiris</emphasis>
</taxonomicName>
share the following characters: (i) body moderate to quite large in size; (ii) spiracular disc-pores associated with each anterior spiracle mainly in a group dorsad and laterad to peritreme, none extending mesad to inner margin of spiracular peritreme; (iii) with or without disc-pores associated with each posterior spiracle but, when present, few; (iv) lobes on abdominal segment VII well developed; (v) lobes of abdominal segment VIII moderately large and obvious, with 1 or more setae; (vi) ante-anal setae present, and (vii) the segmental arrangement of the dorsopleural setae.
</paragraph>
<paragraph id="410E067FFFA5FFB8FF6C6146FE2E7016" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="127">
<emphasis id="73C5DA6DFFA5FFB8FF6C6146FE55769B" box="[199,418,871,896]" italics="true" pageId="5" pageNumber="127">Generic characters</emphasis>
.
<taxonomicName id="86B17DFCFFA5FFB8FE1A6149FDEF769B" box="[433,536,871,896]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FE1A6149FDEF769B" box="[433,536,871,896]" italics="true" pageId="5" pageNumber="127">Kalasiris</emphasis>
</taxonomicName>
is closest to
<taxonomicName id="86B17DFCFFA5FFB8FD056149FC6E769B" authority="Maskell" authorityName="Maskell" box="[686,921,871,896]" class="Insecta" family="Coccidae" genus="Ctenochiton" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FD056149FCC0769B" box="[686,823,871,896]" italics="true" pageId="5" pageNumber="127">Ctenochiton</emphasis>
Maskell
</taxonomicName>
and
<taxonomicName id="86B17DFCFFA5FFB8FC786149FA8A769B" authority="Henderson &amp; Hodgson" authorityName="Henderson &amp; Hodgson" box="[979,1405,871,896]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FC786149FB87769B" box="[979,1136,871,896]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
Henderson &amp; Hodgson
</taxonomicName>
in having abundant multilocular disc-pores medially across all abdominal segments. It also resembles species of
<taxonomicName id="86B17DFCFFA5FFB8FF3C6181FEC376D3" box="[151,308,943,968]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FF3C6181FEC376D3" box="[151,308,943,968]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
</taxonomicName>
in having 2
<typeStatus id="9E0AB8DDFFA5FFB8FE6F619EFE0876DC" box="[452,511,944,967]" pageId="5" pageNumber="127">types</typeStatus>
of ventral tubular duct, small spiracles, abundant marginal setae and large simple pores laterad to the anal plates. However,
<taxonomicName id="86B17DFCFFA5FFB8FDC061FAFD2576F6" box="[619,722,980,1005]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FDC061FAFD2576F6" box="[619,722,980,1005]" italics="true" pageId="5" pageNumber="127">Kalasiris</emphasis>
</taxonomicName>
differs from
<taxonomicName id="86B17DFCFFA5FFB8FCCE61FAFBF576F6" box="[869,1026,980,1005]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FCCE61FAFBF576F6" box="[869,1026,980,1005]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
</taxonomicName>
in lacking tubular ducts medially on the thorax (present in
<taxonomicName id="86B17DFCFFA5FFB8FE3161D9FDC0710B" box="[410,567,1015,1040]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FE3161D9FDC0710B" box="[410,567,1015,1040]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
</taxonomicName>
) and in having the submarginal tubular ducts tending to lie radially (more random in
<taxonomicName id="86B17DFCFFA5FFB8FEBF6632FE46712E" box="[276,433,1052,1077]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FEBF6632FE46712E" box="[276,433,1052,1077]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
</taxonomicName>
). Adult female
<taxonomicName id="86B17DFCFFA5FFB8FDC36632FBFE712E" authority="Maskell" authorityName="Maskell" box="[616,1033,1052,1077]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="leptospermi">
<emphasis id="73C5DA6DFFA5FFB8FDC36632FC65712E" box="[616,914,1052,1077]" italics="true" pageId="5" pageNumber="127">Crystallotesta leptospermi</emphasis>
(Maskell)
</taxonomicName>
is superficially rather similar
<emphasis id="73C5DA6DFFA5FFB8FACF6633FF1D714C" italics="true" pageId="5" pageNumber="127">
to
<taxonomicName id="86B17DFCFFA5FFB8FA296633FF1D714C" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="martini">K. martini</taxonomicName>
</emphasis>
in having: (i) a rather elongate body; (ii) eight-segmented antennae, (iii) macropores more or less restricted to more medial reticulation lines; (iv) quite large reticulation setae on margin, and (v) long stigmatic spines. However
<taxonomicName id="86B17DFCFFA5FFB8FEAC66A6FE5871BB" box="[263,431,1159,1184]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="leptospermi">
<emphasis id="73C5DA6DFFA5FFB8FEAC66A6FE5871BB" box="[263,431,1159,1184]" italics="true" pageId="5" pageNumber="127">C. leptospermi</emphasis>
</taxonomicName>
has a few tubular ducts medially on the thorax, and the distribution of the reticulation plates is different, with only approximately 5 plates between anal plates and anterior margin (
<taxonomicName id="86B17DFCFFA5FFB8FB066683FAD771DF" box="[1197,1312,1197,1220]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="73C5DA6DFFA5FFB8FB066683FAD771DF" box="[1197,1312,1197,1220]" italics="true" pageId="5" pageNumber="127">K. martini</emphasis>
</taxonomicName>
has eight). It is clear that these two genera (
<taxonomicName id="86B17DFCFFA5FFB8FE5466E1FD6B71F3" box="[511,668,1231,1256]" class="Insecta" family="Coccidae" genus="Crystallotesta" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FE5466E1FD6B71F3" box="[511,668,1231,1256]" italics="true" pageId="5" pageNumber="127">Crystallotesta</emphasis>
</taxonomicName>
and
<taxonomicName id="86B17DFCFFA5FFB8FD7A66E1FCCF71F3" box="[721,824,1231,1256]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="genus">
<emphasis id="73C5DA6DFFA5FFB8FD7A66E1FCCF71F3" box="[721,824,1231,1256]" italics="true" pageId="5" pageNumber="127">Kalasiris</emphasis>
</taxonomicName>
) are very close and may need to be synonymised when more species are discovered.
</paragraph>
<paragraph id="410E067FFFA5FFB8FF6C6736FE007063" blockId="5.[151,1437,151,1400]" pageId="5" pageNumber="127">
Name derivation. The species name
<taxonomicName id="86B17DFCFFA5FFB8FDD46737FD25702B" box="[639,722,1305,1328]" class="Insecta" family="Coccidae" genus="Kalasiris" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="127" phylum="Arthropoda" rank="species" species="martini">
<emphasis id="73C5DA6DFFA5FFB8FDD46737FD25702B" box="[639,722,1305,1328]" italics="true" pageId="5" pageNumber="127">martini</emphasis>
</taxonomicName>
refers to Dr. Nicholas Martin, Research Associate, Landcare Research, Auckland,
<collectingCountry id="39A646EFFFA5FFB8FE2C6713FDD7704E" box="[391,544,1340,1365]" name="New Zealand" pageId="5" pageNumber="127">New Zealand</collectingCountry>
, who collected these insects and also many other scale insects in
<collectingCountry id="39A646EFFFA5FFB8FB546713FA6F704E" box="[1279,1432,1340,1365]" name="New Zealand" pageId="5" pageNumber="127">New Zealand</collectingCountry>
, several of them new to science.
</paragraph>
</subSubSection>
</treatment>
</document>