149 lines
17 KiB
XML
149 lines
17 KiB
XML
<document id="65B60953E4A9BDAD9C143C10D5A0A102" ID-DOI="10.1206/3928.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5369854" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268040215" checkinUser="carolina" docAuthor="Hopkins, Melanie J." docDate="2019" docId="B10287DEFF97B058FE6EFD09FCE2F934" docLanguage="en" docName="N3928 highres.pdf" docOrigin="American Museum Novitates 2019 (3928)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2019/issue-3928/3928.1/Phylogenetic-Analysis-and-Revision-of-the-Trilobite-Subfamily-Balnibarbiinae-Olenidae/10.1206/3928.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Cloacaspis ceryx" docType="treatment" docVersion="4" masterDocId="4D3BFFA6FF9CB053FF99FFE8FF9AFFCB" masterDocTitle="Phylogenetic analysis and revision of the trilobite subfamily Balnibarbiinae (Olenidae)" masterLastPageNumber="1" masterPageNumber="1" pageNumber="1" updateTime="1699205220592" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="DE6DCA79F159E6FC417ADC39C5B38BDC" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="B6B2721FAFB3D79D98A2BFA7AC1B9B8C">
|
||
<mods:title id="8CA15C49854C2C3B125C23A58438099C">Phylogenetic analysis and revision of the trilobite subfamily Balnibarbiinae (Olenidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="C14840270DCDD213D573F28B33FBE85A" type="personal">
|
||
<mods:role id="561E567F28DD7D5D71F9D144564A797E">
|
||
<mods:roleTerm id="1C136F3B342CF16F0AF828AF9EF57516">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D62489A0A52B08527E1D43697A4508A5">Hopkins, Melanie J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="76C0BECA5D389DCFAA4C58E2E5A8B08B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8C63D274E25040307718E6BF02187025" type="host">
|
||
<mods:titleInfo id="47328A2C1D86CF40A89F09E08C2D573C">
|
||
<mods:title id="3E8AFF715FD50047DC39A0A038F7B5AC">American Museum Novitates</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="42B97D6A350B140443FDE021F222D863">
|
||
<mods:date id="14661F22FBE0E9E4D819AEA70EF5AD13">2019</mods:date>
|
||
<mods:detail id="D0574F8851820205BE638571F323344E" type="pubDate">
|
||
<mods:number id="3B9580971E17EDB199BDD484EF9D3BCF">2019-05-07</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E124EB5D770B246A4C25E34CDCBB752C" type="volume">
|
||
<mods:number id="1FCFEEDC9B2F31B97789F120A87E9F49">2019</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DBBBAFE403F2B64BB2B7E8FBFEC4232D" type="issue">
|
||
<mods:number id="84BFF06FE63AC516F90C53FA44A1371A">3928</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="87C0B11784B5BB9254B656765B425514" unit="page">
|
||
<mods:start id="0E7813B9649DD23C249B6BC802039428">1</mods:start>
|
||
<mods:end id="267F224DCD3E48F6A855450B3AAD3E29">1</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="05739191F9C840ECADC1A768C5F4CDAD">
|
||
<mods:url id="259353180C440C23551466CBA9B05A8A">https://bioone.org/journals/american-museum-novitates/volume-2019/issue-3928/3928.1/Phylogenetic-Analysis-and-Revision-of-the-Trilobite-Subfamily-Balnibarbiinae-Olenidae/10.1206/3928.1.full</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="632BCEBF1F94F8CBE183A205B5725D17">journal article</mods:classification>
|
||
<mods:identifier id="18ACE7B2CDF3650175D335B6255C9AE9" type="DOI">10.1206/3928.1</mods:identifier>
|
||
<mods:identifier id="74478581205C359B6D7DF38BD67701DC" type="ISSN">0003-0082</mods:identifier>
|
||
<mods:identifier id="E76435DF22B8A121BC308BABFAD230A3" type="Zenodo-Dep">5369854</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="B10287DEFF97B058FE6EFD09FCE2F934" ID-DOI="http://doi.org/10.5281/zenodo.5461574" ID-Zenodo-Dep="5461574" LSID="urn:lsid:plazi:treatment:B10287DEFF97B058FE6EFD09FCE2F934" httpUri="http://treatment.plazi.org/id/B10287DEFF97B058FE6EFD09FCE2F934" pageId="11">
|
||
<subSubSection id="71B16543FF97B058FE6EFD09FCFBFD37" box="[503,865,737,764]" pageId="11" type="nomenclature">
|
||
<paragraph id="391436C8FF97B058FE6EFD09FCFBFD37" blockId="11.[503,865,737,764]" box="[503,865,737,764]" pageId="11">
|
||
<heading id="625C81A4FF97B058FE6EFD09FCFBFD37" box="[503,865,737,764]" centered="true" fontSize="11" level="2" pageId="11" reason="5">
|
||
<taxonomicName id="FEAB4D4BFF97B058FE6EFD09FCFBFD37" authority="(Fortey, 1974)" baseAuthorityName="Fortey" baseAuthorityYear="1974" box="[503,865,737,764]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="ceryx">
|
||
<emphasis id="0BDFEADAFF97B058FE6EFD09FD2EFD30" box="[503,692,737,763]" italics="true" pageId="11">Cloacaspis ceryx</emphasis>
|
||
(
|
||
<bibRefCitation id="5D3A4B39FF97B058FD5DFD0AFCC3FD37" author="Fortey, R. A." box="[708,857,738,764]" pageId="11" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey, 1974</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71B16543FF97B058FDFAFCF1FD6FFCFF" box="[611,757,793,820]" pageId="11" type="description">
|
||
<paragraph id="391436C8FF97B058FDFAFCF1FD6FFCFF" blockId="11.[611,757,793,820]" box="[611,757,793,820]" pageId="11">
|
||
<heading id="625C81A4FF97B058FDFAFCF1FD6FFCFF" box="[611,757,793,820]" centered="true" fontSize="36" level="3" pageId="11" reason="7">
|
||
<figureCitation id="A1902A4DFF97B058FDFAFCF1FD6FFCFF" box="[611,757,793,820]" captionStart="FIGURE 4" captionStartId="10.[132,218,1492,1513]" captionTargetBox="[163,1205,248,1465]" captionTargetId="figure-41@10.[163,719,685,1129]" captionTargetPageId="10" captionText="FIGURE 4. Cloacaspis spp. A. Cloacaspis ekphymosa Fortey, 1974, USNM 144232. Collected from the Lost River Area on Seward Peninsula, Alaska, and previously identified as Triarthrus sp. (Ross, 1965). Arrow pointing to postpalpebral furrow. B. Anterior border of Cloacaspis ceryx anataphra (Fortey, 1974), AMNH- FI-101655. Black arrows pointing to faint anterior border pits on internal mold. C. Cloacaspis ceryx anataphra (Fortey, 1974), AMNH-FI-101655. Collected from PO-18 of the Olenidsletta Member of the Valhallfonna Formation (Kröger et al., 2017). D) Cloacaspis ceryx ceryx (Fortey, 1974), AMNH-FI-101657. Close-up of E, anterior glabellar lobe and F,posterior wing, showing presence of fine granulation of recently prepared surface (white arrow) and lack of granulation on exposed and slightly eroded surface (black arrow). Collected from bed PO-18 of the Olenidsletta Member of the Valhallfonna Formation (Kröger et al., 2017). Scale bars = 1 mm except A and D, where scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5369862" httpUri="https://zenodo.org/record/5369862/files/figure.png" pageId="11">Figure 4B–F</figureCitation>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71B16543FF97B058FF1DFCBBFE73FCA0" box="[132,489,851,876]" pageId="11" type="reference_group">
|
||
<paragraph id="391436C8FF97B058FF1DFCBBFE73FCA0" blockId="11.[132,489,851,876]" box="[132,489,851,876]" pageId="11">
|
||
<treatmentCitationGroup id="19BB11E6FF97B058FF1DFCBBFE73FCA0" box="[132,489,851,876]" pageId="11">
|
||
<treatmentCitation id="B80A10D9FF97B058FF1DFCBBFE7FFCA0" author="Fortey, R. A." box="[132,485,851,876]" page="29" pageId="11" year="1974">
|
||
<taxonomicName id="FEAB4D4BFF97B058FF1DFCBBFE7FFCA0" ID-CoL="KHR9" authority="Fortey, 1974: 29" authorityName="Fortey" authorityPageNumber="29" authorityYear="1974" box="[132,485,851,876]" class="Trilobita" family="Olenidae" genus="Balnibarbi" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="ceryx">
|
||
<emphasis id="0BDFEADAFF97B058FF1DFCBBFEABFCA0" box="[132,305,851,875]" italics="true" pageId="11">Balnibarbi ceryx</emphasis>
|
||
<bibRefCitation id="5D3A4B39FF97B058FEA1FCBCFE7FFCA0" author="Fortey, R. A." box="[312,485,851,876]" pageId="11" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey, 1974: 29</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71B16543FF97B058FF2DFC61FE65FB95" pageId="11" type="materials_examined">
|
||
<paragraph id="391436C8FF97B058FF2DFC61FB8CFC68" blockId="11.[132,1237,905,1791]" box="[180,1046,905,932]" pageId="11">
|
||
<materialsCitation id="89C33C95FF97B058FF2DFC61FB8CFC68" accessionNumber="SMA 84034" box="[180,1046,905,932]" pageId="11" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="E610886AFF97B058FF2DFC61FEA2FC68" box="[180,312,905,931]" pageId="11" type="holotype">HOLOTYPE</typeStatus>
|
||
: Cranidium, SMA 84034 (figured in
|
||
<bibRefCitation id="5D3A4B39FF97B058FD72FC62FCE4FC68" author="Fortey, R. A." box="[747,894,905,932]" pageId="11" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey, 1974</bibRefCitation>
|
||
: pl. 8, fig. 1).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="391436C8FF97B058FF2DFC47FE1DFC25" blockId="11.[132,1237,905,1791]" pageId="11">
|
||
<collectionCode id="5FBAAE0DFF97B058FF2DFC47FEF6FC02" box="[180,364,943,969]" pageId="11">TYPE LOCALITY</collectionCode>
|
||
: Profilstranda, Olenidsletta Member, Valhallfonna Formation, Ny Friesland, Spitsbergen,
|
||
<collectingCountry id="41BC7658FF97B058FE82FC3CFE19FC25" box="[283,387,980,1006]" name="Svalbard and Jan Mayen" pageId="11">Svalbard</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="391436C8FF97B058FF2DFC12FE65FB95" blockId="11.[132,1237,905,1791]" pageId="11">
|
||
EMENDED DIAGNOSIS:
|
||
<taxonomicName id="FEAB4D4BFF97B058FE24FC11FD17FBD8" box="[445,653,1017,1043]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="0BDFEADAFF97B058FE24FC11FDAFFBD8" box="[445,565,1017,1043]" italics="true" pageId="11">Cloacaspis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
with greater (sag., exsag.) preglabellar length than other species of the genus; anterior border comes to a point on midline. Moderately sized pygidium with three axial rings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="71B16543FF97B058FF2DFB82FCE2F934" pageId="11" type="discussion">
|
||
<paragraph id="391436C8FF97B058FF2DFB82FCE2F934" blockId="11.[132,1237,905,1791]" pageId="11">
|
||
DISCUSSION:
|
||
<bibRefCitation id="5D3A4B39FF97B058FED4FB82FE70FB4F" author="Fortey, R. A." box="[333,490,1130,1156]" pageId="11" pagination="1 - 129" refId="ref7470" refString="Fortey, R. A. 1974. The Ordovician trilobites of Spitsbergen. I. Olenidae. Norsk Polarinstitutt Skrifter 160: 1 - 129." type="journal article" year="1974">Fortey (1974)</bibRefCitation>
|
||
distinguished
|
||
<taxonomicName id="FEAB4D4BFF97B058FD0AFB81FC15FB48" authority="ceryx" authorityName="ceryx (Fortey" authorityYear="1974" box="[659,911,1129,1155]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="ceryx">
|
||
<emphasis id="0BDFEADAFF97B058FD0AFB81FC15FB48" box="[659,911,1129,1155]" italics="true" pageId="11">Cloacaspis ceryx ceryx</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="FEAB4D4BFF97B058FC50FB81FF22FB62" baseAuthorityName="Fortey" baseAuthorityYear="1974" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="anataphra">
|
||
<emphasis id="0BDFEADAFF97B058FC50FB81FF22FB62" italics="true" pageId="11">Cloacaspis ceryx anataphra</emphasis>
|
||
</taxonomicName>
|
||
by the latter having a shorter (sag.) preglabellar field relative to the glabella, no visible pits in the anterior border furrow, and a smooth exoskeletal surface. Examination of new and old specimens suggests that the mean preglabellar/glabella ratio is on average shorter in
|
||
<taxonomicName id="FEAB4D4BFF97B058FBECFB32FF62FAD2" baseAuthorityName="Fortey" baseAuthorityYear="1974" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="anataphra">
|
||
<emphasis id="0BDFEADAFF97B058FBECFB32FF62FAD2" italics="true" pageId="11">C. ceryx anataphra</emphasis>
|
||
</taxonomicName>
|
||
, but there is overlap in variation in this variable. In addition, while pits are not visible in the anterior border furrow, they are present and visible on the internal mold (fig. 4B, C). On the few specimens where it was possible to estimate the number of anterior pits, the number is slightly smaller (25–30) compared to
|
||
<taxonomicName id="FEAB4D4BFF97B058FDAFFA87FD4DFA42" authority="ceryx" authorityName="ceryx (Fortey" authorityYear="1974" box="[566,727,1391,1417]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="ceryx">
|
||
<emphasis id="0BDFEADAFF97B058FDAFFA87FD4DFA42" box="[566,727,1391,1417]" italics="true" pageId="11">C. ceryx ceryx</emphasis>
|
||
</taxonomicName>
|
||
(>30). Although granulation is very fine on
|
||
<taxonomicName id="FEAB4D4BFF97B058FF1DFA7DFEBCFA65" authority="ceryx" authorityName="ceryx (Fortey" authorityYear="1974" box="[132,294,1429,1455]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="subSpecies" species="ceryx" subSpecies="ceryx">
|
||
<emphasis id="0BDFEADAFF97B058FF1DFA7DFEBCFA65" box="[132,294,1429,1455]" italics="true" pageId="11">C. ceryx ceryx</emphasis>
|
||
</taxonomicName>
|
||
and its preservation is very sensitive to any erosion of the surface (fig. 4D–F), it does appear that some specimens truly lack surface granulation. Based on this evidence, the two taxa are retained as subspecies, but the species complex is reassigned to the genus
|
||
<taxonomicName id="FEAB4D4BFF97B058FBC4FA37FB4EFA32" authorityName="Fortey" authorityYear="1974" box="[1117,1236,1503,1529]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="0BDFEADAFF97B058FBC4FA37FB4EFA32" box="[1117,1236,1503,1529]" italics="true" pageId="11">Cloacaspis</emphasis>
|
||
</taxonomicName>
|
||
based on the results of the phylogenetic analysis. The species complex is united with other
|
||
<taxonomicName id="FEAB4D4BFF97B058FB3DF9ECFEB6F988" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="0BDFEADAFF97B058FB3DF9ECFF49F988" italics="true" pageId="11">Cloacaspis</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
by having advanced genal spines (char. 34), relatively large palpebral lobes (char. 19), and forward curvature of the palpebral lobe (char. 20). However,
|
||
<taxonomicName id="FEAB4D4BFF97B058FC5FF9A7FB19F9A2" baseAuthorityName="Fortey" baseAuthorityYear="1974" box="[966,1155,1615,1641]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="ceryx">
|
||
<emphasis id="0BDFEADAFF97B058FC5FF9A7FB19F9A2" box="[966,1155,1615,1641]" italics="true" pageId="11">Cloacaspis ceryx</emphasis>
|
||
</taxonomicName>
|
||
shares some characters with some
|
||
<taxonomicName id="FEAB4D4BFF97B058FE26F99CFD15F945" box="[447,655,1652,1678]" class="Trilobita" family="Olenidae" genus="Balnibarbi" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="0BDFEADAFF97B058FE26F99CFDACF945" box="[447,566,1652,1678]" italics="true" pageId="11">Balnibarbi</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, including greater divergence of the anterior dorsal suture (char. 9), relatively wide palpebral lobes (char. 21), relatively short 3P (char. 25) larger number of axial rings (char. 37), and medial indent on posterior margin of pygidium (char. 44). The latter three characters are also shared with
|
||
<taxonomicName id="FEAB4D4BFF97B058FD3FF90CFCEBF935" authorityName="Fortey" authorityYear="1974" box="[678,881,1764,1790]" class="Trilobita" family="Olenidae" genus="Cloacaspis" kingdom="Animalia" order="Ptychopariida" pageId="11" phylum="Arthropoda" rank="species" species="dejecta">
|
||
<emphasis id="0BDFEADAFF97B058FD3FF90CFCEBF935" box="[678,881,1764,1790]" italics="true" pageId="11">Cloacaspis dejecta</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |