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<document id="82C33356FE1F7F2E6A99E312FBE5522D" ID-CLB-Dataset="55746" ID-DOI="10.26107/RBZ-2020-0087" ID-GBIF-Dataset="34e1b6c1-e284-4c8c-893a-5ac15e63a357" ID-ISSN="2345-7600" ID-Zenodo-Dep="5350238" ID-ZooBank="406FF8D0-4551-4396-91BE-82ADAD24641C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630204834083" checkinUser="diego" docAuthor="Hosoishi, Shingo" docDate="2020" docId="956A9222355EFFDE83A5B93DFCA156A2" docLanguage="en" docName="RafflesBZool.68.759-768.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Crematogaster ransonneti Mayr 1868" docType="treatment" docVersion="6" lastPageNumber="761" masterDocId="6953EA5A355FFFDC8010BA20FFA75312" masterDocTitle="Taxonomic review of the Crematogaster ransonneti-group in Asia, with description of a new species from Malaysia (Hymenoptera: Formicidae: Myrmicinae)" masterLastPageNumber="768" masterPageNumber="759" pageNumber="760" updateTime="1699203462450" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2EEC85B173F74E2A076A81C9F0ADB5A7">Taxonomic review of the Crematogaster ransonneti-group in Asia, with description of a new species from Malaysia (Hymenoptera: Formicidae: Myrmicinae)</mods:title>
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<mods:namePart id="6198ECCEE2F035B847FCC3CBB096981E">Hosoishi, Shingo</mods:namePart>
<mods:affiliation id="5578C3D2EE77E0185E6F1C308AFF40A7">Institute of Tropical Agriculture, Kyushu University, Motooka 744, Nishi-ku, Fukuoka, 819 - 0395 Japan</mods:affiliation>
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<treatment id="956A9222355EFFDE83A5B93DFCA156A2" ID-DOI="http://doi.org/10.5281/zenodo.6512378" ID-GBIF-Taxon="194955436" ID-Zenodo-Dep="6512378" LSID="urn:lsid:plazi:treatment:956A9222355EFFDE83A5B93DFCA156A2" httpUri="http://treatment.plazi.org/id/956A9222355EFFDE83A5B93DFCA156A2" lastPageId="2" lastPageNumber="761" pageId="1" pageNumber="760">
<subSubSection id="55D970BF355EFFDD83A5B93DFA845027" box="[949,1315,797,821]" pageId="1" pageNumber="760" type="nomenclature">
<paragraph id="1D7C2334355EFFDD83A5B93DFA845027" blockId="1.[949,1315,797,821]" box="[949,1315,797,821]" pageId="1" pageNumber="760">
<heading id="46349458355EFFDD83A5B93DFA845027" bold="true" box="[949,1315,797,821]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="760" reason="2">
<emphasis id="2FB7FF26355EFFDD83A5B93DFA845027" bold="true" box="[949,1315,797,821]" pageId="1" pageNumber="760">
<taxonomicName id="DAC358B7355EFFDD83A5B93DFB715027" ID-CoL="6BDD5" authorityName="Mayr" authorityYear="1868" box="[949,1238,797,821]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD83A5B93DFB715027" bold="true" box="[949,1238,797,821]" italics="true" pageId="1" pageNumber="760">Crematogaster ransonneti</emphasis>
</taxonomicName>
-group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355EFFDD833DB97DFC1F5507" pageId="1" pageNumber="760" type="description">
<paragraph id="1D7C2334355EFFDD833DB97DFC1F5507" blockId="1.[811,1452,861,1557]" pageId="1" pageNumber="760">
The
<taxonomicName id="DAC358B7355EFFDD8370B97EFBDC5067" authorityName="Mayr" authorityYear="1868" box="[864,1147,862,885]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD8370B97EFBDC5067" box="[864,1147,862,885]" italics="true" pageId="1" pageNumber="760">Crematogaster ransonneti</emphasis>
</taxonomicName>
-group can be distinguished from other Asian
<taxonomicName id="DAC358B7355EFFDD83E3B95EFB4D5087" box="[1011,1258,893,917]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="2FB7FF26355EFFDD83E3B95EFB345087" box="[1011,1171,894,917]" italics="true" pageId="1" pageNumber="760">Crematogaster</emphasis>
species
</taxonomicName>
by the following characters: (1) mandible with four teeth; (2) three-segmented antennal club; (3) deep metanotal groove; (4) propodeal spine well-developed and long (PSL 0.210.63); (5) propodeal spine directed posterolaterally in dorsal view; (6) petiole diamond-shaped and broadest at mid-length; (7) petiole distinctly wider than postpetiole (WI 7392); and (8) postpetiole weakly globule but without distinct longitudinal median sulcus. This species group is similar to the
<taxonomicName id="DAC358B7355EFFDD8346BEBEFC7257A7" authorityName="Forel" authorityYear="1911" box="[854,981,1181,1205]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="fraxatrix">
<emphasis id="2FB7FF26355EFFDD8346BEBEFC7257A7" box="[854,981,1181,1205]" italics="true" pageId="1" pageNumber="760">C. fraxatrix</emphasis>
</taxonomicName>
-group in having a diamond-shaped petiole (
<bibRefCitation id="79525EC5355EFFDD8324BE9DFBEE57C7" author="Hosoishi S &amp; Ogata K" box="[820,1097,1213,1237]" pageId="1" pageNumber="760" pagination="57 - 68" refId="ref4924" refString="Hosoishi S &amp; Ogata K (2014) Description and DNA barcoding of Crematogaster fraxatrix Forel, 1911 and two new closely related species from Cambodia and Indonesia (Hymenoptera: Formicidae). ZooKeys, 374: 57 - 68." type="journal article" year="2014">Hosoishi &amp; Ogata, 2014</bibRefCitation>
), but can be distinguished from the latter by the aforementioned characters (3), (4), and (5). Using a comprehensive molecular phylogeny,
<bibRefCitation id="79525EC5355EFFDD8546BEDDFC245627" author="Blaimer BB" pageId="1" pageNumber="760" pagination="421 - 436" refId="ref4506" refString="Blaimer BB (2012 b) Acrobat ants go global - origin, evolution and systematics of the genus Crematogaster (Hymenoptera: Formicidae). Molecular Phylogenetics and Evolution, 65: 421 - 436." type="journal article" year="2012">Blaimer (2012b)</bibRefCitation>
revealed a sister relationship between
<taxonomicName id="DAC358B7355EFFDD853DBF3EFA0C5627" authorityName="Forel" authorityYear="1911" box="[1325,1451,1309,1333]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="fraxatrix">
<emphasis id="2FB7FF26355EFFDD853DBF3EFA0C5627" box="[1325,1451,1309,1333]" italics="true" pageId="1" pageNumber="760">C. fraxatrix</emphasis>
</taxonomicName>
and
<taxonomicName id="DAC358B7355EFFDD8349BF1EFC605647" authorityName="Emery" authorityYear="1888" box="[857,967,1341,1365]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ferrarii">
<emphasis id="2FB7FF26355EFFDD8349BF1EFC605647" box="[857,967,1341,1365]" italics="true" pageId="1" pageNumber="760">C. ferrarii</emphasis>
</taxonomicName>
(as
<taxonomicName id="DAC358B7355EFFDD83E2BF1EFB275647" authorityName="Emery" authorityYear="1888" box="[1010,1152,1341,1365]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ferrarii">
<emphasis id="2FB7FF26355EFFDD83E2BF1EFBAD5647" box="[1010,1034,1342,1365]" italics="true" pageId="1" pageNumber="760">C.</emphasis>
cf.
<emphasis id="2FB7FF26355EFFDD8421BF1DFB275647" box="[1073,1152,1341,1365]" italics="true" pageId="1" pageNumber="760">ferrarii</emphasis>
</taxonomicName>
). The phylogenetic position of the
<taxonomicName id="DAC358B7355EFFDD8368BF7EFBA95667" authorityName="Mayr" authorityYear="1868" box="[888,1038,1374,1397]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD8368BF7EFBA95667" box="[888,1038,1374,1397]" italics="true" pageId="1" pageNumber="760">C. ransonneti</emphasis>
</taxonomicName>
-group is currently unknown because fresh material of the group was unavailable for me. In the future, molecular phylogenetic analysis using fresh material will help verify the relatedness of the
<taxonomicName id="DAC358B7355EFFDD84C1BF9EFAC356C7" authorityName="Mayr" authorityYear="1868" box="[1233,1380,1470,1493]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD84C1BF9EFAC356C7" box="[1233,1380,1470,1493]" italics="true" pageId="1" pageNumber="760">C. ransonneti</emphasis>
</taxonomicName>
-group with other species, which is currently inferred mostly from morphology.
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355EFFDD833DBC1DFAA85487" pageId="1" pageNumber="760" type="discussion">
<paragraph id="1D7C2334355EFFDD833DBC1DFBE85587" blockId="1.[813,1453,1597,1685]" pageId="1" pageNumber="760">
This species group is distributed throughout parts of South and Southeast Asia (
<figureCitation id="85F83FB1355EFFDD841ABC7DFBEF5567" box="[1034,1096,1629,1653]" captionStart="Fig" captionStartId="2.[136,171,1186,1208]" captionTargetBox="[137,1475,188,1170]" captionTargetId="figure-323@2.[134,1479,184,1174]" captionTargetPageId="2" captionText="Fig. 1. Distribution map of Crematogaster cornuta, C. keris, and C. ransonneti. Open circle = C. cornuta; open square = C. keris; open triangle = C. ransonneti." figureDoi="http://doi.org/10.5281/zenodo.5350240" httpUri="https://zenodo.org/record/5350240/files/figure.png" pageId="1" pageNumber="760">Fig. 1</figureCitation>
), but has not been recorded from the Indochinese Peninsula.
</paragraph>
<paragraph id="1D7C2334355EFFDD833DBC9DFCDF55E7" blockId="1.[813,1452,1725,1878]" pageId="1" pageNumber="760">
<heading id="46349458355EFFDD833DBC9DFCDF55E7" bold="true" centered="true" fontSize="10" level="2" pageId="1" pageNumber="760" reason="2">
<emphasis id="2FB7FF26355EFFDD833DBC9DFCDF55E7" bold="true" pageId="1" pageNumber="760">
List of species included in the
<taxonomicName id="DAC358B7355EFFDD8494BC9DFA0555C7" authorityName="Mayr" authorityYear="1868" box="[1156,1442,1725,1749]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD8494BC9DFA0555C7" bold="true" box="[1156,1442,1725,1749]" italics="true" pageId="1" pageNumber="760">Crematogaster ransonneti</emphasis>
</taxonomicName>
- group.
</emphasis>
</heading>
</paragraph>
<paragraph id="1D7C2334355EFFDD833DBCDEFBEF5407" blockId="1.[813,1452,1725,1878]" box="[813,1096,1789,1813]" pageId="1" pageNumber="760">
<taxonomicName id="DAC358B7355EFFDD833DBCDEFBEF5407" authority="Crawley, 1924" authorityName="Crawley" authorityYear="1924" box="[813,1096,1789,1813]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="2FB7FF26355EFFDD833DBCDEFC065407" box="[813,929,1790,1813]" italics="true" pageId="1" pageNumber="760">C. cornuta</emphasis>
<bibRefCitation id="79525EC5355EFFDD83B9BCDDFBEF5407" author="Crawley WC" box="[937,1096,1789,1813]" pageId="1" pageNumber="760" pagination="380 - 409" refId="ref4667" refString="Crawley WC (1924) Ants from Sumatra. With biological notes by Edward Jacobson. Annals and Magazine of Natural History, Series 9, 13: 380 - 409." type="journal article" year="1924">Crawley, 1924</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1D7C2334355EFFDD833DBD3EFBB45427" blockId="1.[813,1452,1725,1878]" box="[813,1043,1821,1845]" pageId="1" pageNumber="760">
<taxonomicName id="DAC358B7355EFFDD833DBD3EFC255427" authority="Hosoishi, 2020" authorityName="Hosoishi" authorityYear="2020" box="[813,898,1821,1845]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="keris" status="sp. nov.">
<emphasis id="2FB7FF26355EFFDD833DBD3EFC255427" box="[813,898,1821,1845]" italics="true" pageId="1" pageNumber="760">C. keris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3484425D355EFFDD8381BD3EFBB45427" box="[913,1043,1821,1845]" pageId="1" pageNumber="760" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="1D7C2334355EFFDD833DBD1EFBE05447" blockId="1.[813,1452,1725,1878]" box="[813,1095,1853,1878]" pageId="1" pageNumber="760">
<taxonomicName id="DAC358B7355EFFDD833DBD1EFBE05447" authority="Mayr, 1868" authorityName="Mayr" authorityYear="1868" box="[813,1095,1853,1878]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD833DBD1EFC675447" box="[813,960,1854,1877]" italics="true" pageId="1" pageNumber="760">C. ransonneti</emphasis>
<bibRefCitation id="79525EC5355EFFDD83D8BD1EFBE05447" author="Mayr G" box="[968,1095,1853,1878]" pageId="1" pageNumber="760" pagination="287 - 288" refId="ref5033" refString="Mayr G (1868) Crematogaster ransonneti n. sp. Verhandlungen der k. k. Zoologisch-Botanischen Gesellschaft in Wien, 18: 287 - 288." type="journal article" year="1868">Mayr, 1868</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1D7C2334355EFFDD833DBD5DFAA85487" blockId="1.[813,1451,1917,2056]" box="[813,1295,1917,1941]" pageId="1" pageNumber="760">
<emphasis id="2FB7FF26355EFFDD833DBD5DFAA85487" bold="true" box="[813,1295,1917,1941]" pageId="1" pageNumber="760">Key to species based on the worker caste.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355EFFDE833DBDBCFCA156A2" lastPageId="2" lastPageNumber="761" pageId="1" pageNumber="760" type="key">
<key id="69527405355EFFDE833DBDBCFCA156A2" lastPageId="2" lastPageNumber="761" pageId="1" pageNumber="760">
<keyStep id="A6373B91355EFFDD833DBDBCFA0D5B15" pageId="1" pageNumber="760">
<paragraph id="1D7C2334355EFFDD833DBDBCFA0D54DC" blockId="1.[813,1451,1917,2056]" pageId="1" pageNumber="760">
<keyLead id="A6328001355EFFDD833DBDBCFA0D54DC" pageId="1" pageNumber="760">
1. Dorsal surface of head smooth and shiny. Promesonotum smooth and shiny .....................................................
<taxonomicName id="DAC358B7355EFFDD8553BD99FA0D54DC" ID-CoL="6BDD5" authorityName="Mayr" authorityYear="1868" box="[1347,1450,1977,1998]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="760" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355EFFDD8553BD99FA0D54DC" box="[1347,1450,1977,1998]" italics="true" pageId="1" pageNumber="760">ransonneti</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D7C2334355EFFDD833DBDF6FA0D5B15" blockId="1.[813,1451,1917,2056]" pageId="1" pageNumber="760">
<keyLead id="A6328001355EFFDD833DBDF6FA0D5B15" pageId="1" pageNumber="760"> Gena sculptured or striated with longitudinal rugulae. Promesonotum striated with rugulae ......................................2</keyLead>
</paragraph>
</keyStep>
<caption id="49BC73BC355DFFDE8098BE82FEDE57C6" ID-DOI="http://doi.org/10.5281/zenodo.5350240" ID-Zenodo-Dep="5350240" httpUri="https://zenodo.org/record/5350240/files/figure.png" pageId="2" pageNumber="761" startId="2.[136,171,1186,1208]" targetBox="[137,1475,188,1170]" targetPageId="2">
<paragraph id="1D7C2334355DFFDE8098BE82FEDE57C6" blockId="2.[136,1474,1186,1236]" pageId="2" pageNumber="761">
Fig. 1. Distribution map of
<taxonomicName id="DAC358B7355DFFDE818BBE82FDDC57A5" ID-CoL="ZBM8" authorityName="Crawley" authorityYear="1924" box="[411,635,1186,1207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="2FB7FF26355DFFDE818BBE82FDDC57A5" box="[411,635,1186,1207]" italics="true" pageId="2" pageNumber="761">Crematogaster cornuta</emphasis>
</taxonomicName>
,
<taxonomicName id="DAC358B7355DFFDE8299BE82FD7157A5" authorityName="Hosoishi" authorityYear="2020" box="[649,726,1186,1207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="keris">
<emphasis id="2FB7FF26355DFFDE8299BE82FD7157A5" box="[649,726,1186,1207]" italics="true" pageId="2" pageNumber="761">C. keris</emphasis>
</taxonomicName>
, and
<taxonomicName id="DAC358B7355DFFDE8300BE82FC3457A5" ID-CoL="6BDD5" authorityName="Mayr" authorityYear="1868" box="[784,915,1186,1207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355DFFDE8300BE82FC3457A5" box="[784,915,1186,1207]" italics="true" pageId="2" pageNumber="761">C. ransonneti</emphasis>
</taxonomicName>
. Open circle =
<taxonomicName id="DAC358B7355DFFDE8421BE82FB3F57A5" ID-CoL="ZBM8" authorityName="Crawley" authorityYear="1924" box="[1073,1176,1186,1207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="2FB7FF26355DFFDE8421BE82FB3F57A5" box="[1073,1176,1186,1207]" italics="true" pageId="2" pageNumber="761">C. cornuta</emphasis>
</taxonomicName>
; open square =
<taxonomicName id="DAC358B7355DFFDE852ABE82FA2257A5" authorityName="Hosoishi" authorityYear="2020" box="[1338,1413,1186,1207]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="keris">
<emphasis id="2FB7FF26355DFFDE852ABE82FA2257A5" box="[1338,1413,1186,1207]" italics="true" pageId="2" pageNumber="761">C. keris</emphasis>
</taxonomicName>
; open triangle =
<taxonomicName id="DAC358B7355DFFDE80E0BE9FFED457C6" ID-CoL="6BDD5" authorityName="Mayr" authorityYear="1868" box="[240,371,1215,1236]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="ransonneti">
<emphasis id="2FB7FF26355DFFDE80E0BE9FFED457C6" box="[240,371,1215,1236]" italics="true" pageId="2" pageNumber="761">C. ransonneti</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<keyStep id="A6373B91355DFFDE8098BF2CFCA156A2" pageId="2" pageNumber="761">
<paragraph id="1D7C2334355DFFDE8098BF2CFCA15649" blockId="2.[136,777,1291,1457]" pageId="2" pageNumber="761">
<keyLead id="A6328001355DFFDE8098BF2CFCA15649" pageId="2" pageNumber="761">
2. Posterior half of head with longitudinal rugulae. Mesopleuron and lateral surface of propodeum with longitudinal rugulae. Propodeal dorsum with longitudinal rugulae ..............
<taxonomicName id="DAC358B7355DFFDE82ABBF66FCA15649" ID-CoL="ZBM8" authorityName="Crawley" authorityYear="1924" box="[699,774,1350,1371]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="2FB7FF26355DFFDE82ABBF66FCA15649" box="[699,774,1350,1371]" italics="true" pageId="2" pageNumber="761">cornuta</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="1D7C2334355DFFDE8098BF42FCA156A2" blockId="2.[136,777,1291,1457]" pageId="2" pageNumber="761">
<keyLead id="A6328001355DFFDE8098BF42FCA156A2" pageId="2" pageNumber="761">
Posterior half of head smooth. Mesopleuron and lateral surface of propodeum generally smooth. Propodeal dorsum smooth and shiny...................................................................................
<taxonomicName id="DAC358B7355DFFDE82C7BFBBFCA156A2" authorityName="Hosoishi" authorityYear="2020" box="[727,774,1435,1456]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="761" phylum="Arthropoda" rank="species" species="keris">
<emphasis id="2FB7FF26355DFFDE82C7BFBBFCA156A2" box="[727,774,1435,1456]" italics="true" pageId="2" pageNumber="761">keris</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>