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<document id="22E1924AB8618377D0FE1ABE7309D82D" ID-CLB-Dataset="55746" ID-DOI="10.26107/RBZ-2020-0087" ID-GBIF-Dataset="34e1b6c1-e284-4c8c-893a-5ac15e63a357" ID-ISSN="2345-7600" ID-Zenodo-Dep="5350238" ID-ZooBank="406FF8D0-4551-4396-91BE-82ADAD24641C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630204834083" checkinUser="diego" docAuthor="Hosoishi, Shingo" docDate="2020" docId="956A9222355BFFDA811DBC3EFB8854EF" docLanguage="en" docName="RafflesBZool.68.759-768.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Crematogaster keris Hosoishi 2020, new species" docType="treatment" docVersion="6" lastPageNumber="765" masterDocId="6953EA5A355FFFDC8010BA20FFA75312" masterDocTitle="Taxonomic review of the Crematogaster ransonneti-group in Asia, with description of a new species from Malaysia (Hymenoptera: Formicidae: Myrmicinae)" masterLastPageNumber="768" masterPageNumber="759" pageNumber="763" updateTime="1699203462450" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="322A75A4CDC07424DA37D16D9868DA41" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="88C79EB0F7193E116740CD9C21605183">
<mods:title id="048CD0ADF6CCB60EF93D8C68FD029C91">Taxonomic review of the Crematogaster ransonneti-group in Asia, with description of a new species from Malaysia (Hymenoptera: Formicidae: Myrmicinae)</mods:title>
</mods:titleInfo>
<mods:name id="E97F1CFF8E76278D3C89DD6B671EE6F8" type="personal">
<mods:role id="C098FCEA5DF8DAA907BB4F5A318B8A09">
<mods:roleTerm id="6236C7758AD50C0C58AF175E1F801310">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F2178AC821CDC3BF3DA49753FD30A82">Hosoishi, Shingo</mods:namePart>
<mods:affiliation id="D80B73AE66692A04BB9EF07F2658AA0D">Institute of Tropical Agriculture, Kyushu University, Motooka 744, Nishi-ku, Fukuoka, 819 - 0395 Japan</mods:affiliation>
<mods:nameIdentifier id="18BBE53BA2D942112E5914A8FFC3C9A3" type="email">hosoishi@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F977B0B5AAE6CBE9BB6A076DBEB66DE8">text</mods:typeOfResource>
<mods:relatedItem id="9E4E5DC7268262D6C8685C5E0BE48CA9" type="host">
<mods:titleInfo id="A94A511EA8F3D778513ACE5B3682CE2D">
<mods:title id="D9C0B6EA98422D0D06DDF0FE1D534C98">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="6038EA1F1A4003A1B201F655EC1594B5">
<mods:date id="A3A9CF9DB36F46B667BC0137942D75C9">2020</mods:date>
<mods:detail id="51FE90B4C358A61ED01374CA1AB78052" type="pubDate">
<mods:number id="1B245CBF3116950E570D7207316F6DDF">2020-10-12</mods:number>
</mods:detail>
<mods:detail id="631A5DBD6A2649A6A2873C5513190E4D" type="volume">
<mods:number id="8CB4D81A4CBC7D7710118E884749C122">68</mods:number>
</mods:detail>
<mods:extent id="3C198229B62F7DC64F7401D2C4C7F3D6" unit="page">
<mods:start id="15317BEA354A708867C08887E4E6983A">759</mods:start>
<mods:end id="0772F423CF7A916495F5A6AFBD870852">768</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2DDA73EDDF3990377DA382D5C2A65752">journal article</mods:classification>
<mods:identifier id="A6FDFE3CD13328DA1516218603AAA18F" type="CLB-Dataset">55746</mods:identifier>
<mods:identifier id="8E6FACF4ECEB82338FFB1E69F3CE879B" type="DOI">10.26107/RBZ-2020-0087</mods:identifier>
<mods:identifier id="6DFC39A464FFE0C54C7DFDFA705DA2D9" type="GBIF-Dataset">34e1b6c1-e284-4c8c-893a-5ac15e63a357</mods:identifier>
<mods:identifier id="1266EBF7A236BF597046C337A2AA6C37" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="ACB2BC601C046E6673FCCCD76EE1BFF5" type="Zenodo-Dep">5350238</mods:identifier>
<mods:identifier id="8E2A15B1182BF150550C1DD75C29E386" type="ZooBank">406FF8D0-4551-4396-91BE-82ADAD24641C</mods:identifier>
</mods:mods>
<treatment id="956A9222355BFFDA811DBC3EFB8854EF" ID-DOI="http://doi.org/10.5281/zenodo.5449415" ID-GBIF-Taxon="194955438" ID-Zenodo-Dep="5449415" LSID="urn:lsid:plazi:treatment:956A9222355BFFDA811DBC3EFB8854EF" httpUri="http://treatment.plazi.org/id/956A9222355BFFDA811DBC3EFB8854EF" lastPageId="6" lastPageNumber="765" pageId="4" pageNumber="763">
<subSubSection id="55D970BF355BFFD8811DBC3EFD255524" box="[269,642,1565,1590]" pageId="4" pageNumber="763" type="nomenclature">
<paragraph id="1D7C2334355BFFD8811DBC3EFD255524" blockId="4.[269,642,1565,1622]" box="[269,642,1565,1590]" pageId="4" pageNumber="763">
<heading id="46349458355BFFD8811DBC3EFD255524" bold="true" box="[269,642,1565,1590]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="763" reason="2">
<emphasis id="2FB7FF26355BFFD8811DBC3EFD255524" bold="true" box="[269,642,1565,1590]" pageId="4" pageNumber="763">
<taxonomicName id="DAC358B7355BFFD8811DBC3EFE4A5527" authority="Hosoishi, 2020" authorityName="Hosoishi" authorityYear="2020" box="[269,493,1565,1590]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="763" phylum="Arthropoda" rank="species" species="keris" status="sp. nov.">
<emphasis id="2FB7FF26355BFFD8811DBC3EFE4A5527" bold="true" box="[269,493,1565,1590]" italics="true" pageId="4" pageNumber="763">Crematogaster keris</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="3484425D355BFFD881ECBC3EFD255524" box="[508,642,1566,1590]" pageId="4" pageNumber="763" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355BFFD8819BBC1EFDA35544" box="[395,516,1598,1622]" pageId="4" pageNumber="763" type="description">
<paragraph id="1D7C2334355BFFD8819BBC1EFDA35544" blockId="4.[269,642,1565,1622]" box="[395,516,1598,1622]" pageId="4" pageNumber="763">
(
<figureCitation id="85F83FB1355BFFD88183BC1EFE475544" box="[403,480,1598,1622]" captionStart="Fig" captionStartId="5.[113,148,2061,2083]" captionTargetBox="[112,1453,193,2046]" captionTargetId="figure-9@5.[102,1473,190,2056]" captionTargetPageId="5" captionText="Fig. 3. Crematogaster keris, KUEC paratype worker from Malaysia (HW, 1.02; WL, 1.09). A, body in lateral view; B, full-face view of head; C, dorsal view of mesosoma; D, dorsal view of petiole and postpetiole." figureDoi="http://doi.org/10.5281/zenodo.5350246" httpUri="https://zenodo.org/record/5350246/files/figure.png" pageId="4" pageNumber="763">Figs. 3</figureCitation>
,
<figureCitation id="85F83FB1355BFFD881FEBC1EFE5B5544" box="[494,508,1598,1622]" captionStart="Fig" captionStartId="6.[135,170,1471,1493]" captionTargetBox="[137,1473,189,1459]" captionTargetId="figure-8@6.[117,1488,169,1467]" captionTargetPageId="6" captionText="Fig. 4. Mouthparts of C. keris worker. A, labrum in outer view; B, left maxilla in outer view; C, labium in lateral view. Abbreviations: Ca = cardo; Ga = galea; Gls = glossa; Hyp = hypopharynx; Lbp = labial palp; Mxco = maxillary comb; Mxp = maxillary palp; St = stipes; Prgls = paraglossa; Prm = prementum; Sglb = subglossal brush." figureDoi="http://doi.org/10.5281/zenodo.5350248" httpUri="https://zenodo.org/record/5350248/files/figure.png" pageId="4" pageNumber="763">4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355BFFD88098BC5EFDBC54E4" pageId="4" pageNumber="763" type="materials_examined">
<paragraph id="1D7C2334355BFFD88098BC5EFE7955C4" blockId="4.[136,775,1662,1750]" pageId="4" pageNumber="763">
<emphasis id="2FB7FF26355BFFD88098BC5EFEDD5584" bold="true" box="[136,378,1662,1686]" pageId="4" pageNumber="763">Material examined.</emphasis>
<materialsCitation id="ADAB2969355BFFD88196BC5EFE7955C4" ID-GBIF-Occurrence="3760606306" collectingDate="2003-07-06" collectionCode="FRIM" collectorName="H. Kojima" country="Malaysia" location="Endau Rompin National Park" pageId="4" pageNumber="763" specimenCount="1" specimenCount-worker="1" typeStatus="holotype">
<typeStatus id="C2789D96355BFFD88196BC5EFE575584" box="[390,496,1662,1686]" pageId="4" pageNumber="763" type="holotype">Holotype</typeStatus>
(
<specimenCount id="0BC5E8BD355BFFD88214BC5EFDFE5584" box="[516,601,1662,1686]" pageId="4" pageNumber="763" type="worker">worker</specimenCount>
) (
<collectionCode id="7BD2BBF1355BFFD88264BC5EFD1B5584" box="[628,700,1662,1686]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="4" pageNumber="763">FRIM</collectionCode>
) from
<collectingCountry id="65D463A4355BFFD88098BCBEFF4B55A4" box="[136,236,1694,1718]" name="Malaysia" pageId="4" pageNumber="763">Malaysia</collectingCountry>
,
<location id="181C75EF355BFFD880E9BCBEFD9C55A4" LSID="urn:lsid:plazi:treatment:956A9222355BFFDA811DBC3EFB8854EF:181C75EF355BFFD880E9BCBEFD9C55A4" box="[249,571,1694,1718]" country="Malaysia" name="Endau Rompin National Park" pageId="4" pageNumber="763">Endau Rompin National Park</location>
,
<location id="181C75EF355BFFD88256BCBEFD0055A4" LSID="urn:lsid:plazi:treatment:956A9222355BFFDA811DBC3EFB8854EF:181C75EF355BFFD88256BCBEFD0055A4" box="[582,679,1694,1718]" country="Malaysia" name="Station 1" pageId="4" pageNumber="763">Station 1</location>
, coll.
<collectorName id="B03646E2355BFFD882FDBCBEFF7A55C4" pageId="4" pageNumber="763">H. Kojima</collectorName>
,
<date id="697D05F4355BFFD880F9BC9EFECE55C4" box="[233,361,1726,1750]" pageId="4" pageNumber="763" value="2003-07-06">
<collectingDate id="7939FC1C355BFFD880F9BC9EFECE55C4" box="[233,361,1726,1750]" pageId="4" pageNumber="763" value="2003-07-06">6 July 2003</collectingDate>
</date>
(fogging).
</materialsCitation>
</paragraph>
<paragraph id="1D7C2334355BFFD88098BCDEFD035444" blockId="4.[136,774,1790,1878]" pageId="4" pageNumber="763">
<materialsCitation id="ADAB2969355BFFD88098BCDEFD035444" ID-GBIF-Occurrence="3760606303" collectingDate="2003-07-06" collectionCode="BMNH, CASC, FRIM, HNHM, IEBR, ITBC, KUEC, MBBJ, MCZC, MHNG, NHMB, NHMW, THNHM, ZRC" collectorName="H. Kojima" country="Malaysia" location="Endau Rompin National Park" pageId="4" pageNumber="763" specimenCount="14" specimenCount-worker="14" typeStatus="paratype">
<typeStatus id="C2789D96355BFFD88098BCDEFF5F5404" box="[136,248,1790,1814]" pageId="4" pageNumber="763" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="0BC5E8BD355BFFD88117BCDEFE215404" box="[263,390,1790,1814]" pageId="4" pageNumber="763" type="worker">14 workers</specimenCount>
(
<collectionCode id="7BD2BBF1355BFFD88188BCDEFE495404" box="[408,494,1790,1814]" pageId="4" pageNumber="763">BMNH</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD881EEBCDEFDEF5404" box="[510,584,1790,1814]" pageId="4" pageNumber="763">CASC</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD88247BCDEFD3A5407" box="[599,669,1790,1813]" country="Malaysia" httpUri="http://grbio.org/cool/rw2k-chgr" name="Forest Research Institute, Malaysia" pageId="4" pageNumber="763">FRIM</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD882BCBCDEFCA55407" box="[684,770,1790,1813]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="763">HNHM</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD88098BD3EFF6D5427" box="[136,202,1822,1845]" pageId="4" pageNumber="763">IEBR</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD880CABD3EFEBB5424" box="[218,284,1822,1846]" pageId="4" pageNumber="763">ITBC</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD8813CBD3EFEDD5424" box="[300,378,1822,1846]" country="Japan" name="Kyushu University Entomology Collection" pageId="4" pageNumber="763">KUEC</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD8819ABD3EFE725427" box="[394,469,1822,1845]" country="Indonesia" httpUri="http://grbio.org/cool/k5yn-4b5j" name="Museum Zoologicum Bogoriense, Entomology Collection" pageId="4" pageNumber="763">MBBJ</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD881F5BD3EFD915424" box="[485,566,1822,1846]" pageId="4" pageNumber="763">MCZC</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD88256BD3EFD3A5424" box="[582,669,1822,1846]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="763" type="Museum">MHNG</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD882BDBD3EFCA55427" box="[685,770,1822,1845]" country="0" name="Natural History Museum Bucharest" pageId="4" pageNumber="763">NHMB</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD88098BD1EFF455447" box="[136,226,1854,1877]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="763" type="Museum">NHMW</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD880FFBD1EFEF35447" box="[239,340,1854,1877]" pageId="4" pageNumber="763">THNHM</collectionCode>
,
<collectionCode id="7BD2BBF1355BFFD88170BD1EFE3F5444" box="[352,408,1854,1878]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="4" pageNumber="763">ZRC</collectionCode>
), same data as holotype.
</materialsCitation>
</paragraph>
<paragraph id="1D7C2334355BFFD88098BD5EFDBC54E4" blockId="4.[135,775,1918,2038]" pageId="4" pageNumber="763">
<emphasis id="2FB7FF26355BFFD88098BD5EFE7E5484" bold="true" box="[136,473,1918,1942]" pageId="4" pageNumber="763">Non-type material examined.</emphasis>
<materialsCitation id="ADAB2969355BFFD881F2BD5EFEE054A4" ID-GBIF-Occurrence="3760606307" collectingDate="2003-07-06" collectionCode="KUEC" collectorName="H. Kojima" country="Malaysia" location="Endau Rompin National Park" pageId="4" pageNumber="763" specimenCount="20" specimenCount-worker="20">
<specimenCount id="0BC5E8BD355BFFD881F2BD5EFDFB5484" box="[482,604,1918,1942]" pageId="4" pageNumber="763" type="worker">20 workers</specimenCount>
(
<collectionCode id="7BD2BBF1355BFFD8827DBD5EFD1C5484" box="[621,699,1918,1942]" country="Japan" name="Kyushu University Entomology Collection" pageId="4" pageNumber="763">KUEC</collectionCode>
), same data as holotype;
</materialsCitation>
<materialsCitation id="ADAB2969355BFFD88142BDBEFDBC54E4" ID-GBIF-Occurrence="3760606304" collectingDate="1995-08-20" collectionCode="KUEC" collectorName="T. Yumoto" country="Malaysia" location="Lambir National Park" municipality="Miri" pageId="4" pageNumber="763" specimenCount="1" specimenCount-worker="1" stateProvince="Sarawak">
<specimenCount id="0BC5E8BD355BFFD88142BDBEFE1054A4" box="[338,439,1950,1974]" pageId="4" pageNumber="763" type="worker">1 worker</specimenCount>
(
<collectionCode id="7BD2BBF1355BFFD881D8BDBEFDB054A4" box="[456,535,1950,1974]" country="Japan" name="Kyushu University Entomology Collection" pageId="4" pageNumber="763">KUEC</collectionCode>
),
<collectingCountry id="65D463A4355BFFD8823EBDBEFD3354A4" box="[558,660,1950,1974]" name="Malaysia" pageId="4" pageNumber="763">Malaysia</collectingCountry>
,
<collectingRegion id="DF07EDD6355BFFD882B3BDBEFCA454A4" box="[675,771,1950,1974]" pageId="4" pageNumber="763">Sarawak</collectingRegion>
,
<collectingMunicipality id="FD18B94E355BFFD88098BD9EFF1D54C4" box="[136,186,1982,2006]" pageId="4" pageNumber="763">Miri</collectingMunicipality>
,
<location id="181C75EF355BFFD880D3BD9EFE0B54C4" LSID="urn:lsid:plazi:treatment:956A9222355BFFDA811DBC3EFB8854EF:181C75EF355BFFD880D3BD9EFE0B54C4" box="[195,428,1982,2006]" country="Malaysia" municipality="Miri" name="Lambir National Park" pageId="4" pageNumber="763" stateProvince="Sarawak">Lambir National Park</location>
,
<location id="181C75EF355BFFD881A5BD9EFDF754C4" LSID="urn:lsid:plazi:treatment:956A9222355BFFDA811DBC3EFB8854EF:181C75EF355BFFD881A5BD9EFDF754C4" box="[437,592,1982,2006]" country="Malaysia" municipality="Miri" name="Tower Region" pageId="4" pageNumber="763" stateProvince="Sarawak">Tower Region</location>
, coll.
<collectorName id="B03646E2355BFFD8829CBD9EFCA554C7" box="[652,770,1982,2005]" pageId="4" pageNumber="763">T. Yumoto</collectorName>
,
<date id="697D05F4355BFFD88097BDFEFE9254E4" box="[135,309,2014,2038]" pageId="4" pageNumber="763" value="1995-08-20">
<collectingDate id="7939FC1C355BFFD88097BDFEFE9254E4" box="[135,309,2014,2038]" pageId="4" pageNumber="763" value="1995-08-20">20 August 1995</collectingDate>
</date>
(from
<taxonomicName id="DAC358B7355BFFD88196BDFEFDAC54E4" box="[390,523,2014,2038]" class="Magnoliopsida" family="Rubiaceae" genus="Myrmecodia" kingdom="Plantae" order="Gentianales" pageId="4" pageNumber="763" phylum="Tracheophyta" rank="genus">
<emphasis id="2FB7FF26355BFFD88196BDFEFDAC54E4" box="[390,523,2014,2038]" italics="true" pageId="4" pageNumber="763">Myrmecodia</emphasis>
</taxonomicName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355BFFD88353BA9DFA665247" pageId="4" pageNumber="763" type="diagnosis">
<paragraph id="1D7C2334355BFFD88353BA9DFA665247" blockId="4.[834,1474,189,341]" pageId="4" pageNumber="763">
<emphasis id="2FB7FF26355BFFD88353BA9DFBEB53C4" bold="true" box="[835,1100,189,214]" pageId="4" pageNumber="763">Diagnosis of worker.</emphasis>
Anterior portion of head with longitudinal rugulae. Mesonotum with lateral ridge in lateral view. Propodeal spine very long. In dorsal view petiole diamond-shaped, broadest at mid-length. Postpetiole weakly bilobed posteriorly, but without longitudinal median sulcus.
</paragraph>
</subSubSection>
<subSubSection id="55D970BF355BFFDA8353BB5EFE1D5B2F" lastPageId="6" lastPageNumber="765" pageId="4" pageNumber="763" type="description">
<paragraph id="1D7C2334355BFFD88353BB5EFAAB5124" blockId="4.[834,1476,381,566]" pageId="4" pageNumber="763">
<emphasis id="2FB7FF26355BFFD88353BB5EFC5E52A7" bold="true" pageId="4" pageNumber="763">
<heading id="46349458355BFFD88353BB5EFA665284" bold="true" box="[835,1473,381,406]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="763" reason="2">
Measurements and indices:
<typeStatus id="C2789D96355BFFD884B3BB5EFAB65284" box="[1187,1297,382,406]" pageId="4" pageNumber="763" type="holotype">Holotype</typeStatus>
and
<typeStatus id="C2789D96355BFFD88545BB5EFA665284" box="[1365,1473,382,406]" pageId="4" pageNumber="763" type="paratype">paratype</typeStatus>
</heading>
workers (n =15).
</emphasis>
HW 0.77–1.13; HL 0.71–1.07; CI 102–109; SL 0.66–0.88; SI 69–91; EL 0.17–0.24; PW 0.45–0.66; WL 0.77–1.21; PSL 0.24–0.48; PtL 0.21–0.35; PtW 0.22–0.39; PtH 0.14–0.22; PpL 0.13–0.25; PpW 0.20–0.32; PtHI 61–71; PtWI 99–115; PpWI 124–167; WI 80–92.
</paragraph>
<paragraph id="1D7C2334355BFFD88353B87DFC375187" blockId="4.[834,1474,605,661]" pageId="4" pageNumber="763">
<emphasis id="2FB7FF26355BFFD88353B87DFB325164" bold="true" box="[835,1173,605,630]" pageId="4" pageNumber="763">General description of worker.</emphasis>
Workers with polymorphism in size.
</paragraph>
<paragraph id="1D7C2334355BFFD88353B89DFAB050E7" blockId="4.[834,1476,701,1013]" pageId="4" pageNumber="763">Head broader than long in full-face view, posterior margin almost straight, posterior corner rounded in smaller worker, but angulate in larger workers. Frontal carina extending to mid-length of head. Occipital carina developed. Mandible with four teeth, apical and subapical teeth large. Anterior clypeal margin weakly concave in median portion. Compound eyes slightly projecting beyond lateral margins of head in full-face view. Scape exceeding posterior corner of head in smaller worker, but reaching posterior corner of head in larger worker. Antennal club 3-segmented.</paragraph>
<paragraph id="1D7C2334355BFFD88353BE3DFBBB5624" blockId="4.[834,1475,1053,1334]" pageId="4" pageNumber="763">Labrum broader than long; median cleft weakly developed; labral tubercle absent; distal area of the outer face with three to four stout setae. Maxilla with maxillary comb; transverse stipital groove absent; stipe subrectangular; maxillary palp 5-segmented; four apical segments longer than broad. Labium with sclerotised prementum on ventral surface; subglossal brush present; paraglossa with sensory peg at distal end; labial palp 3-segmented; apical segment longer than broad. Palp formula (5, 3).</paragraph>
<paragraph id="1D7C2334355BFFD88353BF7EFB315587" blockId="4.[834,1475,1373,1685]" pageId="4" pageNumber="763">Pronotum with weak ridges on dorsolateral margin. In lateral view, mesonotum posteriorly with indistinct longitudinal ridges. Mesonotum slightly higher than pronotum in lateral view. Metanotal groove straight in dorsal view, forming a deep concavity between mesonotum and propodeum. Metapleural gland opening slit-shaped. Propodeal spiracles dorsoventrally oval, its horizontal diameter slightly larger than vertical diameter, located on posterolateral corner of propodeum in lateral view. Propodeal spines long, directed posterolaterally in dorsal view.</paragraph>
<paragraph id="1D7C2334355BFFD88353BC9DFB795467" blockId="4.[834,1477,1725,1909]" pageId="4" pageNumber="763">Petiole diamond-shaped, broadest at mid-length in dorsal view; spiracle situated at midportion between dorsal and ventral margin of petiole in lateral view, directed laterally. Subpetiolar process undeveloped. Postpetiole weakly bilobed, but without longitudinal median sulcus in dorsal view. Petiole wider than postpetiole in dorsal view.</paragraph>
<paragraph id="1D7C2334355BFFDA8353BDBDFDF6558F" blockId="4.[834,1474,1949,2101]" lastBlockId="6.[136,775,1605,1693]" lastPageId="6" lastPageNumber="765" pageId="4" pageNumber="763">Sparsely hirsute with erect setae. Scape with abundant suberect to decumbent setae. Dorsal face of head with erect to suberect setae. Clypeus with suberect setae; one pair of longer setae directed medially on anteriormost portion. Anterior clypeal margin with two pairs of longer setae, mixed with shorter setae on the sides. Dorsum of promesonotum with long erect setae and short suberect setae. Fourth abdominal tergite with sparse erect to suberect setae.</paragraph>
<caption id="49BC73BC355AFFD98061B22DFCCA5B2D" ID-DOI="http://doi.org/10.5281/zenodo.5350246" ID-Zenodo-Dep="5350246" httpUri="https://zenodo.org/record/5350246/files/figure.png" pageId="5" pageNumber="764" startId="5.[113,148,2061,2083]" targetBox="[112,1453,193,2046]" targetPageId="5">
<paragraph id="1D7C2334355AFFD98061B22DFCCA5B2D" blockId="5.[113,1452,2061,2111]" pageId="5" pageNumber="764">
Fig. 3.
<taxonomicName id="DAC358B7355AFFD980AAB22DFE6A5B30" authority=", KUEC" authorityName="KUEC" box="[186,461,2061,2082]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="764" phylum="Arthropoda" rank="species" species="keris">
<emphasis id="2FB7FF26355AFFD980AAB22DFED85B30" box="[186,383,2061,2082]" italics="true" pageId="5" pageNumber="764">Crematogaster keris</emphasis>
, KUEC
</taxonomicName>
paratype worker from Malaysia (HW, 1.02; WL, 1.09). A, body in lateral view; B, full-face view of head; C, dorsal view of mesosoma; D, dorsal view of petiole and postpetiole.
</paragraph>
</caption>
<caption id="49BC73BC3559FFDA8097BF9FFD59551C" ID-DOI="http://doi.org/10.5281/zenodo.5350248" ID-Zenodo-Dep="5350248" httpUri="https://zenodo.org/record/5350248/files/figure.png" pageId="6" pageNumber="765" startId="6.[135,170,1471,1493]" targetBox="[137,1473,189,1459]" targetPageId="6">
<paragraph id="1D7C23343559FFDA8097BF9FFD59551C" blockId="6.[135,1475,1471,1550]" pageId="6" pageNumber="765">
Fig. 4. Mouthparts of
<taxonomicName id="DAC358B73559FFDA814DBF9FFE0E56C6" box="[349,425,1471,1492]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="765" phylum="Arthropoda" rank="species" species="keris">
<emphasis id="2FB7FF263559FFDA814DBF9FFE0E56C6" box="[349,425,1471,1492]" italics="true" pageId="6" pageNumber="765">C. keris</emphasis>
</taxonomicName>
worker. A, labrum in outer view; B, left maxilla in outer view; C, labium in lateral view. Abbreviations: Ca = cardo; Ga = galea; Gls = glossa; Hyp = hypopharynx; Lbp = labial palp; Mxco = maxillary comb; Mxp = maxillary palp; St = stipes; Prgls = paraglossa; Prm = prementum; Sglb = subglossal brush.
</paragraph>
</caption>
<paragraph id="1D7C23343559FFDA8098BCE5FD9954EF" blockId="6.[135,777,1733,2045]" pageId="6" pageNumber="765">Dorsal surface of head longitudinally rugose on anterior half, but relatively smooth on frons and posterior half; longitudinal rugulae surrounding antennal sockets and on gena. Clypeus with weak longitudinal rugulae. Pronotum striated with feeble rugulae anteriorly. Mesonotum weakly striated with feeble rugulae near lateral ridge. Lateral surface of pronotum smooth and shiny. Mesopleuron smooth in small worker, but weakly sculptured with feeble rugulae in large worker. Lateral surface of propodeum smooth and shiny. Propodeal dorsum and declivity smooth and shiny.</paragraph>
<paragraph id="1D7C23343559FFDA8098B205FE1D5B2F" blockId="6.[136,442,2085,2109]" box="[136,442,2085,2109]" pageId="6" pageNumber="765">Body colour reddish-brown.</paragraph>
</subSubSection>
<subSubSection id="55D970BF3559FFDA8353BC65FA9E55AE" pageId="6" pageNumber="765" type="distribution">
<paragraph id="1D7C23343559FFDA8353BC65FA9E55AE" blockId="6.[834,1476,1605,1725]" pageId="6" pageNumber="765">
<emphasis id="2FB7FF263559FFDA8353BC65FBD2554F" bold="true" box="[835,1141,1605,1629]" pageId="6" pageNumber="765">Distribution and biology.</emphasis>
This species is known from
<collectingCountry id="65D463A43559FFDA8353BC45FC02556F" box="[835,933,1637,1661]" name="Malaysia" pageId="6" pageNumber="765">Malaysia</collectingCountry>
(Peninsula, Borneo) (
<figureCitation id="85F83FB13559FFDA8484BC45FB73556F" box="[1172,1236,1637,1661]" captionStart="Fig" captionStartId="2.[136,171,1186,1208]" captionTargetBox="[137,1475,188,1170]" captionTargetId="figure-323@2.[134,1479,184,1174]" captionTargetPageId="2" captionText="Fig. 1. Distribution map of Crematogaster cornuta, C. keris, and C. ransonneti. Open circle = C. cornuta; open square = C. keris; open triangle = C. ransonneti." figureDoi="http://doi.org/10.5281/zenodo.5350240" httpUri="https://zenodo.org/record/5350240/files/figure.png" pageId="6" pageNumber="765">Fig. 1</figureCitation>
). The
<typeStatus id="C2789D963559FFDA850DBC45FAEC556F" box="[1309,1355,1637,1661]" pageId="6" pageNumber="765">type</typeStatus>
specimens were collected by canopy fogging and the Bornean specimen was collected from the ant-plant
<taxonomicName id="DAC358B73559FFDA84BEBC85FA9455AF" box="[1198,1331,1701,1725]" class="Magnoliopsida" family="Rubiaceae" genus="Myrmecodia" kingdom="Plantae" order="Gentianales" pageId="6" pageNumber="765" phylum="Tracheophyta" rank="genus">
<emphasis id="2FB7FF263559FFDA84BEBC85FA9455AF" box="[1198,1331,1701,1725]" italics="true" pageId="6" pageNumber="765">Myrmecodia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="55D970BF3559FFDA8353BCC5FB3C546F" pageId="6" pageNumber="765" type="etymology">
<paragraph id="1D7C23343559FFDA8353BCC5FB3C546F" blockId="6.[835,1476,1765,1917]" pageId="6" pageNumber="765">
<emphasis id="2FB7FF263559FFDA8353BCC5FC6055EF" bold="true" box="[835,967,1765,1789]" pageId="6" pageNumber="765">Etymology.</emphasis>
The species name ‘keris’ is directly adopted from the Malayan asymmetrical dagger of the same name, a traditional weapon of indigenous Malays, referring to the elongated propodeal spines of the species. The name is treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="55D970BF3559FFDA8353BD85FB8854EF" pageId="6" pageNumber="765" type="discussion">
<paragraph id="1D7C23343559FFDA8353BD85FB8854EF" blockId="6.[835,1473,1957,2045]" pageId="6" pageNumber="765">
<emphasis id="2FB7FF263559FFDA8353BD85FC2B54AF" bold="true" box="[835,908,1957,1981]" pageId="6" pageNumber="765">Notes.</emphasis>
This species is similar to
<taxonomicName id="DAC358B73559FFDA84D0BD85FA9F54AF" authorityName="Crawley" authorityYear="1924" box="[1216,1336,1957,1981]" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="765" phylum="Arthropoda" rank="species" species="cornuta">
<emphasis id="2FB7FF263559FFDA84D0BD85FA9F54AF" box="[1216,1336,1957,1981]" italics="true" pageId="6" pageNumber="765">C. cornuta</emphasis>
</taxonomicName>
, but can be distinguished by the smooth surface of posterior half of head and propodeum.
</paragraph>
</subSubSection>
</treatment>
</document>
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