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<document id="55BFC429883A1ED69F1B35E9EA0A4E4C" ID-CLB-Dataset="56755" ID-DOI="10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FFE46A64FF849CE11D55B972" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Philetor brachypterus" docType="treatment" docVersion="16" lastPageNumber="790" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="789" updateTime="1718993610009" updateUser="carolina">
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<mods:titleInfo id="6B77E1C4C7C55265F6B0FE09026F53F6">
<mods:title id="EA435D526FAC9664A169959A3E0820AE">Vespertilionidae</mods:title>
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<mods:name id="7F7594FA56366968ECC8DA4D5AAA1FCA" type="personal">
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<mods:namePart id="447E32B0AEAE50EF8701A4EE0E97B286">Don E. Wilson</mods:namePart>
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<mods:namePart id="063703D1A41D199913B5A6AFC4831A21">Russell A. Mittermeier</mods:namePart>
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<mods:publisher id="72172691A8A13337E467CFAF6CF5BFBC">Lynx Edicions</mods:publisher>
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<mods:title id="B7518F8EB6594989FF5C8ADD7024EBBD">Handbook of the Mammals of the World – Volume 9 Bats</mods:title>
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<mods:classification id="E73AD4AD3F1CC2583C70B5862D27D5E6">book chapter</mods:classification>
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<treatment id="4C3D87E8FFE46A64FF849CE11D55B972" ID-DOI="http://doi.org/10.5281/zenodo.6581228" ID-GBIF-Taxon="195628099" ID-Zenodo-Dep="6581228" LSID="urn:lsid:plazi:treatment:4C3D87E8FFE46A64FF849CE11D55B972" httpUri="http://treatment.plazi.org/id/4C3D87E8FFE46A64FF849CE11D55B972" lastPageId="32" lastPageNumber="790" pageId="31" pageNumber="789">
<subSubSection id="8C8E6575FFE46A5BFF849CE11EB1B1D4" box="[120,177,2744,2790]" pageId="31" pageNumber="789" type="multiple">
<paragraph id="C42B36FEFFE46A5BFF849CE11EB1B1D4" blockId="31.[118,1168,2744,2874]" box="[120,177,2744,2790]" pageId="31" pageNumber="789">
<heading id="9F638192FFE46A5BFF849CE11EB1B1D4" box="[120,177,2744,2790]" pageId="31" pageNumber="789">
<figureCitation id="5CAF2A7BFFE46A5BFF849CE11EB1B1D4" box="[120,177,2744,2790]" captionStart="Plate 57: Vespertilionidae" captionStartId="27.[123,155,3200,3221]" captionTargetBox="[12,2751,19,3659]" captionTargetPageId="26" captionText="49. Eurasian Particolored Bat (Vespertilio murinus), 50. Asian Particolored Bat (Vespertilio sinensis), 51. Pygmy Bamboo Bat (Tylonycteris pygmaea), 52. Indomalayan Lesser Bamboo Bat (Tylonycteris fulvida), 53. Sunda Lesser Bamboo Bat (Tylonycteris pachypus), 54. Tonkin Greater Bamboo Bat (Tylonycteris tonkinensis), 55. Malayan Greater Bamboo Bat (Tylonycteris malayana), 56. Sumatran Greater Bamboo Bat (Tylonycteris robustula), 57. Yok Don Helmeted Bat (Cassistrellus yokdonensis), 58. Surat Helmeted Bat (Cassustrellus dimissus), 59. Rohu’s Bat (Phuletor brachypterus), 60. Western False Pipistrelle (Falsistrellus mackenziei), 61. Eastern False Pipistrelle (Falsistrellus tasmaniensis), 62. Yellow-lipped Cave Bat (Vespadelus douglasorum), 63. Northern Cave Bat (Vespadelus caurinus), 64. Finlayson’s Cave Bat (Vespadelus finlaysoni), 65. Eastern Cave Bat (Vespadelus troughtoni), 66. Inland Forest Bat (Vespadelus baverstocki), 67. Eastern Forest Bat (Vespadelus pumilus), 68. Little Forest Bat (Vespadelus vulturnus), 69. Large Forest Bat (Vespadelus darlingtoni), 70. Southern Forest Bat (Vespadelus regulus), 71. Large-eared Pied Bat (Chalinolobus dwyeri), 72. Little Pied Bat (Chalinolobus picatus), 73. Hoary Wattled Bat (Chalinolobus nigrogriseus), 74. Gould’s Wattled Bat (Chalinolobus gouldii), 75. New Caledonian Wattled Bat (Chalinolobus neocaledonicus), 76. Chocolate Wattled Bat (Chalinolobus morio), 77. New Zealand Long-tailed Bat (Chalinolobus tuberculatus)" figureDoi="http://doi.org/10.5281/zenodo.6398506" httpUri="https://zenodo.org/record/6398506/files/figure.png" pageId="31" pageNumber="789">59.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFF3E9CE11FA6B1D4" box="[194,422,2744,2790]" pageId="31" pageNumber="789" type="vernacular_names">
<paragraph id="C42B36FEFFE46A5BFF3E9CE11FA6B1D4" blockId="31.[118,1168,2744,2874]" box="[194,422,2744,2790]" pageId="31" pageNumber="789">
<heading id="9F638192FFE46A5BFF3E9CE11FA6B1D4" box="[194,422,2744,2790]" pageId="31" pageNumber="789">
<vernacularName id="4A9746D0FFE46A5BFF3E9CE11FA6B1D4" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[194,422,2744,2790]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="eng" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Rohu’s Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFE1B9CE11D5CB1D4" box="[487,860,2744,2790]" pageId="31" pageNumber="789" type="nomenclature">
<paragraph id="C42B36FEFFE46A5BFE1B9CE11D5CB1D4" blockId="31.[118,1168,2744,2874]" box="[487,860,2744,2790]" pageId="31" pageNumber="789">
<heading id="9F638192FFE46A5BFE1B9CE11D5CB1D4" box="[487,860,2744,2790]" pageId="31" pageNumber="789">
<taxonomicName id="03944D7DFFE46A5BFE1B9CE11D5CB1D4" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[487,860,2744,2790]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">
<emphasis id="F6E0EAECFFE46A5BFE1B9CE11D5CB1D4" box="[487,860,2744,2790]" italics="true" pageId="31" pageNumber="789">Philetor brachypterus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFF8B9CA01A8FB007" pageId="31" pageNumber="789" type="vernacular_names">
<paragraph id="C42B36FEFFE46A5BFF8B9CA01A7FB03C" blockId="31.[118,1168,2744,2874]" box="[119,1151,2809,2830]" pageId="31" pageNumber="789">
<heading id="9F638192FFE46A5BFF8B9CA01A7FB03C" box="[119,1151,2809,2830]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFF8B9CA01EC3B03C" bold="true" box="[119,195,2809,2830]" pageId="31" pageNumber="789">French:</emphasis>
<vernacularName id="4A9746D0FFE46A5BFF369CA01FBBB03C" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[202,443,2809,2830]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="fra" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Vespertilion brachyptere</vernacularName>
/
<emphasis id="F6E0EAECFFE46A5BFE2C9CA01C2AB03C" bold="true" box="[464,554,2809,2830]" pageId="31" pageNumber="789">German:</emphasis>
<vernacularName id="4A9746D0FFE46A5BFDC99CA01CE4B03C" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[565,740,2809,2830]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="deu" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Rohu-Fledermaus</vernacularName>
/
<emphasis id="F6E0EAECFFE46A5BFD059CA01D54B03C" bold="true" box="[761,852,2809,2830]" pageId="31" pageNumber="789">Spanish:</emphasis>
<vernacularName id="4A9746D0FFE46A5BFCA79CA01A7FB03C" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[859,1151,2809,2830]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="esp" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">
<vernacularName id="4A9746D0FFE46A5BFCA79CA01DEEB03C" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[859,1006,2809,2830]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="esp" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Vespertilionido</vernacularName>
de alas cortas
</vernacularName>
</heading>
</paragraph>
<paragraph id="C42B36FEFFE46A5BFF8B9D791A8FB007" blockId="31.[118,1168,2744,2874]" box="[119,1167,2848,2869]" pageId="31" pageNumber="789">
<heading id="9F638192FFE46A5BFF8B9D791A8FB007" box="[119,1167,2848,2869]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFF8B9D791F6FB007" bold="true" box="[119,367,2848,2869]" pageId="31" pageNumber="789">Other common names:</emphasis>
<vernacularName id="4A9746D0FFE46A5BFE849D791C83B007" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[376,643,2848,2869]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="eng" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Narrow-winged Brown Bat</vernacularName>
,
<vernacularName id="4A9746D0FFE46A5BFD6D9D791D93B007" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[657,915,2848,2869]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="eng" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Narrow-winged Pipistrelle</vernacularName>
,
<vernacularName id="4A9746D0FFE46A5BFC5D9D791A8FB007" ID-CoL="76XYZ" authority="(Temminck, 1840)" baseAuthorityName="Temminck" baseAuthorityYear="1840" box="[929,1167,2848,2869]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" language="eng" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Short-winged Pipistrelle</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFD219D3E1D95B09E" pageId="31" pageNumber="789" type="reference_group">
<paragraph id="C42B36FEFFE46A5BFD219D3E1D95B09E" blockId="31.[732,1325,2919,3342]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFD219D3E1D78B0B6" bold="true" box="[733,888,2919,2948]" pageId="31" pageNumber="789">Taxonomy.</emphasis>
<taxonomicName id="03944D7DFFE46A5BFC669D3E1D90B09E" ID-CoL="7G2SW" authority="Temminck, 1840" authorityName="Temminck" authorityYear="1840" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Vespertilio brachypterus Temminck, 1840</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFC599DD61DD2B0E1" pageId="31" pageNumber="789" type="materials_examined">
<paragraph id="C42B36FEFFE46A5BFC599DD61DD2B0E1" blockId="31.[732,1325,2919,3342]" pageId="31" pageNumber="789">
<materialsCitation id="74FC3CA3FFE46A5BFC599DD61DD2B0E1" ID-GBIF-Occurrence="3783319370" pageId="31" pageNumber="789">
“le district de Padang,”
<collectingRegion id="0650F81CFFE46A5BFAFE9DD61D30B0E1" country="Indonesia" name="Sumatera Utara" pageId="31" pageNumber="789">Sumatra</collectingRegion>
,
<collectingCountry id="BC83766EFFE46A5BFCBD9DEB1DCEB0E1" box="[833,974,2994,3027]" name="Indonesia" pageId="31" pageNumber="789">Indonesia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFD209D80170BB803" pageId="31" pageNumber="789" type="discussion">
<paragraph id="C42B36FEFFE46A5BFD209D80170BB803" blockId="31.[732,1325,2919,3342]" lastBlockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
A close relationship of
<taxonomicName id="03944D7DFFE46A5BFBEA9D801A77B0C8" authorityName="Thomas" authorityYear="1902" box="[1046,1143,3033,3066]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Philetor</taxonomicName>
to
<taxonomicName id="03944D7DFFE46A5BFB5E9D801B2DB0C8" authorityName="Peters" authorityYear="1872" box="[1186,1325,3033,3066]" class="Mammalia" family="Vespertilionidae" genus="Tylonycteris" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Tylonycteris</taxonomicName>
was already suspected by some earlier authors; this was confirmed by the DNA-barcode study of C. M. Francis and colleagues in 2010, and in subsequent phylogenetic analyses. More recent phylogenetic studies by M. Ruedi and colleagues in 2017 and T. Gorfol and colleagues in 2018 and 2019 show
<taxonomicName id="03944D7DFFE46A5BFF3E9B4C1F23B604" authorityName="Thomas" authorityYear="1902" box="[194,291,3349,3382]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Philetor</taxonomicName>
to be closely related to
<taxonomicName id="03944D7DFFE46A5BFD9D9B4C1CF5B604" box="[609,757,3349,3382]" class="Mammalia" family="Vespertilionidae" genus="Cassistrellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Cassistrellus</taxonomicName>
and
<taxonomicName id="03944D7DFFE46A5BFCC79B4C1DC7B604" authorityName="Peters" authorityYear="1872" box="[827,967,3349,3382]" class="Mammalia" family="Vespertilionidae" genus="Tylonycteris" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Tylonycteris</taxonomicName>
, the three forming a separate clade. The genus Philetorwas long considered to include only one named form, P. rohui, described by O. Thomas in 1902 from New
<collectingCountry id="BC83766EFFE46A5BFCF99B311D6BB6B7" box="[773,875,3432,3461]" name="Guinea-Bissau" pageId="31" pageNumber="789">Guinea</collectingCountry>
. In 1966, J. E. Hill demonstrated that the Peninsular Malaysian form described by F. N. Chasen in 1940 as
<taxonomicName id="03944D7DFFE46A5BF6AC974917C2BA03" box="[2384,2498,272,305]" class="Mammalia" family="Vespertilionidae" genus="Eptesicus" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Eptesicus</taxonomicName>
verecundus also belonged to
<taxonomicName id="03944D7DFFE46A5BF966971918F9BA6B" authorityName="Thomas" authorityYear="1902" box="[1690,1785,320,345]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Philetor</taxonomicName>
, and he treated it as a race of P. hui. In 1971, Hill showed that
<taxonomicName id="03944D7DFFE46A5BFA45970618D1BAB2" authorityName="Temminck" authorityYear="1840" box="[1465,1745,351,384]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">Vespertilio brachypterus</taxonomicName>
of Sumatra, at that time classified as a species of Pipustrellus, was also referable to
<taxonomicName id="03944D7DFFE46A5BF99997D218C4BA9A" authorityName="Thomas" authorityYear="1902" box="[1637,1732,395,424]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Philetor</taxonomicName>
, he treated it as a third race, and as it was the oldest name, it became the correct name for the species. Despite Hill's recognition of three subspecies, the actual pattern of variation has proved too complex to fit this taxonomy;it had been suggested that variations in the shape of the narial emargination (Uor V-shaped gap) justified the subspecies, but in 1983 K. F. Koopman regarded this feature as unreliable. In 2017, V. C. Lim and colleagues performed NJ analyses of the barcoding gene, and suggested cryptic species among Peninsular Malaysian and/or Bornean specimens. In a morphological study in 2017, U. Saikia and colleagues reidentified specimens supposedly of
<taxonomicName id="03944D7DFFE46A5BFA2194981898B9D0" authorityName="Temminck" authorityYear="1840" box="[1501,1688,705,738]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="brachypterus">P. brachypterus</taxonomicName>
from
<collectingCountry id="BC83766EFFE46A5BF9119498193FB9D0" box="[1773,1855,705,738]" name="Nepal" pageId="31" pageNumber="789">Nepal</collectingCountry>
and Sikkim as
<taxonomicName id="03944D7DFFE46A5BF7EF949816CEB9D0" baseAuthorityName="Thomas" baseAuthorityYear="1915" box="[2067,2254,705,738]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="joffrei">Hypsugo joffrei</taxonomicName>
; morphological data suggest that
<taxonomicName id="03944D7DFFE46A5BFA0994B01856B838" authorityName="Thomas" authorityYear="1902" box="[1525,1622,745,778]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Philetor</taxonomicName>
has a close evolutionary relationship with
<taxonomicName id="03944D7DFFE46A5BF75E94B0172EB838" authorityName="Peters" authorityYear="1872" box="[2210,2350,745,778]" class="Mammalia" family="Vespertilionidae" genus="Tylonycteris" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Tylonycteris</taxonomicName>
and
<taxonomicName id="03944D7DFFE46A5BF68994B017E6B838" baseAuthorityName="Thomas" baseAuthorityYear="1915" box="[2421,2534,745,778]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="species" species="joffrei">H. joffrei</taxonomicName>
. The internal taxonomy ofthis species is in need of revision. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA8B95611610BF2C" pageId="31" pageNumber="789" type="distribution">
<caption id="90EB6676FFE46A5BFA8B95611610BF2C" ID-DOI="http://doi.org/10.5281/zenodo.6397896" ID-Zenodo-Dep="6397896" httpUri="https://zenodo.org/record/6397896/files/figure.png" inLine="true" pageId="31" pageNumber="789" targetBox="[117,709,2926,3341]" targetPageId="31">
<paragraph id="C42B36FEFFE46A5BFA8B95611610BF2C" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA8B95611827B86B" bold="true" box="[1399,1575,824,857]" pageId="31" pageNumber="789">Distribution.</emphasis>
<collectingCountry id="BC83766EFFE46A5BF9CA956118D4B86B" box="[1590,1748,824,857]" name="Philippines" pageId="31" pageNumber="789">Philippines</collectingCountry>
(S Luzon,
<collectingRegion id="0650F81CFFE46A5BF88295611636B86B" box="[1918,2102,824,857]" country="Philippines" name="Catanduanes" pageId="31" pageNumber="789">Catanduanes</collectingRegion>
, Panay, Samar,
<collectingRegion id="0650F81CFFE46A5BF6E095611769B86B" box="[2332,2409,824,857]" country="Philippines" name="Southern Leyte" pageId="31" pageNumber="789">Leyte</collectingRegion>
, Negros, and Mindanao Is), Peninsular
<collectingCountry id="BC83766EFFE46A5BF90B953A196FB8B2" box="[1783,1903,867,896]" name="Malaysia" pageId="31" pageNumber="789">Malaysia</collectingCountry>
, CW Sumatra, N &amp; NW Borneo, Sulawesi, New
<collectingCountry id="BC83766EFFE46A5BFA8B95DF1BDEB895" box="[1399,1502,902,935]" name="Guinea-Bissau" pageId="31" pageNumber="789">Guinea</collectingCountry>
and the
<collectingCountry id="BC83766EFFE46A5BF99B95DF19A0B895" box="[1639,1952,902,935]" name="Papua New Guinea" pageId="31" pageNumber="789">Bismarck Archipelago</collectingCountry>
(
<collectingRegion id="0650F81CFFE46A5BF84695DF166CB895" box="[1978,2156,902,935]" country="Papua New Guinea" name="New Ireland" pageId="31" pageNumber="789">New Ireland</collectingRegion>
and
<collectingRegion id="0650F81CFFE46A5BF74095DF1765B895" box="[2236,2405,902,935]" country="Papua New Guinea" name="West New Britain" pageId="31" pageNumber="789">New Britain</collectingRegion>
Is). A record from Java is erroneous, and a record from Bangka I does not appear to be authentic. Previous records from
<collectingCountry id="BC83766EFFE46A5BF94B95831901B8C5" box="[1719,1793,986,1015]" name="India" pageId="31" pageNumber="789">India</collectingCountry>
(
<collectingRegion id="0650F81CFFE46A5BF8EA95831979B8C5" box="[1814,1913,986,1015]" country="India" name="Sikkim" pageId="31" pageNumber="789">Sikkim</collectingRegion>
) and C &amp; E
<collectingCountry id="BC83766EFFE46A5BF7D69583167DB8C5" box="[2090,2173,986,1015]" name="Nepal" pageId="31" pageNumber="789">Nepal</collectingCountry>
are misidentifications, now attributable to Joftre’s Pipistrelle (
<taxonomicName id="03944D7DFFE46A5BF8C795A419ADBF2C" box="[1851,1965,1021,1054]" class="Mammalia" family="Vespertilionidae" genus="Hypsugo" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Hypsugo</taxonomicName>
joffrer).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA8A9270173AB38D" pageId="31" pageNumber="789" type="description">
<paragraph id="C42B36FEFFE46A5BFA8A9270173AB38D" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA8A92701870BF74" bold="true" box="[1398,1648,1065,1094]" pageId="31" pageNumber="789">Descriptive notes.</emphasis>
Head-body 52-64 mm, tail 30-38 mm, ear 13-16 mm, hindfoot 8-10 mm, forearm 30-38 mm; weight 8-13 g. Males are larger than females in almost all dimensions. Fur is short and dense, the upperparts reddish to dark brown, with pale gray-brown bases and dark tips; underparts are paler, red brown to gray brown. Head is ratherflat, with no noticeable forehead. Sparsely haired muzzle is short and broad, and appears swollen. Nostrils are widely separated. Ears are short, triangular and of moderate size; posterior margin attaches to underside of throat; there is a small antitragal lobe at base of anterior margin; inner surface of ear sparsely haired; tragusis short, blunt, and fleshy. Eyes are minute, with no facial hairs. Wings are narrow, with fifth digit shortened, its tip reaching only to halfway along first phalanx of fourth finger; these are attached to the lower end of the tibiae. Males and females have unusually elaborate genitalia: elongated penis of male lacks a prepuce, and has a cushion-like, bristly pad at end of main shaft, followed by a narrow shaft supporting a broadened triangular structure; vulva of female is very small and is longitudinally oriented, rather than horizontally as in other bats, and there are five separate pads around the vulva; particularly striking is the deep dark brown color of part of the penis tip in
<taxonomicName id="03944D7DFFE46A5BF835902D1628BDA7" authorityName="Thomas" authorityYear="1902" box="[1993,2088,1652,1685]" class="Mammalia" family="Vespertilionidae" genus="Philetor" kingdom="Animalia" order="Chiroptera" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Philetor</taxonomicName>
, a feature not described before. Skull is short and somewhat spherical in appearance, with conspicuous peg-like projections over anterior edge of orbit. Skull slopes evenly in profile; rostrum is short, deep and broad. The indistinct supraorbital ridges terminate in very well-developed supraorbital processes; basioccipital pits are deep; braincase bulbous; sagittal crest and lambdoid ridges are slightly developed. Zygomata are extremely weak; interorbital region is broad. Dental formulais12/3,C1/1,P 1/2, M 3/3 (x2) = 32. I? is elongated and narrow basally with two cusps (bicuspidate), I” is smaller and conical, and separated from C'by a narrow diastema; C' has broad anterior face and well-defined secondary cusp posteriorly; P? is absent. There is a known case of an individual from Sulawesi (Bogani Nani Wartabone National Park) possessing a small P?; more material is needed to evaluate this unusual feature. P* is short,its length about half its width; crowded between C' and M'. Lower incisors are tricuspidate and separated from each other; lower molars are nyctalodont. Condylo-basal length 12-9-15 mm; maxillary tooth row 4-6-5 mm. Chromosomal complement has 2n = 26 and FN = 48 (Ulu Gombak, Peninsular
<collectingCountry id="BC83766EFFE46A5BF74F9EC7172DB38D" box="[2227,2349,2206,2239]" name="Malaysia" pageId="31" pageNumber="789">Malaysia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA889E9C189AB2C9" pageId="31" pageNumber="789" type="biology_ecology">
<paragraph id="C42B36FEFFE46A5BFA889E9C189AB2C9" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA889E9C1BE4B3D4" bold="true" box="[1396,1508,2245,2278]" pageId="31" pageNumber="789">Habitat.</emphasis>
Both pristine forest and disturbed areas. In the
<collectingCountry id="BC83766EFFE46A5BF75B9E9C1747B3D4" box="[2215,2375,2245,2278]" name="Philippines" pageId="31" pageNumber="789">Philippines</collectingCountry>
, the species occurs in primary and disturbed lowland forests from sea level to
<quantity id="036C9B1BFFE46A5BF6FB9EB41770B23C" box="[2311,2416,2285,2318]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="31" pageNumber="789" unit="m" value="1450.0">1450 m</quantity>
. In
<collectingCountry id="BC83766EFFE46A5BF6579EB41422B23C" box="[2475,2594,2285,2318]" name="Malaysia" pageId="31" pageNumber="789">Malaysia</collectingCountry>
, it can be found in peat-swamp forest. Elsewhere in the region it has been recorded in degraded forest adjacent to good lowland and montane forest. On Mindanao (Mount Kitanglad) netted in disturbed lowland forest at
<quantity id="036C9B1BFFE46A5BF7EA9F3A166CB2B6" box="[2070,2156,2403,2436]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="789" unit="m" value="800.0">800 m</quantity>
, and in residual montane forest at
<quantity id="036C9B1BFFE46A5BFA609FD21805B29E" box="[1436,1541,2443,2476]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="31" pageNumber="789" unit="m" value="1450.0">1450 m</quantity>
. In New
<collectingCountry id="BC83766EFFE46A5BF9709FD218F3B29E" box="[1676,1779,2443,2476]" name="Guinea-Bissau" pageId="31" pageNumber="789">Guinea</collectingCountry>
found in primary and disturbed tropical moist forests. It can be found in coconut plantations adjacent to forest. It ranges from sea level up to elevations of
<quantity id="036C9B1BFFE46A5BF9D29F831896B2C9" box="[1582,1686,2522,2555]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="31" pageNumber="789" unit="m" value="2100.0">2100 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA889C5B183EB1AA" pageId="31" pageNumber="789" type="food_feeding">
<paragraph id="C42B36FEFFE46A5BFA889C5B183EB1AA" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA889C5B1884B111" bold="true" box="[1396,1668,2562,2595]" pageId="31" pageNumber="789">Food and Feeding.</emphasis>
Rohu’s Bat appears to forage in relatively open areas and typically flies at high speed. It forages in the forest understory and subcanopy, and over grassland. Stomachs of three specimens contained insects including
<taxonomicName id="03944D7DFFE46A5BF6B09C0817E9B140" box="[2380,2537,2641,2674]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="789" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
and
<taxonomicName id="03944D7DFFE46A5BFA889C2E183AB1AA" box="[1396,1594,2679,2712]" class="Insecta" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="789" phylum="Arthropoda" rank="order">Hymenoptera</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA889CF91427B022" pageId="31" pageNumber="789" type="breeding">
<paragraph id="C42B36FEFFE46A5BFA889CF91427B022" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA889CF91BFAB1F3" bold="true" box="[1396,1530,2720,2753]" pageId="31" pageNumber="789">Breeding.</emphasis>
Pregnant females with a single embryo were found from late May to mid-June in Borneo and March/April in Peninsular
<collectingCountry id="BC83766EFFE46A5BF7D79C9E16A3B1DA" box="[2091,2211,2759,2792]" name="Malaysia" pageId="31" pageNumber="789">Malaysia</collectingCountry>
. On Mindanao (Mount Kitanglad) an adult female netted in April was pregnant with a single near-term embryo.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA8E9D4F19F3B779" pageId="31" pageNumber="789" type="activity">
<paragraph id="C42B36FEFFE46A5BFA8E9D4F19F3B779" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA8E9D4F185EB005" bold="true" box="[1394,1630,2838,2871]" pageId="31" pageNumber="789">Activity patterns.</emphasis>
Rohu’s Bat leaves the day roost at dusk. It roosts in tree hollows and holes 1-5-4-5 m aboveground. Bats sometimes roost in coconut palms in New
<collectingCountry id="BC83766EFFE46A5BF6479D671423B06D" box="[2491,2595,2878,2911]" name="Guinea-Bissau" pageId="31" pageNumber="789">Guinea</collectingCountry>
, but not in South Asia. In
<collectingCountry id="BC83766EFFE46A5BF9229D3C19EFB0B4" box="[1758,2031,2917,2950]" name="Papua New Guinea" pageId="31" pageNumber="789">Papua New Guinea</collectingCountry>
, one specimen was taken from a hollow tree that had previously been occupied by a Lesser Tree Mouse (
<taxonomicName id="03944D7DFFE46A5BF6CE9DD517C8B09F" authorityName="Thomas" authorityYear="1888" box="[2354,2504,2956,2989]" class="Mammalia" family="Muridae" genus="Chiruromys" kingdom="Animalia" order="Rodentia" pageId="31" pageNumber="789" phylum="Chordata" rank="genus">Chiruromys</taxonomicName>
vates). On
<collectingRegion id="0650F81CFFE46A5BFA509DED1856B0E7" box="[1452,1622,2996,3029]" country="Papua New Guinea" name="West New Britain" pageId="31" pageNumber="789">New Britain</collectingRegion>
, most bats were found in hollow trees in rainforest. There appear to be no roosting records from caves. Echolocation call from Mount Makiling, Laguna Province, Luzon, had a mean frequency of 30-6 kHz. In New
<collectingCountry id="BC83766EFFE46A5BF7379A5A1731B716" box="[2251,2353,3075,3108]" name="Guinea-Bissau" pageId="31" pageNumber="789">Guinea</collectingCountry>
heard making an insect-like call from a hole in a coconut palm.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA8F9A0B176FB7DB" pageId="31" pageNumber="789" type="biology_ecology">
<paragraph id="C42B36FEFFE46A5BFA8F9A0B176FB7DB" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA8F9A0B164DB741" bold="true" box="[1395,2125,3154,3187]" pageId="31" pageNumber="789">Movements, Home range and Social organization.</emphasis>
Rohu’s Bats forage in relatively open areas and typically fly at high speed, having correspondingly narrow, stubby wings. Up to 55 individuals have been collected from a single tree. On
<collectingRegion id="0650F81CFFE46A5BF68B9AF91422B7F3" box="[2423,2594,3232,3265]" country="Papua New Guinea" name="West New Britain" pageId="31" pageNumber="789">New Britain</collectingRegion>
, bats were found in mixed groups or segregated male and female groups.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFE46A5BFA8F9AB61737B6B5" pageId="31" pageNumber="789" type="conservation">
<paragraph id="C42B36FEFFE46A5BFA8F9AB61737B6B5" blockId="31.[1389,2606,272,3472]" pageId="31" pageNumber="789">
<emphasis id="F6E0EAECFFE46A5BFA8F9AB618D3B622" bold="true" box="[1395,1747,3311,3344]" pageId="31" pageNumber="789">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List in view of its wide distribution and expected large population. Rohu’s Bat occurs in a number of protected areas, and tolerates a degree of habitat modification. There appear to be no major threats. Surveys and research are needed overits entire range.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFDB6A64FF53974E1D55B972" pageId="32" pageNumber="790" type="bibRefCitation_list">
<paragraph id="C42B36FEFFDB6A64FF53974E1D55B972" blockId="32.[174,1379,279,580]" pageId="32" pageNumber="790">
<emphasis id="F6E0EAECFFDB6A64FF53974E1F48BA1E" bold="true" box="[175,328,279,300]" pageId="32" pageNumber="790">Bibliography.</emphasis>
Amador et al. (2018), Bates &amp; Harrison (1997), Boitani et al. (2006), Bonaccorso (1998), Brass (1964), Chasen (1940), Corbet &amp; Hill (1992), Flannery (1995a, 1995b), Francis (2008a), Francis et al. (2010), Gor fol &amp; Csorba (2018), Gorfol et al. (2019), Heaney, Balete, Alviola et al. (2012), Heaney, Balete, Dolar et al. (1998), Heaney, Balete &amp; Rickart (2016), Heaney, Tabaranza et al. (2006), Heller &amp; Volleth (1989), Hill (1966, 1971b), Hill &amp; Harrison (1987), Hill &amp; Rozendaal (1989), Kock (1981), Koopman (1982b, 1983), Lim, B.L. et al. (1972), Lim, V.C. et al. (2017),
<collectingRegion id="0650F81CFFDB6A64FE8D97811FCEBAC3" box="[369,462,472,497]" country="United Kingdom" name="Medway" pageId="32" pageNumber="790">Medway</collectingRegion>
(1983), Miller (1907), Payne et al. (1985), Phillipps &amp; Phillipps (2016), Rosell-Ambal, Tabaranza, Bonaccorso et al. (2008), Ruedi,
<collectingRegion id="0650F81CFFDB6A64FD6397A61CCFB92A" box="[671,719,511,536]" country="Hungary" name="Eger" pageId="32" pageNumber="790">Eger</collectingRegion>
et al. (2017), Saikia et al. (2017), Sediock (2001), Simmons (2005), Tate (1942b), Thomas (1902b), Volleth &amp; Heller (1994a).
</paragraph>
</subSubSection>
</treatment>
</document>
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