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<document id="602E50C3BCCB3CE2CFDA929150CE69AC" ID-CLB-Dataset="56755" ID-DOI="10.5281/zenodo.6397752" ID-GBIF-Dataset="45351c32-25dd-422c-bdb2-00e73deb4943" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6397752" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1648655544658" checkinUser="conny" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="4C3D87E8FFBE6A02FA879B4E19A7BFFD" docLanguage="en" docName="hbmw_9_Vespertilionidae_716.pdf.imf" docOrigin="Handbook of the Mammals of the World – Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Arielulus cuprosus Francis &amp; Hill 1984" docType="treatment" docVersion="15" lastPageNumber="828" masterDocId="B004FF90FFFB6A44FFFC96591E00BB32" masterDocTitle="Vespertilionidae" masterLastPageNumber="981" masterPageNumber="716" pageNumber="827" updateTime="1718993610009" updateUser="carolina">
<mods:mods id="83FEEE197CCB16840595E13E6FC07B2D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4E8F0C6DC9F1E3B76BDB47F5F0FAAAEE">
<mods:title id="942685829ED9A37AA31AE0612FC7E61C">Vespertilionidae</mods:title>
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<mods:namePart id="5E1E2DA032B7EDFAB1CD07350E5E70FE">Don E. Wilson</mods:namePart>
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<mods:namePart id="E5DCC80F4796EDCAF4B23BB6615BD60D">Russell A. Mittermeier</mods:namePart>
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<mods:dateOther id="19508ED55EA63A472694F052EDD54F31" type="pubDate">2019-10-31</mods:dateOther>
<mods:publisher id="4CD378950A17B180C7D1379EEAAED7D2">Lynx Edicions</mods:publisher>
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<mods:title id="97C9FDE52CF6192789B49FA37987559B">Handbook of the Mammals of the World – Volume 9 Bats</mods:title>
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<treatment id="4C3D87E8FFBE6A02FA879B4E19A7BFFD" ID-DOI="http://doi.org/10.5281/zenodo.6578484" ID-GBIF-Taxon="195628478" ID-Zenodo-Dep="6578484" LSID="urn:lsid:plazi:treatment:4C3D87E8FFBE6A02FA879B4E19A7BFFD" httpUri="http://treatment.plazi.org/id/4C3D87E8FFBE6A02FA879B4E19A7BFFD" lastPageId="70" lastPageNumber="828" pageId="69" pageNumber="827">
<subSubSection id="8C8E6575FFBE6A01FA879B4E1BCBB677" box="[1403,1483,3351,3397]" pageId="69" pageNumber="827" type="multiple">
<paragraph id="C42B36FEFFBE6A01FA879B4E1BCBB677" blockId="69.[1399,2427,3351,3477]" box="[1403,1483,3351,3397]" pageId="69" pageNumber="827">
<heading id="9F638192FFBE6A01FA879B4E1BCBB677" box="[1403,1483,3351,3397]" pageId="69" pageNumber="827">
<figureCitation id="5CAF2A7BFFBE6A01FA879B4E1BCBB677" box="[1403,1483,3351,3397]" captionStart="Plate 60: Vespe" captionStartId="61.[119,149,3300,3325]" captionTargetBox="[12,2753,18,3658]" captionTargetPageId="60" captionText="118. Cape Serotine (Neoromicia capensis), 119. Malagasy Serotine (Neoromicia matroka), 120. Isalo Serotine (Neoromicia malagasyensis), 121. Roberts’s Serotine (Neoromicia robertsi), 122. Stanley’s Serotine (Neoromicia stanleyi), 123. Zulu Serotine (Neoromicia zuluensis), 124. Tiny Serotine (Neoromicia guineensis), 125. Somali Serotine (Neoromicia somalica), 126. Banana Serotine (Neoromicia nanus), 127. Rendall’s Serotine (Neoromicia rendalli), 128. Dark-brown Serotine (Neoromicia brunnea), 129. Rosevear’s Serotine (Neoromicia roseveari), 130. Isabelline Serotine (Neoromicia isabella), 131. Grandidier’s Serotine (Neoromicia grandidieri), 132. White-winged Serotine (Neoromicia tenuipinnis), 133. Heller’s Serotine (Neoromicia helios), 134. De Winton’s LLong-eared Bat (Laephotis wintoni), 135. Angolan Long-eared Bat (Laephotis angolensis), 136. Botswanan Long-eared Bat (Laephotis botswanae), 137. Namib Long-eared Bat (Laephotis namibensis), 138. Sind Bat (Rhyneptesicus nasutus), 139. Collared Sprite (Thainycteris aureocollaris), 140. Necklace Sprite (Thainycleris torquatus), 141. Bronze Sprite (Arielulus circumdatus), 142. Social Sprite (Arelulus societatis), 143. Coppery Sprite (Arielulus cuprosus)" figureDoi="http://doi.org/10.5281/zenodo.6398519" httpUri="https://zenodo.org/record/6398519/files/figure.png" pageId="69" pageNumber="827">143.</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBE6A01FA219B4E1917B677" box="[1501,1815,3351,3397]" pageId="69" pageNumber="827" type="vernacular_names">
<paragraph id="C42B36FEFFBE6A01FA219B4E1917B677" blockId="69.[1399,2427,3351,3477]" box="[1501,1815,3351,3397]" pageId="69" pageNumber="827">
<heading id="9F638192FFBE6A01FA219B4E1917B677" box="[1501,1815,3351,3397]" pageId="69" pageNumber="827">
<vernacularName id="4A9746D0FFBE6A01FA219B4E1917B677" ID-CoL="GMM9" authorityName="Francis &amp; Hill" authorityYear="1984" box="[1501,1815,3351,3397]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" language="eng" order="Chiroptera" pageId="69" pageNumber="827" phylum="Chordata" rank="species" species="cuprosus">Coppery Sprite</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBE6A01F8B79B4E16A8B677" box="[1867,2216,3351,3397]" pageId="69" pageNumber="827" type="nomenclature">
<paragraph id="C42B36FEFFBE6A01F8B79B4E16A8B677" blockId="69.[1399,2427,3351,3477]" box="[1867,2216,3351,3397]" pageId="69" pageNumber="827">
<heading id="9F638192FFBE6A01F8B79B4E16A8B677" box="[1867,2216,3351,3397]" pageId="69" pageNumber="827">
<taxonomicName id="03944D7DFFBE6A01F8B79B4E16A8B677" ID-CoL="GMM9" authorityName="Francis &amp; Hill" authorityYear="1984" box="[1867,2216,3351,3397]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" order="Chiroptera" pageId="69" pageNumber="827" phylum="Chordata" rank="species" species="cuprosus">
<emphasis id="F6E0EAECFFBE6A01F8B79B4E16A8B677" box="[1867,2216,3351,3397]" italics="true" pageId="69" pageNumber="827">Arielulus cuprosus</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBE6A01FA859B0E1936B6A1" pageId="69" pageNumber="827" type="vernacular_names">
<paragraph id="C42B36FEFFBE6A01FA859B0E177AB65E" blockId="69.[1399,2427,3351,3477]" box="[1401,2426,3415,3436]" pageId="69" pageNumber="827">
<heading id="9F638192FFBE6A01FA859B0E177AB65E" box="[1401,2426,3415,3436]" pageId="69" pageNumber="827">
<emphasis id="F6E0EAECFFBE6A01FA859B0E1BC4B65E" bold="true" box="[1401,1476,3415,3436]" pageId="69" pageNumber="827">French:</emphasis>
<vernacularName id="4A9746D0FFBE6A01FA329B0E188DB65E" ID-CoL="GMM9" authorityName="Francis &amp; Hill" authorityYear="1984" box="[1486,1677,3415,3436]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" language="fra" order="Chiroptera" pageId="69" pageNumber="827" phylum="Chordata" rank="species" species="cuprosus">Farfadet de Bornéo</vernacularName>
/
<emphasis id="F6E0EAECFFBE6A01F95E9B0E18FCB65E" bold="true" box="[1698,1788,3415,3436]" pageId="69" pageNumber="827">German:</emphasis>
<vernacularName id="4A9746D0FFBE6A01F8FA9B0E163DB65E" ID-CoL="GMM9" authorityName="Francis &amp; Hill" authorityYear="1984" box="[1798,2109,3415,3436]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" language="deu" order="Chiroptera" pageId="69" pageNumber="827" phylum="Chordata" rank="species" species="cuprosus">Kupferfarbene Elfenfledermaus</vernacularName>
/
<emphasis id="F6E0EAECFFBE6A01F7AE9B0E16ADB65E" bold="true" box="[2130,2221,3415,3436]" pageId="69" pageNumber="827">Spanish:</emphasis>
<vernacularName id="4A9746D0FFBE6A01F7499B0E177AB65E" ID-CoL="GMM9" authorityName="Francis &amp; Hill" authorityYear="1984" box="[2229,2426,3415,3436]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" language="esp" order="Chiroptera" pageId="69" pageNumber="827" phylum="Chordata" rank="species" species="cuprosus">Arielulus de Borneo</vernacularName>
</heading>
</paragraph>
<paragraph id="C42B36FEFFBE6A01FA849B271936B6A1" blockId="69.[1399,2427,3351,3477]" box="[1400,1846,3454,3475]" pageId="69" pageNumber="827">
<heading id="9F638192FFBE6A01FA849B271936B6A1" box="[1400,1846,3454,3475]" pageId="69" pageNumber="827">
<emphasis id="F6E0EAECFFBE6A01FA849B27186FB6A1" bold="true" box="[1400,1647,3454,3475]" pageId="69" pageNumber="827">Other common names:</emphasis>
<vernacularName id="4A9746D0FFBE6A01F9849B271936B6A1" ID-CoL="GMM9" authorityName="Francis &amp; Hill" authorityYear="1984" box="[1656,1846,3454,3475]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" language="eng" order="Chiroptera" pageId="69" pageNumber="827" phylum="Chordata" rank="species" species="cuprosus">Coppery Pipistrelle</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FCE7977F1AB7BAA0" pageId="70" pageNumber="828" type="materials_examined">
<paragraph id="C42B36FEFFBD6A02FCE7977F1DDDBA59" blockId="70.[794,1388,294,717]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FCE7977F1DB6BA71" bold="true" box="[795,950,294,323]" pageId="70" pageNumber="828">Taxonomy.</emphasis>
<taxonomicName id="03944D7DFFBD6A02FC2A977F1DD8BA59" authority="Francis &amp; Hill, 1984" authorityName="Francis &amp; Hill" authorityYear="1984" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="cuprosus">Pipistrellus cuprosus Francis &amp; Hill, 1984</taxonomicName>
,
</paragraph>
<paragraph id="C42B36FEFFBD6A02FC0D97131AB7BAA0" blockId="70.[794,1388,294,717]" pageId="70" pageNumber="828">
<materialsCitation id="74FC3CA3FFBD6A02FC0D97131AB7BAA0" ID-GBIF-Occurrence="3783319756" latitude="5.866667" longitude="117.933334" pageId="70" pageNumber="828">
*Sepilok,
<collectingRegion id="0650F81CFFBD6A02FB7C97131AD5BA59" box="[1152,1237,330,363]" country="Malaysia" name="Sabah" pageId="70" pageNumber="828">Sabah</collectingRegion>
,
<geoCoordinate id="A1A05039FFBD6A02FB1397131B62BA59" box="[1263,1378,330,363]" degrees="5" direction="north" minutes="52" orientation="latitude" pageId="70" pageNumber="828" precision="925" value="5.866667">5°52’ N</geoCoordinate>
,
<geoCoordinate id="A1A05039FFBD6A02FCE3972C1DA5BAA0" box="[799,933,373,402]" degrees="117" direction="east" minutes="56" orientation="longitude" pageId="70" pageNumber="828" precision="925" value="117.933334">117°56’ E</geoCoordinate>
,” Borneo,
<collectingCountry id="BC83766EFFBD6A02FBC0972C1AB3BAA0" box="[1084,1203,373,402]" name="Malaysia" pageId="70" pageNumber="828">Malaysia</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FCE097C01B37B9FF" pageId="70" pageNumber="828" type="discussion">
<paragraph id="C42B36FEFFBD6A02FCE097C01B37B9FF" blockId="70.[794,1388,294,717]" pageId="70" pageNumber="828">
Previously considered a lowland form of
<taxonomicName id="03944D7DFFBD6A02FCE097991DD3BAD3" authorityName="Temminck" authorityYear="1840" box="[796,979,448,481]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="circumdatus">A. circumdatus</taxonomicName>
. In 2018, in a study by L.. I. Amador and colleagues,
<taxonomicName id="03944D7DFFBD6A02FBB197BE1AE1B93A" authorityName="Francis &amp; Hill" authorityYear="1984" box="[1101,1249,487,520]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="cuprosus">A. cuprosus</taxonomicName>
clustered with
<taxonomicName id="03944D7DFFBD6A02FC9B94561A9BB902" authorityName="Kock &amp; Storch" authorityYear="1996" box="[871,1179,527,560]" class="Mammalia" family="Vespertilionidae" genus="Thainycteris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="aureocollaris">Thainycteris aureocollaris</taxonomicName>
(under
<taxonomicName id="03944D7DFFBD6A02FAE494561D3FB965" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="genus">Arielulus</taxonomicName>
) and
<taxonomicName id="03944D7DFFBD6A02FC6F946F1A3FB965" authorityName="Peters" authorityYear="1868" box="[915,1087,566,599]" class="Mammalia" family="Vespertilionidae" genus="Hesperoptenus" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="genus">Hesperoptenus</taxonomicName>
tomes: in a clade related to
<taxonomicName id="03944D7DFFBD6A02FC9B943B1A74B94D" baseAuthorityName="Dobson" baseAuthorityYear="1877" box="[871,1140,610,639]" class="Mammalia" family="Vespertilionidae" genus="Rhyneptesicus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="nasutus">Rhyneptesicus nasutus</taxonomicName>
and Glauconycterts, further work is needed on the relationships of the three
<taxonomicName id="03944D7DFFBD6A02FC3C94E91A2FB9FF" box="[960,1071,688,717]" class="Mammalia" family="Vespertilionidae" genus="Arielulus" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="genus">Arielulus</taxonomicName>
species. Monotypic.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FF49948E1B01B859" pageId="70" pageNumber="828" type="distribution">
<caption id="90EB6676FFBD6A02FF49948E1B01B859" ID-DOI="http://doi.org/10.5281/zenodo.6398073" ID-Zenodo-Dep="6398073" httpUri="https://zenodo.org/record/6398073/files/figure.png" inLine="true" pageId="70" pageNumber="828" targetBox="[180,771,301,715]" targetPageId="70">
<paragraph id="C42B36FEFFBD6A02FF49948E1B01B859" blockId="70.[180,1385,727,1229]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FF49948E1F65B9C6" bold="true" box="[181,357,727,756]" pageId="70" pageNumber="828">Distribution.</emphasis>
Known from a few localities in Borneo (
<collectingRegion id="0650F81CFFBD6A02FC52948E1A02B9C6" box="[942,1026,727,756]" country="Malaysia" name="Sabah" pageId="70" pageNumber="828">Sabah</collectingRegion>
and
<collectingCountry id="BC83766EFFBD6A02FBB3948E1AC7B9C6" box="[1103,1223,727,756]" name="Malaysia" pageId="70" pageNumber="828">Sarawak</collectingCountry>
), including Sepilok Forest Reserve, Batu Punggul, Mt Mulu, Similajau National Park, Lanjak Entimau Wildlife Sanctuary, Mt Penrisen, Kubah National Park, and Samunsam Wildlife Sanctuary. A record of
<taxonomicName id="03944D7DFFBD6A02FE0B95171CADB859" box="[503,685,846,875]" class="Mammalia" family="Vespertilionidae" genus="Pipistrellus" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="undetermined">Pipistrellus sp.</taxonomicName>
from Kalimantan may also be this species.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FF489528193FB94D" pageId="70" pageNumber="828" type="description">
<paragraph id="C42B36FEFFBD6A02FF489528193FB94D" blockId="70.[180,1385,727,1229]" lastBlockId="70.[1453,2664,298,1234]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FF4895281FADB8A0" bold="true" box="[180,429,881,914]" pageId="70" pageNumber="828">Descriptive notes.</emphasis>
Head-body
<quantity id="036C9B1BFFBD6A02FD9895281CC4B8A0" box="[612,708,881,914]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="70" pageNumber="828" unit="mm" value="57.0">57 mm</quantity>
(one specimen), tail 35-42 mm, ear 11-12 mm, hindfoot 7-9 mm, forearm 34-5-37 mm; weight 5-3-6-5 g. The Coppery Sprite resembles the Social Sprite (
<taxonomicName id="03944D7DFFBD6A02FE1295991C8CB8D3" authorityName="Hill" authorityYear="1972" box="[494,652,960,993]" class="Pinopsida" family="Cupressaceae" genus="Arelulus" kingdom="Plantae" order="Pinales" pageId="70" pageNumber="828" phylum="Tracheophyta" rank="species" species="societatis">A. societatis</taxonomicName>
). It has distinctive long, thick fur dorsally, with black or blackish-brown hair bases, and reddish-orange or coppery tips; pelage over crown of head is similarly tipped, but slightly paler and more yellowish than on back; underparts are dark brown or brown, hairs tipped with yellowish white on upper chest near shoulders, elsewhere with grayish white. Flight membranes are black; fur extends on dorsal surface of uropatagium but forearms and tibiae are not haired. Ears and tragus are dark brown with buffy rims; ears are moderately large, with distinct lobe at base near eye. Baculum is very small, with short, slender shaft, tip slightly upcurved, shaft supported by paired basal flanges separated posteriorly by shallow, rounded emargination; baculum is like that of the Social Sprite, but smaller;it differs from that of the Bronze Sprite (
<taxonomicName id="03944D7DFFBD6A02F97B97C01949BA88" authorityName="Temminck" authorityYear="1840" box="[1671,1865,409,442]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="70" pageNumber="828" phylum="Chordata" rank="species" species="circumdatus">A. circumdatus</taxonomicName>
) in its higher, more angular upper margin and more angular basal lobes. Skull is small, with rounded, globular and inflated braincase; postorbital region is wide, supraorbital ridges are weakly developed. Dentition is like that in the Social Sprite, but smaller. P? is very tiny (sometimes lacking) and displaced inward so that C' and P* are touching. Condylo-basal lengths are 12-9-13 mm, maxillary tooth row lengths are 4-8—4-9 mm.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA4C94DF175EB9FC" pageId="70" pageNumber="828" type="biology_ecology">
<paragraph id="C42B36FEFFBD6A02FA4C94DF175EB9FC" blockId="70.[1453,2664,298,1234]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA4C94DF181FB995" bold="true" box="[1456,1567,646,679]" pageId="70" pageNumber="828">Habitat.</emphasis>
Lowland dipterocarp and montane forest. Found in understory oftall forest, along the edge of a steep cliff, and in a forest gap at high elevation.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA4C948D19A4B9C7" box="[1456,1956,724,757]" pageId="70" pageNumber="828" type="food_feeding">
<paragraph id="C42B36FEFFBD6A02FA4C948D19A4B9C7" blockId="70.[1453,2664,298,1234]" box="[1456,1956,724,757]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA4C948D18B5B9C7" bold="true" box="[1456,1717,724,757]" pageId="70" pageNumber="828">Food and Feeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA4C95591925B82F" box="[1456,1829,768,797]" pageId="70" pageNumber="828" type="breeding">
<paragraph id="C42B36FEFFBD6A02FA4C95591925B82F" blockId="70.[1453,2664,298,1234]" box="[1456,1829,768,797]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA4C95591836B82F" bold="true" box="[1456,1590,768,797]" pageId="70" pageNumber="828">Breeding.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA5295711453B85E" pageId="70" pageNumber="828" type="activity">
<paragraph id="C42B36FEFFBD6A02FA5295711453B85E" blockId="70.[1453,2664,298,1234]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA5295711898B877" bold="true" box="[1454,1688,808,837]" pageId="70" pageNumber="828">Activity patterns.</emphasis>
The Coppery Sprite is nocturnal. It is probably a canopy species. Mistnetted over a river in Lanjak Entimau and Samunsam wildlife sanctuaries (
<collectingCountry id="BC83766EFFBD6A02F63795161445B85E" box="[2507,2629,847,876]" name="Malaysia" pageId="70" pageNumber="828">Sarawak</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA4C952B175BB8A1" box="[1456,2395,882,915]" pageId="70" pageNumber="828" type="biology_ecology">
<paragraph id="C42B36FEFFBD6A02FA4C952B175BB8A1" blockId="70.[1453,2664,298,1234]" box="[1456,2395,882,915]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA4C952B166CB8A1" bold="true" box="[1456,2156,882,915]" pageId="70" pageNumber="828">Movements, Home range and Social organization.</emphasis>
No information.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA4C95FB1989BF03" pageId="70" pageNumber="828" type="conservation">
<paragraph id="C42B36FEFFBD6A02FA4C95FB1989BF03" blockId="70.[1453,2664,298,1234]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA4C95FB1910B889" bold="true" box="[1456,1808,930,955]" pageId="70" pageNumber="828">Status and Conservation.</emphasis>
Classified as Vulnerable on The IUCN Red List, as its area of occupancy is estimated to be less than
<quantity id="036C9B1BFFBD6A02F8209598164CB8D0" box="[2012,2124,961,994]" metricMagnitude="6" metricUnit="m" metricValue="2.0" pageId="70" pageNumber="828" unit="km" value="2000.0">2000 km</quantity>
?2. Less than 20 individuals have been captured at only eight locations in
<collectingRegion id="0650F81CFFBD6A02F86895B019E6BF38" box="[1940,2022,1001,1034]" country="Malaysia" name="Sabah" pageId="70" pageNumber="828">Sabah</collectingRegion>
and
<collectingCountry id="BC83766EFFBD6A02F7D395B016A2BF38" box="[2095,2210,1001,1034]" name="Malaysia" pageId="70" pageNumber="828">Sarawak</collectingCountry>
. Extent and quality of its habitat seem to be in continuous decline.
</paragraph>
</subSubSection>
<subSubSection id="8C8E6575FFBD6A02FA4C926619A7BFFD" pageId="70" pageNumber="828" type="bibRefCitation_list">
<paragraph id="C42B36FEFFBD6A02FA4C926619A7BFFD" blockId="70.[1453,2664,298,1234]" pageId="70" pageNumber="828">
<emphasis id="F6E0EAECFFBD6A02FA4C92661846BF6A" bold="true" box="[1456,1606,1087,1112]" pageId="70" pageNumber="828">Bibliography.</emphasis>
Abdullah et al. (2010), Amador et al. (2018), Benda (2010a), Boitani et al. (2006), Corbet &amp; Hill (1992), Heller &amp;Volleth (1989), Hill &amp; Francis (1984), Hill &amp; Harrison (1987), Khan, Swier et al. (2008), Khan, Tahir et al. (2017), Kumaran et al. (2011), McArthur (2016b) Mohd-Azlan et al.(2008) Payne et a/.(1985), Simmons (2005), Suyanto &amp; Struebig (2007), Tingga et al. (2008).
</paragraph>
</subSubSection>
</treatment>
</document>
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