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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.974.54794" ID-GBIF-Dataset="5f0b0760-7a15-49c0-9e80-fc6747f8046e" ID-PMC="PMC7557337" ID-Pensoft-Pub="1313-2970-974-31" ID-Pensoft-UUID="20CAFA6BA01359369DA3F27F90E22660" ID-PubMed="33110378" ID-ZooBank="9B8888660F074DECAE7B88DFB0A4621C" ModsDocID="1313-2970-974-31" checkinTime="1602130124706" checkinUser="pensoft" docAuthor="Wood, Thomas James, Michez, Denis, Cejas, Diego, Lhomme, Patrick &amp; Rasmont, Pierre" docDate="2020" docId="2CA14BAE677E5F5D9D1749202FF5D7F8" docLanguage="en" docName="ZooKeys 974: 31-92" docOrigin="ZooKeys 974" docSource="http://dx.doi.org/10.3897/zookeys.974.54794" docTitle="Andrena (Poliandrena) breviceps Wood 2020, sp. nov." docType="treatment" docUuid="8F5F1214-8FDB-404D-BB5C-E9C0F7E8031B" docUuidSource="ZooBank" docVersion="4" id="20CAFA6BA01359369DA3F27F90E22660" lastPageNumber="31" masterDocId="20CAFA6BA01359369DA3F27F90E22660" masterDocTitle="An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species" masterLastPageNumber="92" masterPageNumber="31" pageNumber="31" updateTime="1668169591898" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>An update and revision of the Andrena fauna of Morocco (Hymenoptera, Apoidea, Andrenidae) with the description of eleven new North African species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wood, Thomas James</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5653-224X</mods:nameIdentifier>
<mods:nameIdentifier type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Michez, Denis</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-8880-1838</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cejas, Diego</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lhomme, Patrick</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium &amp; International Center of Agricultural Research in the Dry Areas, Rabat, Morocco</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6735-9104</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rasmont, Pierre</mods:namePart>
<mods:affiliation>Laboratoire de Zoologie, Universite de Mons, 7000, Mons, Belgium</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>974</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>31</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.974.54794</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.974.54794</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-974-31</mods:identifier>
<mods:identifier type="ZooBank">9B8888660F074DECAE7B88DFB0A4621C</mods:identifier>
<mods:identifier type="Pensoft-UUID">20CAFA6BA01359369DA3F27F90E22660</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="168243726" LSID="urn:lsid:zoobank.org:act:8F5F1214-8FDB-404D-BB5C-E9C0F7E8031B" httpUri="http://treatment.plazi.org/id/2CA14BAE677E5F5D9D1749202FF5D7F8" lastPageNumber="31" pageId="0" pageNumber="31">
<subSubSection pageId="0" pageNumber="31" type="nomenclature">
<paragraph pageId="0" pageNumber="31">
<taxonomicName LSID="2CA14BAE-677E-5F5D-9D17-49202FF5D7F8" authority="Wood" authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Poliandrena) breviceps" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="breviceps" status="sp. nov." subGenus="Poliandrena">Andrena (Poliandrena) breviceps Wood</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="31">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">Figures 83-86</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="materials_examined">
<paragraph pageId="0" pageNumber="31">Material.</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Holotype</emphasis>
: Morocco:
<normalizedToken originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
, 10 km N Erfoud, 10.iv.1995, 1♀, leg. Ma. Halada. Deposited in the
<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
.
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Paratypes</emphasis>
: Morocco:
<normalizedToken originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
, 10 km N Erfoud, 10.iv.1995, 2♀, leg. Ma. Halada,
<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
;
<normalizedToken originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
, 20 km E Agdz, 20.iv.1995, 2♀, leg. Mi. Halada,
<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
;
<normalizedToken originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
, Tagounite, 60 km S Zagora, 23.iv.1995, 1♀, leg. Ma. Halada,
<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
. Paratypes are deposited at the
<normalizedToken originalValue="OÖLM">OOELM</normalizedToken>
with a female retained in the personal collection of TJW.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="diagnosis">
<paragraph pageId="0" pageNumber="31">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="31">
The subgenus
<taxonomicName class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="(Poliandrena)" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="subGenus" subGenus="Poliandrena">Poliandrena</taxonomicName>
Warncke, 1968 is
<taxonomicName class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="currently" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="currently">currently</taxonomicName>
unsatisfactorily defined (females keying out in three places in
<bibRefCitation author="Warncke, K" journalOrPublisher="Memorias e Estudos do Museu Zoologico da Universidade de Coimbra" pageId="0" pageNumber="31" pagination="1 - 110" refId="B47" refString="Warncke, K, 1968a. Die Untergattungen der westpalaearktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 307: 1 - 110" title="Die Untergattungen der westpalaearktischen Bienengattung Andrena F." volume="307" year="1968 a">
<normalizedToken originalValue="Warnckes">Warncke's</normalizedToken>
1968a
</bibRefCitation>
key) and will probably be broken up in the future as it has been shown to be strongly polyphyletic (
<bibRefCitation DOI="https://doi.org/10.1101/2020.06.09.103556" author="Pisanty, G" journalOrPublisher="Gounouilhou, Bordeaux" pageId="0" pageNumber="31" refId="B28" refString="Pisanty, G, Richter, R, Martin, T, Dettman, J, Cardinal, S, 2020. Molecular phylogeny and historical biogeography of andrenine bees (Hymenoptera: Andrenidae). bioRxiv. https://doi.org/10.1101/2020.06.09.103556" title="Molecular phylogeny and historical biogeography of andrenine bees (Hymenoptera: Andrenidae). bioRxiv." url="https://doi.org/10.1101/2020.06.09.103556" year="2020">Pisanty et al. 2020</bibRefCitation>
). It predominantly contains Mediterranean species that have heads that are short and broad (Fig.
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">84</figureCitation>
), short facial foveae, strongly punctate metasomas (Fig.
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">86</figureCitation>
), and often show a propodeal triangle that is defined by an external carina and internal rugosity (but not honeycomb-areolate,
<bibRefCitation author="Warncke, K" journalOrPublisher="Memorias e Estudos do Museu Zoologico da Universidade de Coimbra" pageId="0" pageNumber="31" pagination="1 - 110" refId="B47" refString="Warncke, K, 1968a. Die Untergattungen der westpalaearktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 307: 1 - 110" title="Die Untergattungen der westpalaearktischen Bienengattung Andrena F." volume="307" year="1968 a">Warncke 1968a</bibRefCitation>
). As such, members of the
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Poliandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Poliandrena</emphasis>
</taxonomicName>
are often recognised by their similarity to each other rather than by a single character
<emphasis italics="true" pageId="0" pageNumber="31">per se</emphasis>
, in other words a wastebasket taxon.
</paragraph>
<paragraph pageId="0" pageNumber="31">
<taxonomicName authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena breviceps" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="breviceps">
<emphasis italics="true" pageId="0" pageNumber="31">Andrena breviceps</emphasis>
</taxonomicName>
is small relative to the rest of the
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Poliandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Poliandrena</emphasis>
</taxonomicName>
, comparable in size to
<taxonomicName lsidName="A. marsae" pageId="0" pageNumber="31" rank="species" species="marsae">
<emphasis italics="true" pageId="0" pageNumber="31">A. marsae</emphasis>
</taxonomicName>
Schmiedeknecht, 1900 and
<taxonomicName lsidName="A. laurivora" pageId="0" pageNumber="31" rank="species" species="laurivora">
<emphasis italics="true" pageId="0" pageNumber="31">A. laurivora</emphasis>
</taxonomicName>
Warncke, 1974, two of the smallest
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Poliandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Poliandrena</emphasis>
</taxonomicName>
, but
<taxonomicName lsidName="A. breviceps" pageId="0" pageNumber="31" rank="species" species="breviceps">
<emphasis italics="true" pageId="0" pageNumber="31">A. breviceps</emphasis>
</taxonomicName>
can easily be separated from them by the colour of the tergites which are dark brown (red in
<taxonomicName lsidName="A. marsae" pageId="0" pageNumber="31" rank="species" species="marsae">
<emphasis italics="true" pageId="0" pageNumber="31">A. marsae</emphasis>
</taxonomicName>
, dark metallic green-blue in
<taxonomicName lsidName="A. laurivora" pageId="0" pageNumber="31" rank="species" species="laurivora">
<emphasis italics="true" pageId="0" pageNumber="31">A. laurivora</emphasis>
</taxonomicName>
) and the sculpturing of the scutum which is shiny and (relatively within the
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Poliandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Poliandrena</emphasis>
</taxonomicName>
) sparsely punctate, punctures separated by 2-3 puncture diameters (Fig.
<figureCitation captionStart="Figures 7176" captionStartId="F12" captionText="Figures 71 - 76. Andrena (Notandrena) acutidentis sp. nov. 71 male profile 72 male face 73 male gena 74 male dorsum 75 male tergites 76 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures71-76" httpUri="https://binary.pensoft.net/fig/460653" pageId="0" pageNumber="31">75</figureCitation>
, separated by less than
<normalizedToken originalValue="½">1/2</normalizedToken>
a puncture diameter in
<taxonomicName lsidName="A. marsae" pageId="0" pageNumber="31" rank="species" species="marsae">
<emphasis italics="true" pageId="0" pageNumber="31">A. marsae</emphasis>
</taxonomicName>
and by one puncture diameter in
<taxonomicName lsidName="A. laurivora" pageId="0" pageNumber="31" rank="species" species="laurivora">
<emphasis italics="true" pageId="0" pageNumber="31">A. laurivora</emphasis>
</taxonomicName>
, underlying integument shagreened, with a metallic glint). It is also similar to the larger
<taxonomicName lsidName="A. relata" pageId="0" pageNumber="31" rank="species" species="relata">
<emphasis italics="true" pageId="0" pageNumber="31">A. relata</emphasis>
</taxonomicName>
Warncke, 1967 which is newly recorded for Morocco (see below) because of their similar dark tergites and general appearance. However, it can also be separated using the same scutal punctation character (in
<taxonomicName lsidName="A. relata" pageId="0" pageNumber="31" rank="species" species="relata">
<emphasis italics="true" pageId="0" pageNumber="31">A. relata</emphasis>
</taxonomicName>
punctures dense laterally, centrally separated by at most two puncture diameters) and also by the clypeus where
<taxonomicName lsidName="A. breviceps" pageId="0" pageNumber="31" rank="species" species="breviceps">
<emphasis italics="true" pageId="0" pageNumber="31">A. breviceps</emphasis>
</taxonomicName>
has a central longitudinal impunctate line that is not present in
<taxonomicName lsidName="A. relata" pageId="0" pageNumber="31" rank="species" species="relata">
<emphasis italics="true" pageId="0" pageNumber="31">A. relata</emphasis>
</taxonomicName>
. Overall, the shiny and relatively sparsely punctate scutum (Fig.
<figureCitation captionStart="Figures 7176" captionStartId="F12" captionText="Figures 71 - 76. Andrena (Notandrena) acutidentis sp. nov. 71 male profile 72 male face 73 male gena 74 male dorsum 75 male tergites 76 male genitalia." figureDoi="10.3897/zookeys.974.54794.figures71-76" httpUri="https://binary.pensoft.net/fig/460653" pageId="0" pageNumber="31">75</figureCitation>
) in combination with its small size should allow separation from other
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Poliandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Poliandrena</emphasis>
</taxonomicName>
from North Africa.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="description">
<paragraph pageId="0" pageNumber="31">Description.</paragraph>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Female</emphasis>
: Body length 8.5-9 mm (Fig.
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">83</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Head</emphasis>
: Black, clearly wider than long (Fig.
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">84</figureCitation>
). Clypeus broad, arched, with large punctures, punctures separated by one puncture diameter except for a central slightly raised impunctate line, underlying surface uneven, slightly but irregularly raised between punctures, shining. Process of labrum broadly trapezoidal, fore margin weakly emarginate. Gena as wide as width of compound eye. Gena, vertex, face, and scape with moderately dense white hairs, longest of these not exceeding length of the scape. Foveae moderately broad, occupying 2/3 of distance between the of compound eye and lateral ocellus, of normal length, not extending below the level of the antennal insertions. Antennae bright, scape and pedicel dark, A3 apically marked with orange, A4-12 predominantly orange, A3 exceeding A4+5, shorter than A4+5+6. Ocelloccipital distance short,
<normalizedToken originalValue="½">1/2</normalizedToken>
width of lateral ocellus.
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Mesosoma</emphasis>
: Scutum and scutellum moderately punctured, punctures separated by 1-2 puncture diameters, underlying surface shiny, very weakly shagreened laterally and anteriorly (Fig.
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">85</figureCitation>
). Margins of scutum and scutellum with short, dense whitish hairs, these extending only very sparsely onto the disc. Episternum and propodeum microreticulate, dull, propodeum with weak rugosity, propodeal triangle defined with small but clear slightly raised carina, propodeal triangle with sparse and weak rugosity centrally. Episternum and propodeum with white hairs, longest not exceeding length of the scape. Legs dark, tarsi lightened brown, pubescence whitish to brownish. Femoral and tibial scopa white. Wings hyaline, venation and stigma brown. Nervulus antefurcal.
<emphasis bold="true" italics="true" pageId="0" pageNumber="31">Metasoma</emphasis>
: Tergites brownish, margins slightly depressed, lightened white to yellow, apically translucent (Fig.
<figureCitation captionStart="Figures 8386" captionStartId="F14" captionText="Figures 83 - 86. Andrena (Poliandrena) breviceps sp. nov. 83 female profile 84 female head 85 female dorsum 86 female tergites." figureDoi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31">86</figureCitation>
). Tergites densely, evenly, and finely punctate, punctures separated by one puncture diameter, underlying surface very weakly shagreened, shining. T1 with two small lateral hair patches of white hair on apical margin, T2-4 apically with complete white hair bands obscuring underlying surface. T5-6 with golden hairs flanking pygidial plate. Pygidial plate rounded triangular, flat, without raised margin.
</paragraph>
<caption doi="10.3897/zookeys.974.54794.figures83-86" httpUri="https://binary.pensoft.net/fig/460655" pageId="0" pageNumber="31" start="Figures 8386" startId="F14">
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Figures 83-86.</emphasis>
<taxonomicName authorityName="Wood" authorityYear="2020" class="Insecta" family="Andrenidae" genus="Andrena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Andrena (Poliandrena) breviceps" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="species" species="breviceps" subGenus="Poliandrena">Andrena (Poliandrena) breviceps</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="31">83</emphasis>
female profile
<emphasis bold="true" pageId="0" pageNumber="31">84</emphasis>
female head
<emphasis bold="true" pageId="0" pageNumber="31">85</emphasis>
female dorsum
<emphasis bold="true" pageId="0" pageNumber="31">86</emphasis>
female tergites.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="31">
<emphasis bold="true" pageId="0" pageNumber="31">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="distribution">
<paragraph pageId="0" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="31">
The eastern Moroccan desert in the province of
<normalizedToken originalValue="Drâa-Tafilalet">Draa-Tafilalet</normalizedToken>
(Fig.
<figureCitation captionStart="Figure 145" captionStartId="F24" captionText="Figure 145. Distribution of sampling locations of newly described species A Andrena hebescens (circles), Andrena semiadesus (squares) B Andrena tenebricorpus (diamond), Andrena niveofacies (circle), Andrena triangulivalvis (triangle) C Andrena sparsipunctata (circles), Andrena breviceps (diamonds) D Andrena acutidentis (squares), Andrena farinosoides (triangles), Andrena nigriclypeus (circle). Relief is indicated by shading, measured in meters above sea level." figureDoi="10.3897/zookeys.974.54794.figure145" httpUri="https://binary.pensoft.net/fig/460665" pageId="0" pageNumber="31">145c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="floral preferences">
<paragraph pageId="0" pageNumber="31">Floral preferences.</paragraph>
<paragraph pageId="0" pageNumber="31">None recorded.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="31" type="etymology">
<paragraph pageId="0" pageNumber="31">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="31">
The name
<emphasis italics="true" pageId="0" pageNumber="31">brevi</emphasis>
(short) +
<emphasis italics="true" pageId="0" pageNumber="31">ceps</emphasis>
(head) was chosen because of the particularly short and wide head of this species, even within the
<taxonomicName authorityName="Warncke" authorityYear="1968" class="Insecta" family="Andrenidae" genus="Poliandrena" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Poliandrena" order="Hymenoptera" pageId="0" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="31">Poliandrena</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>