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<document id="620FBCDA43334FC5D45C110DE9DBAF68" ID-CLB-Dataset="46273" ID-DOI="10.11646/zootaxa.3774.4.4" ID-GBIF-Dataset="08d36794-9df4-40f1-bda3-6d2d01ce1722" ID-ISSN="1175-5326" ID-Zenodo-Dep="227391" ID-ZooBank="61E51B03-4DF4-4A04-9F6C-EE1965FC2F42" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460393412826" checkinUser="plazi" docAuthor="Constantinescu, Ioana Cristina, Chişamera, Gabriel, Mukhim, Khlur B. &amp; Adam, Costică" docDate="2014" docId="1F5C2C08FF88F82EFF62FB97FEB925BA" docLanguage="en" docName="zt03774p366.pdf" docOrigin="Zootaxa 3774 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Timalinyssus actinodurae Constantinescu, sp. nov." docType="treatment" docVersion="10" lastPageNumber="358" masterDocId="E3655470FF89F829FFF5FFCAFF892005" masterDocTitle="Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India)" masterLastPageNumber="366" masterPageNumber="351" pageNumber="352" updateTime="1698298487732" updateUser="ExternalLinkService">
<mods:mods id="D56FE2FA1D0507CE20670C384F7BAF6A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="5972E423E61B3EB029B3C27C2946CB14">Two new feather mite species of the family Pteronyssidae (Acarina: Analgoidea) from Meghalaya (Northeast India)</mods:title>
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<mods:name id="41C66091C437418F66ABD06AAAA23AC9" type="personal">
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<mods:namePart id="57BCB151DB40126DB4C149057E4BB07F">Constantinescu, Ioana Cristina</mods:namePart>
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<mods:name id="9570F750B49FA4561F8CCCCC16134096" type="personal">
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<mods:namePart id="93D1136D310502CA43083E0FC23E8AF5">Chişamera, Gabriel</mods:namePart>
</mods:name>
<mods:name id="5B892F76211954D5381BD9C1F6B65B8A" type="personal">
<mods:role id="B3913E4717E143D33F80F1CF3B339DE3">
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<mods:namePart id="8F1BDC0D87F68EB5FA1E35DC34850936">Mukhim, Khlur B.</mods:namePart>
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<mods:name id="3E128BE23784A997DDD76D09BF7F09A0" type="personal">
<mods:role id="C3212057ABCF6ED7D91F8F67A45E84E9">
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<mods:namePart id="E16365B808D74A228114FD295CF26A0B">Adam, Costică</mods:namePart>
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<mods:title id="21B394B76A0C68275CEA9523BDCE4689">Zootaxa</mods:title>
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<mods:part id="EDC2E4D1628DC876FE2B54AA763B5D61">
<mods:date id="82BC04031B16C689B78D0547F319985C">2014</mods:date>
<mods:detail id="649C3386FACE9DBD6E5CE02173E7E81F" type="volume">
<mods:number id="4427A8EBF202FE24120F105852E6A550">3774</mods:number>
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<mods:detail id="4F7B9B1952CE0C32C5434532D3DADFD0" type="issue">
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<mods:identifier id="3786EC4159A2F4A842EEA641FDC19A8A" type="DOI">10.11646/zootaxa.3774.4.4</mods:identifier>
<mods:identifier id="417B64D776ADD3D753B4FD1D7BC1518A" type="GBIF-Dataset">08d36794-9df4-40f1-bda3-6d2d01ce1722</mods:identifier>
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<treatment id="1F5C2C08FF88F82EFF62FB97FEB925BA" ID-DOI="http://doi.org/10.5281/zenodo.6134466" ID-GBIF-Taxon="119417410" ID-Zenodo-Dep="6134466" LSID="urn:lsid:plazi:treatment:1F5C2C08FF88F82EFF62FB97FEB925BA" httpUri="http://treatment.plazi.org/id/1F5C2C08FF88F82EFF62FB97FEB925BA" lastPageId="7" lastPageNumber="358" pageId="1" pageNumber="352">
<subSubSection id="DFEFCE95FF88F828FF62FB97FE9B249F" pageId="1" pageNumber="352" type="nomenclature">
<paragraph id="974A9D1EFF88F828FF62FB97FD562473" blockId="1.[151,735,1117,1178]" box="[151,735,1117,1142]" pageId="1" pageNumber="352">
<heading id="CC022A72FF88F828FF62FB97FD562473" bold="true" box="[151,735,1117,1142]" fontSize="10" level="2" pageId="1" pageNumber="352" reason="2">
<emphasis id="A581410CFF88F828FF62FB97FD562473" bold="true" box="[151,735,1117,1142]" pageId="1" pageNumber="352">
<taxonomicName id="50F5E69DFF88F828FF62FB97FDF72473" authority="Constantinescu" authorityName="Constantinescu" box="[151,638,1117,1142]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF88F828FF62FB97FE492473" bold="true" box="[151,448,1117,1142]" italics="true" pageId="1" pageNumber="352">Timalinyssus actinodurae</emphasis>
Constantinescu
</taxonomicName>
<taxonomicNameLabel id="BEB2FC77FF88F828FD70FB97FD562473" box="[645,735,1117,1142]" pageId="1" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="974A9D1EFF88F828FF62FB4BFE9B249F" blockId="1.[151,735,1117,1178]" box="[151,274,1153,1178]" pageId="1" pageNumber="352">
(
<figureCitation id="0FCE819BFF88F828FF6AFB4BFE81249F" box="[159,264,1153,1178]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1877,1899]" captionStartId-1="3.[151,250,1877,1899]" captionStartId-2="4.[151,250,1794,1816]" captionStartId-3="5.[151,250,1793,1815]" captionStartId-4="6.[151,250,1877,1899]" captionTargetBox-0="[224,1362,193,1855]" captionTargetBox-1="[229,1357,193,1856]" captionTargetBox-4="[218,1368,193,1855]" captionTargetId-0="figure@2.[224,1362,193,1856]" captionTargetId-1="figure@3.[229,1358,193,1856]" captionTargetId-4="figure@6.[218,1368,193,1856]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-4="6" captionText-0="FIGURE 1. Timalinyssus actinodurae sp. nov., male idiosoma, dorsal view." captionText-1="FIGURE 2. Timalinyssus actinodurae sp. nov., male idiosoma, ventral view." captionText-2="FIGURE 3. Timalinyssus actinodurae sp. nov., female idiosoma, dorsal view." captionText-3="FIGURE 4. Timalinyssus actinodurae sp. nov., female idiosoma, ventral view." captionText-4="FIGURE 5. Timalinyssus actinodurae sp. nov., dorsal view of male legs. A—tarsus I, B—tarsus II, C—tibia and tarsus IV, D—tarsus III." httpUri-0="https://zenodo.org/record/227392/files/figure.png" httpUri-1="https://zenodo.org/record/227393/files/figure.png" httpUri-4="https://zenodo.org/record/227394/files/figure.png" pageId="1" pageNumber="352">Figs. 15</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DFEFCE95FF88F82EFF62FB03FCD2221D" lastPageId="7" lastPageNumber="358" pageId="1" pageNumber="352" type="description">
<paragraph id="974A9D1EFF88F828FF62FB03FBFE27D3" blockId="1.[151,1436,1225,2006]" pageId="1" pageNumber="352">
<emphasis id="A581410CFF88F828FF62FB03FEA024E7" bold="true" box="[151,297,1225,1250]" pageId="1" pageNumber="352">Description.</emphasis>
MALE (
<typeStatus id="484E23BCFF88F828FE78FB03FE7A24E7" box="[397,499,1225,1250]" pageId="1" pageNumber="352" type="holotype">holotype</typeStatus>
, range measurements for 2
<typeStatus id="484E23BCFF88F828FCDCFB00FC1D24E7" box="[809,916,1226,1250]" pageId="1" pageNumber="352" type="paratype">paratypes</typeStatus>
in parantheses,
<figureCitation id="0FCE819BFF88F828FBBCFB03FB3D24E7" box="[1097,1204,1225,1250]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1877,1899]" captionStartId-1="3.[151,250,1877,1899]" captionTargetBox-0="[224,1362,193,1855]" captionTargetBox-1="[229,1357,193,1856]" captionTargetId-0="figure@2.[224,1362,193,1856]" captionTargetId-1="figure@3.[229,1358,193,1856]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Timalinyssus actinodurae sp. nov., male idiosoma, dorsal view." captionText-1="FIGURE 2. Timalinyssus actinodurae sp. nov., male idiosoma, ventral view." httpUri-0="https://zenodo.org/record/227392/files/figure.png" httpUri-1="https://zenodo.org/record/227393/files/figure.png" pageId="1" pageNumber="352">Figs. 12</figureCitation>
). Idiosoma 590 long (575580), 350 wide (310335). Prodorsal shield length 165 (160165), width 130 (125140), not fused with scapular shields. Distance between bases of setae
<emphasis id="A581410CFF88F828FD31FAD8FD52252C" box="[708,731,1298,1321]" italics="true" pageId="1" pageNumber="352">se</emphasis>
115 (115120), posterior margin concave, lateral margins with incisions near posterior angles. Setae
<emphasis id="A581410CFF88F828FDB5FAFCFDD2254B" box="[576,603,1334,1358]" italics="true" pageId="1" pageNumber="352">c2</emphasis>
short, filiform, about 30 (2426) in length, situated on medial margins of humeral shields. Setae
<emphasis id="A581410CFF88F828FE69FA90FE3F2577" box="[412,438,1370,1394]" italics="true" pageId="1" pageNumber="352">c3</emphasis>
enlarged in basal part and filiform in apical part, 150 (160175) in length. Hysteronotal shield with straight anterior margin, anterior angles rounded, with strongly sclerotized median band stretching from level of trochanters IV to supranal concavity; length from anterior margin to bases of setae
<emphasis id="A581410CFF88F828FB49FA68FB6A25BF" box="[1212,1251,1442,1466]" italics="true" pageId="1" pageNumber="352">ps1</emphasis>
330 (325330), width at anterior margin 105 (105110). Area between prodorsal and hysteronotal shields wide, distance along midline between these shields 80 (8095). Width of opisthosoma at level of setae
<emphasis id="A581410CFF88F828FBA4FA20FBEF2607" box="[1105,1126,1514,1538]" italics="true" pageId="1" pageNumber="352">f2</emphasis>
90 (8590). Opisthosomal lobes small, nearly rectangular, with slightly acute inner angle and blunt lateral angle. Terminal cleft U-shaped, length 25 (2025), supranal concavity opens posteriorly. Setae
<emphasis id="A581410CFF88F828FC89F9F8FC1E264F" box="[892,919,1586,1610]" italics="true" pageId="1" pageNumber="352">e1</emphasis>
and openings
<emphasis id="A581410CFF88F828FBB3F9FBFBD2264F" box="[1094,1115,1585,1610]" italics="true" pageId="1" pageNumber="352">gl</emphasis>
approximately at the same transverse level. Lengths of dorsal setae:
<emphasis id="A581410CFF88F828FD99F99FFD24266B" box="[620,685,1621,1646]" italics="true" pageId="1" pageNumber="352">c2-d2</emphasis>
175 (170175),
<emphasis id="A581410CFF88F828FC9EF99FFC22266B" box="[875,939,1621,1646]" italics="true" pageId="1" pageNumber="352">d2-e2</emphasis>
180 (170175),
<emphasis id="A581410CFF88F828FB9CF99FFB2D266B" box="[1129,1188,1621,1646]" italics="true" pageId="1" pageNumber="352">d2-gl</emphasis>
95 (7090),
<emphasis id="A581410CFF88F828FACDF99FFAFE266B" box="[1336,1399,1621,1646]" italics="true" pageId="1" pageNumber="352">e2-h2</emphasis>
70 (6270),
<emphasis id="A581410CFF88F828FF08F9B3FEB72697" box="[253,318,1657,1682]" italics="true" pageId="1" pageNumber="352">h2-h2</emphasis>
60 (5060),
<emphasis id="A581410CFF88F828FE3BF9B3FD872697" box="[462,526,1657,1682]" italics="true" pageId="1" pageNumber="352">h3-h3</emphasis>
45 (4546),
<emphasis id="A581410CFF88F828FD6AF9B3FD622697" box="[671,747,1657,1682]" italics="true" pageId="1" pageNumber="352">ps1-h3</emphasis>
8 (67). Transventral sclerite shaped as a thin Y, length along midline 10 (812), width of neck part 8 (68). Epiandrum shaped as an inverted U, fused with posterior end of transventral sclerite, tips of epiandrum extending considerably beyond base of genital apparatus. Length of genital apparatus 24 (2524), width at base 13 (1416), aedeagus minute. Setae
<emphasis id="A581410CFF88F828FC35F92CFC4726F8" box="[960,974,1766,1789]" italics="true" pageId="1" pageNumber="352">g</emphasis>
situated on base of genital arch. Setae
<emphasis id="A581410CFF88F828FA75F92CFA1226FB" box="[1408,1435,1766,1790]" italics="true" pageId="1" pageNumber="352">4a</emphasis>
and posterior pair of genital papillae at level of midlength of genital arch. Anal suckers ovate, their size excluding surrounding membrane: longer diameter 14 (1012), shorter diameter 8 (67). Adanal shield shaped as an inverted Y, narrow, not completely encircling anal field and not fused posteriorly. Ventral measurements:
<emphasis id="A581410CFF88F828FB24F89BFA9B276F" box="[1233,1298,1873,1898]" italics="true" pageId="1" pageNumber="352">4b-4a</emphasis>
66 (6466),
<emphasis id="A581410CFF88F828FF62F8BCFF43278B" box="[151,202,1910,1934]" italics="true" pageId="1" pageNumber="352">4a-g</emphasis>
28 (2526),
<emphasis id="A581410CFF88F828FEAFF8BCFE38278B" box="[346,433,1910,1934]" italics="true" pageId="1" pageNumber="352">ps3-ps3</emphasis>
24 (2430),
<emphasis id="A581410CFF88F828FDB7F8BFFD07278B" box="[578,654,1909,1934]" italics="true" pageId="1" pageNumber="352">ps3-h3</emphasis>
42 (3840). Tarsus
<date id="E34BBBDEFF88F828FC9BF8BCFC24278B" box="[878,941,1909,1934]" pageId="1" pageNumber="352" value="1968-03">III 68</date>
(7076) in length, with acute apical process and 2 small teeth on dorsal longitudinal crest, macroseta
<emphasis id="A581410CFF88F828FCCAF850FCC327B4" box="[831,842,1946,1969]" italics="true" pageId="1" pageNumber="352">r</emphasis>
with very thick basal part, setae
<emphasis id="A581410CFF88F828FB29F850FB6E27B4" box="[1244,1255,1946,1969]" italics="true" pageId="1" pageNumber="352">s</emphasis>
represented by macroseta about 1/3 of macroseta
<emphasis id="A581410CFF88F828FDE3F874FDAB27D0" box="[534,546,1982,2005]" italics="true" pageId="1" pageNumber="352">r,</emphasis>
other setae filiform, shorter than segment (
<figureCitation id="0FCE819BFF88F828FBFCF877FBC227D3" box="[1033,1099,1981,2006]" captionStart="FIGURE 5" captionStartId="6.[151,250,1877,1899]" captionTargetBox="[218,1368,193,1855]" captionTargetId="figure@6.[218,1368,193,1856]" captionTargetPageId="6" captionText="FIGURE 5. Timalinyssus actinodurae sp. nov., dorsal view of male legs. A—tarsus I, B—tarsus II, C—tibia and tarsus IV, D—tarsus III." httpUri="https://zenodo.org/record/227394/files/figure.png" pageId="1" pageNumber="352">Fig. 5</figureCitation>
D).
</paragraph>
<caption id="C38ACD96FF8BF82BFF62F89FFC15276E" box="[151,924,1877,1899]" httpUri="https://zenodo.org/record/227392/files/figure.png" pageId="2" pageNumber="353" targetBox="[224,1362,193,1855]" targetPageId="2">
<paragraph id="974A9D1EFF8BF82BFF62F89FFC15276E" blockId="2.[151,924,1877,1899]" box="[151,924,1877,1899]" pageId="2" pageNumber="353">
<emphasis id="A581410CFF8BF82BFF62F89FFE85276E" bold="true" box="[151,268,1877,1899]" pageId="2" pageNumber="353">FIGURE 1</emphasis>
.
<taxonomicName id="50F5E69DFF8BF82BFEECF89FFD95276E" box="[281,540,1877,1899]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF8BF82BFEECF89FFD95276E" box="[281,540,1877,1899]" italics="true" pageId="2" pageNumber="353">Timalinyssus actinodurae</emphasis>
</taxonomicName>
<emphasis id="A581410CFF8BF82BFDD6F89CFDFA276E" bold="true" box="[547,627,1878,1899]" pageId="2" pageNumber="353">
<taxonomicNameLabel id="BEB2FC77FF8BF82BFDD6F89CFDFA276E" box="[547,627,1878,1899]" pageId="2" pageNumber="353" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male idiosoma, dorsal view.
</paragraph>
</caption>
<caption id="C38ACD96FF8AF82AFF62F89FFC2C276E" box="[151,933,1877,1899]" httpUri="https://zenodo.org/record/227393/files/figure.png" pageId="3" pageNumber="354" targetBox="[229,1357,193,1856]" targetPageId="3">
<paragraph id="974A9D1EFF8AF82AFF62F89FFC2C276E" blockId="3.[151,933,1877,1899]" box="[151,933,1877,1899]" pageId="3" pageNumber="354">
<emphasis id="A581410CFF8AF82AFF62F89FFE85276E" bold="true" box="[151,268,1877,1899]" pageId="3" pageNumber="354">FIGURE 2</emphasis>
.
<taxonomicName id="50F5E69DFF8AF82AFEECF89FFD95276E" box="[281,540,1877,1899]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="3" pageNumber="354" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF8AF82AFEECF89FFD95276E" box="[281,540,1877,1899]" italics="true" pageId="3" pageNumber="354">Timalinyssus actinodurae</emphasis>
</taxonomicName>
<emphasis id="A581410CFF8AF82AFDD6F89CFDFA276E" bold="true" box="[547,627,1878,1899]" pageId="3" pageNumber="354">
<taxonomicNameLabel id="BEB2FC77FF8AF82AFDD6F89CFDFA276E" box="[547,627,1878,1899]" pageId="3" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male idiosoma, ventral view.
</paragraph>
</caption>
<caption id="C38ACD96FF8DF82DFF62F8C8FC39271D" box="[151,944,1794,1816]" pageId="4" pageNumber="355">
<paragraph id="974A9D1EFF8DF82DFF62F8C8FC39271D" blockId="4.[151,944,1794,1816]" box="[151,944,1794,1816]" pageId="4" pageNumber="355">
<emphasis id="A581410CFF8DF82DFF62F8C8FE852712" bold="true" box="[151,268,1794,1816]" pageId="4" pageNumber="355">FIGURE 3</emphasis>
.
<taxonomicName id="50F5E69DFF8DF82DFEECF8C8FD95271D" box="[281,540,1794,1816]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="4" pageNumber="355" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF8DF82DFEECF8C8FD95271D" box="[281,540,1794,1816]" italics="true" pageId="4" pageNumber="355">Timalinyssus actinodurae</emphasis>
</taxonomicName>
<emphasis id="A581410CFF8DF82DFDD6F8C9FDFA271D" bold="true" box="[547,627,1795,1816]" pageId="4" pageNumber="355">
<taxonomicNameLabel id="BEB2FC77FF8DF82DFDD6F8C9FDFA271D" box="[547,627,1795,1816]" pageId="4" pageNumber="355" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female idiosoma, dorsal view.
</paragraph>
</caption>
<caption id="C38ACD96FF8CF82CFF62F8CBFC312712" box="[151,952,1793,1815]" pageId="5" pageNumber="356">
<paragraph id="974A9D1EFF8CF82CFF62F8CBFC312712" blockId="5.[151,952,1793,1815]" box="[151,952,1793,1815]" pageId="5" pageNumber="356">
<emphasis id="A581410CFF8CF82CFF62F8CBFE852713" bold="true" box="[151,268,1793,1815]" pageId="5" pageNumber="356">FIGURE 4</emphasis>
.
<taxonomicName id="50F5E69DFF8CF82CFEECF8CBFD952712" box="[281,540,1793,1815]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="5" pageNumber="356" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF8CF82CFEECF8CBFD952712" box="[281,540,1793,1815]" italics="true" pageId="5" pageNumber="356">Timalinyssus actinodurae</emphasis>
</taxonomicName>
<emphasis id="A581410CFF8CF82CFDD6F8CBFDFA2713" bold="true" box="[547,627,1793,1814]" pageId="5" pageNumber="356">
<taxonomicNameLabel id="BEB2FC77FF8CF82CFDD6F8CBFDFA2713" box="[547,627,1793,1814]" pageId="5" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female idiosoma, ventral view.
</paragraph>
</caption>
<caption id="C38ACD96FF8FF82FFF62F89FFEAD278F" httpUri="https://zenodo.org/record/227394/files/figure.png" pageId="6" pageNumber="357" targetBox="[218,1368,193,1855]" targetPageId="6">
<paragraph id="974A9D1EFF8FF82FFF62F89FFEAD278F" blockId="6.[151,1436,1877,1930]" pageId="6" pageNumber="357">
<emphasis id="A581410CFF8FF82FFF62F89FFE87276E" bold="true" box="[151,270,1877,1899]" pageId="6" pageNumber="357">FIGURE 5</emphasis>
.
<taxonomicName id="50F5E69DFF8FF82FFEE8F89FFDAB276E" box="[285,546,1877,1899]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="6" pageNumber="357" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF8FF82FFEE8F89FFDAB276E" box="[285,546,1877,1899]" italics="true" pageId="6" pageNumber="357">Timalinyssus actinodurae</emphasis>
</taxonomicName>
<emphasis id="A581410CFF8FF82FFDDEF89CFDF4276E" bold="true" box="[555,637,1878,1899]" pageId="6" pageNumber="357">
<taxonomicNameLabel id="BEB2FC77FF8FF82FFDDEF89CFDF4276E" box="[555,637,1878,1899]" pageId="6" pageNumber="357" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, dorsal view of male legs. A—tarsus I, B—tarsus II, C—tibia and tarsus IV, D—tarsus III.
</paragraph>
</caption>
<paragraph id="974A9D1EFF8EF82EFF32FF52FCD2221D" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="358">
FEMALE (1
<typeStatus id="484E23BCFF8EF82EFEA8FF52FE4920B5" box="[349,448,152,176]" pageId="7" pageNumber="358" type="paratype">paratype</typeStatus>
,
<figureCitation id="0FCE819BFF8EF82EFE39FF5DFDB320B5" box="[460,570,151,176]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,250,1794,1816]" captionStartId-1="5.[151,250,1793,1815]" captionText-0="FIGURE 3. Timalinyssus actinodurae sp. nov., female idiosoma, dorsal view." captionText-1="FIGURE 4. Timalinyssus actinodurae sp. nov., female idiosoma, ventral view." pageId="7" pageNumber="358">Figs. 34</figureCitation>
). Idiosoma 555 long, 225 wide. Prodorsal shield: posterior angles fused with scapular shields, posterior margin straight, length of shield 155, width 138, setae
<emphasis id="A581410CFF8EF82EFBE9FF77FBBA20D1" box="[1052,1075,189,212]" italics="true" pageId="7" pageNumber="358">se</emphasis>
separated by 130. Setae
<emphasis id="A581410CFF8EF82EFAB2FF76FAE820D1" box="[1351,1377,188,212]" italics="true" pageId="7" pageNumber="358">c2</emphasis>
hairlike, about 20 long, situated on striated tegument. Hysteronotal shield almost rectangular, with anterior margin concave, anterior angles acute and extending laterally, length (from anterior margin to lobar apices) 325, width at anterior margin 150. Area between prodorsal and hysteronotal shields wide, distance along midline between these shields 80. Posterior end of opisthosoma with 1 pair of widely separated opisthosomal lobes bearing bases of setae
<emphasis id="A581410CFF8EF82EFF62FEA5FF3A218D" box="[151,179,367,392]" italics="true" pageId="7" pageNumber="358">h3</emphasis>
. Opisthosomal lobes short, attenuate apically, strongly bent medially, lateral margin with blunt-angular membrane. Length of terminal cleft 32 width at lobar bases 58. Setae
<emphasis id="A581410CFF8EF82EFC53FE5EFC4921A9" box="[934,960,404,428]" italics="true" pageId="7" pageNumber="358">e1</emphasis>
and openings
<emphasis id="A581410CFF8EF82EFB91FE5EFBF321A8" box="[1124,1146,404,429]" italics="true" pageId="7" pageNumber="358">gl</emphasis>
situated approximately at same transverse level. Dorsal measurements
<emphasis id="A581410CFF8EF82EFD79FE7DFD4521D5" box="[652,716,439,464]" italics="true" pageId="7" pageNumber="358">c2-d2</emphasis>
120,
<emphasis id="A581410CFF8EF82EFCFEFE7DFCC221D5" box="[779,843,439,464]" italics="true" pageId="7" pageNumber="358">d2-e2</emphasis>
125,
<emphasis id="A581410CFF8EF82EFC7CFE7DFC4021D5" box="[905,969,439,464]" italics="true" pageId="7" pageNumber="358">e2-h3</emphasis>
75,
<emphasis id="A581410CFF8EF82EFC0FFE7DFBBC21D5" box="[1018,1077,439,464]" italics="true" pageId="7" pageNumber="358">d2-gl</emphasis>
135,
<emphasis id="A581410CFF8EF82EFB81FE7DFB2421D5" box="[1140,1197,439,464]" italics="true" pageId="7" pageNumber="358">e1-gl</emphasis>
25,
<emphasis id="A581410CFF8EF82EFB2BFE7DFAA321D5" box="[1246,1322,439,464]" italics="true" pageId="7" pageNumber="358">h2-ps1</emphasis>
50,
<emphasis id="A581410CFF8EF82EFAAEFE7DFA1521D5" box="[1371,1436,439,464]" italics="true" pageId="7" pageNumber="358">h2-h2</emphasis>
170,
<emphasis id="A581410CFF8EF82EFF3BFE16FE8621F0" box="[206,271,476,501]" italics="true" pageId="7" pageNumber="358">h3-h3</emphasis>
110. Epigynium approximately semicircular, 50 long, 110 wide. Apodemes of egg-laying opening short, extending to midlevel of trochanters III. Epimerites IVa absent.
</paragraph>
</subSubSection>
<subSubSection id="DFEFCE95FF8EF82EFF32FDEEFD22257D" pageId="7" pageNumber="358" type="materials_examined">
<paragraph id="974A9D1EFF8EF82EFF32FDEEFABB2280" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="358">
<emphasis id="A581410CFF8EF82EFF32FDEEFEFD2238" bold="true" box="[199,372,548,573]" pageId="7" pageNumber="358">
<typeStatus id="484E23BCFF8EF82EFF32FDEEFE8B2238" box="[199,258,548,573]" pageId="7" pageNumber="358">Type</typeStatus>
material
</emphasis>
. Male
<typeStatus id="484E23BCFF8EF82EFE24FDEEFDBA2238" box="[465,563,548,573]" pageId="7" pageNumber="358" type="holotype">holotype</typeStatus>
(ANA091),
<specimenCount id="81F35697FF8EF82EFD25FDEEFCAB2238" box="[720,802,548,573]" pageId="7" pageNumber="358" type="male">2 male</specimenCount>
(ANA093) and
<specimenCount id="81F35697FF8EF82EFC1BFDEEFBDF2238" box="[1006,1110,548,573]" pageId="7" pageNumber="358" type="female">1 female</specimenCount>
(ANA092)
<typeStatus id="484E23BCFF8EF82EFB19FDEFFADE2238" box="[1260,1367,549,573]" pageId="7" pageNumber="358" type="paratype">paratypes</typeStatus>
from
<taxonomicName id="50F5E69DFF8EF82EFF62FD8DFDC52265" authority="Hodgson" authorityName="Hodgson" box="[151,588,583,608]" class="Aves" family="Leiothrichidae" genus="Actinodura" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="358" phylum="Chordata" rank="species" species="cyanouroptera">
<emphasis id="A581410CFF8EF82EFF62FD8DFE4F2265" box="[151,454,583,608]" italics="true" pageId="7" pageNumber="358">Actinodura cyanouroptera</emphasis>
(Hodgson)
</taxonomicName>
(
<taxonomicName id="50F5E69DFF8EF82EFD95FD8DFC8A2265" box="[608,771,583,608]" class="Aves" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="358" phylum="Chordata" rank="order">Passeriformes</taxonomicName>
:
<taxonomicName id="50F5E69DFF8EF82EFCE0FD8DFC372265" box="[789,958,583,608]" class="Aves" family="Leiothrichidae" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="358" phylum="Chordata" rank="family">Leiothrichidae</taxonomicName>
);
<emphasis id="A581410CFF8EF82EFC23FD8DFBBB2265" bold="true" box="[982,1074,583,608]" pageId="7" pageNumber="358">
<collectingCountry id="EFE2DD8EFF8EF82EFC23FD8DFBA42265" box="[982,1069,583,608]" name="India" pageId="7" pageNumber="358">INDIA</collectingCountry>
:
</emphasis>
Meghalaya, East Jaintia Hills District, the region of Khahnar village,
<geoCoordinate id="F2C1FBD9FF8EF82EFDA5FDA6FD732280" box="[592,762,620,645]" direction="north" orientation="latitude" pageId="7" pageNumber="358" precision="1" value="25.365917">25°21'57.30&quot;N</geoCoordinate>
,
<geoCoordinate id="F2C1FBD9FF8EF82EFCF0FDA6FC252280" box="[773,940,620,645]" direction="east" orientation="longitude" pageId="7" pageNumber="358" precision="1" value="92.614365">92°36'51.72&quot;E</geoCoordinate>
;
<date id="E34BBBDEFF8EF82EFC4DFDA6FB0B2281" box="[952,1154,620,645]" pageId="7" pageNumber="358" value="2013-02-20">20 February 2013</date>
, coll. C. Adam.
</paragraph>
<paragraph id="974A9D1EFF8EF82EFF32FD45FD22257D" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="358">
<emphasis id="A581410CFF8EF82EFF32FD45FE4722AD" bold="true" box="[199,462,655,680]" pageId="7" pageNumber="358">Differential diagnosis.</emphasis>
Of the five previously known species,
<taxonomicName id="50F5E69DFF8EF82EFC79FD45FB2522AD" box="[908,1196,655,680]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="actinodurae" status="sp. nov.">
<emphasis id="A581410CFF8EF82EFC79FD45FB2522AD" box="[908,1196,655,680]" italics="true" pageId="7" pageNumber="358">Timalinyssus actinodurae</emphasis>
</taxonomicName>
<emphasis id="A581410CFF8EF82EFB41FD45FA8622AD" bold="true" box="[1204,1295,655,680]" pageId="7" pageNumber="358">
<taxonomicNameLabel id="BEB2FC77FF8EF82EFB41FD45FA8622AD" box="[1204,1295,655,680]" pageId="7" pageNumber="358" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is closest to
<taxonomicName id="50F5E69DFF8EF82EFF62FD7FFE7B22C9" authority="Mironov, 2001" authorityName="Mironov" authorityYear="2001" box="[151,498,692,717]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="A581410CFF8EF82EFF62FD7FFEB222C9" box="[151,315,692,716]" italics="true" pageId="7" pageNumber="358">T. formosanus</emphasis>
Mironov, 2001
</taxonomicName>
from
<taxonomicName id="50F5E69DFF8EF82EFDB4FD7EFB4622C8" authority="Ogilvie-Grant" authorityName="Ogilvie-Grant" box="[577,1231,692,717]" class="Aves" family="Leiothrichidae" genus="Actinodura" kingdom="Animalia" order="Passeriformes" pageId="7" pageNumber="358" phylum="Chordata" rank="species" species="morrisoniana">
<emphasis id="A581410CFF8EF82EFDB4FD7EFCEC22C9" box="[577,869,692,717]" italics="true" pageId="7" pageNumber="358">Actinodura morrisoniana</emphasis>
Ogilvie-Grant (Leiothrichidae)
</taxonomicName>
(Mironov 2001). Males in both species have tarsus III moderately elongated, with an acute apical process and teeth on the longitudinal dorsal crest, the epiandrum shaped as an inverted U with tips extending considerably beyond the base of the genital apparatus, a similar shape of the genital apparatus, and the adanal shield shaped as an inverted Y. Males of the new species clearly differ from those of
<taxonomicName id="50F5E69DFF8EF82EFCF7FC8FFC2B2359" box="[770,930,836,860]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="A581410CFF8EF82EFCF7FC8FFC2B2359" box="[770,930,836,860]" italics="true" pageId="7" pageNumber="358">T. formosanus</emphasis>
</taxonomicName>
due to the following features: the prodorsal shield is free from the scapular shields, the hysteronotal shield has a straight anterior margin and heavily sclerotized median band, the anterior angles of the hysteronotal shield are rounded, the setae
<emphasis id="A581410CFF8EF82EFB10FC46FA8523A1" box="[1253,1292,908,932]" italics="true" pageId="7" pageNumber="358">ps1</emphasis>
are situated almost apically, near the inner angle of the lobes, the transventral sclerite is present, the setae
<emphasis id="A581410CFF8EF82EFB4EFC7BFB4F23CD" box="[1211,1222,945,968]" italics="true" pageId="7" pageNumber="358">s</emphasis>
are represented by macrosetae, and the setae
<emphasis id="A581410CFF8EF82EFE49FC1EFE4323E8" box="[444,458,980,1005]" italics="true" pageId="7" pageNumber="358">d</emphasis>
are hair-like. In males of
<emphasis id="A581410CFF8EF82EFD19FC1FFC1B23E9" box="[748,914,980,1004]" italics="true" pageId="7" pageNumber="358">
<taxonomicName id="50F5E69DFF8EF82EFD19FC1FFC0723E9" box="[748,910,980,1004]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">T. formosanus</taxonomicName>
,
</emphasis>
the prodorsal shield is fused with the scapular shields, the hysteronotal shield has the anterior margin slightly sinuous, its anterior angles are acute and a sclerotised band in median band is absent, the setae
<emphasis id="A581410CFF8EF82EFD0CFBD6FC962431" box="[761,799,1052,1076]" italics="true" pageId="7" pageNumber="358">ps1</emphasis>
are situated clearly distant from the inner angle of the lobes, the transventral sclerite is absent, the setae
<emphasis id="A581410CFF8EF82EFD2FFBF5FD61245D" box="[730,744,1087,1112]" italics="true" pageId="7" pageNumber="358">d</emphasis>
are represented by macrosetae, and the setae
<emphasis id="A581410CFF8EF82EFB08FB8BFA81245D" box="[1277,1288,1089,1112]" italics="true" pageId="7" pageNumber="358">s</emphasis>
are filiform. Females in both species have the prodorsal shield fused with the scapular shields and clearly differ from
<taxonomicName id="50F5E69DFF8EF82EFA72FBAFFE9424A5" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="A581410CFF8EF82EFA72FBAFFA1D2479" box="[1415,1428,1125,1148]" italics="true" pageId="7" pageNumber="358">T</emphasis>
.
<emphasis id="A581410CFF8EF82EFF62FB42FE9424A5" box="[151,285,1160,1184]" italics="true" pageId="7" pageNumber="358">formosanus</emphasis>
</taxonomicName>
, as well from the other three previously known females of the genus (
<taxonomicName id="50F5E69DFF8EF82EFB8CFB43FAFF24A5" authority="Mironov" authorityName="Mironov" box="[1145,1398,1159,1184]" class="Arachnida" family="Pteronyssidae" genus="Mouchetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="oliferae">
<emphasis id="A581410CFF8EF82EFB8CFB43FB7B24A5" box="[1145,1266,1159,1184]" italics="true" pageId="7" pageNumber="358">T. oliferae</emphasis>
(Mironov)
</taxonomicName>
,
<taxonomicName id="50F5E69DFF8EF82EFA72FB43FE5624C0" authority="Wang and Wang" authorityName="Wang and Wang" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="longitarsus">
<emphasis id="A581410CFF8EF82EFA72FB43FE9C24C0" italics="true" pageId="7" pageNumber="358">T. longitarsus</emphasis>
Wang and Wang
</taxonomicName>
,
<taxonomicName id="50F5E69DFF8EF82EFE19FB67FD7824C0" authority="Mironov" authorityName="Mironov" box="[492,753,1196,1221]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="curvilobus">
<emphasis id="A581410CFF8EF82EFE19FB67FD0924C0" box="[492,640,1196,1221]" italics="true" pageId="7" pageNumber="358">T. curvilobus</emphasis>
Mironov
</taxonomicName>
), due to the shape of the opisthosomal lobes that are short, apically attenuate, strongly bent medially, and with blunt-angular lateral membrane. In females of
<emphasis id="A581410CFF8EF82EFB00FB1BFA1524ED" box="[1269,1436,1232,1256]" italics="true" pageId="7" pageNumber="358">
<taxonomicName id="50F5E69DFF8EF82EFB00FB1BFA1124ED" box="[1269,1432,1232,1256]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">T. formosanus</taxonomicName>
,
</emphasis>
and also in
<emphasis id="A581410CFF8EF82EFED5FB3FFE152508" box="[288,412,1268,1293]" italics="true" pageId="7" pageNumber="358">
<taxonomicName id="50F5E69DFF8EF82EFED5FB3FFE112508" box="[288,408,1268,1293]" class="Arachnida" family="Pteronyssidae" genus="Mouchetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="oliferae">T. oliferae</taxonomicName>
,
</emphasis>
the opisthosomal lobes are long and roughly ovoid, while in
<taxonomicName id="50F5E69DFF8EF82EFB9FFB3FFB772508" box="[1130,1278,1268,1293]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="curvilobus">
<emphasis id="A581410CFF8EF82EFB9FFB3FFB772508" box="[1130,1278,1268,1293]" italics="true" pageId="7" pageNumber="358">T. curvilobus</emphasis>
</taxonomicName>
the lobes are indistinct and only short cone-like extensions are present, and, in females of
<taxonomicName id="50F5E69DFF8EF82EFC04FAD3FB022535" box="[1009,1163,1303,1328]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="longitarsus">
<emphasis id="A581410CFF8EF82EFC04FAD3FB022535" box="[1009,1163,1303,1328]" italics="true" pageId="7" pageNumber="358">T. longitarsus</emphasis>
</taxonomicName>
, they do not have lobes. Furthermore, females of
<taxonomicName id="50F5E69DFF8EF82EFE5BFAF7FDDB2550" box="[430,594,1340,1365]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="actinodurae">
<emphasis id="A581410CFF8EF82EFE5BFAF7FDDB2550" box="[430,594,1340,1365]" italics="true" pageId="7" pageNumber="358">T. actinodurae</emphasis>
</taxonomicName>
differ from
<taxonomicName id="50F5E69DFF8EF82EFD29FAF7FCF22551" box="[732,891,1340,1364]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="A581410CFF8EF82EFD29FAF7FD632551" box="[732,746,1341,1364]" italics="true" pageId="7" pageNumber="358">T</emphasis>
.
<emphasis id="A581410CFF8EF82EFD0DFAF6FCF22551" box="[760,891,1340,1364]" italics="true" pageId="7" pageNumber="358">formosanus</emphasis>
</taxonomicName>
due to the absence of epimerites IVa (rudiments of epimerites IVa are present in
<taxonomicName id="50F5E69DFF8EF82EFE09FAABFD12257D" box="[508,667,1376,1400]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="A581410CFF8EF82EFE09FAABFD12257D" box="[508,667,1376,1400]" italics="true" pageId="7" pageNumber="358">T. formosanus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="DFEFCE95FF8EF82EFF32FA4EFEB925BA" pageId="7" pageNumber="358" type="etymology">
<paragraph id="974A9D1EFF8EF82EFF32FA4EFEB925BA" blockId="7.[151,1437,151,1472]" pageId="7" pageNumber="358">
<emphasis id="A581410CFF8EF82EFF32FA4EFEC72598" bold="true" box="[199,334,1412,1437]" pageId="7" pageNumber="358">Etymology.</emphasis>
The specific epithet
<taxonomicName id="50F5E69DFF8EF82EFDC2FA4EFD372598" box="[567,702,1412,1437]" class="Arachnida" family="Pteronyssidae" genus="Timalinyssus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="7" pageNumber="358" phylum="Arthropoda" rank="species" species="actinodurae">
<emphasis id="A581410CFF8EF82EFDC2FA4EFD372598" box="[567,702,1412,1437]" italics="true" pageId="7" pageNumber="358">actinodurae</emphasis>
</taxonomicName>
refers to the generic name of the host species and is a noun in the genitive case.
</paragraph>
</subSubSection>
</treatment>
</document>