160 lines
17 KiB
XML
160 lines
17 KiB
XML
<document id="B3555D29267183CCFEF21B9A6C070D53" ID-DOI="10.11646/zootaxa.4048.4.6" ID-GBIF-Dataset="12dd6a18-8481-4dc9-8727-8cfbfc138f7a" ID-ISSN="1175-5326" ID-Zenodo-Dep="237775" ID-ZooBank="FE00B390-0362-42F7-B6F8-67D35141E014" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461348369585" checkinUser="plazi" docAuthor="Sinclair, Bradley J." docDate="2015" docId="1F0D87A0FFDBFF84FF6DC199AD91F818" docLanguage="en" docName="zt04048p564.pdf" docOrigin="Zootaxa 4048 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Asymphyloptera havasu Sinclair, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="558" masterDocId="E334FFD8FFDEFF81FFFAC367AF58FFA2" masterDocTitle="New World species of Asymphyloptera Collin (Diptera: Empididae: Clinocerinae)" masterLastPageNumber="564" masterPageNumber="553" pageNumber="558" updateTime="1698621086257" updateUser="plazi">
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<mods:titleInfo id="3C8C2240953A59CAEA810B1297C257A1">
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<mods:title id="0DF621C0B08EDB4696CD0651C2E05364">New World species of Asymphyloptera Collin (Diptera: Empididae: Clinocerinae)</mods:title>
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<mods:name id="9A7BF2A963A94E61B7001741446F0D23" type="personal">
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<mods:namePart id="E7F5CEE4CFCFD819BCCF0D13C0DBAF24">Sinclair, Bradley J.</mods:namePart>
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<mods:typeOfResource id="4582FBD60A774630F1838B7E99963AAC">text</mods:typeOfResource>
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<mods:title id="DAC05456961A6517E651DE1006921884">Zootaxa</mods:title>
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<mods:part id="0CABA9A89D0A51D6E63676DF4356C355">
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<mods:date id="F7607B24822A838E654962EDCA230745">2015</mods:date>
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<mods:detail id="B37FD0BA327D3AEF6E56D00303C11129" type="volume">
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<mods:number id="1CC07FA4E9CA9C389E61581C86CEA509">4048</mods:number>
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<mods:number id="D400868B210738C49347E40A2EAB4256">4</mods:number>
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<mods:classification id="E46DECAEB13449C145334839CD9B11C7">journal article</mods:classification>
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<mods:identifier id="6B35C318343149E419077FEFC7F0CB85" type="DOI">10.11646/zootaxa.4048.4.6</mods:identifier>
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<treatment id="1F0D87A0FFDBFF84FF6DC199AD91F818" ID-DOI="http://doi.org/10.5281/zenodo.6113497" ID-GBIF-Taxon="119645591" ID-Zenodo-Dep="6113497" LSID="urn:lsid:plazi:treatment:1F0D87A0FFDBFF84FF6DC199AD91F818" httpUri="http://treatment.plazi.org/id/1F0D87A0FFDBFF84FF6DC199AD91F818" lastPageNumber="558" pageId="5" pageNumber="558">
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<subSubSection id="DFBE653DFFDBFF84FF6DC199AFA1FC98" pageId="5" pageNumber="558" type="nomenclature">
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<paragraph id="971B36B6FFDBFF84FF6DC199AD45FCBA" blockId="5.[151,541,766,826]" box="[151,541,766,792]" pageId="5" pageNumber="558">
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<heading id="CC5381DAFFDBFF84FF6DC199AD45FCBA" bold="true" box="[151,541,766,792]" fontSize="11" level="1" pageId="5" pageNumber="558" reason="1">
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<emphasis id="A5D0EAA4FFDBFF84FF6DC199AD45FCBA" bold="true" box="[151,541,766,792]" pageId="5" pageNumber="558">
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<taxonomicName id="50A44D35FFDBFF84FF6DC199AEE0FCBA" ID-CoL="9HTVZ" box="[151,440,766,792]" class="Insecta" family="Empididae" genus="Asymphyloptera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="havasu" status="sp. nov.">
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<emphasis id="A5D0EAA4FFDBFF84FF6DC199AEE0FCBA" bold="true" box="[151,440,766,792]" italics="true" pageId="5" pageNumber="558">Asymphyloptera havasu</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BEE357DFFFDBFF84FE45C199AD45FCBA" box="[447,541,766,792]" pageId="5" pageNumber="558" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="971B36B6FFDBFF84FF6DC046AFA1FC98" blockId="5.[151,541,766,826]" box="[151,249,801,826]" pageId="5" pageNumber="558">
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(
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<figureCitation id="0F9F2A33FFDBFF84FF65C046AFA9FC98" box="[159,241,801,826]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1900,1923]" captionTargetBox="[192,1402,203,1862]" captionTargetId="figure@7.[179,1408,193,1879]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Male terminalia of Asymphyloptera, lateral views. 13, A. dominica sp. nov.; 14, A. havasu sp. nov.; 15, A. lutea sp. nov.; 16, A. mexicana sp. nov. Scale bars 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." httpUri="https://zenodo.org/record/237779/files/figure.png" pageId="5" pageNumber="558">Fig. 14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="DFBE653DFFDBFF84FF6DC00EAE52FBDC" pageId="5" pageNumber="558" type="materials_examined">
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<paragraph id="971B36B6FFDBFF84FF6DC00EAFA1FBB0" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF6DC00EAE19FC20" bold="true" box="[151,321,873,898]" pageId="5" pageNumber="558">
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<typeStatus id="481F8814FFDBFF84FF6DC00EAF8AFC20" box="[151,210,873,898]" pageId="5" pageNumber="558">Type</typeStatus>
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material
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</emphasis>
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.
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<emphasis id="A5D0EAA4FFDBFF84FEAEC00EAEABFC20" bold="true" box="[340,499,873,898]" pageId="5" pageNumber="558">
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<typeStatus id="481F8814FFDBFF84FEAEC00EAEABFC20" box="[340,499,873,898]" pageId="5" pageNumber="558" type="holotype">HOLOTYPE</typeStatus>
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</emphasis>
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, ♂ labelled: “
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<collectingCountry id="EFB37626FFDBFF84FD66C00DAD82FC20" box="[668,730,874,898]" name="United States of America" pageId="5" pageNumber="558">USA</collectingCountry>
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: AZ:Grand Canyon NP/ CR#7, Elves Canyon [Chasm]/
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<date id="E31A1076FFDBFF84FA88C00DAE5BFC04" pageId="5" pageNumber="558" value="1998-05-10" valueMax="1998-05-21" valueMin="1998-05-10">10– 21.v.1998</date>
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/ J.F. MacDonald”; “
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<typeStatus id="481F8814FFDBFF84FE13C0E9AD27FC04" box="[489,639,910,934]" pageId="5" pageNumber="558" type="holotype">HOLOTYPE</typeStatus>
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/
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<taxonomicName id="50A44D35FFDBFF84FD74C0EAAC10FC04" box="[654,840,909,934]" class="Insecta" family="Empididae" genus="Asymphyloptera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="genus">Asymphyloptera</taxonomicName>
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/
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<taxonomicName id="50A44D35FFDBFF84FCADC0EAACFCFC04" box="[855,932,909,934]" class="Insecta" family="Empididae" genus="Asymphyloptera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="havasu">havasu</taxonomicName>
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/ Sinclair [red label]” (
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<collectionCode id="F1B5AE73FFDBFF84FB65C0E9AB86FC04" box="[1183,1246,910,934]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="5" pageNumber="558">CNC</collectionCode>
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).
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<emphasis id="A5D0EAA4FFDBFF84FB0AC0EAAACAFC04" bold="true" box="[1264,1426,909,934]" pageId="5" pageNumber="558">
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<typeStatus id="481F8814FFDBFF84FB0AC0EAAACAFC04" box="[1264,1426,909,934]" pageId="5" pageNumber="558" type="paratype">PARATYPES</typeStatus>
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</emphasis>
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:
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<emphasis id="A5D0EAA4FFDBFF84FF6DC0D6AF97FC68" bold="true" box="[151,207,945,970]" pageId="5" pageNumber="558">
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<collectingCountry id="EFB37626FFDBFF84FF6DC0D6AF97FC68" box="[151,207,945,970]" name="United States of America" pageId="5" pageNumber="558">USA</collectingCountry>
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</emphasis>
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.
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<emphasis id="A5D0EAA4FFDBFF84FF27C0D6AE65FC68" bold="true" box="[221,317,945,970]" pageId="5" pageNumber="558">Arizona</emphasis>
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: Same data as
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<typeStatus id="481F8814FFDBFF84FE13C0D6AD13FC68" box="[489,587,945,970]" pageId="5" pageNumber="558" type="holotype">holotype</typeStatus>
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(
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<specimenCount id="81A2FD3FFFDBFF84FDA1C0D5ADDFFC6B" box="[603,647,946,970]" pageId="5" pageNumber="558" type="generic">2 ♂</specimenCount>
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, 2 ♀,
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<collectionCode id="F1B5AE73FFDBFF84FD35C0D5AC56FC68" box="[719,782,946,970]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="5" pageNumber="558">CNC</collectionCode>
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); Grand Canyon NP, CR#15, Fern Canyon,
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<date id="E31A1076FFDBFF84FB07C0D5AAC0FC68" box="[1277,1432,946,970]" pageId="5" pageNumber="558" value="1998-05-10" valueMax="1998-05-21" valueMin="1998-05-10">10–21.v.1998</date>
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, J.F. MacDonald (
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<specimenCount id="81A2FD3FFFDBFF84FEA1C0B1AEDCFC4F" box="[347,388,982,1006]" pageId="5" pageNumber="558" type="generic">1 ♂</specimenCount>
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, 2 ♀,
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<collectionCode id="F1B5AE73FFDBFF84FE31C0B1AD51FC4C" box="[459,521,982,1006]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="5" pageNumber="558">CNC</collectionCode>
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); Grand Canyon NP, CR#3, Saddle Canyon,
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<date id="E31A1076FFDBFF84FBFBC0B1ABC4FC4C" box="[1025,1180,982,1006]" pageId="5" pageNumber="558" value="1998-05-10" valueMax="1998-05-21" valueMin="1998-05-10">10–21.v.1998</date>
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, J.F. MacDonald (
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<specimenCount id="81A2FD3FFFDBFF84FA96C0B1AACFFC4F" box="[1388,1431,982,1006]" pageId="5" pageNumber="558" type="generic">1 ♂</specimenCount>
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,
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<collectionCode id="F1B5AE73FFDBFF84FF6DC09DAFB5FBB0" LSID="urn:lsid:biocol.org:col:34871" box="[151,237,1018,1042]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="558">USNM</collectionCode>
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).
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</paragraph>
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<paragraph id="971B36B6FFDBFF84FF3DC77AAE52FBDC" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF3DC77AAE0EFB94" bold="true" box="[199,342,1053,1078]" pageId="5" pageNumber="558">Recognition</emphasis>
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. This mostly dark species may be distinguished by pale fore coxa (viewed anteriorly) and dark pleura; narrow, tapered surstylus with pointed apex; apex of phallus expanded, membranous and strongly bent anteriorly.
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</paragraph>
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</subSubSection>
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<subSubSection id="DFBE653DFFDBFF84FF3DC7EEAEE8F91C" pageId="5" pageNumber="558" type="description">
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<paragraph id="971B36B6FFDBFF84FF3DC7EEAEA7FB4B" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF3DC7EEAE09FB00" bold="true" box="[199,337,1161,1186]" pageId="5" pageNumber="558">Description</emphasis>
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.
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<emphasis id="A5D0EAA4FFDBFF84FEA5C7EEAEC3FB00" bold="true" box="[351,411,1161,1186]" pageId="5" pageNumber="558">Male</emphasis>
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. Postpedicel globular with long, slender apical extension, arising subapically. Ocellar setae stout, similar in length to postpedicel extension. Labrum slightly shorter than clypeus; palpus brown, subequal to labrum, tapered to slender apex.
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</paragraph>
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<paragraph id="971B36B6FFDBFF84FF3DC792AEBEFA60" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">Pleura dark brown with thin bluish pruinescence. Chaetotaxy very stout; 1 pprn short, very slender; 2 npl, lower slender and shorter; 4 dc, anterior dc offset; 2 apical sctl, slightly longer than prescutellar dc, lacking lateral sctl. Legs brown, fore coxae pale yellowish-brown (anterior view); fore and mid femora with antero- and posteroventral rows of pale, slender setae, subequal in length to width of femur; fore tibia with biserial row of erect setae. Wing length 1.7–2.0 mm; crossvein h distinct; base of wing with slender setulae along posterior margin. Halter pale, yellowish-brown.</paragraph>
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<paragraph id="971B36B6FFDBFF84FF3DC6AAABB5F9D4" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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Male terminalia (
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<figureCitation id="0F9F2A33FFDBFF84FE66C6AAAEA2FA44" box="[412,506,1485,1510]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1900,1923]" captionTargetBox="[192,1402,203,1862]" captionTargetId="figure@7.[179,1408,193,1879]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Male terminalia of Asymphyloptera, lateral views. 13, A. dominica sp. nov.; 14, A. havasu sp. nov.; 15, A. lutea sp. nov.; 16, A. mexicana sp. nov. Scale bars 0.1 mm. Abbreviations: pgt—postgonite; ph—phallus; sur—surstylus." httpUri="https://zenodo.org/record/237779/files/figure.png" pageId="5" pageNumber="558">Fig. 14</figureCitation>
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): Cercus with anterior lobe slightly produced; posterior lobe long, slender. Hypandrium prolonged as broad, paired postgonites; apical margin of postgonite not extended anteriorly, apex pointed. Epandrium narrow, with elongate setae on apical half. Surstylus shorter than cercus, slender, arched, tapered to pointed apex; not extending beyond phallus. Phallus straight; apex expanded, membranous, strongly bent anteriorly; apical margin hooked anteriorly; ejaculatory apodeme narrow, short, not expanded.
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</paragraph>
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<paragraph id="971B36B6FFDBFF84FF3DC5E6AEE8F91C" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF3DC5E6AE46F938" bold="true" box="[199,286,1665,1690]" pageId="5" pageNumber="558">Female</emphasis>
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. Similar to male except as follows: lacking long ventral setae on fore and mid femora and fore tibia. Terminalia not dissected.
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</paragraph>
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</subSubSection>
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<subSubSection id="DFBE653DFFDBFF84FF3DC5AEACC0F8A4" pageId="5" pageNumber="558" type="etymology">
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<paragraph id="971B36B6FFDBFF84FF3DC5AEACC0F8A4" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF3DC5AEAE1EF940" bold="true" box="[199,326,1737,1762]" pageId="5" pageNumber="558">Etymology</emphasis>
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. Named after the Havasupai, a native American tribe that have lived for centuries in the Grand Canyon region;
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<taxonomicName id="50A44D35FFDBFF84FEB1C589AEF9F8A7" box="[331,417,1774,1797]" class="Insecta" family="Empididae" genus="Asymphyloptera" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="563" phylum="Arthropoda" rank="species" species="havasu" status="sp. nov.">
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<emphasis id="A5D0EAA4FFDBFF84FEB1C589AEF9F8A7" box="[331,417,1774,1797]" italics="true" pageId="5" pageNumber="558">Havasu</emphasis>
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</taxonomicName>
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means “blue-green water” and
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<emphasis id="A5D0EAA4FFDBFF84FCF8C589AC7DF8A7" box="[770,805,1774,1797]" italics="true" pageId="5" pageNumber="558">pai</emphasis>
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“people”.
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</paragraph>
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</subSubSection>
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<subSubSection id="DFBE653DFFDBFF84FF3DC476ADE3F8EC" pageId="5" pageNumber="558" type="distribution">
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<paragraph id="971B36B6FFDBFF84FF3DC476ADE3F8EC" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF3DC476AE01F888" bold="true" box="[199,345,1809,1834]" pageId="5" pageNumber="558">Distribution</emphasis>
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. Known from several seepages along the Colorado River, collected by John MacDonald during a survey in Grand Canyon National Park (
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<figureCitation id="0F9F2A33FFDBFF84FDA0C452ADF6F8EC" box="[602,686,1845,1870]" captionStart="FIGURE 17" captionStartId="8.[151,250,1990,2012]" captionTargetBox="[264,1319,193,1957]" captionTargetId="figure@8.[264,1319,193,1957]" captionTargetPageId="8" captionText="FIGURE 17. Distribution of Asymphyloptera in the New World." httpUri="https://zenodo.org/record/237780/files/figure.png" pageId="5" pageNumber="558">Fig. 17</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="DFBE653DFFDBFF84FF3DC43EAD91F818" pageId="5" pageNumber="558" type="discussion">
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<paragraph id="971B36B6FFDBFF84FF3DC43EAD91F818" blockId="5.[151,1437,873,1978]" pageId="5" pageNumber="558">
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<emphasis id="A5D0EAA4FFDBFF84FF3DC43EAE6CF8D0" bold="true" box="[199,308,1881,1906]" pageId="5" pageNumber="558">Remarks</emphasis>
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. This species was collected in deeply shaded madicolous habitats on the canyon walls, formed by cold streams flowing down narrow slot canyons (
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<figureCitation id="0F9F2A33FFDBFF84FD09C41AAC3FF834" box="[755,871,1917,1942]" captionStart="FIGURES 4 – 7" captionStartId="2.[151,264,1831,1854]" captionTargetBox="[151,1436,841,1810]" captionTargetId="figure@2.[151,1436,841,1811]" captionTargetPageId="2" captionText="FIGURES 4 – 7. Examples of Asymphyloptera habitats. 4, type locality of A. cajanuma sp. nov.: upper montane rainforest or cloud forest at Cajanuma, Ecuador (~ 3000 m) (specimen collected in Malaise trap set over small stream) (photograph M. Pollet); 5, type locality of A. chiricahua sp. nov.: East Turkey Creek, Arizona, USA (specimens collected on wet over hanging rocks); 6, Saddle Canyon, Grand Canyon, Arizona, USA (specimens aspirated off seepage on right) (photograph J. F. MacDonald); 7, type locality of A. havasu sp. nov.: Elves Chasm, Grand Canyon, Arizona, USA (specimens aspirated off seepage on left) (photograph Flickr)." httpUri="https://zenodo.org/record/237777/files/figure.png" pageId="5" pageNumber="558">Figs 6, 7</figureCitation>
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). These sites are characterized by mosses and maidenhair ferns (MacDonald pers. comm. 2015).
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |