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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.907.36563" ID-GBIF-Dataset="69422977-aa4d-4fe3-a450-8ef54d6fa905" ID-PMC="PMC7002455" ID-Pensoft-Pub="1313-2970-907-1" ID-Pensoft-UUID="9920267E73CF5E00B644DED1F101D965" ID-PubMed="32063727" ID-ZooBank="DBF339E5EBBC461994388359C769473F" ModsDocID="1313-2970-907-1" checkinTime="1580398733216" checkinUser="pensoft" docAuthor="Leger, Theo, Kehlmaier, Christian, Vairappan, Charles S. &amp; Nuss, Matthias" docDate="2020" docId="1D67A747BF035E8082D28B0E052D6243" docLanguage="en" docName="ZooKeys 907: 1-99" docOrigin="ZooKeys 907" docSource="http://dx.doi.org/10.3897/zookeys.907.36563" docTitle="Hoploscopa mallyi Leger &amp; Nuss 2020, sp. nov." docType="treatment" docUuid="13552C8E-ED5E-447B-859A-8402DD48BA72" docUuidSource="ZooBank" docVersion="4" id="9920267E73CF5E00B644DED1F101D965" lastPageNumber="1" masterDocId="9920267E73CF5E00B644DED1F101D965" masterDocTitle="Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding" masterLastPageNumber="99" masterPageNumber="1" pageNumber="1" updateTime="1668168367651" updateUser="ExternalLinkService">
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<mods:title>Twenty-six new species of Hoploscopa (Lepidoptera, Crambidae) from South-East Asia revealed by morphology and DNA barcoding</mods:title>
</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Leger, Theo</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kehlmaier, Christian</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Vairappan, Charles S.</mods:namePart>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Nuss, Matthias</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>907</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.907.36563</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.907.36563</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-907-1</mods:identifier>
<mods:identifier type="ZooBank">DBF339E5EBBC461994388359C769473F</mods:identifier>
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<treatment ID-GBIF-Taxon="161926070" LSID="urn:lsid:zoobank.org:act:13552C8E-ED5E-447B-859A-8402DD48BA72" httpUri="http://treatment.plazi.org/id/1D67A747BF035E8082D28B0E052D6243" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="1D67A747-BF03-5E80-82D2-8B0E052D6243" authority="Léger &amp; Nuss" authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa mallyi" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="mallyi" status="sp. nov.">
Hoploscopa mallyi
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; Nuss
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figures 3445" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">Figs 37</figureCitation>
<figureCitation captionStart="Figures 7681" captionStartId="F10" captionText="Figures 76 - 81. Male genitalia features of Hoploscopa. 76 Hoploscopa mallyi sp. nov., paratype, TL 339 ♂ 77 Hoploscopa gracilis sp. nov., paratype, TL 539 ♂ 78 Hoploscopa agtuuganonensis sp. nov., paratype, TL 675 ♂ 79 Hoploscopa boleta sp. nov., holotype, TL 440 ♂ 80 Hoploscopa pseudometacrossa sp. nov., paratype, TL 441 ♂ (uncus apex missing) 81 Hoploscopa metacrossa, TL 443 ♂." figureDoi="10.3897/zookeys.907.36563.figures76-81" httpUri="https://binary.pensoft.net/fig/375765" pageId="0" pageNumber="1">, 76</figureCitation>
<figureCitation captionStart="Figures 115118" captionStartId="F19" captionText="Figures 115 - 118. Female genitalia features of Hoploscopa. 115 Hoploscopa sumatrensis sp. nov., paratype, TL 530 ♀ 116 Hoploscopa mallyi sp. nov., paratype, TL 513 ♀ 117 Hoploscopa gracilis sp. nov., paratype, TL 527 ♀ 118 Hoploscopa agtuuganonensis sp. nov., holotype, TL 616 ♀." figureDoi="10.3897/zookeys.907.36563.figures115-118" httpUri="https://binary.pensoft.net/fig/375774" pageId="0" pageNumber="1">, 116</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: ♀. &quot;Borneo, Mt Kinabalu | Headquarter, 1600m | 10-13.XI.2006, L[icht]F[ang] | leg. W. Mey &amp; K. Ebert&quot;; &quot;Liwagu River | 1500m, Turm&quot;; &quot;DNA voucher | Lepidoptera | M. Nuss 2007 | [vertically written:] no. 124&quot;; &quot;TL366 | ♀&quot;. Deposited in BORN.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 4 ♂, 2 ♀. Malaysia: 1 ♂, 1 ♀ (genitalia on slides TL533 ♂ and TL340 ♀ respectively), Sabah, Kinabalu National Park Headquarters, 250 m from Pandanus Trail starting point,
<geoCoordinate degrees="6" direction="north" minutes="0" orientation="latitude" precision="15" seconds="34" value="6.009444">6°0'34&quot;N</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="32" orientation="longitude" precision="15" seconds="20" value="116.53889">116°32'20&quot;E</geoCoordinate>
1640 m, UV light, 07.vi.2015, leg. T.
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; R. Mally; 2 ♂ (1 with genitalia on slide TL339 ♂), Sabah, Mesilau logging site, 400 m before entrance to Mesilau Nature Reserve,
<geoCoordinate degrees="6" direction="north" minutes="2" orientation="latitude" precision="15" seconds="22" value="6.0394444">6°2'22&quot;N</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="35" orientation="longitude" precision="15" seconds="54" value="116.598335">116°35'54&quot;E</geoCoordinate>
, 1930 m, UV light, 02.vi.2015, T.
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; R. Mally; 1 ♀ (DNA voucher 3197, genitalia on slide TL513 ♀), Sabah, Kinabalu Park Headquarters, ca. 300 m from starting point of Kiau View Trail,
<geoCoordinate degrees="6" direction="north" minutes="0" orientation="latitude" precision="15" seconds="25" value="6.0069447">6°0'25&quot;N</geoCoordinate>
,
<geoCoordinate degrees="116" direction="east" minutes="32" orientation="longitude" precision="15" seconds="21" value="116.53917">116°32'21&quot;E</geoCoordinate>
, 1660 m, UV light, 06.vi.2015, T.
<normalizedToken originalValue="Léger">Leger</normalizedToken>
&amp; R. Mally (MTD); 1 ♂, Borneo, Mt Kinabalu, Mesilau, 2000 m, 14-17.xi.2006, W. Mey &amp; K. Ebert (MFNB).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The forewings of
<taxonomicName lsidName="H. mallyi" pageId="0" pageNumber="1" rank="species" species="mallyi">
<emphasis italics="true" pageId="0" pageNumber="1">H. mallyi</emphasis>
</taxonomicName>
sp. nov. display pale yellow crescent-shaped median discoidal stigma filled with red, together with median cubital patch forming a Y. In male genitalia, the gnathos projection is long, slender, reaching circa 2/3 of the uncus length, and the juxta is ogive-shaped, with a slightly indented apex. In female genitalia, the ductus bursae is long, broad, curved twice, and the corpus bursae is small, globular, with one short curved thorn.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="similar species">
<paragraph pageId="0" pageNumber="1">Similar species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa agtuuganonensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="agtuuganonensis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa agtuuganonensis</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName lsidName="H. gracilis" pageId="0" pageNumber="1" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">H. gracilis</emphasis>
</taxonomicName>
sp. nov. In
<taxonomicName lsidName="H. agtuuganonensis" pageId="0" pageNumber="1" rank="species" species="agtuuganonensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. agtuuganonensis</emphasis>
</taxonomicName>
sp. nov., median discoidal stigma together with cubital and dorsal patches forms a band progressively narrowing toward dorsum (median cubital patch reduced to small streak, median dorsal patch absent in
<taxonomicName lsidName="H. mallyi" pageId="0" pageNumber="1" rank="species" species="mallyi">
<emphasis italics="true" pageId="0" pageNumber="1">H. mallyi</emphasis>
</taxonomicName>
sp. nov.).
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa gracilis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa gracilis</emphasis>
</taxonomicName>
sp. nov. displays wing pattern similar to
<taxonomicName lsidName="H. mallyi" pageId="0" pageNumber="1" rank="species" species="mallyi">
<emphasis italics="true" pageId="0" pageNumber="1">H. mallyi</emphasis>
</taxonomicName>
sp. nov. and can be best separated by examination of genitalia: the uncus and valva are slenderer, the valva apex is pointed (rounded in
<taxonomicName lsidName="H. mallyi" pageId="0" pageNumber="1" rank="species" species="mallyi">
<emphasis italics="true" pageId="0" pageNumber="1">H. mallyi</emphasis>
</taxonomicName>
sp. nov.) and the juxta displays a broad duck beak-shaped apex. In female genitalia of
<taxonomicName lsidName="H. gracilis" pageId="0" pageNumber="1" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">H. gracilis</emphasis>
</taxonomicName>
sp. nov., the ductus bursae shows a narrow loop at posterior 1/3, and the thorn is less curved.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
.
</emphasis>
Antennae dorsally with brown scales. Proboscis brown to pale brown scaled. Maxillary palpi brown, base and inner side pale yellow. Labial palpi brown, ventral base and inner side pale yellow.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
(Fig.
<figureCitation captionStart="Figures 3445" captionStartId="F4" captionText="Figures 34 - 45. Forewing of Hoploscopa species. 34 Hoploscopa ignitamaculae sp. nov., paratype, ♂, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 30 - 31. iii. 2000 (A. Kallies &amp; C. Zorn) (genitalia on slide TL 509 ♂) 35 Hoploscopa albomaculata sp. nov., paratype, ♀, Indonesia, North Sulawesi, Danau Mooat, east of Kotamobagu, 1000 m, 25 - 26. iii. 2000, (A. Kallies &amp; C. Zorn) (genitalia on slide TL 540 ♀) 36 Hoploscopa sumatrensis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 6 - 26. i. 1996 (A. Kallies) (genitalia on slide TL 755 ♂) 37 Hoploscopa mallyi sp. nov., paratype, ♂, Malaysia, Sabah, Mt Kinabalu, Mesilau, 2000 m, 14 - 17. xi. 2006 (W. Mey &amp; K. Ebert) 38 Hoploscopa gracilis sp. nov., holotype, ♂, Indonesia, Sumatra-Holzweg, 25 km SSW-Pematangsiantar, road to Prapat, 13. ii. 1996 (A. Kallies) (genitalia on slide TL 754 ♂) 39 Hoploscopa agtuuganonensis sp. nov., holotype, ♀, Philippines, Mindanao, Mt Agtuuganon, 1050 m, 28. v- 7. vi. 1996 (W. Mey) (genitalia on slide TL 616 ♀) 40 Hoploscopa boleta sp. nov., paratype, ♀, Papua New Guinea, Morobe, Mt Kaindi, 2360 m, 3. x. 1992 (V. O. Becker) (genitalia on slide TL 448 ♀) 41 Hoploscopa pseudometacrossa sp. nov., holotype, ♀, Papua New Guinea, W. Hhl, Prv, near Mt. Hagen, Kuk Ag, Res. Sta., 1600 m, 19 - 20. viii. 1983 (Scott E. &amp; Pamela Miller) (genitalia on slide TL 689 ♀) 42 Hoploscopa metacrossa Hampson, ♀, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller). 43 Hoploscopa jubata sp. nov., paratype, ♂, Papua New Guinea, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 44 Hoploscopa jubata sp. nov., paratype, ♀, Morobe Province, near Bulolo, Mt Susu National Reserve, Araucaria forest, 975 m, 27 - 28. viii. 1983 (S. Miller) 45 Hindwing scent organ in Hoploscopa metacrossa. Paratype, ♂, Papua New Guinea, Morobe Province, Wau, 1200 m, 8 - 14. xii. 1976 (G. F. Hevel &amp; R. E. Dietz) (genitalia on slide TL 443 ♂)." figureDoi="10.3897/zookeys.907.36563.figures34-45" httpUri="https://binary.pensoft.net/fig/375759" pageId="0" pageNumber="1">37</figureCitation>
). Collar pale yellow. Forewing length: 10-12 mm (♂), 11-12 mm (♀); forewing ground colour brown; basal longitudinal red streak basally edged with pale yellow; costal field red; median discoidal stigma crescent-shaped, pale yellow, filled with red; median cubital patch streak-like, pale yellow, together with discoidal stigma forming a Y; postmedian patch triangular, red, basal edge partially pale yellow, with pale yellow blotch at costa; postmedian area suffused with pale yellow and iridescent scales; subterminal field marked with red; fringes brown with pale yellow dots. Hindwing pale brown. Forelegs brown. Midlegs brown; femur and tibia distally pale yellow. Hindlegs with femur brown; tibia pale yellow speckled with brown; tarsi pale yellow to pale brown.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Male genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 2) (Fig.
<figureCitation captionStart="Figures 7681" captionStartId="F10" captionText="Figures 76 - 81. Male genitalia features of Hoploscopa. 76 Hoploscopa mallyi sp. nov., paratype, TL 339 ♂ 77 Hoploscopa gracilis sp. nov., paratype, TL 539 ♂ 78 Hoploscopa agtuuganonensis sp. nov., paratype, TL 675 ♂ 79 Hoploscopa boleta sp. nov., holotype, TL 440 ♂ 80 Hoploscopa pseudometacrossa sp. nov., paratype, TL 441 ♂ (uncus apex missing) 81 Hoploscopa metacrossa, TL 443 ♂." figureDoi="10.3897/zookeys.907.36563.figures76-81" httpUri="https://binary.pensoft.net/fig/375765" pageId="0" pageNumber="1">76</figureCitation>
). Uncus slender, medially narrowed, apex spatulate. Gnathos projection slender, ca. 2/3 of uncus length, apex blunt. Valva ventral margin gently bent dorsad on distal 1/3, dorsal margin convex, apex rounded. Juxta ogival, with base quadrangular, gently narrowing toward apex, apex narrow, notched. Saccus triangular, pointing dorsad.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Female genitalia</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
= 3) (Fig.
<figureCitation captionStart="Figures 115118" captionStartId="F19" captionText="Figures 115 - 118. Female genitalia features of Hoploscopa. 115 Hoploscopa sumatrensis sp. nov., paratype, TL 530 ♀ 116 Hoploscopa mallyi sp. nov., paratype, TL 513 ♀ 117 Hoploscopa gracilis sp. nov., paratype, TL 527 ♀ 118 Hoploscopa agtuuganonensis sp. nov., holotype, TL 616 ♀." figureDoi="10.3897/zookeys.907.36563.figures115-118" httpUri="https://binary.pensoft.net/fig/375774" pageId="0" pageNumber="1">116</figureCitation>
). Anterior apophyses with dorsal bump at posterior 1/3. Antrum sclerotisation forming a short ring. Ductus bursae long, broad, forming two large curves. Corpus bursae small, globular, reticulate, sclerotised between thorn and corpus opening, medially with broad faintly sclerotised band. Thorn sabre-like, curved, with small dents pointing toward thorn base, basally with conspicuous outwardly projected extension.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Known from the slopes of Mount Kinabalu on Borneo, at altitudes between 1,600 m and 2,000 m.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="phylogenetic relationships">
<paragraph pageId="0" pageNumber="1">Phylogenetic relationships.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is recovered in a clade with
<taxonomicName lsidName="H. agtuuganonensis" pageId="0" pageNumber="1" rank="species" species="agtuuganonensis">
<emphasis italics="true" pageId="0" pageNumber="1">H. agtuuganonensis</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="H. gracilis" pageId="0" pageNumber="1" rank="species" species="gracilis">
<emphasis italics="true" pageId="0" pageNumber="1">H. gracilis</emphasis>
</taxonomicName>
sp. nov. in the ML analysis of the COI barcode (BS = 92). The morphology of these species is in agreement with the molecular findings: in male genitalia, gnathos projection is long and slender; in female genitalia, the antrum sclerotisation is short, the ductus bursae is relatively broad, looped or with conspicuous curves, and the corpus bursae is small, globular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">This species is dedicated to our colleague and friend Richard Mally, an eminent lepidopterologist.</paragraph>
<caption doi="10.3897/zookeys.907.36563.figures58-63" httpUri="https://binary.pensoft.net/fig/375762" pageId="0" pageNumber="1" start="Figures 5863" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 58-63.</emphasis>
Male genitalia features of
<taxonomicName authorityName="Meyrick" authorityYear="1886" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">58</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa titika" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="titika">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa titika</emphasis>
</taxonomicName>
sp. nov., holotype, TL505 ♂ (apical part of juxta attached to phallus)
<emphasis bold="true" pageId="0" pageNumber="1">59</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa pangrangoensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pangrangoensis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa pangrangoensis</emphasis>
</taxonomicName>
sp. nov., paratype, TL659 ♂
<emphasis bold="true" pageId="0" pageNumber="1">60</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa isarogensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="isarogensis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa isarogensis</emphasis>
</taxonomicName>
sp. nov., paratype, TL762 ♂
<emphasis bold="true" pageId="0" pageNumber="1">61</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa ypsilon" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ypsilon">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa ypsilon</emphasis>
</taxonomicName>
sp. nov., paratype, TL624 ♂
<emphasis bold="true" pageId="0" pageNumber="1">62</emphasis>
<taxonomicName authorityName="Leger &amp; Nuss" authorityYear="2020" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa danaoensis" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="danaoensis">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa danaoensis</emphasis>
</taxonomicName>
sp. nov., holotype, TL632 ♂
<emphasis bold="true" pageId="0" pageNumber="1">63</emphasis>
<taxonomicName baseAuthorityName="Rotschild" baseAuthorityYear="1915" class="Insecta" family="Crambidae" genus="Hoploscopa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hoploscopa quadripuncta" order="Lepidoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="quadripuncta">
<emphasis italics="true" pageId="0" pageNumber="1">Hoploscopa quadripuncta</emphasis>
</taxonomicName>
, holotype,
<taxonomicName family="Pyralidae" lsidName="" pageId="0" pageNumber="1" rank="family">Pyralidae</taxonomicName>
Brit. Mus. Slide N° BMNH20257 ♂.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>