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<document id="F60A8FF0387D4D6637F4D77EBA8A308E" ID-CLB-Dataset="2878" ID-DOI="10.11646/zootaxa.5082.5.3" ID-GBIF-Dataset="b829a58c-42a2-44f1-b0ad-5346289588a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5796363" ID-ZooBank="9F4AE75E-4AE6-40A9-A7BB-868894FAF85B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1640070984751" checkinUser="plazi" docAuthor="Budashkin, Yuriy &amp; Richter, Ignác" docDate="2021" docId="13265852FF9FC006FF26FB2FFDFDF8A7" docLanguage="en" docName="Zootaxa.5082.5.3.pdf" docOrigin="Zootaxa 5082 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aethes kindermanniana subsp. macedonica Richter &amp; Budashkin 2021, ssp. nov." docType="treatment" docVersion="7" lastPageNumber="451" masterDocId="EF1F202AFF96C00CFFB1FF84FFCAF90A" masterDocTitle="New records of Cochylini moths (Lepidoptera: Tortricidae) from the Balkans" masterLastPageNumber="456" masterPageNumber="441" pageNumber="450" updateTime="1699472860831" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D55881D82A0A94854B17C83BAA9D61A0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D178F207E0CD6E5E604018B197EF0978">
<mods:title id="B231CE15D1FC86BAD5053F101A129E36">New records of Cochylini moths (Lepidoptera: Tortricidae) from the Balkans</mods:title>
</mods:titleInfo>
<mods:name id="84986D384DBED0C70DBD5F7D8FF14E03" type="personal">
<mods:role id="DD6A125EA1BB6947F5B1B3A99DC5DD52">
<mods:roleTerm id="E9CF9B9FBBBE7AA787401BE7C4E16441">Author</mods:roleTerm>
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<mods:namePart id="2243D4A6BAEC633969E8CEF69AF2C3DB">Budashkin, Yuriy</mods:namePart>
<mods:nameIdentifier id="69A23AD95E7678E9CE47CAFA6ABC3B55" type="ORCID">0000-0001-7824-8338</mods:nameIdentifier>
<mods:affiliation id="E09A437588C70927D287BF669478BC78">T. I. Vyazemsky Karadag Scientific Station-Nature Reserve-Branch of FIC “ A. O. Kovalevsky Institute of Biology of the South Seas ” RAS, 298188, Kurortnoye, Pheodosia, Crimea Republic, Russia. budashkin @ ukr. net; https: // orcid. org / 0000 - 0001 - 7824 - 8338</mods:affiliation>
<mods:nameIdentifier id="BC105CDF1E899EE3E3C5A4C1D8437DDD" type="email">budashkin@ukr.net</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="8A2BAE9B86E9E5A717A13E99B5D88784">Richter, Ignác</mods:namePart>
<mods:nameIdentifier id="B3A6FB0044D081665FE27FAC1066F4BC" type="ORCID">0000-0001-8122-8709</mods:nameIdentifier>
<mods:affiliation id="CB4A788F3DBB4F2363B03DBCF74726C8">Malá Čausa, 289, Slovakia. ignac. richter @ gmail. com; https: // orcid. org / 0000 - 0001 - 81122 - 8709</mods:affiliation>
<mods:nameIdentifier id="8910D91AA15F3F7331FAF8787AB68B1B" type="email">ignac.richter@gmail.com</mods:nameIdentifier>
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<mods:title id="64C39F9855124839EF8E35CE90F3DF0F">Zootaxa</mods:title>
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<mods:part id="E0FB144247CC923D30BCD2A4BBAD7810">
<mods:date id="E8070987443B96F5A7B444A16C597ABF">2021</mods:date>
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<mods:number id="6A75FE3938113DA7B392EE684A031574">2021-12-21</mods:number>
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<mods:number id="3733AABF18D5C0C0320F62A452F7E2FE">5082</mods:number>
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<mods:number id="FFB790185350F44A16CA6755DAD7C268">5</mods:number>
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<mods:identifier id="727BA71D60B27626226C2829683EE3A0" type="DOI">10.11646/zootaxa.5082.5.3</mods:identifier>
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<treatment id="13265852FF9FC006FF26FB2FFDFDF8A7" ID-DOI="http://doi.org/10.5281/zenodo.5796450" ID-GBIF-Taxon="191698100" ID-Zenodo-Dep="5796450" LSID="urn:lsid:plazi:treatment:13265852FF9FC006FF26FB2FFDFDF8A7" httpUri="http://treatment.plazi.org/id/13265852FF9FC006FF26FB2FFDFDF8A7" lastPageId="10" lastPageNumber="451" pageId="9" pageNumber="450">
<subSubSection id="D395BACFFF9FC005FF26FB2FFC14FDCF" box="[151,990,1195,1222]" pageId="9" pageNumber="450" type="nomenclature">
<paragraph id="9B30E944FF9FC005FF26FB2FFC14FDCF" blockId="9.[151,990,1195,1257]" box="[151,990,1195,1222]" pageId="9" pageNumber="450">
<heading id="C0785E28FF9FC005FF26FB2FFC14FDCF" bold="true" box="[151,990,1195,1222]" fontSize="11" level="1" pageId="9" pageNumber="450" reason="1">
<emphasis id="A9FB3556FF9FC005FF26FB2FFC14FDCF" bold="true" box="[151,990,1195,1222]" pageId="9" pageNumber="450">
<taxonomicName id="5C8F92C7FF9FC005FF26FB2FFCA2FDCF" authority="Richter &amp; Budashkin" authorityName="Richter &amp; Budashkin" authorityYear="2021" box="[151,872,1195,1222]" class="Insecta" family="Tortricidae" genus="Aethes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="subSpecies" species="kindermanniana" status="subsp. nov." subSpecies="macedonica">
<emphasis id="A9FB3556FF9FC005FF26FB2FFD98FDCC" bold="true" box="[151,594,1195,1222]" italics="true" pageId="9" pageNumber="450">Aethes kindermanniana macedonica</emphasis>
Richter &amp; Budashkin
</taxonomicName>
,
<taxonomicNameLabel id="B2C8882DFF9FC005FCC4FB2FFC14FDCF" box="[885,990,1195,1221]" pageId="9" pageNumber="450" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D395BACFFF9FC005FF26FB4BFEE5FDE3" box="[151,303,1231,1257]" pageId="9" pageNumber="450" type="description">
<paragraph id="9B30E944FF9FC005FF26FB4BFEE5FDE3" blockId="9.[151,990,1195,1257]" box="[151,303,1231,1257]" pageId="9" pageNumber="450">
(
<figureCitation id="03B4F5C1FF9FC005FF2EFB4BFEECFDE3" box="[159,294,1231,1257]" captionStart="FIGURES 18–21" captionStartId="10.[151,264,1339,1363]" captionTargetBox="[151,1429,453,1309]" captionTargetId="figure-117@10.[151,1435,453,1316]" captionTargetPageId="10" captionText="FIGURES 18–21. Fig. 18. Adult of Aethes kindermanniana (Croatia). Fig. 19. Male genitalia of Aethes kindermanniana (Croatia). Fig. 20. Adult of Aethes kindermanniana macedonica ssp. nov. (North Macedonia). Fig. 21. Male genitalia of Aethes kindermanniana macedonica ssp. nov. (North Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5796371" httpUri="https://zenodo.org/record/5796371/files/figure.png" pageId="9" pageNumber="450">Figs. 20, 21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D395BACFFF9FC005FF26FA93FBB4FC5F" pageId="9" pageNumber="450" type="materials_examined">
<paragraph id="9B30E944FF9FC005FF26FA93FBB4FC5F" blockId="9.[151,1437,1303,2014]" pageId="9" pageNumber="450">
<emphasis id="A9FB3556FF9FC005FF26FA93FE8FFC3B" bold="true" box="[151,325,1303,1329]" pageId="9" pageNumber="450">Type material.</emphasis>
<materialsCitation id="2BE7E319FF9FC005FEFDFA93FF0EFC5C" ID-GBIF-Occurrence="3421208308" collectingDate="2019-07-30" collectionCode="R" county="North" location="Asan Gjura" municipality="Galicica Nacional" pageId="9" pageNumber="450" specimenCode="GP 30461" specimenCount="1" specimenCount-male="1" stateProvince="Dytiki Makedonia" typeStatus="holotype">
<typeStatus id="443457E6FF9FC005FEFDFA93FE7EFC3B" box="[332,436,1303,1329]" pageId="9" pageNumber="450" type="holotype">Holotype</typeStatus>
<specimenCount id="8D8922CDFF9FC005FE0AFA9CFE18FC38" box="[443,466,1304,1330]" pageId="9" pageNumber="450" type="male">♂</specimenCount>
,
<emphasis id="A9FB3556FF9FC005FE6FFA93FD7AFC3B" bold="true" box="[478,688,1303,1329]" pageId="9" pageNumber="450">
<collectingCounty id="725191C8FF9FC005FE6FFA93FDECFC3B" box="[478,550,1303,1329]" pageId="9" pageNumber="450">North</collectingCounty>
<collectingRegion id="594B27A6FF9FC005FD9DFA93FD7AFC3B" box="[556,688,1303,1329]" country="Greece" name="Dytiki Makedonia" pageId="9" pageNumber="450">Macedonia</collectingRegion>
</emphasis>
,
<collectingMunicipality id="7B54733EFF9FC005FD0EFA93FC42FC3B" box="[703,904,1303,1329]" pageId="9" pageNumber="450">Galičica Nacional</collectingMunicipality>
park,
<location id="9E50BF9FFF9FC005FC7CFA93FB9BFC3B" LSID="urn:lsid:plazi:treatment:13265852FF9FC006FF26FB2FFDFDF8A7:9E50BF9FFF9FC005FC7CFA93FB9BFC3B" box="[973,1105,1303,1329]" county="North" municipality="Galicica Nacional" name="Asan Gjura" pageId="9" pageNumber="450" stateProvince="Dytiki Makedonia">Asan Gjura</location>
,
<date id="EF31CF84FF9FC005FBEAFA93FB11FC3B" box="[1115,1243,1303,1329]" pageId="9" pageNumber="450" value="2019-07-30">
<collectingDate id="FF75366CFF9FC005FBEAFA93FB11FC3B" box="[1115,1243,1303,1329]" pageId="9" pageNumber="450" value="2019-07-30">30.vii.2019</collectingDate>
</date>
(I.
<collectionCode id="FD9E7181FF9FC005FB4AFA93FAD9FC3B" box="[1275,1299,1303,1329]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="450">R</collectionCode>
.).
<specimenCode id="CB29413FFF9FC005FA9DFA93FA56FC3B" box="[1324,1436,1303,1329]" collectionCode="GP" country="Brazil" name="Instituto de Geociencias, Universidade de Sao Paulo" pageId="9" pageNumber="450">GP 30461</specimenCode>
IgR
</materialsCitation>
.
<materialsCitation id="2BE7E319FF9FC005FF7EFAB8FBB0FC5F" ID-GBIF-Occurrence="3421208361" box="[207,1146,1339,1366]" collectingDate="2019-07-30" collectionCode="R, TMLF" county="North" location="Asan Gjura" municipality="Galicica Nacional" pageId="9" pageNumber="450" specimenCount="2" specimenCount-male="2" stateProvince="Dytiki Makedonia" typeStatus="paratype">
<typeStatus id="443457E6FF9FC005FF7EFAB8FEF6FC5C" box="[207,316,1340,1366]" pageId="9" pageNumber="450" type="paratype">Paratypes</typeStatus>
<specimenCount id="8D8922CDFF9FC005FEF2FABFFEA3FC5F" box="[323,361,1339,1365]" pageId="9" pageNumber="450" type="male">2♂</specimenCount>
, same data as for the holotype (I.
<collectionCode id="FD9E7181FF9FC005FD6BFABFFD3BFC5F" box="[730,753,1339,1365]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="450">R</collectionCode>
). DNA Barcode
<collectionCode id="FD9E7181FF9FC005FC1DFAB8FC3DFC5C" box="[940,1015,1340,1366]" pageId="9" pageNumber="450">TMLF</collectionCode>
Lep 28719
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D395BACFFF9FC005FF76FADBFA47FF27" pageId="9" pageNumber="450" type="diagnosis">
<paragraph id="9B30E944FF9FC005FF76FADBFA47FF27" blockId="9.[151,1437,1303,2014]" pageId="9" pageNumber="450">
<emphasis id="A9FB3556FF9FC005FF76FADBFE88FC73" bold="true" box="[199,322,1375,1401]" pageId="9" pageNumber="450">Diagnosis.</emphasis>
The new subspecies is close to
<taxonomicName id="5C8F92C7FF9FC005FD6EFADBFE97FC97" authority="kindermanniana (Treitschke, 1830) (Treitschke 1830)" authorityName="kindermanniana (Treitschke, 1830) (Treitschke" authorityYear="1830" baseAuthorityName="Treitschke" baseAuthorityYear="1830" class="Insecta" family="Tortricidae" genus="Aethes" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="450" phylum="Arthropoda" rank="subSpecies" species="kindermanniana" subSpecies="kindermanniana">
<emphasis id="A9FB3556FF9FC005FD6EFADBFB72FC73" box="[735,1208,1375,1401]" italics="true" pageId="9" pageNumber="450">Aethes kindermanniana kindermanniana</emphasis>
(
<bibRefCitation id="FF1E94B5FF9FC005FB7FFADBFA5EFC73" author="Treitschke, F." box="[1230,1428,1375,1401]" pageId="9" pageNumber="450" refId="ref11478" refString="Treitschke, F. (1830) Die Schmetterlinge von Europa. Vol. 8. Ernst Fleischer, Leipzig, VIII + 312 pp." type="book" year="1830">Treitschke, 1830</bibRefCitation>
) (
<bibRefCitation id="FF1E94B5FF9FC005FF2FFA07FE9FFC97" author="Treitschke, F." box="[158,341,1411,1437]" pageId="9" pageNumber="450" refId="ref11478" refString="Treitschke, F. (1830) Die Schmetterlinge von Europa. Vol. 8. Ernst Fleischer, Leipzig, VIII + 312 pp." type="book" year="1830">Treitschke 1830</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="03B4F5C1FF9FC005FEDAFA07FE23FC97" box="[363,489,1411,1437]" captionStart="FIGURES 18–21" captionStartId="10.[151,264,1339,1363]" captionTargetBox="[151,1429,453,1309]" captionTargetId="figure-117@10.[151,1435,453,1316]" captionTargetPageId="10" captionText="FIGURES 18–21. Fig. 18. Adult of Aethes kindermanniana (Croatia). Fig. 19. Male genitalia of Aethes kindermanniana (Croatia). Fig. 20. Adult of Aethes kindermanniana macedonica ssp. nov. (North Macedonia). Fig. 21. Male genitalia of Aethes kindermanniana macedonica ssp. nov. (North Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5796371" httpUri="https://zenodo.org/record/5796371/files/figure.png" pageId="9" pageNumber="450">Figs 18, 19</figureCitation>
) by habitus and the structure of male genitalia, but differs by larger size, more diffuse and lighter pattern of the forewing. Additionally, hindwing is lighter and thinner, socii are less prickly, the acantae on the apex of the median process of transtilla are larger, and the medioventral process of the aedeagus more slender. Although the new subspecies is insufficiently supported by molecular data (0.31% distance from the nominate subspecies), the morphological differences suggest the existence of a separate subspecies from North
<collectingCountry id="E398A9D4FF9FC005FAB8F997FA43FF27" box="[1289,1417,1555,1581]" name="Macedonia" pageId="9" pageNumber="450">Macedonia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="D395BACFFF9FC006FF76F9B3FC7FF848" lastPageId="10" lastPageNumber="451" pageId="9" pageNumber="450" type="description">
<paragraph id="9B30E944FF9FC005FF76F9B3FF05FE9F" blockId="9.[151,1437,1303,2014]" pageId="9" pageNumber="450">
<emphasis id="A9FB3556FF9FC005FF76F9B3FE93FF5B" bold="true" box="[199,345,1591,1617]" pageId="9" pageNumber="450">Description.</emphasis>
Adult (
<figureCitation id="03B4F5C1FF9FC005FE1BF9B3FE37FF5B" box="[426,509,1591,1617]" captionStart="FIGURES 18–21" captionStartId="10.[151,264,1339,1363]" captionTargetBox="[151,1429,453,1309]" captionTargetId="figure-117@10.[151,1435,453,1316]" captionTargetPageId="10" captionText="FIGURES 18–21. Fig. 18. Adult of Aethes kindermanniana (Croatia). Fig. 19. Male genitalia of Aethes kindermanniana (Croatia). Fig. 20. Adult of Aethes kindermanniana macedonica ssp. nov. (North Macedonia). Fig. 21. Male genitalia of Aethes kindermanniana macedonica ssp. nov. (North Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5796371" httpUri="https://zenodo.org/record/5796371/files/figure.png" pageId="9" pageNumber="450">Fig. 20</figureCitation>
). Sexual dimorphism unknown. Wingspan
<quantity id="5C7744A1FF9FC005FC66F9B3FBE2FF58" box="[983,1064,1591,1618]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="450" unit="mm" value="13.0">13 mm</quantity>
. The basic colour of head, tegulae and thorax white to pale yellow. Patagia, anterior part of tegulae and posterior part of thorax deep yellow. Scales on patagia and on posterior part of thorax long and erect. The labial palpi of medium length, thick. Segment 3 much shorter than segment 2. Upper side dirty white, lateral side ochreous–yellow. Segment 3 brown, white at the tip. Scapus deep yellow, antennae brown, circled dirty yellow, densely covered with small hairs in males. Forewing medium wide, triangular, with a relatively well pronounced tornal angle. Its main colour dirty white. Forewing pattern blurred, consists of olive and dark olive spots and bands. Median band most prominent, noticeably widening towards costa, preapical band significantly shorter, runs from tornal angle of wing almost perpendicular to its costal margin. Cilia line olive or dark olive, cilia yellowish white or dirty white. Hindwing light grey, with dirty white cilia.
</paragraph>
<paragraph id="9B30E944FF9FC006FF76F81BFAC8F9DF" blockId="9.[151,1437,1303,2014]" lastBlockId="10.[151,1436,151,429]" lastPageId="10" lastPageNumber="451" pageId="9" pageNumber="450">
<emphasis id="A9FB3556FF9FC005FF76F81BFEB3FEB3" bold="true" box="[199,377,1951,1977]" pageId="9" pageNumber="450">Male genitalia.</emphasis>
(
<figureCitation id="03B4F5C1FF9FC005FE38F81BFE16FEB3" box="[393,476,1951,1977]" captionStart="FIGURES 18–21" captionStartId="10.[151,264,1339,1363]" captionTargetBox="[151,1429,453,1309]" captionTargetId="figure-117@10.[151,1435,453,1316]" captionTargetPageId="10" captionText="FIGURES 18–21. Fig. 18. Adult of Aethes kindermanniana (Croatia). Fig. 19. Male genitalia of Aethes kindermanniana (Croatia). Fig. 20. Adult of Aethes kindermanniana macedonica ssp. nov. (North Macedonia). Fig. 21. Male genitalia of Aethes kindermanniana macedonica ssp. nov. (North Macedonia)." figureDoi="http://doi.org/10.5281/zenodo.5796371" httpUri="https://zenodo.org/record/5796371/files/figure.png" pageId="9" pageNumber="450">Fig. 21</figureCitation>
). Tegumen wide and short. Socii thin, with few spines on underside of their proximal parts. The median process of transtilla wide and short, rounded trapezoidal, with two large spines at apex. Valva rather narrow, noticeably narrowed towards apex. Aedeagus wide and short, with 3 spiny lobes on underside of its tube and on lateral sides, with a long and rather thin median process directed caudad. Cornuti absent.
</paragraph>
<paragraph id="9B30E944FF9CC006FF76FF5BFD17F817" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="451">
<emphasis id="A9FB3556FF9CC006FF76FF5BFE49F9F3" bold="true" box="[199,387,223,249]" pageId="10" pageNumber="451">Molecular data.</emphasis>
BIN BOLD: BOLD AAF5176 (n=2). The differences from barcoded specimens from
<collectingCountry id="E398A9D4FF9CC006FA84FF5BFA52F9F3" box="[1333,1432,223,249]" name="Bulgaria" pageId="10" pageNumber="451">Bulgaria</collectingCountry>
,
<collectingCountry id="E398A9D4FF9CC006FF26FE87FECAF817" box="[151,256,259,285]" name="Germany" pageId="10" pageNumber="451">Germany</collectingCountry>
,
<collectingCountry id="E398A9D4FF9CC006FEBDFE87FEA8F817" box="[268,354,259,285]" name="Finland" pageId="10" pageNumber="451">Finland</collectingCountry>
and
<collectingCountry id="E398A9D4FF9CC006FE29FE87FE38F817" box="[408,498,259,285]" name="Norway" pageId="10" pageNumber="451">Norway</collectingCountry>
is 0.31% p-distance.
</paragraph>
<paragraph id="9B30E944FF9CC006FF76FEA2FC7FF848" blockId="10.[151,1436,151,429]" box="[199,949,294,322]" pageId="10" pageNumber="451">
<emphasis id="A9FB3556FF9CC006FF76FEA2FEF2F84A" bold="true" box="[199,312,294,320]" pageId="10" pageNumber="451">Bionomy.</emphasis>
Bionomy and early stages are unknown. Habitat (
<figureCitation id="03B4F5C1FF9CC006FCDBFEA3FC63F848" box="[874,937,295,322]" captionStart="FIGURES 1–9" captionStartId="7.[151,264,1830,1854]" captionTargetBox="[225,1363,181,1795]" captionTargetId="figure-19@7.[225,1363,181,1806]" captionTargetPageId="7" captionText="FIGURES 1–9. Fig. 1. Adult of Phtheochroa amasiana (North Macedonia). Fig. 2. Male genitalia of Phtheochroa amasiana (North Macedonia). Fig. 3. Adult of Cochylimorpha erlebachi (Croatia). Fig. 4. Female genitalia of Cochylimorpha erlebachi (Croatia). Fig. 5. Adult of Aethes larissae sp. nov. holotype, (North Macedonia). Figs 6-7. Head of Aethes larissae sp. nov. holotype, (North Macedonia). Fig. 8. Male genitalia of Aethes larissae sp. nov. holotype, (North Macedonia). Fig. 9. Habitat of Aethes larissae sp. nov. and Aethes kindermanniana macedonica ssp. nov. (North Macedonia Galičica Nacional Park, Asan Gjura,)." figureDoi="http://doi.org/10.5281/zenodo.5796365" httpUri="https://zenodo.org/record/5796365/files/figure.png" pageId="10" pageNumber="451">Fig.9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D395BACFFF9CC006FF76FECEFCC8F86F" box="[199,770,330,357]" pageId="10" pageNumber="451" type="distribution">
<paragraph id="9B30E944FF9CC006FF76FECEFCC8F86F" blockId="10.[151,1436,151,429]" box="[199,770,330,357]" pageId="10" pageNumber="451">
<emphasis id="A9FB3556FF9CC006FF76FECEFEABF86E" bold="true" box="[199,353,330,356]" pageId="10" pageNumber="451">Distribution.</emphasis>
Known from North
<collectingCountry id="E398A9D4FF9CC006FDF6FECFFD09F86F" box="[583,707,331,357]" name="Macedonia" pageId="10" pageNumber="451">Macedonia</collectingCountry>
only.
</paragraph>
</subSubSection>
<subSubSection id="D395BACFFF9CC006FF76FEEBFDFDF8A7" pageId="10" pageNumber="451" type="etymology">
<paragraph id="9B30E944FF9CC006FF76FEEBFDFDF8A7" blockId="10.[151,1436,151,429]" pageId="10" pageNumber="451">
<emphasis id="A9FB3556FF9CC006FF76FEEBFE84F883" bold="true" box="[199,334,367,393]" pageId="10" pageNumber="451">Etymology.</emphasis>
The name of this new subspecies is topotypical—given following the name of the country North
<collectingCountry id="E398A9D4FF9CC006FF26FE17FEDCF8A7" box="[151,278,403,429]" name="Macedonia" pageId="10" pageNumber="451">Macedonia</collectingCountry>
, where it was discovered.
</paragraph>
</subSubSection>
</treatment>
</document>
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