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<document id="BF209380772DCFC15AC368C46B06B82D" ID-DOI="10.11646/zootaxa.3697.1.1" ID-GBIF-Dataset="b2711d98-f97a-4771-bb6b-ff472a2f5a80" ID-ISSN="1175-5326" ID-Zenodo-Dep="284383" ID-ZooBank="492264BB-E919-447D-9D67-C226DE21A0CE" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460660988741" checkinUser="plazi" docAuthor="Reemer, Menno" docDate="2013" docId="03B4BF12FFD3FFBBFF60F99D4024957C" docLanguage="en" docName="zt03697p088.pdf" docOrigin="Zootaxa 3697 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Stipomorpha elcopala Reemer, 2013, spec. nov." docType="treatment" docVersion="7" lastPageNumber="48" masterDocId="FF8DC76AFFFDFF94FFF7FFAB43079029" masterDocTitle="Taxonomic exploration of Neotropical Microdontinae (Diptera: Syrphidae) mimicking stingless bees" masterLastPageNumber="88" masterPageNumber="1" pageNumber="47" updateTime="1698347572540" updateUser="plazi">
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<mods:title id="0E99A1BDC7C57CF90C4D4269FE0686D3">Taxonomic exploration of Neotropical Microdontinae (Diptera: Syrphidae) mimicking stingless bees</mods:title>
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<mods:namePart id="8D59CE268F49E6285720DC8843F9E7B0">Reemer, Menno</mods:namePart>
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<mods:date id="9B9CC7EE276B1AB98205A347BC750581">2013</mods:date>
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<mods:number id="3533CF584191DA65B591130545BDD52A">3697</mods:number>
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<treatment id="03B4BF12FFD3FFBBFF60F99D4024957C" ID-DOI="http://doi.org/10.5281/zenodo.5624911" ID-GBIF-Taxon="127665218" ID-Zenodo-Dep="5624911" LSID="urn:lsid:plazi:treatment:03B4BF12FFD3FFBBFF60F99D4024957C" httpUri="http://treatment.plazi.org/id/03B4BF12FFD3FFBBFF60F99D4024957C" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<subSubSection id="C3075D8FFFD3FFBAFF60F99D4268965B" pageId="46" pageNumber="47" type="nomenclature">
<paragraph id="8BA20E04FFD3FFBAFF60F99D411A9679" blockId="46.[151,541,1590,1650]" box="[151,541,1590,1616]" pageId="46" pageNumber="47">
<heading id="D0EAB968FFD3FFBAFF60F99D411A9679" bold="true" box="[151,541,1590,1616]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B969D216FFD3FFBAFF60F99D411A9679" bold="true" box="[151,541,1590,1616]" pageId="46" pageNumber="47">
<taxonomicName id="4C1D7587FFD3FFBAFF60F99D42999679" ID-CoL="8F6HK" box="[151,414,1590,1616]" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="elcopala" status="sp. nov.">
<emphasis id="B969D216FFD3FFBAFF60F99D42999679" bold="true" box="[151,414,1590,1616]" italics="true" pageId="46" pageNumber="47">Stipomorpha elcopala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A6F6DFFD3FFBAFE52F99D411A9679" box="[421,541,1590,1616]" pageId="46" pageNumber="47" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA20E04FFD3FFBAFF60F9F24268965B" blockId="46.[151,541,1590,1650]" box="[151,367,1625,1650]" pageId="46" pageNumber="47">
<figureCitation id="13261281FFD3FFBAFF60F9F24235965B" box="[151,306,1625,1650]" captionStart="FIGURES 130 141. 130 133" captionStartId="45.[151,264,1850,1872]" captionTargetBox="[185,1407,218,1801]" captionTargetId="figure@45.[151,1436,193,1828]" captionTargetPageId="45" captionText="FIGURES 130 141. 130 133. Stipomorpha crematogastri female (holotype). 130. head frontal; 131. head lateral; 132. puparium dorsal; 133. puparium lateral. 134 137. Stipomorpha dichromata female (holotype). 134. habitus dorsal; 135. habitus lateral; 136. head frontal; 137. head lateral. 138 141. Stipomorpha elcopala male (holotype). 138. head frontal; 139. habitus lateral; 140. habitus dorsal; 141. wing." httpUri="https://zenodo.org/record/284398/files/figure.png" pageId="46" pageNumber="47">Figs 138141</figureCitation>
,
<figureCitation id="13261281FFD3FFBAFEC9F9F1426C965B" box="[318,363,1626,1650]" captionStart="FIGURES 222 232. 222 224" captionStartId="73.[151,264,1766,1788]" captionTargetBox="[188,1407,227,1712]" captionTargetId="figure@73.[151,1436,193,1745]" captionTargetPageId="73" captionText="FIGURES 222 232. 222 224. Stipomorpha wheeleri. 222. male (paratype), head frontal. 223. female (holotype), habitus dorsal; 224. female (holotype); habitus lateral. 225 227. Stipomorpha zophera male (holotype). 225. habitus dorsal; 226. habitus lateral; 227. head frontal. 228 229. Stipomorpha, tergites 1 &amp; 2 dorsal. 228. S. mendax; 229. S. mixta. 230 232. Stipomorpha, male genitalia lateral. 230. S. apicula (holotype); 231. S. elcopala (holotype); 232. S. fallax (holotype)." httpUri="https://zenodo.org/record/284406/files/figure.png" pageId="46" pageNumber="47">231</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFD3FFBAFF60F90A41619760" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BA20E04FFD3FFBAFF60F90A42B7972B" blockId="46.[151,1436,1697,2010]" pageId="46" pageNumber="47">
<emphasis id="B969D216FFD3FFBAFF60F90A42BA9693" bold="true" box="[151,445,1697,1722]" pageId="46" pageNumber="47">
Studied
<typeStatus id="54A6B0A6FFD3FFBAFF0AF90A42379693" box="[253,304,1697,1722]" pageId="46" pageNumber="47">type</typeStatus>
specimens.
</emphasis>
<typeStatus id="54A6B0A6FFD3FFBAFE3FF90941679693" box="[456,608,1698,1722]" pageId="46" pageNumber="47" type="holotype">HOLOTYPE</typeStatus>
.
<collectingCountry id="F30A4E94FFD3FFBAFD98F909401C9693" box="[623,795,1698,1722]" name="Costa Rica" pageId="46" pageNumber="47">COSTA RICA</collectingCountry>
.
<collectingCountry id="F30A4E94FFD3FFBAFCDDF90A40609693" box="[810,871,1697,1722]" name="Malta" pageId="46" pageNumber="47">Male</collectingCountry>
. Label 1: “
<collectingCountry id="F30A4E94FFD3FFBAFC19F909479B9693" box="[1006,1180,1698,1722]" name="Costa Rica" pageId="46" pageNumber="47">COSTA RICA</collectingCountry>
. Cartago, Cordillera / central, Pejivalle, Humo, El Copal /
<quantity id="4CE5A3E1FFD3FFBAFDC7F96E418296F4" box="[560,645,1733,1758]" metricMagnitude="3" metricUnit="m" metricValue="1.026" pageId="46" pageNumber="47" unit="m" value="1026.0">1026 m</quantity>
asl. Malaise trap. /
<geoCoordinate id="EE2968C3FFD3FFBAFC94F96E470596F7" box="[867,1026,1733,1758]" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="1" value="9.782194">09°4655.9”N</geoCoordinate>
<geoCoordinate id="EE2968C3FFD3FFBAFBFEF96E47AA96F7" box="[1033,1197,1733,1758]" direction="west" orientation="longitude" pageId="46" pageNumber="47" precision="1" value="-83.752556">83°4509.2”W</geoCoordinate>
/
<date id="FFA328C4FFD3FFBAFB33F96D465896F7" box="[1220,1375,1734,1758]" pageId="46" pageNumber="47" value="2010-09-09" valueMax="2010-09-10" valueMin="2010-09-09">9-10.IX.2010</date>
. Leg. J.T. Smit.”. Coll.
<taxonomicName id="4C1D7587FFD3FFBAFEADF94142AC972B" box="[346,427,1770,1794]" class="Insecta" family="Syrphidae" genus="Mermerizon" kingdom="Animalia" order="Diptera" pageId="46" pageNumber="44" phylum="Arthropoda" rank="species" species="inbio">INBIO</taxonomicName>
.
</paragraph>
<paragraph id="8BA20E04FFD3FFBAFF30F8A541619760" blockId="46.[151,1436,1697,2010]" pageId="46" pageNumber="47">
<typeStatus id="54A6B0A6FFD3FFBAFF30F8A54251970F" box="[199,342,1806,1830]" pageId="46" pageNumber="47" type="paratype">PARATYPE</typeStatus>
.
<collectingCountry id="F30A4E94FFD3FFBAFE9FF8A54100970F" box="[360,519,1806,1830]" name="Honduras" pageId="46" pageNumber="47">HONDURAS</collectingCountry>
.
<collectingCountry id="F30A4E94FFD3FFBAFDEDF8A64150970F" box="[538,599,1805,1830]" name="Malta" pageId="46" pageNumber="47">Male</collectingCountry>
. Label 1: “
<collectingCountry id="F30A4E94FFD3FFBAFD1EF8A6405B970F" box="[745,860,1805,1830]" name="Honduras" pageId="46" pageNumber="47">Honduras</collectingCountry>
: Yoro, Palo de / Comba,
<geoCoordinate id="EE2968C3FFD3FFBAFB50F8A6460E970F" box="[1191,1289,1805,1830]" direction="north" orientation="latitude" pageId="46" pageNumber="47" precision="925" value="15.183333">15°11N</geoCoordinate>
,
<geoCoordinate id="EE2968C3FFD3FFBAFAEAF8A54683970F" box="[1309,1412,1806,1830]" direction="west" orientation="longitude" pageId="46" pageNumber="47" precision="925" value="-87.65">87°39W</geoCoordinate>
/
<date id="FFA328C4FFD3FFBAFF60F89A42119763" box="[151,278,1841,1866]" pageId="46" pageNumber="47" value="1995-09-29">29.IX.1995</date>
/ leg. R. Cave”. Coll. MZLU.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFD3FFBBFF30F8FE42A693EE" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="description">
<paragraph id="8BA20E04FFD3FFBAFF30F8FE41309747" blockId="46.[151,1436,1697,2010]" box="[199,567,1877,1902]" pageId="46" pageNumber="47">
<emphasis id="B969D216FFD3FFBAFF30F8FE42559747" bold="true" box="[199,338,1877,1902]" pageId="46" pageNumber="47">Description</emphasis>
(based on
<typeStatus id="54A6B0A6FFD3FFBAFE3CF8FE41289747" box="[459,559,1877,1902]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BA20E04FFD3FFBAFF30F8D2412797B8" blockId="46.[151,1436,1697,2010]" box="[199,544,1913,1938]" pageId="46" pageNumber="47">
<emphasis id="B969D216FFD3FFBAFF30F8D2425497BB" bold="true" box="[199,339,1913,1938]" pageId="46" pageNumber="47">Adult male.</emphasis>
Body size:
<quantity id="4CE5A3E1FFD3FFBAFE20F8D2411B97B8" box="[471,540,1913,1938]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="46" pageNumber="47" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
<paragraph id="8BA20E04FFD3FFBBFF30F83646419134" blockId="46.[151,1436,1697,2010]" lastBlockId="47.[151,1437,151,1365]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="B969D216FFD3FFBAFF30F8364209979F" bold="true" box="[199,270,1949,1974]" pageId="46" pageNumber="47">Head.</emphasis>
Face occupying 1/4 of head width in frontal view; black with narrow yellow lateral margins; entirely white pilose. Gena black. Occiput black; black pilose dorsally, white pilose ventrally. Oral cavity with slightly produced lateral margins. Frons black; short black pilose, except for bare triangular part posterior to lunula. Vertex black; black pilose. Eye bare. Antennal fossa about as wide as high. Antenna black; scape and pedicel black pilose; antennal ratio 3:1:6; basoflagellomere parallel-sided with narrowly rounded apex, with small sensory pit at 3/4 from base; arista slender, about 3/4 of length of basoflagellomere, appearing bare under low magnification.
</paragraph>
<paragraph id="8BA20E04FFD2FFBBFF30FE8C47ED91F9" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FE8C42219169" bold="true" box="[199,294,295,320]" pageId="47" pageNumber="48">Thorax.</emphasis>
Black. Postpronotum, scutum and postalar callus short black pilose, except for two small patches of white pile along transverse suture and two small white pilose patches anterior to scutellum. Scutellum subrectangular, without calcars; white pilose. Anepisternum a little convex, without sulcus, black pilose anterodorsally and along posterior margin. Anepimeron black pilose dorsally, white pilose ventrally. Katepisternum white pilose dorsally. Katepimeron bare. Calypter brownish yellow, halter blackish.
</paragraph>
<paragraph id="8BA20E04FFD2FFBBFF30FE7747319214" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">
Wing: hyaline, with very faint dark cloud between apex of costal cell and vena
<taxonomicName id="4C1D7587FFD2FFBBFBC2FE77477B91DC" box="[1077,1148,476,501]" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="spuria">spuria</taxonomicName>
, and with yellowish veins bordering and posterior to pterostigma; microtrichose except bare basally on cell r1 along vein RS, on posterior 1/2 of cell br, basal 5/6 of cell bm, anterior 1/2 of cell cup and mediobasal 1/5 of alula.
</paragraph>
<paragraph id="8BA20E04FFD2FFBBFF30FDE341219281" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">Legs: brownish black, except fifth tarsomeres of all legs yellow. Legs black pilose, except anterior four tibiae posteriorly pale pilose and hind-tibia pale pilose on basal 2/3; pile on hind-tibia about as long as width of tibia. Coxae and trochanters black pilose.</paragraph>
<paragraph id="8BA20E04FFD2FFBBFF30FD1F42BD938D" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FD1F424492E4" bold="true" box="[199,323,692,717]" pageId="47" pageNumber="48">Abdomen.</emphasis>
Reddish, except lateral margins of tergites 1 and 2 blackish brown. Tergite 2 wider than thorax, widest point at around half the lentgh; tergites 3 and 4 strongly narrowing. Antetergite very large. Tergite 1 laterally mixed black and yellow pilose, bare medially. Tergite 2 black pilose laterally, yellow pilose anterolaterally and sublaterally; almost bare medially. Tergites 3 and 4 fused, without a visible suture; sparseley black pilose medially. Sternite 1 bare, separated from sternite 2 by a membrane of about the width of sternite 1. Sternite 2 bare, laterally more than twice as wide as medially, separated from sternite 3 by a membrane of twice the median width of sternite 2. Genitalia as in fig. 231.
</paragraph>
<paragraph id="8BA20E04FFD2FFBBFF30FC0442A693EE" blockId="47.[151,1437,151,1365]" box="[199,417,943,968]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FC04422293E1" bold="true" box="[199,293,943,968]" pageId="47" pageNumber="48">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFD2FFBBFF30FC7F400C9439" pageId="47" pageNumber="48" type="etymology">
<paragraph id="8BA20E04FFD2FFBBFF30FC7F400C9439" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FC7F424993C4" bold="true" box="[199,334,980,1005]" pageId="47" pageNumber="48">Etymology.</emphasis>
The name of this species refers to its
<typeStatus id="54A6B0A6FFD2FFBBFD03FC7E402393C4" box="[756,804,981,1005]" pageId="47" pageNumber="48">type</typeStatus>
locality: El Copal, a nature reserve in the Central Valley of
<collectingCountry id="F30A4E94FFD2FFBBFF42FC5342349439" box="[181,307,1015,1040]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
. The name is to be treated as an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFD2FFBBFF30FBB741E99519" pageId="47" pageNumber="48" type="diagnosis">
<paragraph id="8BA20E04FFD2FFBBFF30FBB7422294CE" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FBB74246941C" bold="true" box="[199,321,1052,1077]" pageId="47" pageNumber="48">Diagnosis.</emphasis>
Although the black thorax and legs in combination with the reddish abdomen remind of
<taxonomicName id="4C1D7587FFD2FFBBFF60FB9441069471" authority="Curran" authorityName="Curran" box="[151,513,1087,1112]" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="apicula">
<emphasis id="B969D216FFD2FFBBFF60FB94428B9471" box="[151,396,1087,1112]" italics="true" pageId="47" pageNumber="48">Stipomorpha apicula</emphasis>
(Curran)
</taxonomicName>
, this species is morphologically most similar to
<taxonomicName id="4C1D7587FFD2FFBBFB84FB9442089454" authority="Shannon" authorityName="Shannon" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="lacteipennis">
<emphasis id="B969D216FFD2FFBBFB84FB94469B9471" box="[1139,1436,1087,1112]" italics="true" pageId="47" pageNumber="48">Stipomorpha lacteipennis</emphasis>
(Shannon)
</taxonomicName>
. These two species share similar length ratios of the antennal segments, a pilose posterior margin of the anepisternum, a pilose dorsal part of the katepimeron, a partially bare alula, a notably triangular abdomen, and similar morphology of the male genitalia. However, the reddish abdomen readily separates
<taxonomicName id="4C1D7587FFD2FFBBFB4BFB06463B94EC" box="[1212,1340,1196,1221]" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="elcopala">
<emphasis id="B969D216FFD2FFBBFB4BFB06463B94EC" box="[1212,1340,1196,1221]" italics="true" pageId="47" pageNumber="48">S. elcopala</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1D7587FFD2FFBBFA70FB06421994C1" class="Insecta" family="Syrphidae" genus="Stipomorpha" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="lacteipennis">
<emphasis id="B969D216FFD2FFBBFA70FB06421994C1" italics="true" pageId="47" pageNumber="48">S. lacteipennis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA20E04FFD2FFBBFF30FB5F41E99519" blockId="47.[151,1437,151,1365]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FB5F42169524" bold="true" box="[199,273,1268,1293]" pageId="47" pageNumber="48">Notes.</emphasis>
The
<typeStatus id="54A6B0A6FFD2FFBBFEBDFB5E42AC9524" box="[330,427,1269,1293]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
differs from the
<typeStatus id="54A6B0A6FFD2FFBBFD9EFB5F41CC9524" box="[617,715,1268,1293]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
in its larger size (
<quantity id="4CE5A3E1FFD2FFBBFC63FB5F40EB9525" box="[916,1004,1268,1293]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="47" pageNumber="48" unit="mm" value="6.5">6.5 mm</quantity>
) and the entirely yellow tergites 1 and 2 (lateral margins not blackish brown as in
<typeStatus id="54A6B0A6FFD2FFBBFD8BFABC41E49519" box="[636,739,1303,1328]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C3075D8FFFD2FFBBFF30FA974024957C" box="[199,803,1340,1365]" pageId="47" pageNumber="48" type="distribution">
<paragraph id="8BA20E04FFD2FFBBFF30FA974024957C" blockId="47.[151,1437,151,1365]" box="[199,803,1340,1365]" pageId="47" pageNumber="48">
<emphasis id="B969D216FFD2FFBBFF30FA974267957C" bold="true" box="[199,352,1340,1365]" pageId="47" pageNumber="48">Distribution.</emphasis>
Known from
<collectingCountry id="F30A4E94FFD2FFBBFE09FA97417F957C" box="[510,632,1340,1365]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
and
<collectingCountry id="F30A4E94FFD2FFBBFD59FA974018957C" box="[686,799,1340,1365]" name="Honduras" pageId="47" pageNumber="48">Honduras</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>