279 lines
36 KiB
XML
279 lines
36 KiB
XML
<document ID-CLB-Dataset="20724" ID-DOI="http://dx.doi.org/10.3897/dez.69.79931" ID-GBIF-Dataset="e58ae50b-2b62-4c1c-b1fd-09b0f6aa001d" ID-Pensoft-Pub="1860-1324-1-19" ID-Pensoft-UUID="024B3AF3339C5F8CAA644EA4F78EA082" ID-ZooBank="02E8488BDDA7464CABC239424200939E" ModsDocID="1860-1324-69-1-19" checkinTime="1644448212315" checkinUser="pensoft" docAuthor="Schmidt, Joachim, Scholz, Stephan & Will, Kipling" docDate="2022" docId="F42684FF1B2F5D3291B7385129F63ED1" docLanguage="en" docName="DeutEntomolZeit 69(1): 19-44" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-02-09" docSource="http://dx.doi.org/10.3897/dez.69.79931" docTitle="Quasicalathus conservans Schmidt & Will 2022, sp. nov." docType="treatment" docUuid="1EAF9667-E896-445B-ADED-0DA2B055DF2B" docUuidSource="ZooBank" docVersion="6" id="024B3AF3339C5F8CAA644EA4F78EA082" lastPageNumber="19" masterDocId="024B3AF3339C5F8CAA644EA4F78EA082" masterDocTitle="Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3 D imaging" masterLastPageNumber="44" masterPageNumber="19" pageNumber="19" updateTime="1732797172808" updateUser="ExternalLinkService">
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<mods:titleInfo id="F217C4C14525D3AC6218F47547C4910F">
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<mods:title id="CC17939B5CF74E2AB5CBFA94C269BC9B">Character analysis and descriptions of Eocene sphodrine fossils (Coleoptera, Carabidae) using light microscopy, micro-CT scanning, and 3 D imaging</mods:title>
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<mods:name id="ADAA02502C1487EC7A4188243A9720AB" type="personal">
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<mods:roleTerm id="0AC634E4F4F77CA27ABE6C505FCB466B">Author</mods:roleTerm>
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<mods:namePart id="0EB90A1279DF6AE8A5FC98972523B391">Schmidt, Joachim</mods:namePart>
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<mods:affiliation id="AAE2FFC9B3296F6B671BB7D442624216">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitaetsplatz 2, 18055 Rostock, German</mods:affiliation>
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<mods:nameIdentifier id="115289C107744E9B214ADA94F1D985CF" type="email">agonumschmidt@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="50900A175D87A9CF71B0BEB513B119F4">Author</mods:roleTerm>
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<mods:namePart id="CB5D82FA6A8E61A32C0DC0D5B842E1E4">Scholz, Stephan</mods:namePart>
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<mods:affiliation id="76FCB07A967AFFD59F1339D6607714A7">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitaetsplatz 2, 18055 Rostock, German</mods:affiliation>
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<mods:name id="89E242F8427B33F76D6EDAA9F167C992" type="personal">
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<mods:roleTerm id="018FC3A2AEADBB6B5C031FF8A6129524">Author</mods:roleTerm>
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<mods:namePart id="DAE7DFBE9712ED273C16A5A20494A39F">Will, Kipling</mods:namePart>
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<mods:nameIdentifier id="1C25C0B24778A679E522E2703C4ED767" type="ORCID">https://orcid.org/0000-0002-7056-9011</mods:nameIdentifier>
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<mods:affiliation id="BF875EC915A771A9EA1175A17C4443A7">Essig Museum of Entomology, 1101 Valley Life Sciences Building, # 4780 University of California, Berkeley, CA 94720 - 4780, USA</mods:affiliation>
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<mods:typeOfResource id="9E778DC1C2A150FAD9B5C44472B89A5A">text</mods:typeOfResource>
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<mods:title id="2DFB9DC590A6F85B5E3AB4A6AF639285">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="386CA500D7C8C2E8F0482DFF5C0A1DC8">2022</mods:date>
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<mods:number id="5AA244585D6822239A7E5FA2F3FF96DE">2022-02-09</mods:number>
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<mods:detail id="AB1682921B6AD11C9DD5FE7F8A6A6650" type="volume">
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<mods:number id="519C6BE75C7C2991818EC9C0EC382417">69</mods:number>
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<mods:number id="538052E9B7CB274103D9D15B778054D6">1</mods:number>
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<mods:start id="EBDB4D911EE0CCA5A8A7F95C86FE0D24">19</mods:start>
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<mods:end id="4B11DFB13C9AFDC5FD473755A66D1C4D">44</mods:end>
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<mods:url id="30AA0859B5838812A362993B7ABFF045">http://dx.doi.org/10.3897/dez.69.79931</mods:url>
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<mods:classification id="530D4690BFD66E27B44E5891019475F7">journal article</mods:classification>
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<mods:identifier id="F65E8097E472AC263D8F3F4F2817FC2D" type="DOI">http://dx.doi.org/10.3897/dez.69.79931</mods:identifier>
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<mods:identifier id="E6D45EA9D07C4951AE926DD0EE2E1B39" type="Pensoft-Pub">1860-1324-1-19</mods:identifier>
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<mods:identifier id="4B9A358ABFE7B7C8AE69F052FA58553E" type="Pensoft-UUID">024B3AF3339C5F8CAA644EA4F78EA082</mods:identifier>
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<subSection id="C5775D80DCBBDF2A03D02D8E99CFC82A" pageId="0" pageNumber="19" type="results">
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<treatment id="F42684FF1B2F5D3291B7385129F63ED1" ID-GBIF-Taxon="193323625" LSID="urn:lsid:zoobank.org:act:1EAF9667-E896-445B-ADED-0DA2B055DF2B" httpUri="http://treatment.plazi.org/id/F42684FF1B2F5D3291B7385129F63ED1" lastPageNumber="19" pageId="0" pageNumber="19" scope_family="Carabidae" scope_order="Coleoptera">
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<subSubSection id="1710DF34158F9944FCB743C6DA810C24" pageId="0" pageNumber="19" type="nomenclature">
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<paragraph id="4FEDB00E5ABEACB82373A4FC1D84290D" pageId="0" pageNumber="19">
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<taxonomicName id="A4A923D96F6C2BA2F126641FEE455361" LSID="http://zoobank.org1EAF9667-E896-445B-ADED-0DA2B055DF2B" authority="Schmidt & Will" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans" status="sp. nov.">Quasicalathus conservans Schmidt & Will</taxonomicName>
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<taxonomicNameLabel id="5A6E0E0A4F921EB6ECF35FBD659CA53E" pageId="0" pageNumber="19">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="2B9B33ADDA2B10C4987143C5C166BDC0" pageId="0" pageNumber="19" type="materials_examined">
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<paragraph id="B216CEAC2C5ED745F7CC14E85D4B2EE5" pageId="0" pageNumber="19">Material studied.</paragraph>
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<paragraph id="97864569A337DEDB2CEEE78097F27EED" pageId="0" pageNumber="19">
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Male in Rovno amber, with specimen label data "
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<emphasis id="D1276E46AEB3746DCE97C1C65D20152E" bold="true" pageId="0" pageNumber="19">SDEI-Amb-002529</emphasis>
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", deposited in the Senckenberg Deutsches Entomologisches Institut,
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<normalizedToken id="A72524F6BAE1FF563B662E6AB26C97E9" originalValue="Müncheberg">Muencheberg</normalizedToken>
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,
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<collectingCountry id="14A2893D945CB17B850B8E2B1EBB6D82" name="Germany">Germany</collectingCountry>
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. Size of the amber piece 20
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<normalizedToken id="9486496ED7228EA91966BA3597062769" originalValue="×">x</normalizedToken>
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10
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<normalizedToken id="CDF6DA98C58A4E646577F917D66D0C73" originalValue="×">x</normalizedToken>
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<quantity id="9B75999AD44AA39D3F36A836CAE54C26" metricMagnitude="-3" metricUnit="m" metricValue="8.5" unit="mm" value="8.5">8.5 mm</quantity>
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.
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</paragraph>
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<paragraph id="3E349C078AA215214CF3BE7FDC52FC26" pageId="0" pageNumber="19">
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Preservation status: The amber is clear but pervaded by numerous small air bubbles and extensive flowlines attached to the embedded fossil that is therefore only partly visible using light microscopy (Figs
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<figureCitation id="D49D56DF4D493664C326B3C1D5CA12B8" captionStart="Figures 31–33" captionStartId="F6" captionText="Figures 31 - 33. Quasicalathus, light microscopic images of the holotypes of Q. agonicollis sp. nov. (31.) and Q. conservans sp. nov. (32, 33.). 31. Ventral side of head showing chaetotaxy of mentum; 32. General view of the amber piece with fossil in dorsal view; 32. Left lateral view. Abbreviations: ce - compound eye; el - elytron; ems - external seta of submentum; gu - gula; ims - internal seta of submentum; msf - mesofemur; mt - mentum; mtf - metafemur; prf - profemur; pt - pronotum." figureDoi="10.3897/dez.69.79931.figures31-33" httpUri="https://binary.pensoft.net/fig/643956" pageId="0" pageNumber="19">32</figureCitation>
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,
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<figureCitation id="B37769D99DA4A18A5271B407E9DDA7D2" captionStart="Figures 31–33" captionStartId="F6" captionText="Figures 31 - 33. Quasicalathus, light microscopic images of the holotypes of Q. agonicollis sp. nov. (31.) and Q. conservans sp. nov. (32, 33.). 31. Ventral side of head showing chaetotaxy of mentum; 32. General view of the amber piece with fossil in dorsal view; 32. Left lateral view. Abbreviations: ce - compound eye; el - elytron; ems - external seta of submentum; gu - gula; ims - internal seta of submentum; msf - mesofemur; mt - mentum; mtf - metafemur; prf - profemur; pt - pronotum." figureDoi="10.3897/dez.69.79931.figures31-33" httpUri="https://binary.pensoft.net/fig/643956" pageId="0" pageNumber="19">33</figureCitation>
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). The exoskeleton of the fossil, including most parts of the aedeagus, is well preserved and, therefore, important diagnostic characters could be reconstructed using micro-CT (Figs
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<figureCitation id="2EDD9356193697F663B09AFB30BCE8ED" captionStart="Figures 78–80" captionStartId="F15" captionText="Figures 78 - 80. Quasicalathus conservans sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 78. Dorsal aspect; 79. Left lateral aspect (aed - aedeagus); 80. Ventral aspect; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software." figureDoi="10.3897/dez.69.79931.figures78-80" httpUri="https://binary.pensoft.net/fig/643965" pageId="0" pageNumber="19">78</figureCitation>
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-
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<figureCitation id="ED3976A3AFBCF5CB3451DA9443096132" captionStart="Figures 81–89" captionStartId="F16" captionText="Figures 81 - 89. Quasicalathus conservans sp. nov., volume rendering of the holotype. 81. Head, dorsal aspect (the arrows point to the insertions of the supraorbital setae); 82. Head, ventral aspect; 83. Left external part of metathorax, ventral view; 84. Submentum (the arrows point to the insertions of the four lateral setae); 85. Posterior part of prosternum and procoxae; 86. Posterior part of metasternum and metacoxae; 87 - 89. Preserved remains of the aedeagus (87. Right lateral aspect; 88. Dorsal aspect; 89. Left lateral aspect). Abbreviations: bb - basal bulb of aedeagal median lobe; ce - compound eye; cxp - metacoxal plate; eph - partly evaginated lobes of endophallus; gu - gula; mem - metepimeron; mes - metepisternum; mtt - mentum tooth; mv - metaventrite; pcx - procoxa; pmr - preserved distal part of right paramere of aedeagal median lobe; psp - prosternal process; sc - scutellum; sps - setae of sensory pit; tl - terminal lamella of aedeagal median lobe." figureDoi="10.3897/dez.69.79931.figures81-89" httpUri="https://binary.pensoft.net/fig/643966" pageId="0" pageNumber="19">89</figureCitation>
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).
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</paragraph>
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<paragraph id="7A17640601317296E408E887C5E10428" pageId="0" pageNumber="19">Syninclusions: One stellate hair, few tiny dust particles.</paragraph>
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</subSubSection>
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<subSubSection id="47062BB518F2F079D0E5FF257F3AE43A" pageId="0" pageNumber="19" type="description">
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<paragraph id="EF2F7794B70383BF6F88600FF1BBC27C" pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph id="B69316B6B6B07DEFF59BEA7E816C1A7A" pageId="0" pageNumber="19">
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Measurements see Table
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<tableCitation id="AA2B3CF5081450AC1CCDA1EB2CE32A01" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Measurements [µm]." httpUri="http://table.plazi.org/id/C19891A363A03B4DD93DC20ECFC36EE4" pageId="0" pageNumber="19" tableUuid="C19891A363A03B4DD93DC20ECFC36EE4">2</tableCitation>
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.
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</paragraph>
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<paragraph id="7A50F82520448E14FCF7B55EF8CABA1B" pageId="0" pageNumber="19">Standardized body length: 7.3 mm.</paragraph>
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<paragraph id="252803ADBA20A5607C2CB2B645EE1BC0" pageId="0" pageNumber="19">Proportions: A3L/HL = 0.44;</paragraph>
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<paragraph id="CAFEB290400A3BEE1BCFD4A76C23DFF0" pageId="0" pageNumber="19">EyL/ HW(-) = 0.72;</paragraph>
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<paragraph id="92CADF363D0E2C08F9F8C54D1EB8502E" pageId="0" pageNumber="19">PW/HW(+) = 1.40;</paragraph>
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<paragraph id="5941AF75C2324408B6CAC27D5146919F" pageId="0" pageNumber="19">PW/PL = 1.27;</paragraph>
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<paragraph id="013A82A9C2F0D9AC8D7CCC498E0FAA3B" pageId="0" pageNumber="19">PW/PWb = 1.11;</paragraph>
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<paragraph id="A0418EE70A2F26B659296954F14054D7" pageId="0" pageNumber="19">PWb/PWa = 1.46;</paragraph>
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<paragraph id="6EA089928F1D2D386DC997E946C4ADF6" pageId="0" pageNumber="19">EW/PW = 1.64;</paragraph>
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<paragraph id="564436DA9FCB931D25CBB5FA1A8148A1" pageId="0" pageNumber="19">EL/EW = 1.51;</paragraph>
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<paragraph id="382AE2F88D797DD2C53C8573E52327C4" pageId="0" pageNumber="19">EpL/EpW = 1.43;</paragraph>
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<paragraph id="8459972544DBF81EDB7BF58CE1E06DBF" pageId="0" pageNumber="19">EL/FL = 2.58;</paragraph>
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<paragraph id="8C8A539F82DCD58F9CD0050383C7512F" pageId="0" pageNumber="19">EL/AedL = 3.64.</paragraph>
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<paragraph id="0DF8B740249A580DC581E466F38A0B25" pageId="0" pageNumber="19">
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Head: As described in
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<taxonomicName id="B1B3E2ABCDA8BC3E3177930534CF5B7F" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
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<emphasis id="CB07447867A11572782B7ADA78DD0043" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="B0155381F07705BA340ECEC9BAFF646A" pageId="0" pageNumber="19">
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Prothorax: Pronotal lateral margin moderately narrowed toward base, slightly concave before laterobasal angles, angles slightly obtuse (Fig.
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<figureCitation id="6456A720D58C15C84107B50B42267B2F" captionStart="Figures 78–80" captionStartId="F15" captionText="Figures 78 - 80. Quasicalathus conservans sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 78. Dorsal aspect; 79. Left lateral aspect (aed - aedeagus); 80. Ventral aspect; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software." figureDoi="10.3897/dez.69.79931.figures78-80" httpUri="https://binary.pensoft.net/fig/643965" pageId="0" pageNumber="19">78</figureCitation>
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). Prosternal process with traces of a lateral bead (Fig.
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<figureCitation id="4299C955179F7581DCAE49DA0ECAD698" captionStart="Figures 81–89" captionStartId="F16" captionText="Figures 81 - 89. Quasicalathus conservans sp. nov., volume rendering of the holotype. 81. Head, dorsal aspect (the arrows point to the insertions of the supraorbital setae); 82. Head, ventral aspect; 83. Left external part of metathorax, ventral view; 84. Submentum (the arrows point to the insertions of the four lateral setae); 85. Posterior part of prosternum and procoxae; 86. Posterior part of metasternum and metacoxae; 87 - 89. Preserved remains of the aedeagus (87. Right lateral aspect; 88. Dorsal aspect; 89. Left lateral aspect). Abbreviations: bb - basal bulb of aedeagal median lobe; ce - compound eye; cxp - metacoxal plate; eph - partly evaginated lobes of endophallus; gu - gula; mem - metepimeron; mes - metepisternum; mtt - mentum tooth; mv - metaventrite; pcx - procoxa; pmr - preserved distal part of right paramere of aedeagal median lobe; psp - prosternal process; sc - scutellum; sps - setae of sensory pit; tl - terminal lamella of aedeagal median lobe." figureDoi="10.3897/dez.69.79931.figures81-89" httpUri="https://binary.pensoft.net/fig/643966" pageId="0" pageNumber="19">85</figureCitation>
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). In all other characters as described for the new genus, above.
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</paragraph>
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<paragraph id="5019CD06B57A1253AD9C514BE746DF71" pageId="0" pageNumber="19">
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Pterothorax: Elytra with basal margin markedly concave and humerus markedly protruded anteriorly; basal margin forming a slightly obtuse angle (ca. 115°) with the lateral margin (Fig.
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<figureCitation id="DCC8F0F1A06CE58036C707DBEBBA416F" captionStart="Figures 78–80" captionStartId="F15" captionText="Figures 78 - 80. Quasicalathus conservans sp. nov., volume rendering of the holotype using different grey scales of the Amira software. 78. Dorsal aspect; 79. Left lateral aspect (aed - aedeagus); 80. Ventral aspect; the aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software." figureDoi="10.3897/dez.69.79931.figures78-80" httpUri="https://binary.pensoft.net/fig/643965" pageId="0" pageNumber="19">78</figureCitation>
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). Elytral striae moderately deeply engraved, intervals moderately convex. In all other characters as described for the new genus, above.
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</paragraph>
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<paragraph id="701C17B96F5CB766CE9C726E77F3CB22" pageId="0" pageNumber="19">
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Aedeagus: Length of median lobe about 1.33 mm; median lobe terminal lamella moderately long, almost evenly narrowed from base to apex (Fig.
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<figureCitation id="EE5DE9F9BBDABA0B114EC3B6F407D062" captionStart="Figures 81–89" captionStartId="F16" captionText="Figures 81 - 89. Quasicalathus conservans sp. nov., volume rendering of the holotype. 81. Head, dorsal aspect (the arrows point to the insertions of the supraorbital setae); 82. Head, ventral aspect; 83. Left external part of metathorax, ventral view; 84. Submentum (the arrows point to the insertions of the four lateral setae); 85. Posterior part of prosternum and procoxae; 86. Posterior part of metasternum and metacoxae; 87 - 89. Preserved remains of the aedeagus (87. Right lateral aspect; 88. Dorsal aspect; 89. Left lateral aspect). Abbreviations: bb - basal bulb of aedeagal median lobe; ce - compound eye; cxp - metacoxal plate; eph - partly evaginated lobes of endophallus; gu - gula; mem - metepimeron; mes - metepisternum; mtt - mentum tooth; mv - metaventrite; pcx - procoxa; pmr - preserved distal part of right paramere of aedeagal median lobe; psp - prosternal process; sc - scutellum; sps - setae of sensory pit; tl - terminal lamella of aedeagal median lobe." figureDoi="10.3897/dez.69.79931.figures81-89" httpUri="https://binary.pensoft.net/fig/643966" pageId="0" pageNumber="19">88</figureCitation>
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); terminal lamella almost straight, with tip very slightly bent ventrally (Figs
|
||
<figureCitation id="5602B1E2C314BE9072F9151CC77A325E" captionStart="Figures 81–89" captionStartId="F16" captionText="Figures 81 - 89. Quasicalathus conservans sp. nov., volume rendering of the holotype. 81. Head, dorsal aspect (the arrows point to the insertions of the supraorbital setae); 82. Head, ventral aspect; 83. Left external part of metathorax, ventral view; 84. Submentum (the arrows point to the insertions of the four lateral setae); 85. Posterior part of prosternum and procoxae; 86. Posterior part of metasternum and metacoxae; 87 - 89. Preserved remains of the aedeagus (87. Right lateral aspect; 88. Dorsal aspect; 89. Left lateral aspect). Abbreviations: bb - basal bulb of aedeagal median lobe; ce - compound eye; cxp - metacoxal plate; eph - partly evaginated lobes of endophallus; gu - gula; mem - metepimeron; mes - metepisternum; mtt - mentum tooth; mv - metaventrite; pcx - procoxa; pmr - preserved distal part of right paramere of aedeagal median lobe; psp - prosternal process; sc - scutellum; sps - setae of sensory pit; tl - terminal lamella of aedeagal median lobe." figureDoi="10.3897/dez.69.79931.figures81-89" httpUri="https://binary.pensoft.net/fig/643966" pageId="0" pageNumber="19">87</figureCitation>
|
||
,
|
||
<figureCitation id="30AFA7319A9FE259D6678CA47D179DC8" captionStart="Figures 81–89" captionStartId="F16" captionText="Figures 81 - 89. Quasicalathus conservans sp. nov., volume rendering of the holotype. 81. Head, dorsal aspect (the arrows point to the insertions of the supraorbital setae); 82. Head, ventral aspect; 83. Left external part of metathorax, ventral view; 84. Submentum (the arrows point to the insertions of the four lateral setae); 85. Posterior part of prosternum and procoxae; 86. Posterior part of metasternum and metacoxae; 87 - 89. Preserved remains of the aedeagus (87. Right lateral aspect; 88. Dorsal aspect; 89. Left lateral aspect). Abbreviations: bb - basal bulb of aedeagal median lobe; ce - compound eye; cxp - metacoxal plate; eph - partly evaginated lobes of endophallus; gu - gula; mem - metepimeron; mes - metepisternum; mtt - mentum tooth; mv - metaventrite; pcx - procoxa; pmr - preserved distal part of right paramere of aedeagal median lobe; psp - prosternal process; sc - scutellum; sps - setae of sensory pit; tl - terminal lamella of aedeagal median lobe." figureDoi="10.3897/dez.69.79931.figures81-89" httpUri="https://binary.pensoft.net/fig/643966" pageId="0" pageNumber="19">89</figureCitation>
|
||
). In all other characters as described for the new genus, above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D7DEB1838750921015830C6807D664DE" pageId="0" pageNumber="19" type="differential diagnosis">
|
||
<paragraph id="F9599E5ACAB9A9D338D8B3A2037B9849" pageId="0" pageNumber="19">Differential diagnosis.</paragraph>
|
||
<paragraph id="5BFB674D8A014B2DB39ECC60BDC3A708" pageId="0" pageNumber="19">
|
||
In external characters,
|
||
<taxonomicName id="E5AA74AFC3813868F133328A0DA47C1A" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="C1676C444716BAB65A3A3C5ACF4EDAE1" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. appears identical to
|
||
<taxonomicName id="43F48AD9CD70F71317A183A3AF0B62B6" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="079030CF3605CCFFC4E4A482C6335BDA" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
, however, it can be distinguished by the male genitalia. The new species differs by the proportionally larger aedeagus (EL/AedL = 3.64 instead of 4.05 in
|
||
<taxonomicName id="97C8DA5384842215E494CBADB43111B3" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="6CBCBC22CC0FBBA70F665396B791113E" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
) and by the shape of the median lobe terminal lamella that is nearly evenly narrowed toward the apex if viewed in dorsal aspect and almost straight if viewed laterally (Figs
|
||
<figureCitation id="BA6655423789264E553CC8B4328E69C1" captionStart="Figures 39–46" captionStartId="F8" captionText="Figures 39 - 46. Quasicalathus elpis (Ortuno and Arillo 2009), volume rendering of specimen " Groehn 7814 " using different grey scales of the Amira software. 39. Dorsal aspect; 40. Lateral aspect. The displaced aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software in Figures 39 and 40; 41. Head (the arrows point to the insertions of the supraorbital setae); 42. Pronotum (the arrows point to the insertions of the lateral setae); 43 - 46. Remains of the aedeagus in right lateral aspect (43.); Left lateral aspect (44.); Left lateral aspect (45.); Dorsal aspect (46.). The distal margins of the styloid apophysis of the right paramere in Fig. 43 and the lobate apophysis of the left paramere in Fig. 45 are highlighted by red dotted lines. Abbreviations: bb - basal bulb of aedeagal median lobe; os - distal ostium; pml - left paramere; pmr - right paramere; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures39-46" httpUri="https://binary.pensoft.net/fig/643958" pageId="0" pageNumber="19">43</figureCitation>
|
||
,
|
||
<figureCitation id="1BFA0672998C172935F37EB6195AB731" captionStart="Figures 39–46" captionStartId="F8" captionText="Figures 39 - 46. Quasicalathus elpis (Ortuno and Arillo 2009), volume rendering of specimen " Groehn 7814 " using different grey scales of the Amira software. 39. Dorsal aspect; 40. Lateral aspect. The displaced aedeagus (highlighted by red colour) was separated by the segmentation function of Amira software in Figures 39 and 40; 41. Head (the arrows point to the insertions of the supraorbital setae); 42. Pronotum (the arrows point to the insertions of the lateral setae); 43 - 46. Remains of the aedeagus in right lateral aspect (43.); Left lateral aspect (44.); Left lateral aspect (45.); Dorsal aspect (46.). The distal margins of the styloid apophysis of the right paramere in Fig. 43 and the lobate apophysis of the left paramere in Fig. 45 are highlighted by red dotted lines. Abbreviations: bb - basal bulb of aedeagal median lobe; os - distal ostium; pml - left paramere; pmr - right paramere; tl - terminal lamella of median lobe." figureDoi="10.3897/dez.69.79931.figures39-46" httpUri="https://binary.pensoft.net/fig/643958" pageId="0" pageNumber="19">44</figureCitation>
|
||
versus 87, 89).
|
||
<taxonomicName id="FF0919F053D55A258C969B6621D65EF0" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="FDB69386AED35116BC6EE901D46905DA" italics="true" pageId="0" pageNumber="19">Quasicalathus conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from the above-described
|
||
<taxonomicName id="ADC5D4598CBA49F54FEB49AD81CC9DA4" class="Insecta" kingdom="Animalia" lsidName="Q. agonicollis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="agonicollis">
|
||
<emphasis id="191C39B538128F0DBF5E5790A92D2E3E" italics="true" pageId="0" pageNumber="19">Q. agonicollis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by larger body (SBL> 7 mm), less obtuse laterobasal pronotal angles, the more markedly concave elytral basal margin, more markedly projected humeri, the less obtuse humeral angle (<120°), and by the larger aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7210089EDCB0F35B626FC7AA18693D08" pageId="0" pageNumber="19" type="remarks">
|
||
<paragraph id="FE45D18D454602D96884AEF8B8722ADA" pageId="0" pageNumber="19">Remarks.</paragraph>
|
||
<paragraph id="4943FF3335A3DAF51BC9B1C6E825C872" pageId="0" pageNumber="19">
|
||
<taxonomicName id="7ECE80229143438AEF33C3ACD416F358" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="B5091D017762EB3C7E5D3CA59F628A0F" italics="true" pageId="0" pageNumber="19">Quasicalathus conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="31180967B31B0F2566641BEEAE20359A" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="6AA4C31983675E878E586BB5D27C7E2E" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
are, based on current knowledge, indistinguishable in their external characters but differ clearly in the shape and proportions of the male genitalia.
|
||
<bibRefCitation id="BB6F619B9F66D299DD799F2C5FAA3346" author="Ball, GE" journalOrPublisher="Transactions of the American Entomological Society" pageId="0" pageNumber="19" pagination="412 - 533" refId="B4" refString="Ball, GE, Negre, J, 1972. The taxonomy of the Nearctic species of the genus Calathus Bonelli (Coleoptera: Carabidae: Agonini). Transactions of the American Entomological Society 98: 412 - 533" title="The taxonomy of the Nearctic species of the genus Calathus Bonelli (Coleoptera: Carabidae: Agonini)." volume="98" year="1972">
|
||
Ball and
|
||
<normalizedToken id="991CAEC0981431886527DA1FFBC87124" originalValue="Nègre">Negre</normalizedToken>
|
||
(1972)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="2AA9B75EFF070C4B4973E5F3F94D57C2" author="Schmidt, J" editor="Hartmann, M" journalOrPublisher="Vol. VI. Verein der Freunde und Foerderer des Naturkundemuseums Erfurt e. V., Erfurt" pageId="0" pageNumber="19" pagination="297 - 318" refId="B30" refString="Schmidt, J, 2018. Notes on the taxonomy of Calathus Bonelli, 1810, with special reference to the C. heinertzi group from the Nepal Himalaya (Insecta: Coleoptera: Carabidae: Sphodrini). In: Hartmann, M, Barclay, MVL, Weipert, J, Eds., Biodiversity and Natural Heritage of the Himalaya. Vol. VI. Verein der Freunde und Foerderer des Naturkundemuseums Erfurt e.V., Erfurt: 297 - 318" title="Notes on the taxonomy of Calathus Bonelli, 1810, with special reference to the C. heinertzi group from the Nepal Himalaya (Insecta: Coleoptera: Carabidae: Sphodrini)." volumeTitle="Biodiversity and Natural Heritage of the Himalaya." year="2018">Schmidt (2018)</bibRefCitation>
|
||
detailed a similar situation found among many species pairs of Mexican and Himalayan
|
||
<taxonomicName id="27DFCEB6D0F17291DE2D4515CC536930" class="Insecta" family="Carabidae" genus="Calathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calathus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="F4B4C786E9DB672C57C7DF3F4A1A2916" italics="true" pageId="0" pageNumber="19">Calathus</emphasis>
|
||
</taxonomicName>
|
||
that can only be separated by features of the male genitalia. We hypothesize that
|
||
<taxonomicName id="4117A19085EFC0D56F5759E42ECA7C33" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="12A7144F1110C951951D588E03170E7F" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="02154409D90C010EA3044D21C7F6DABF" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="8C127C65ED4033556CF9050F2694857B" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. were most likely allopatric species, distributed in geographically separated parts of the Eocene amber forests of northern Europe. Under this hypothesis,
|
||
<taxonomicName id="9E08DE83677247ACAF958F63FA89442A" class="Insecta" kingdom="Animalia" lsidName="Q. elpis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="elpis">
|
||
<emphasis id="4D3A8534116571DDF810ED6FF8571A96" italics="true" pageId="0" pageNumber="19">Q. elpis</emphasis>
|
||
</taxonomicName>
|
||
was endemic to the more western part of the Eocene forest and was thus fossilized in Baltic amber, while
|
||
<taxonomicName id="FD50F4B3B8FFEFF268E0293921EC232A" class="Insecta" kingdom="Animalia" lsidName="Q. conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="D82FF2468FCE976DEE69F7720B4E3E19" italics="true" pageId="0" pageNumber="19">Q. conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. was endemic to the more eastern part of the area and therefore fossilized in Rovno amber. This assumption is consistent with the faunistic data showing that beetle fossils of Baltic and Rovno amber deposits have only few species in common (less than 13%;
|
||
<bibRefCitation id="5434B6A3B311A727579D3CDACEABF7F6" DOI="https://doi.org/10.11646/zootaxa.5016.2.5" author="Matalin, AV" journalOrPublisher="Zootaxa" pageId="0" pageNumber="19" pagination="243 - 256" refId="B22" refString="Matalin, AV, Perkovsky, EE, Vasilenko, DV, 2021. First record of tiger beetles (Coleoptera: Cicindelidae) from Rovno amber, with the description of a new genus and species. Zootaxa 5016 (2): 243 - 256, DOI: https://doi.org/10.11646/zootaxa.5016.2.5" title="First record of tiger beetles (Coleoptera: Cicindelidae) from Rovno amber, with the description of a new genus and species." url="https://doi.org/10.11646/zootaxa.5016.2.5" volume="5016" year="2021">Matalin et al. 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C511C09DC83B27B54F304EBB0B8771CD" doi="10.3897/dez.69.79931.figures81-89" httpUri="https://binary.pensoft.net/fig/643966" pageId="0" pageNumber="19" start="Figures 81–89" startId="F16">
|
||
<paragraph id="60034732F32FEF6DF898C0BF08288032" pageId="0" pageNumber="19">
|
||
<emphasis id="7A385809CE2BEAB6D6ABA0932477704C" bold="true" pageId="0" pageNumber="19">Figures 81-89.</emphasis>
|
||
<taxonomicName id="F4A0E9529C1EA9CBEE210208970AF8EE" authorityName="Schmidt & Will" authorityYear="2022" class="Insecta" family="Carabidae" genus="Quasicalathus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Quasicalathus conservans" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="conservans">
|
||
<emphasis id="082E835DAACD6F5DFA6DB88B04C09203" italics="true" pageId="0" pageNumber="19">Quasicalathus conservans</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., volume rendering of the holotype.
|
||
<emphasis id="F61B6319C169B930FD4CD0BA52E3E72C" bold="true" pageId="0" pageNumber="19">81.</emphasis>
|
||
Head, dorsal aspect (the arrows point to the insertions of the supraorbital setae);
|
||
<emphasis id="AFC303DE068CB9B79F648486C243FCBA" bold="true" pageId="0" pageNumber="19">82.</emphasis>
|
||
Head, ventral aspect;
|
||
<emphasis id="EC16E3AEC3913B588F14DB73E7555F0C" bold="true" pageId="0" pageNumber="19">83.</emphasis>
|
||
Left external part of metathorax, ventral view;
|
||
<emphasis id="5C30DB732B6634E44CF452A2C7E5BAAC" bold="true" pageId="0" pageNumber="19">84.</emphasis>
|
||
Submentum (the arrows point to the insertions of the four lateral setae);
|
||
<emphasis id="177EE2979AD84D2B37FA3E126A52D7B8" bold="true" pageId="0" pageNumber="19">85.</emphasis>
|
||
Posterior part of prosternum and procoxae;
|
||
<emphasis id="E4D1C4EAE6DA2EB45A9DAD792A89CE1F" bold="true" pageId="0" pageNumber="19">86.</emphasis>
|
||
Posterior part of metasternum and metacoxae;
|
||
<emphasis id="42285EA868226578E8B360DE724C5750" bold="true" pageId="0" pageNumber="19">87-89.</emphasis>
|
||
Preserved remains of the aedeagus (
|
||
<emphasis id="9D3A4EB2F4829ED1CD0873A2870C3D11" bold="true" pageId="0" pageNumber="19">87.</emphasis>
|
||
Right lateral aspect;
|
||
<emphasis id="BAD94640FC5E4CDAB60DCFE04A106279" bold="true" pageId="0" pageNumber="19">88.</emphasis>
|
||
Dorsal aspect;
|
||
<emphasis id="1FB0FC9D9112BFF3E92F0F53F9BEC94B" bold="true" pageId="0" pageNumber="19">89.</emphasis>
|
||
Left lateral aspect). Abbreviations:
|
||
<emphasis id="690754A92F49C230C4A92A1D3D2D4329" bold="true" pageId="0" pageNumber="19">bb</emphasis>
|
||
- basal bulb of aedeagal median lobe;
|
||
<emphasis id="E885713EC683A1DD3FDD373405613356" bold="true" pageId="0" pageNumber="19">ce</emphasis>
|
||
- compound eye;
|
||
<emphasis id="82154CAE9D4B6163EA8B8D3F19CDE30C" bold="true" pageId="0" pageNumber="19">cxp</emphasis>
|
||
- metacoxal plate;
|
||
<emphasis id="BF76E6F75FE77431E378C625780B6F8D" bold="true" pageId="0" pageNumber="19">eph</emphasis>
|
||
- partly evaginated lobes of endophallus;
|
||
<emphasis id="49114567DD790980FF5982B208E1467E" bold="true" pageId="0" pageNumber="19">gu</emphasis>
|
||
- gula;
|
||
<emphasis id="78124DC1B5DE670EE10C4EF38EF07A39" bold="true" pageId="0" pageNumber="19">mem</emphasis>
|
||
- metepimeron;
|
||
<emphasis id="81FDFE3580C23F261476B33D1DC3F512" bold="true" pageId="0" pageNumber="19">mes</emphasis>
|
||
- metepisternum;
|
||
<emphasis id="D21DA2A0C4B43E91559F510ADA09B7CD" bold="true" pageId="0" pageNumber="19">mtt</emphasis>
|
||
- mentum tooth;
|
||
<emphasis id="81986EF66B37484B0DCCE6063E2DDCB7" bold="true" pageId="0" pageNumber="19">mv</emphasis>
|
||
- metaventrite;
|
||
<emphasis id="830055D13D17F619B8C13314A0A50E2B" bold="true" pageId="0" pageNumber="19">pcx</emphasis>
|
||
- procoxa;
|
||
<emphasis id="02AA096D587CFE505464100D439D7C69" bold="true" pageId="0" pageNumber="19">pmr</emphasis>
|
||
-preserved distal part of right paramere of aedeagal median lobe;
|
||
<emphasis id="B133D4AC808210713569688EC7DD2407" bold="true" pageId="0" pageNumber="19">psp</emphasis>
|
||
- prosternal process;
|
||
<emphasis id="29D960D79C5C93709BC4791B77176377" bold="true" pageId="0" pageNumber="19">sc</emphasis>
|
||
- scutellum;
|
||
<emphasis id="6AB88C37BA3C71BE2A0D67557BEA7351" bold="true" pageId="0" pageNumber="19">sps</emphasis>
|
||
- setae of sensory pit;
|
||
<emphasis id="60FA2CC07084C879F30A3B28C8535264" bold="true" pageId="0" pageNumber="19">tl</emphasis>
|
||
- terminal lamella of aedeagal median lobe.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |