173 lines
20 KiB
XML
173 lines
20 KiB
XML
<document id="9CFB4DECCD231BF0C45D590D13132A5B" ID-DOI="10.11646/zootaxa.5523.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="13933848" ID-ZooBank="D8F096C8-5701-4FE8-B17A-77CA0126BB60" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1728985594790" checkinUser="plazi" docAuthor="Sciaky, Riccardo" docDate="2024" docId="9D4DF639FFF8F976A5BBDD46A13FAF16" docLanguage="en" docName="zootaxa.5523.1.1.pdf" docOrigin="Zootaxa 5523 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5523.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stomis davidi Sciaky, 2024, sp. nov." docType="treatment" docUuid="BAB92514-84C9-43AB-847D-9F2FD7E0CA2A" docUuidSource="ZooBank" docVersion="2" lastPageNumber="14" masterDocId="61748E41FFF5F97BA52CDE5CA544A816" masterDocTitle="Review of the Chinese Stomis, with description of nine new taxa and a new key (Carabidae: Pterostichini)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="14" updateTime="1728986179687" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="87AB95A1BD210802010301C5FCEF82E9">Review of the Chinese Stomis, with description of nine new taxa and a new key (Carabidae: Pterostichini)</mods:title>
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<mods:namePart id="0AA8003546B767200DB339F2F50423D0">Sciaky, Riccardo</mods:namePart>
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<mods:title id="39C37A8A1E1D4830DDD5E4A285A4E4F4">Zootaxa</mods:title>
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<mods:part id="E0DF7E57D163E324568B8EF6363474C0">
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<mods:date id="6A9C34DB875CE0650B682A1A82403827">2024</mods:date>
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<mods:number id="19DAEB18099F4720F0DDD8CA7ABD08E8">2024-10-14</mods:number>
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<mods:identifier id="57DAEEF828F800450C53A24BBDBCD6C0" type="DOI">10.11646/zootaxa.5523.1.1</mods:identifier>
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<treatment id="9D4DF639FFF8F976A5BBDD46A13FAF16" ID-DOI="http://doi.org/10.5281/zenodo.13933920" ID-Zenodo-Dep="13933920" LSID="urn:lsid:zoobank.org:act:BAB92514-84C9-43AB-847D-9F2FD7E0CA2A" httpUri="http://treatment.plazi.org/id/9D4DF639FFF8F976A5BBDD46A13FAF16" lastPageNumber="14" pageId="13" pageNumber="14">
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<subSubSection id="5DFE14A4FFF8F976A5BBDD46A60CAB4F" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="155B472FFFF8F976A5BBDD46A4DAAB22" blockId="13.[151,840,793,857]" box="[151,414,793,820]" pageId="13" pageNumber="14">
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<heading id="4E13F043FFF8F976A5BBDD46A4DAAB22" bold="true" box="[151,414,793,820]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="27909B3DFFF8F976A5BBDD46A4DAAB22" bold="true" box="[151,414,793,820]" pageId="13" pageNumber="14">
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<taxonomicName id="D2E43CACFFF8F976A5BBDD46A47DAB22" authorityName="Sciaky" authorityYear="2024" box="[151,313,793,820]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="davidi" status="sp. nov.">
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<emphasis id="27909B3DFFF8F976A5BBDD46A47DAB22" bold="true" box="[151,313,793,820]" italics="true" pageId="13" pageNumber="14">Stomis davidi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3CA32646FFF8F976A46DDD46A4DAAB22" box="[321,414,794,820]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="155B472FFFF8F976A5BBDD62A60CAB4F" blockId="13.[151,840,793,857]" box="[151,840,830,857]" pageId="13" pageNumber="14">https://zoobank.org/bab92514-84c9-43ab-847d-9f2fd7e0ca2a</paragraph>
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</subSubSection>
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<subSubSection id="5DFE14A4FFF8F976A5BBDDD9A44BAC42" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="155B472FFFF8F976A5BBDDD9A70FABFE" blockId="13.[151,1437,901,1792]" pageId="13" pageNumber="14">
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<emphasis id="27909B3DFFF8F976A5BBDDD9A44EAB89" bold="true" box="[151,266,901,927]" pageId="13" pageNumber="14">Diagnosis</emphasis>
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. A
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<taxonomicName id="D2E43CACFFF8F976A418DDDAA43BABB6" authorityName="Clairville" authorityYear="1806" box="[308,383,902,928]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="27909B3DFFF8F976A418DDDAA43BABB6" box="[308,383,902,928]" italics="true" pageId="13" pageNumber="14">Stomis</emphasis>
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</taxonomicName>
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of large size (
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<quantity id="D21CEACAFFF8F976A70FDDDAA7C9ABB6" box="[547,653,902,928]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="13" pageNumber="14" unit="mm" value="11.1">11.1 mm</quantity>
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), related to
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<taxonomicName id="D2E43CACFFF8F976A639DDDBA15FABB6" authority="Sciaky, 1998" authorityName="Sciaky" authorityYear="1998" box="[789,1051,902,928]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="titanus">
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<emphasis id="27909B3DFFF8F976A639DDDBA63AABB6" box="[789,894,903,928]" italics="true" pageId="13" pageNumber="14">P. titanus</emphasis>
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<bibRefCitation id="71753ADEFFF8F976A6AADDDAA15FABB6" author="Sciaky, R." box="[902,1051,902,928]" pageId="13" pageNumber="14" pagination="21 - 59" refId="ref22785" refString="Sciaky, R. (1998) Taxonomic review of the genus Stomis, with revision of the Chinese species (Coleoptera, Carabidae). Memorie della Societa entomologica italiana, 76, 21 - 59." type="journal article" year="1998">Sciaky, 1998</bibRefCitation>
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</taxonomicName>
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, but different in the narrower and more convex body, stronger greenish hue, smaller pronotum with more abrupt constriction before base, striae more superficial and less evidently punctate.
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</paragraph>
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<paragraph id="155B472FFFF8F976A5EBDDADA7ACAC1A" blockId="13.[151,1437,901,1792]" box="[199,744,1009,1036]" pageId="13" pageNumber="14">
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<emphasis id="27909B3DFFF8F976A5EBDDADA426AC1D" bold="true" box="[199,354,1009,1035]" pageId="13" pageNumber="14">Type-locality</emphasis>
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:
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<collectingCountry id="6DF307BFFFF8F976A45DDDAEA4F3AC1A" box="[369,439,1010,1036]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
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,
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<collectingRegion id="D72089CDFFF8F976A4EEDDAEA75BAC1A" box="[450,543,1010,1036]" country="China" name="Sichuan" pageId="13" pageNumber="14">Sichuan</collectingRegion>
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, Yading Nat. Res.
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</paragraph>
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<paragraph id="155B472FFFF8F976A5EBDA49A44BAC42" blockId="13.[151,1437,901,1792]" pageId="13" pageNumber="14">
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<emphasis id="27909B3DFFF8F976A5EBDA49A40AAC39" bold="true" box="[199,334,1045,1071]" pageId="13" pageNumber="14">Type-series</emphasis>
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.
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<materialsCitation id="A58C4D72FFF8F976A475DA4AA44BAC42" collectingDate="2006-06-24" collectingDateMax="2006-06-28" collectingDateMin="2006-06-24" collectionCode="CW" collectorName="Yading Nat. Res., N" country="China" elevation="4000" location="Sichuan" longitude="28.45" pageId="13" pageNumber="14" specimenCount="2" specimenCount-female="2" stateProvince="Sichuan" typeStatus="holotype">
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<typeStatus id="CA5FF98DFFF8F976A475DA4AA485AC26" box="[345,449,1046,1072]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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<specimenCount id="03E28CA6FFF8F976A4EADA4AA499AC26" box="[454,477,1046,1072]" count="1" pageId="13" pageNumber="14" type="female">♀</specimenCount>
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:
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<collectingCountry id="6DF307BFFFF8F976A4CBDA4AA769AC26" box="[487,557,1046,1072]" name="China" pageId="13" pageNumber="14">China</collectingCountry>
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,
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<collectingRegion id="D72089CDFFF8F976A719DA4AA7D6AC26" box="[565,658,1046,1072]" country="China" name="Sichuan" pageId="13" pageNumber="14">Sichuan</collectingRegion>
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,
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<collectorName id="B81122F9FFF8F976A7B5DA4AA635AC26" box="[665,881,1046,1072]" pageId="13" pageNumber="14">Yading Nat. Res., N</collectorName>
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<geoCoordinate id="70D021E8FFF8F976A659DA4AA69DAC26" box="[885,985,1046,1072]" degrees="28" direction="east" minutes="27" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="28.45">28° 27’ E</geoCoordinate>
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100° 20’,
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<quantity id="D21CEACAFFF8F976A164DA4AA1A6AC26" box="[1096,1250,1046,1072]" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="13" pageNumber="14" unit="m" value="4000.0" valueMax="4200.0" valueMin="3800.0">
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<elevation id="9EC9A01CFFF8F976A164DA4AA1A6AC26" box="[1096,1250,1046,1072]" metricMagnitude="3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="13" pageNumber="14" unit="m" value="4000.0" valueMax="4200.0" valueMin="3800.0">3800–4200 m</elevation>
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</quantity>
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,
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<date id="615A61EFFFF8F976A1C7DA4AA0DDAC26" box="[1259,1433,1046,1072]" pageId="13" pageNumber="14" value="2006-06-24" valueMax="2006-06-28" valueMin="2006-06-24">
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<collectingDate id="711E9807FFF8F976A1C7DA4AA0DDAC26" box="[1259,1433,1046,1072]" pageId="13" pageNumber="14" value="2006-06-24" valueMax="2006-06-28" valueMin="2006-06-24">24–28.VI.2006</collectingDate>
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</date>
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:
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<specimenCount id="03E28CA6FFF8F976A5BBDA66A585AC42" box="[151,193,1082,1108]" count="1" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
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(
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<collectionCode id="73F5DFEAFFF8F976A5FCDA66A447AC42" box="[208,259,1082,1108]" pageId="13" pageNumber="14">CW</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5DFE14A4FFF8F976A5EBDA02A710AC8A" pageId="13" pageNumber="14" type="etymology">
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<paragraph id="155B472FFFF8F976A5EBDA02A710AC8A" blockId="13.[151,1437,901,1792]" pageId="13" pageNumber="14">
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<emphasis id="27909B3DFFF8F976A5EBDA02A40CAC6E" bold="true" box="[199,328,1118,1144]" pageId="13" pageNumber="14">Etymology</emphasis>
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. This species is also dedicated to David Wrase, an excellent specialist of
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<taxonomicName id="D2E43CACFFF8F976A154DA02A1AEAC6E" authorityName="Latreille" authorityYear="1802" box="[1144,1258,1118,1144]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
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<taxonomicName id="D2E43CACFFF8F976A1DCDA02A01EAC6E" authorityName="Bonelli" authorityYear="1810" box="[1264,1370,1118,1144]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="tribe" tribe="Harpalini">Harpalini</taxonomicName>
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and a great collector of interesting
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<taxonomicName id="D2E43CACFFF8F976A4F7DADEA714AC8A" authorityName="Latreille" authorityYear="1802" box="[475,592,1154,1180]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="5DFE14A4FFF8F976A5EBDAF9A13FAF16" pageId="13" pageNumber="14" type="description">
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<paragraph id="155B472FFFF8F976A5EBDAF9A766AE5A" blockId="13.[151,1437,901,1792]" pageId="13" pageNumber="14">
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<emphasis id="27909B3DFFF8F976A5EBDAF9A415ACA9" bold="true" box="[199,337,1189,1215]" pageId="13" pageNumber="14">Description</emphasis>
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. Habitus as in
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<figureCitation id="8DDF5BAAFFF8F976A4D5DAFAA708ACD6" box="[505,588,1190,1216]" captionStart="FIGURES 17–20" captionStartId="22.[151,264,1943,1967]" captionTargetBox="[299,1289,184,1916]" captionTargetId="figure-18@22.[296,1292,181,1919]" captionTargetPageId="22" captionText="FIGURES 17–20. Habitus of: 17, S. titanus HT (11.9 mm); 18, S. davidi HT (11.3 mm); 19, S. robustus HT (11.8 mm); (20) S. schoenmanni from Yunnan, Ailao Shan (11.2 mm)." figureDoi="http://doi.org/10.5281/zenodo.13933858" httpUri="https://zenodo.org/record/13933858/files/figure.png" pageId="13" pageNumber="14">Fig. 18</figureCitation>
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. Length
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<quantity id="D21CEACAFFF8F976A782DAFAA650ACD6" box="[686,788,1190,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="13" pageNumber="14" unit="mm" value="11.1">11.1 mm</quantity>
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. integuments rather opaque for the strong microsculpture, dark reddish brown, with a vague greenish reflection on elytra, body rather convex; legs and antennae reddish brown, rather long and slender; head very narrow, with shallow collar constriction right behind the eyes; mandibles very long, almost twice as long as the rest of the head; the right one with an evident notch almost in the middle; eyes very small and flat; antennae relatively long, reaching the elytral base with the 6th article. Pronotum almost as broad as long, rather convex at sides; lateral margins feebly dilated in the anterior half, constricted in regular curve towards the base, then gently sinuate in the posterior fifth; anterior margin advanced at middle; anterior angles hardly projecting forward, very small; base almost straight; basal foveae rather well-defined, linear, impunctate; basal angles almost right; lateral gutter very narrow all along. Elytra slightly oval, slightly dilated in the posterior half, internal striae visible and finely punctate but rather shallow, then gradually shallower laterally; very finely punctate, intervals almost flat; shoulders very markedly constricted, with an evident humeral tooth. Ratios: El/Pl = 2.70; Ew/Pw = 1.56; El/Ew = 1.62.
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</paragraph>
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<paragraph id="155B472FFFF8F976A5EBD809A13FAF16" blockId="13.[151,1437,901,1792]" pageId="13" pageNumber="14">
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<emphasis id="27909B3DFFF8F976A5EBD809A471AE79" bold="true" box="[199,309,1621,1647]" pageId="13" pageNumber="14">Affinities</emphasis>
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. The occurrence of the notch in the right mandible and the large size indicates this species is systematically close to
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<taxonomicName id="D2E43CACFFF8F976A489D826A7F5AE82" authority="Sciaky, 1998" authorityName="Sciaky" authorityYear="1998" box="[421,689,1658,1684]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="titanus">
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<emphasis id="27909B3DFFF8F976A489D826A754AE82" box="[421,528,1658,1684]" italics="true" pageId="13" pageNumber="14">S. titanus</emphasis>
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||
<bibRefCitation id="71753ADEFFF8F976A736D826A7F5AE82" author="Sciaky, R." box="[538,689,1658,1684]" pageId="13" pageNumber="14" pagination="21 - 59" refId="ref22785" refString="Sciaky, R. (1998) Taxonomic review of the genus Stomis, with revision of the Chinese species (Coleoptera, Carabidae). Memorie della Societa entomologica italiana, 76, 21 - 59." type="journal article" year="1998">Sciaky, 1998</bibRefCitation>
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||
</taxonomicName>
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, but it is clearly distinct from it. It differs from
|
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<taxonomicName id="D2E43CACFFF8F976A1C9D826A014AE82" authorityName="Sciaky" authorityYear="1998" box="[1253,1360,1658,1684]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="titanus">
|
||
<emphasis id="27909B3DFFF8F976A1C9D826A014AE82" box="[1253,1360,1658,1684]" italics="true" pageId="13" pageNumber="14">S. titanus</emphasis>
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</taxonomicName>
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in the narrower and more convex body, stronger greenish hue, paler legs, smaller pronotum with more abrupt constriction before base, striae more superficial and less evidently punctate. Also this species can be inserted in the
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<taxonomicName id="D2E43CACFFF8F976A007D89EA0D7AECA" authorityName="Jedlicka" authorityYear="1932" box="[1323,1427,1730,1756]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="chinensis">
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<emphasis id="27909B3DFFF8F976A007D89EA0D7AECA" box="[1323,1427,1730,1756]" italics="true" pageId="13" pageNumber="14">chinensis</emphasis>
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</taxonomicName>
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- group, containing also the species
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<taxonomicName id="D2E43CACFFF8F976A735D8BAA72CAF16" authorityName="Sciaky" authorityYear="1998" box="[537,616,1766,1792]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="farkaci">
|
||
<emphasis id="27909B3DFFF8F976A735D8BAA72CAF16" box="[537,616,1766,1792]" italics="true" pageId="13" pageNumber="14">farkaci</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D2E43CACFFF8F976A75AD8BAA79CAF16" authorityName="Sciaky" authorityYear="1998" box="[630,728,1766,1792]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="facchinii">
|
||
<emphasis id="27909B3DFFF8F976A75AD8BAA79CAF16" box="[630,728,1766,1792]" italics="true" pageId="13" pageNumber="14">facchinii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D2E43CACFFF8F976A7CAD8BAA60AAF16" authorityName="Jedlicka" authorityYear="1932" box="[742,846,1766,1792]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="chinensis">
|
||
<emphasis id="27909B3DFFF8F976A7CAD8BAA60AAF16" box="[742,846,1766,1792]" italics="true" pageId="13" pageNumber="14">chinensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D2E43CACFFF8F976A670D8BBA6E3AF16" authorityName="Sciaky" authorityYear="2024" box="[860,935,1767,1792]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis id="27909B3DFFF8F976A670D8BBA6E3AF16" box="[860,935,1767,1792]" italics="true" pageId="13" pageNumber="14">wrasei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D2E43CACFFF8F976A699D8BBA145AF16" authorityName="Sciaky" authorityYear="1998" box="[949,1025,1767,1792]" class="Insecta" family="Carabidae" genus="Stomis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="titanus">
|
||
<emphasis id="27909B3DFFF8F976A699D8BBA145AF16" box="[949,1025,1767,1792]" italics="true" pageId="13" pageNumber="14">titanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D2E43CACFFF8F976A114D8BBA131AF16" box="[1080,1141,1767,1792]" class="Insecta" family="Carabidae" genus="Stromis" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gigas">
|
||
<emphasis id="27909B3DFFF8F976A114D8BBA131AF16" box="[1080,1141,1767,1792]" italics="true" pageId="13" pageNumber="14">gigas</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |