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<document id="EA4720F6D8590ADDABF170A61A0B50EE" ID-DOI="10.5281/zenodo.5360916" ID-ISSN="2345-7600" ID-Zenodo-Dep="5360916" ID-ZooBank="5AC21789-E1C1-4B48-813D-667D7638893C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630235294695" checkinUser="valdenar" docAuthor="Uy, Christine Jewel C., Malicky, Hans &amp; Bae, Yeon Jae" docDate="2018" docId="862D87A3FFB0FF9BFC3EA921FB0A7A82" docLanguage="en" docName="66rbz664-703.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Amphipsyche petiolata Ulmer 1930" docType="treatment" docVersion="4" lastPageNumber="669" masterDocId="7A14FFDBFFB5FF9EFF84AD57FF997C5E" masterDocTitle="Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia" masterLastPageNumber="703" masterPageNumber="664" pageNumber="669" updateTime="1699194071636" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0BB1F88B3BD59322800F52BA77627604">Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia</mods:title>
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<mods:namePart id="9E8B33D58473C326AE5B28DDE8EEAFB8">Uy, Christine Jewel C.</mods:namePart>
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<mods:namePart id="14FBECF144379AD34D01F76046AD1426">Malicky, Hans</mods:namePart>
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<treatment id="862D87A3FFB0FF9BFC3EA921FB0A7A82" ID-DOI="http://doi.org/10.5281/zenodo.5456983" ID-Zenodo-Dep="5456983" LSID="urn:lsid:plazi:treatment:862D87A3FFB0FF9BFC3EA921FB0A7A82" httpUri="http://treatment.plazi.org/id/862D87A3FFB0FF9BFC3EA921FB0A7A82" lastPageNumber="669" pageId="5" pageNumber="669">
<subSubSection id="469E653EFFB0FF9BFC3EA921FAD278D1" box="[954,1355,1142,1167]" pageId="5" pageNumber="669" type="nomenclature">
<paragraph id="0E3B36B5FFB0FF9BFC3EA921FAD278D1" blockId="5.[954,1355,1142,1199]" box="[954,1355,1142,1167]" pageId="5" pageNumber="669">
<heading id="557381D9FFB0FF9BFC3EA921FAD278D1" bold="true" box="[954,1355,1142,1167]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="669" reason="2">
<taxonomicName id="C9844D36FFB0FF9BFC3EA921FAD278D1" ID-CoL="66FJS" authority="Ulmer, 1930" authorityName="Ulmer" authorityYear="1930" box="[954,1355,1142,1167]" class="Insecta" family="Hydropsychidae" genus="Amphipsyche" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="669" phylum="Arthropoda" rank="species" species="petiolata">
<emphasis id="3CF0EAA7FFB0FF9BFC3EA921FAD278D1" bold="true" box="[954,1355,1142,1167]" pageId="5" pageNumber="669">
<emphasis id="3CF0EAA7FFB0FF9BFC3EA921FB2F78D0" bold="true" box="[954,1206,1142,1166]" italics="true" pageId="5" pageNumber="669">Amphipsyche petiolata</emphasis>
<bibRefCitation id="6A154B44FFB0FF9BFB3AA920FAD278D1" author="Ulmer G" box="[1214,1355,1143,1167]" pageId="5" pageNumber="669" pagination="373 - 498" refId="ref27626" refString="Ulmer G (1930) Trichopteren von den Philippinen und von den Sunda-Inseln. Treubia, 11: 373 - 498." type="journal article" year="1930">Ulmer, 1930</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFB0FF9BFBBEA9C0FB5278F1" box="[1082,1227,1175,1199]" pageId="5" pageNumber="669" type="description">
<paragraph id="0E3B36B5FFB0FF9BFBBEA9C0FB5278F1" blockId="5.[954,1355,1142,1199]" box="[1082,1227,1175,1199]" pageId="5" pageNumber="669">
(
<figureCitation id="96BF2A30FFB0FF9BFBC6A9C0FB5D78F1" box="[1090,1220,1175,1199]" captionStart="Figs" captionStartId="4.[113,157,1448,1470]" captionTargetBox="[114,1452,188,1433]" captionTargetId="figure-9@4.[99,1467,176,1446]" captionTargetPageId="4" captionText="Figs. 1736. Amphipsyche gratiosa. 17, right forewing; Male genitalia: 18, lateral; 19, segment X dorsal; 20, phallus lateral; 21, phallus tip. Amphipsyche magna. Male genitalia: 22, lateral; 24, phallus lateral; 25, phallus tip; 23, thorax. Amphipsyche meridiana. 26, right forewing; Male genitalia: 27, lateral; 28, segment X dorsal; 29, phallus lateral; 30, phallus tip. Amphipsyche parva. Male genitalia: 31, lateral; 32, phallus lateral; 33, phallus tip. Amphipsyche petiolata. 34, lateral; 35, phallus lateral; 36, phallus tip. Scale: 17, 26 = 2 mm, 1825, 2736 = 0.25 mm (2225, 2736 redrawn from Barnard, 1984)." figureDoi="http://doi.org/10.5281/zenodo.5360922" httpUri="https://zenodo.org/record/5360922/files/figure.png" pageId="5" pageNumber="669">Figs. 3436</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFB0FF9BFCC7A981FA177925" pageId="5" pageNumber="669" type="reference_group">
<paragraph id="0E3B36B5FFB0FF9BFCC7A981FA907956" blockId="5.[835,1474,1238,1404]" pageId="5" pageNumber="669">
<treatmentCitationGroup id="2E94119BFFB0FF9BFCC7A981FA907956" pageId="5" pageNumber="669">
<treatmentCitation id="8F2510A4FFB0FF9BFCC7A981FB5678B5" author="Ulmer G" box="[835,1231,1238,1260]" page="434" pageId="5" pageNumber="669" year="1930">
<taxonomicName id="C9844D36FFB0FF9BFCC7A981FB5678B5" ID-CoL="66FJS" authority="Ulmer, 1930: 434" authorityName="Ulmer" authorityPageNumber="434" authorityYear="1930" box="[835,1231,1238,1260]" class="Insecta" family="Hydropsychidae" genus="Amphipsyche" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="669" phylum="Arthropoda" rank="species" species="petiolata">
<emphasis id="3CF0EAA7FFB0FF9BFCC7A981FB8778B5" box="[835,1054,1238,1259]" italics="true" pageId="5" pageNumber="669">Amphipsyche petiolata</emphasis>
<bibRefCitation id="6A154B44FFB0FF9BFBA1A981FB5678B5" author="Ulmer G" box="[1061,1231,1238,1260]" pageId="5" pageNumber="669" pagination="373 - 498" refId="ref27626" refString="Ulmer G (1930) Trichopteren von den Philippinen und von den Sunda-Inseln. Treubia, 11: 373 - 498." type="journal article" year="1930">Ulmer, 1930: 434</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="D13F8817FFB0FF9BFB5EA981FAA478B5" box="[1242,1341,1238,1259]" pageId="5" pageNumber="669" type="lectotype">Lectotype</typeStatus>
female;
<collectingRegion id="CC40F857FFB0FF9BFA15A981FA2678B5" box="[1425,1471,1238,1259]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="669">Java</collectingRegion>
; ZMUH; designated by
<treatmentCitation id="8F2510A4FFB0FF9BFBC8A9A5FB677956" author="Ulmer G" box="[1100,1278,1266,1288]" page="197" pageId="5" pageNumber="669" year="1951">
<bibRefCitation id="6A154B44FFB0FF9BFBC8A9A5FB677956" author="Ulmer G" box="[1100,1278,1266,1288]" pageId="5" pageNumber="669" pagination="1 - 528" refId="ref27648" refString="Ulmer G (1951) Kocherfliegen (Trichopteren) von den Sunda-Inseln (Teil I). Archiv fur Hydrobiologie. Supplement, 19: 1 - 528." type="journal article" year="1951">Ulmer, 1951: 197</bibRefCitation>
</treatmentCitation>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="0E3B36B5FFB0FF9BFCC7A858FA15791F" blockId="5.[835,1474,1238,1404]" pageId="5" pageNumber="669">
<treatmentCitationGroup id="2E94119BFFB0FF9BFCC7A858FA15791F" pageId="5" pageNumber="669">
<taxonomicName id="C9844D36FFB0FF9BFCC7A858FB51797B" authority="Banks, 1931: 395" authorityName="Banks" authorityPageNumber="395" authorityYear="1931" box="[835,1224,1295,1317]" class="Insecta" family="Hydropsychidae" genus="Amphipsyche" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="669" phylum="Arthropoda" rank="species" species="minima">
<emphasis id="3CF0EAA7FFB0FF9BFCC7A858FB8A797B" box="[835,1043,1295,1317]" italics="true" pageId="5" pageNumber="669">Amphipsyche minima</emphasis>
<materialsCitation id="BEEC3CE8FFB0FF9BFB98A858FB51797B" box="[1052,1224,1295,1317]" collectionCode="BMNH" country="Malaysia" location="West" pageId="5" pageNumber="669" specimenCount="1" typeStatus="lectotype">Banks, 1931: 395</materialsCitation>
</taxonomicName>
[
<typeStatus id="D13F8817FFB0FF9BFB52A847FAA0797B" box="[1238,1337,1296,1317]" pageId="5" pageNumber="669" type="lectotype">Lectotype</typeStatus>
female;
<location id="0B5B606EFFB0FF9BFA15A847FA5B797B" LSID="urn:lsid:plazi:treatment:862D87A3FFB0FF9BFC3EA921FB0A7A82:0B5B606EFFB0FF9BFA15A847FA5B797B" box="[1425,1474,1296,1317]" country="Malaysia" name="West" pageId="5" pageNumber="669">West</location>
<collectingCountry id="76937625FFB0FF9BFCEDA87BFC5A791C" box="[873,963,1324,1346]" name="Malaysia" pageId="5" pageNumber="669">Malaysia</collectingCountry>
, BMNH; synonymised by
<treatmentCitation id="8F2510A4FFB0FF9BFB48A87BFA18791F" author="Barnard PC" box="[1228,1409,1323,1346]" page="99" pageId="5" pageNumber="669" year="1984">
<bibRefCitation id="6A154B44FFB0FF9BFB48A87BFA18791F" author="Barnard PC" box="[1228,1409,1323,1346]" pageId="5" pageNumber="669" pagination="71 - 130" refId="ref25573" refString="Barnard PC (1984) Macronematinae caddisflies of the genus Amphipsyche (Trichoptera: Hydropsychidae). Bulletin of the British Museum (Natural History), Entomology, 48 (2): 71 - 130." type="journal article" year="1984">Barnard, 1984: 99</bibRefCitation>
</treatmentCitation>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="0E3B36B5FFB0FF9BFCC7A81EFA177925" blockId="5.[835,1474,1238,1404]" pageId="5" pageNumber="669">
<treatmentCitationGroup id="2E94119BFFB0FF9BFCC7A81EFA177925" pageId="5" pageNumber="669">
<treatmentCitation id="8F2510A4FFB0FF9BFCC7A81EFA8A7900" author="Kimmins DE" box="[835,1299,1353,1375]" page="387" pageId="5" pageNumber="669" year="1955">
<taxonomicName id="C9844D36FFB0FF9BFCC7A81EFA8A7900" authority="Kimmins, 1955: 387" authorityName="Kimmins" authorityPageNumber="387" authorityYear="1955" box="[835,1299,1353,1375]" class="Insecta" family="Hydropsychidae" genus="Amphipsyche" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="669" phylum="Arthropoda" rank="species" species="pubescens">
<emphasis id="3CF0EAA7FFB0FF9BFCC7A81EFBAE7900" box="[835,1079,1353,1374]" italics="true" pageId="5" pageNumber="669">Amphipsyche pubescens</emphasis>
<bibRefCitation id="6A154B44FFB0FF9BFBC5A81EFA8A7900" author="Kimmins DE" box="[1089,1299,1353,1375]" pageId="5" pageNumber="669" pagination="374 - 442" refId="ref26171" refString="Kimmins DE (1955) Results of the Oxford University expedition to Sarawak, 1932. Order Trichoptera. The Sarawak Museum Journal, 6: 374 - 442." type="journal article" year="1955">Kimmins, 1955: 387</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="D13F8817FFB0FF9BFAA6A81EFA187901" box="[1314,1409,1353,1375]" pageId="5" pageNumber="669" type="holotype">Holotype</typeStatus>
male; Borneo; NHMUK; synonymised by
<treatmentCitation id="8F2510A4FFB0FF9BFB4AA831FA1A7925" author="Barnard PC" box="[1230,1411,1381,1404]" page="99" pageId="5" pageNumber="669" year="1984">
<bibRefCitation id="6A154B44FFB0FF9BFB4AA831FA1A7925" author="Barnard PC" box="[1230,1411,1381,1404]" pageId="5" pageNumber="669" pagination="71 - 130" refId="ref25573" refString="Barnard PC (1984) Macronematinae caddisflies of the genus Amphipsyche (Trichoptera: Hydropsychidae). Bulletin of the British Museum (Natural History), Entomology, 48 (2): 71 - 130." type="journal article" year="1984">Barnard, 1984: 99</bibRefCitation>
</treatmentCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFB0FF9BFCC7A8F4FAFB79A2" pageId="5" pageNumber="669" type="diagnosis">
<paragraph id="0E3B36B5FFB0FF9BFCC7A8F4FAFB79A2" blockId="5.[835,1474,1443,1532]" pageId="5" pageNumber="669">
<emphasis id="3CF0EAA7FFB0FF9BFCC7A8F4FC2E79E5" bold="true" box="[835,951,1443,1467]" pageId="5" pageNumber="669">Diagnosis.</emphasis>
<emphasis id="3CF0EAA7FFB0FF9BFC38A8F3FBC579E2" box="[956,1116,1444,1468]" italics="true" pageId="5" pageNumber="669">Male genitalia.</emphasis>
Inferior appendages unsegmented (
<figureCitation id="96BF2A30FFB0FF9BFCCFA893FC027982" box="[843,923,1476,1500]" captionStart="Figs" captionStartId="4.[113,157,1448,1470]" captionTargetBox="[114,1452,188,1433]" captionTargetId="figure-9@4.[99,1467,176,1446]" captionTargetPageId="4" captionText="Figs. 1736. Amphipsyche gratiosa. 17, right forewing; Male genitalia: 18, lateral; 19, segment X dorsal; 20, phallus lateral; 21, phallus tip. Amphipsyche magna. Male genitalia: 22, lateral; 24, phallus lateral; 25, phallus tip; 23, thorax. Amphipsyche meridiana. 26, right forewing; Male genitalia: 27, lateral; 28, segment X dorsal; 29, phallus lateral; 30, phallus tip. Amphipsyche parva. Male genitalia: 31, lateral; 32, phallus lateral; 33, phallus tip. Amphipsyche petiolata. 34, lateral; 35, phallus lateral; 36, phallus tip. Scale: 17, 26 = 2 mm, 1825, 2736 = 0.25 mm (2225, 2736 redrawn from Barnard, 1984)." figureDoi="http://doi.org/10.5281/zenodo.5360922" httpUri="https://zenodo.org/record/5360922/files/figure.png" pageId="5" pageNumber="669">Fig. 34</figureCitation>
). Phallus apex, globose with bifurcate membranous process directing towards apex laterally (
<figureCitation id="96BF2A30FFB0FF9BFA87A8B3FACC79A2" box="[1283,1365,1508,1532]" captionStart="Figs" captionStartId="4.[113,157,1448,1470]" captionTargetBox="[114,1452,188,1433]" captionTargetId="figure-9@4.[99,1467,176,1446]" captionTargetPageId="4" captionText="Figs. 1736. Amphipsyche gratiosa. 17, right forewing; Male genitalia: 18, lateral; 19, segment X dorsal; 20, phallus lateral; 21, phallus tip. Amphipsyche magna. Male genitalia: 22, lateral; 24, phallus lateral; 25, phallus tip; 23, thorax. Amphipsyche meridiana. 26, right forewing; Male genitalia: 27, lateral; 28, segment X dorsal; 29, phallus lateral; 30, phallus tip. Amphipsyche parva. Male genitalia: 31, lateral; 32, phallus lateral; 33, phallus tip. Amphipsyche petiolata. 34, lateral; 35, phallus lateral; 36, phallus tip. Scale: 17, 26 = 2 mm, 1825, 2736 = 0.25 mm (2225, 2736 redrawn from Barnard, 1984)." figureDoi="http://doi.org/10.5281/zenodo.5360922" httpUri="https://zenodo.org/record/5360922/files/figure.png" pageId="5" pageNumber="669">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFB0FF9BFCC7AB74FB3C7A62" box="[835,1189,1571,1596]" pageId="5" pageNumber="669" type="distribution">
<paragraph id="0E3B36B5FFB0FF9BFCC7AB74FB3C7A62" blockId="5.[835,1189,1571,1596]" box="[835,1189,1571,1596]" pageId="5" pageNumber="669">
<emphasis id="3CF0EAA7FFB0FF9BFCC7AB74FC4F7A65" bold="true" box="[835,982,1571,1595]" pageId="5" pageNumber="669">Distribution.</emphasis>
Borneo,
<collectingCountry id="76937625FFB0FF9BFBB8AB73FB387A62" box="[1084,1185,1572,1596]" name="Malaysia" pageId="5" pageNumber="669">Malaysia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="469E653EFFB0FF9BFCC7AB33FB0A7A82" pageId="5" pageNumber="669" type="discussion">
<paragraph id="0E3B36B5FFB0FF9BFCC7AB33FB0A7A82" blockId="5.[835,1474,1636,1756]" pageId="5" pageNumber="669">
<emphasis id="3CF0EAA7FFB0FF9BFCC7AB33FC2B7A22" bold="true" box="[835,946,1636,1660]" pageId="5" pageNumber="669">Remarks.</emphasis>
<bibRefCitation id="6A154B44FFB0FF9BFC33AB33FBC37A22" author="Barnard PC" box="[951,1114,1636,1660]" pageId="5" pageNumber="669" pagination="71 - 130" refId="ref25573" refString="Barnard PC (1984) Macronematinae caddisflies of the genus Amphipsyche (Trichoptera: Hydropsychidae). Bulletin of the British Museum (Natural History), Entomology, 48 (2): 71 - 130." type="journal article" year="1984">Barnard (1984)</bibRefCitation>
clearly described the species. The male genitalia (
<figureCitation id="96BF2A30FFB0FF9BFC76ABD3FBE17AC2" box="[1010,1144,1668,1692]" captionStart="Figs" captionStartId="4.[113,157,1448,1470]" captionTargetBox="[114,1452,188,1433]" captionTargetId="figure-9@4.[99,1467,176,1446]" captionTargetPageId="4" captionText="Figs. 1736. Amphipsyche gratiosa. 17, right forewing; Male genitalia: 18, lateral; 19, segment X dorsal; 20, phallus lateral; 21, phallus tip. Amphipsyche magna. Male genitalia: 22, lateral; 24, phallus lateral; 25, phallus tip; 23, thorax. Amphipsyche meridiana. 26, right forewing; Male genitalia: 27, lateral; 28, segment X dorsal; 29, phallus lateral; 30, phallus tip. Amphipsyche parva. Male genitalia: 31, lateral; 32, phallus lateral; 33, phallus tip. Amphipsyche petiolata. 34, lateral; 35, phallus lateral; 36, phallus tip. Scale: 17, 26 = 2 mm, 1825, 2736 = 0.25 mm (2225, 2736 redrawn from Barnard, 1984)." figureDoi="http://doi.org/10.5281/zenodo.5360922" httpUri="https://zenodo.org/record/5360922/files/figure.png" pageId="5" pageNumber="669">Figs. 3436</figureCitation>
) were redrawn from
<bibRefCitation id="6A154B44FFB0FF9BFAEFABD3FC137AE2" author="Barnard PC" pageId="5" pageNumber="669" pagination="71 - 130" refId="ref25573" refString="Barnard PC (1984) Macronematinae caddisflies of the genus Amphipsyche (Trichoptera: Hydropsychidae). Bulletin of the British Museum (Natural History), Entomology, 48 (2): 71 - 130." type="journal article" year="1984">Barnard (1984</bibRefCitation>
:
<figureCitation id="96BF2A30FFB0FF9BFC1DABF3FBB17AE2" box="[921,1064,1700,1724]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="9.[134,179,1293,1315]" captionStartId-1="11.[136,180,2006,2028]" captionTargetBox-0="[136,1473,190,1277]" captionTargetBox-1="[157,1457,191,1990]" captionTargetId-0="figure-8@9.[110,1494,162,1376]" captionTargetId-1="figure-86@11.[133,1477,181,1997]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="Figs. 5969. Macrostemum caliptera. 59, right forewing. Macrostemum centrotum. 60, right forewing; Male genitalia: 61, lateral; 62, segment X dorsal; 63, phallus lateral; 64, phallus tip. Macrostemum dairiana. 65, right forewing; Male genitalia: 66, lateral; 67, segment X dorsal; 68, phallus lateral; 69, phallus tip. Scale: 59, 60, 65 = 2 mm; 6164, 6669 = 0.02 mm (59 redrawn from Banks, 1931c)." captionText-1="Figs. 7084. Macrostemum dione. 70, right forewing; Male genitalia: 71, lateral; 72, segment X dorsal; 73, phallus lateral; 74, phallus tip. Macrostemum distinguendum. 75, right fore- and hind wing; Male genitalia: 76, lateral; 77, segment X dorsal; 78, phallus lateral; 79, phallus tip. Macrostemum dohrni. 80, right forewing; Male genitalia: 81, lateral; 82, segment X dorsal; 83, phallus lateral; 84, phallus tip. Scale: 70, 75, 80 = 2 mm; 7174, 7679, 8184 = 0.02 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5360928" figureDoi-1="http://doi.org/10.5281/zenodo.5360930" httpUri-0="https://zenodo.org/record/5360928/files/figure.png" httpUri-1="https://zenodo.org/record/5360930/files/figure.png" pageId="5" pageNumber="669">Figs. 6973</figureCitation>
). Diagnosis was also based on the description by
<bibRefCitation id="6A154B44FFB0FF9BFC62AB93FB167A82" author="Barnard PC" box="[998,1167,1732,1756]" pageId="5" pageNumber="669" pagination="71 - 130" refId="ref25573" refString="Barnard PC (1984) Macronematinae caddisflies of the genus Amphipsyche (Trichoptera: Hydropsychidae). Bulletin of the British Museum (Natural History), Entomology, 48 (2): 71 - 130." type="journal article" year="1984">Barnard (1984)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>