505 lines
109 KiB
XML
505 lines
109 KiB
XML
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<mods:title id="F8CBFCAF28659995DFADA1E4371CFA11">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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<mods:affiliation id="D48DB0A5B920D9F3CD7158C0693EEA19">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:date id="8C519AA0556170DA7AD2B78EE7F8810C">2023</mods:date>
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Genus
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<taxonomicName id="53B84D69EB6DED40FDFDFB30606F82CC" ID-CoL="8KWHT" authority="Latreille, 1829" authorityName="Latreille" authorityYear="1829" box="[536,840,1183,1207]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="A6CCEAF8EB6DED40FDFDFB3061BB82CC" bold="true" box="[536,668,1183,1207]" italics="true" pageId="5" pageNumber="498">Cyclosomus</emphasis>
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<bibRefCitation id="F0294B1BEB6DED40FD46FB0F606F82CC" author="LATREILLE, P. A." box="[675,840,1184,1207]" pageId="5" pageNumber="498" refId="ref23782" refString="LATREILLE, P. A. 1829. Les crustaces, les arachnides et les insectes, distribues en familles naturelles, ouvrage format les tomes 4 et 5 de celui de M. le Baron Cuvier sur le regne animal (deuxieme edition). Tome premier. Deterville, Paris, xxvii + 584 pp." type="book" year="1829">Latreille, 1829</bibRefCitation>
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</emphasis>
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</paragraph>
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<treatmentCitation id="151910FBEB6DED40FF9EFB6A62B682A0" author="FABRICIUS, J. C." box="[123,401,1221,1243]" page="221" pageId="5" pageNumber="498" year="1790">
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<taxonomicName id="53B84D69EB6DED40FF9EFB6A62B682A0" ID-CoL="8HFVS" authority="Fabricius, 1790: 221" authorityName="Fabricius" authorityPageNumber="221" authorityYear="1790" box="[123,401,1221,1243]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="A6CCEAF8EB6DED40FF9EFB6A63EC82A0" box="[123,203,1221,1243]" italics="true" pageId="5" pageNumber="498">Scolytus</emphasis>
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<bibRefCitation id="F0294B1BEB6DED40FF35FB6A62B682A0" author="FABRICIUS, J. C." box="[208,401,1221,1243]" pageId="5" pageNumber="498" pagination="213 - 228" refId="ref23357" refString="FABRICIUS, J. C. 1790. Nova insectorum genera. Skrivter af Naturhistorie-Selskabet 1: 213 - 228." type="journal article" year="1790">Fabricius, 1790:221</bibRefCitation>
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, in part [junior homonym of
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<taxonomicName id="53B84D69EB6DED40FD47FB6A60AC82A0" ID-CoL="8HFVS" authority="Geoffroy, 1762" authorityName="Geoffroy" authorityYear="1762" box="[674,907,1221,1243]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="A6CCEAF8EB6DED40FD47FB6A61D582A0" box="[674,754,1221,1243]" italics="true" pageId="5" pageNumber="498">Scolytus</emphasis>
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<bibRefCitation id="F0294B1BEB6DED40FD1DFB6A60AC82A0" author="GEOFFROY, E. L." box="[760,907,1221,1243]" pageId="5" pageNumber="498" refId="ref23463" refString="GEOFFROY, E. L. 1762. Histoire abregee des insectes qui se trouvent aux environs de Paris; dans laquelle ces animaux sont ranges suivant un ordre methodique. Tome premier. Durand, Paris. xxviii + 523 pp. + 10 pls." type="book" year="1762">Geoffroy, 1762</bibRefCitation>
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</taxonomicName>
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] (
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species
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<emphasis id="A6CCEAF8EB6DED40FBF2FB6A67B182A0" box="[1047,1174,1221,1243]" italics="true" pageId="5" pageNumber="498">
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<taxonomicName id="53B84D69EB6DED40FBF2FB6A674D82A0" ID-CoL="8KVGQ" authorityName="Linnaeus" authorityYear="1758" box="[1047,1130,1221,1243]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="genus">Carabus</taxonomicName>
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<emphasis id="A6CCEAF8EB6DED40FF4EFB4D63F88283" box="[171,223,1250,1272]" italics="true" pageId="5" pageNumber="498">limbatus</emphasis>
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</emphasis>
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<bibRefCitation id="F0294B1BEB6DED40FF00FB4D625A8283" author="FABRICIUS, J. C." box="[229,381,1250,1272]" pageId="5" pageNumber="498" refId="ref23424" refString="FABRICIUS, J. C. 1801. Systema eleutheratorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiliae. xxiv + 506 pp." type="book" year="1801">Fabricius, 1801</bibRefCitation>
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, designated by
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<bibRefCitation id="F0294B1BEB6DED40FDF1FB4D61FE8283" author="LATREILLE, P. A." box="[532,729,1250,1272]" pageId="5" pageNumber="498" refId="ref23732" refString="LATREILLE, P. A. 1810. Considerations generales sur l'ordre naturel des animaux composant les classes des crustaces, des arachnides, et des insectes; avec un tableau methodique de leurs genres, disposes. en familles. F. Schoell, Paris, 444 pp." type="book" year="1810">Latreille (1810:426)</bibRefCitation>
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.
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<bibRefCitation id="F0294B1BEB6DED40FD06FB4D60978283" author="FABRICIUS, J. C." box="[739,944,1250,1272]" pageId="5" pageNumber="498" refId="ref23378" refString="FABRICIUS, J. C. 1792. Entomologia systematica emendate et aucta, secundum classes, ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Tome I. Pars I. Proft, Hafniae, xx + 330 pp." type="book" year="1792">Fabricius (1792:180)</bibRefCitation>
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;
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<bibRefCitation id="F0294B1BEB6DED40FC5EFB4C675A8283" author="BOUSQUET, Y." box="[955,1149,1250,1272]" pageId="5" pageNumber="498" pagination="1 - 1722" refId="ref23148" refString="BOUSQUET, Y. 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, North of Mexico. Zookeys 245: 1 - 1722." type="journal article" year="2012">Bousquet (2012:89)</bibRefCitation>
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</treatmentCitationGroup>
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.
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||
</paragraph>
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||
<paragraph id="940736EAEB6DED40FF9EFB506704836E" blockId="5.[123,1174,1183,1725]" box="[123,1059,1279,1301]" pageId="5" pageNumber="498">
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<treatmentCitationGroup id="B4A811C4EB6DED40FF9EFB506704836E" box="[123,1059,1279,1301]" pageId="5" pageNumber="498">
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<treatmentCitation id="151910FBEB6DED40FF9EFB506289836E" author="LATREILLE, P. A." box="[123,430,1279,1301]" page="394" pageId="5" pageNumber="498" year="1829">
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<taxonomicName id="53B84D69EB6DED40FF9EFB506289836E" ID-CoL="8KWHT" authority="Latreille, 1829: 394" authorityName="Latreille" authorityPageNumber="394" authorityYear="1829" box="[123,430,1279,1301]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="genus">
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<emphasis id="A6CCEAF8EB6DED40FF9EFB5063C8836E" box="[123,239,1279,1301]" italics="true" pageId="5" pageNumber="498">Cyclosomus</emphasis>
|
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<bibRefCitation id="F0294B1BEB6DED40FF10FB506289836E" author="LATREILLE, P. A." box="[245,430,1279,1301]" pageId="5" pageNumber="498" refId="ref23782" refString="LATREILLE, P. A. 1829. Les crustaces, les arachnides et les insectes, distribues en familles naturelles, ouvrage format les tomes 4 et 5 de celui de M. le Baron Cuvier sur le regne animal (deuxieme edition). Tome premier. Deterville, Paris, xxvii + 584 pp." type="book" year="1829">Latreille, 1829:394</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
|
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(
|
||
<typeStatus id="4B038848EB6DED40FE5EFB5062C1836E" box="[443,486,1279,1301]" pageId="5" pageNumber="498">type</typeStatus>
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species
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<taxonomicName id="53B84D69EB6DED40FDDDFB5060AE836E" ID-CoL="QWMS" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[568,905,1279,1301]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="species" species="flexuosus">
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<emphasis id="A6CCEAF8EB6DED40FDDDFB5061CC836E" box="[568,747,1279,1301]" italics="true" pageId="5" pageNumber="498">Carabus flexuosus</emphasis>
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<bibRefCitation id="F0294B1BEB6DED40FD14FB5060AE836E" author="FABRICIUS, J. C." box="[753,905,1279,1301]" pageId="5" pageNumber="498" refId="ref23315" refString="FABRICIUS, J. C. 1775. Systema entomologica, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Libraria Kortii, Flensburgi et Lipsiae, xxxii + 832 pp." type="book" year="1775">Fabricius, 1775</bibRefCitation>
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</taxonomicName>
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||
, by monotypy).
|
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB6DED40FF4EFA8B61D08026" pageId="5" pageNumber="498" type="diagnosis">
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<paragraph id="940736EAEB6DED40FF4EFA8B61D08026" blockId="5.[123,1174,1183,1725]" pageId="5" pageNumber="498">
|
||
<emphasis id="A6CCEAF8EB6DED40FF4EFA8B62388340" bold="true" box="[171,287,1316,1339]" pageId="5" pageNumber="498">Diagnosis.</emphasis>
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Adults of
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<taxonomicName id="53B84D69EB6DED40FE75FA8B61438347" box="[400,612,1316,1340]" pageId="5" pageNumber="498">
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<emphasis id="A6CCEAF8EB6DED40FE75FA8B61368347" box="[400,529,1316,1340]" italics="true" pageId="5" pageNumber="498">Cyclosomus</emphasis>
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species
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</taxonomicName>
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can be distinguished from those of other Asian carabid species by the following combination of character states: Body medium-sized, BL =
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<quantity id="53409B0FEB6DED40FBCCFAEB675D8320" box="[1065,1146,1348,1372]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="498" unit="mm" value="6.5">6.5 mm</quantity>
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to
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<quantity id="53409B0FEB6DED40FF9EFACB63F48307" box="[123,211,1380,1404]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="5" pageNumber="498" unit="mm" value="9.5">9.5 mm</quantity>
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; body form broadly ovoid (e.g.,
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<figureCitation id="0C832A6FEB6DED40FDDAFACB61A68307" box="[575,641,1380,1404]" captionStart="FIGURE" captionStartId="4.[171,183,907,926]" captionTargetBox="[123,1172,236,882]" captionTargetId="figure-357@4.[120,1177,216,892]" captionTargetPageId="4" captionText="FIGURE 3. A. Illustration of measurements recorded. EL = elytral length; EW = elytral width; HL = head length; PL.= pronotal width; PWA = width between apices of apical angles of pronotum; PWM = width across widest part of pronotum. B. Illustration of terms used for describing the dark color pattern in Cyclosomus species (example image is of a widespread African species, Cyclosomus buqueti Kolbe). bdb = basal dark band; ldb = longitudinal dark band; lpb = lateral pale band of pronotum; mdb = middle transverse dark band; pas = preapical dark spot." figureDoi="http://doi.org/10.5281/zenodo.11512960" httpUri="https://zenodo.org/record/11512960/files/figure.png" pageId="5" pageNumber="498">Fig. 3</figureCitation>
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||
),
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||
<taxonomicName id="53B84D69EB6DED40FD7CFACB602B8307" authorityName="Latreille" authorityYear="1802" box="[665,780,1380,1404]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="498" phylum="Arthropoda" rank="genus">
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||
<emphasis id="A6CCEAF8EB6DED40FD7CFACB602B8307" box="[665,780,1380,1404]" italics="true" pageId="5" pageNumber="498">Omophron</emphasis>
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</taxonomicName>
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||
-like; head with two pairs of supraoribital setae; labrum with anterior margin deeply emarginate; mentum with medial tooth present, apically emarginate; pronotum (
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<figureCitation id="0C832A6FEB6DED40FE2AFA0B613C83C7" box="[463,539,1444,1468]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="5" pageNumber="498">Fig. 14</figureCitation>
|
||
) very broad, trapezoidal, almost as wide as elytra at humeral angles, distinctly widest at or near base; elytra broadly rounded apically, elytral basic color pattern as in
|
||
<figureCitation id="0C832A6FEB6DED40FF01FA4B62118387" box="[228,310,1508,1532]" captionStart="FIGURE" captionStartId="4.[171,183,907,926]" captionTargetBox="[123,1172,236,882]" captionTargetId="figure-357@4.[120,1177,216,892]" captionTargetPageId="4" captionText="FIGURE 3. A. Illustration of measurements recorded. EL = elytral length; EW = elytral width; HL = head length; PL.= pronotal width; PWA = width between apices of apical angles of pronotum; PWM = width across widest part of pronotum. B. Illustration of terms used for describing the dark color pattern in Cyclosomus species (example image is of a widespread African species, Cyclosomus buqueti Kolbe). bdb = basal dark band; ldb = longitudinal dark band; lpb = lateral pale band of pronotum; mdb = middle transverse dark band; pas = preapical dark spot." figureDoi="http://doi.org/10.5281/zenodo.11512960" httpUri="https://zenodo.org/record/11512960/files/figure.png" pageId="5" pageNumber="498">Fig. 3B</figureCitation>
|
||
; middle and hind tibiae with medial spurs long and serrulate (
|
||
<figureCitation id="0C832A6FEB6DED40FC29FA4B67058387" box="[972,1058,1508,1532]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="5" pageNumber="498">Fig. 2A</figureCitation>
|
||
); front tarsomeres 1 to 3 markedly expanded laterally (
|
||
<figureCitation id="0C832A6FEB6DED40FD81F9AB61C48067" box="[612,739,1540,1564]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="5" pageNumber="498">Figs. 2B, C</figureCitation>
|
||
) in both males and females; middle tarsomeres 1 to 3 broad and with pads of adhesive squamosetae (Stork 1980) ventrally in males (
|
||
<figureCitation id="0C832A6FEB6DED40FB85F98A639B8026" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="5" pageNumber="498">Figs. 2D, E</figureCitation>
|
||
), narrower and without ventral setal pads in females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCA26561EB6DED40FF4EF9C9627880C6" pageId="5" pageNumber="498" type="biology_ecology">
|
||
<paragraph id="940736EAEB6DED40FF4EF9C9627880C6" blockId="5.[123,1174,1183,1725]" pageId="5" pageNumber="498">
|
||
<emphasis id="A6CCEAF8EB6DED40FF4EF9C9623C8006" bold="true" box="[171,283,1638,1661]" pageId="5" pageNumber="498">Diversity:</emphasis>
|
||
At present, six species are recognized in the Afrotropical Region and seven species in the Asian fauna. We here recognize nine
|
||
<taxonomicName id="53B84D69EB6DED40FDB1F92A600E80E6" box="[596,809,1669,1693]" pageId="5" pageNumber="498">
|
||
<emphasis id="A6CCEAF8EB6DED40FDB1F92A61F280E6" box="[596,725,1669,1693]" italics="true" pageId="5" pageNumber="498">Cyclosomus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
as occurring in Asia, with two of these new to science.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCA26561EB6EED4DFECFFF7467A882CB" lastPageId="8" lastPageNumber="501" pageId="6" pageNumber="499" type="key">
|
||
<paragraph id="940736EAEB6EED43FECFFF7460C28689" blockId="6.[298,997,219,243]" box="[298,997,219,243]" pageId="6" pageNumber="499">
|
||
<heading id="CF4F8186EB6EED43FECFFF7460C28689" bold="true" box="[298,997,219,243]" centered="true" fontSize="10" level="2" pageId="6" pageNumber="499" reason="5">
|
||
<emphasis id="A6CCEAF8EB6EED43FECFFF7460C28689" bold="true" box="[298,997,219,243]" pageId="6" pageNumber="499">
|
||
Key for Identification of Adults of
|
||
<taxonomicName id="53B84D69EB6EED43FD53FF7460B68689" box="[694,913,219,243]" pageId="6" pageNumber="499">
|
||
<emphasis id="A6CCEAF8EB6EED43FD53FF74601D8688" bold="true" box="[694,826,219,243]" italics="true" pageId="6" pageNumber="499">Cyclosomus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of Asia
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6EED43FF4EFEA267A88493" blockId="6.[123,1174,269,745]" pageId="6" pageNumber="499">
|
||
Both male and female adults of
|
||
<taxonomicName id="53B84D69EB6EED43FDEEFEA261C5875E" box="[523,738,269,293]" pageId="6" pageNumber="499">
|
||
<emphasis id="A6CCEAF8EB6EED43FDEEFEA261AB875E" box="[523,652,269,293]" italics="true" pageId="6" pageNumber="499">Cyclosomus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from the region can be identified using the key below. We have avoided gender-specific features as much as possible in creating the key, and where features unique to males are used, they are presented only as secondary in importance. However, these features can be very helpful in confirming the identification of males. Male specimens of all species included in the key have small pads of adhesive setae on the ventral surfaces of front tarsomeres 1 to 3. Females have no such pads. Although in most carabid beetle species these front tarsomeres are also distinctly broader in males than in females, in the Asian
|
||
<taxonomicName id="53B84D69EB6EED43FBD2FE6163DC847C" pageId="6" pageNumber="499">
|
||
<emphasis id="A6CCEAF8EB6EED43FBD2FE616381847C" italics="true" pageId="6" pageNumber="499">Cyclosomus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
these differences are very slight and therefore difficult to assess. Contributing to this difficulty is the fact that both males and females have the basal three tarsomeres asymmetrically expanded laterally as flat plates that undoubtedly aid in digging into the sandy substrate they inhabit. More reliable features for distinguishing the sexes are middle tarsomeres 1 to 3, which are distinctly broader in males than in female and have ventral adhesive setae, which are absent from females. We recommend that users of the key review the cautions given in the section entitled “Color and color patterns” under Materials and Methods above because the elytral color pattern is highly varied within some species and similar among at least some members of different species.
|
||
</paragraph>
|
||
<keyStep id="2F4C2E4FEB6EED43FF9EFCA467A9823F" pageId="6" pageNumber="499">
|
||
<paragraph id="940736EAEB6EED43FF9EFCA467A9859F" blockId="6.[123,1174,779,1092]" pageId="6" pageNumber="499">
|
||
<keyLead id="2F4995DFEB6EED43FF9EFCA467A9859F" pageId="6" pageNumber="499">
|
||
<key id="E02961DBEB6EED43FF9EFCA467A9859F" pageId="6" pageNumber="499">
|
||
1 Pronotum (
|
||
<figureCitation id="0C832A6FEB6EED43FEC1FCA462A18558" box="[292,390,779,803]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="6" pageNumber="499">Fig. 14A</figureCitation>
|
||
) with anterior angles narrow, distinctly acute, lateral margins slightly to distinctly sinuate near anterior angles, disk dark piceous to black, with lateral pale bands well-defined and very narrow; median lobe of male genitalia with shaft evenly arcuate and tapered to a thick apical lamella, slightly ventrally deflected in lateral aspect (
|
||
<figureCitation id="0C832A6FEB6EED43FC71FCC460D085F8" box="[916,1015,875,899]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="6" pageNumber="499">Fig. 16A</figureCitation>
|
||
), shaft slightly broadened for most of its length, then tapered to base of short, narrow, smoothly rounded apical lamella in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB6EED43FE38FC04611885B8" box="[477,575,939,963]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="6" pageNumber="499">Fig. 17A</figureCitation>
|
||
).......................................................................
|
||
<taxonomicName id="53B84D69EB6EED43FDC8FC6367A9859F" authority="Kavanaugh & Cueva-Dabkoski" authorityName="Kavanaugh & Cueva-Dabkoski" box="[557,1166,972,996]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="A6CCEAF8EB6EED43FDC8FC636010859F" box="[557,823,972,996]" italics="true" pageId="6" pageNumber="499">Cyclosomus acutangulus</emphasis>
|
||
Kavanaugh & Cueva-Dabkoski
|
||
</taxonomicName>
|
||
</key>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6EED43FF9EFC4367A9823F" blockId="6.[123,1174,779,1092]" pageId="6" pageNumber="499">
|
||
<keyLead id="2F4995DFEB6EED43FF9EFC4367A9823F" pageId="6" pageNumber="499">
|
||
1’ Pronotum (
|
||
<figureCitation id="0C832A6FEB6EED43FEC0FC436259827F" box="[293,382,1004,1028]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="6" pageNumber="499">Figs. 14</figureCitation>
|
||
B-I) with anterior angles broader, lateral margins not sinuate or (in very few specimens) faintly sinuate near anterior angles, lateral pale bands wider, well-defined or not, male genitalia as in
|
||
<figureCitation id="0C832A6FEB6EED43FE56FB83612E823F" box="[435,521,1068,1092]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="6" pageNumber="499">Figs. 16</figureCitation>
|
||
B-H, 17B-H.................................... 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2F4C2E4FEB6EED43FF9EFBF867A982D4" pageId="6" pageNumber="499">
|
||
<paragraph id="940736EAEB6EED43FF9EFBF867A982F4" blockId="6.[123,1174,1111,1200]" pageId="6" pageNumber="499">
|
||
<keyLead id="2F4995DFEB6EED43FF9EFBF867A982F4" pageId="6" pageNumber="499">
|
||
2 (1’)Elytra without (
|
||
<figureCitation id="0C832A6FEB6EED43FE86FBF862D58214" box="[355,498,1111,1135]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="6" pageNumber="499">Figs. 7A, 7C</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6EED43FE1AFBF8612C8214" box="[511,523,1111,1135]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="6" pageNumber="499">8</figureCitation>
|
||
-10,13) a preapical spot or with only a small and faint (
|
||
<figureCitation id="0C832A6FEB6EED43FB8FFBF863EB82F4" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="6" pageNumber="499">Fig. 7D</figureCitation>
|
||
) spot (seen in some specimens from inland parts of Indochina)................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6EED43FF9EFB3767A982D4" blockId="6.[123,1174,1111,1200]" box="[123,1166,1176,1200]" pageId="6" pageNumber="499">
|
||
<keyLead id="2F4995DFEB6EED43FF9EFB3767A982D4" box="[123,1166,1176,1200]" pageId="6" pageNumber="499">
|
||
2’ Elytra (
|
||
<figureCitation id="0C832A6FEB6EED43FF1EFB37626082CB" box="[251,327,1176,1200]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="6" pageNumber="499">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6EED43FEB6FB37624582CB" box="[339,354,1176,1200]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="6" pageNumber="499">6</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6EED43FE8AFB3762AD82CB" box="[367,394,1176,1200]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="6" pageNumber="499">11</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6EED43FE73FB37629482CB" box="[406,435,1176,1200]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="6" pageNumber="499">12</figureCitation>
|
||
,) with distinct preapical dark spot......................... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="940736EAEB6EED43FF9EFB6C67A88067" blockId="6.[123,1174,1219,1725]" pageId="6" pageNumber="499">
|
||
<keyLead id="2F4995DFEB6EED43FF9EFB6C67A88067" pageId="6" pageNumber="499">
|
||
<keyStep id="2F4C2E4FEB6EED43FF9EFB6C67A88067" pageId="6" pageNumber="499">
|
||
3 (2)Elytral epipleuron with long setae over entire length, visible from dorsal aspect in most specimens (
|
||
<figureCitation id="0C832A6FEB6EED43FF1CFB4C627C8280" box="[249,347,1251,1275]" captionStart="FIGURE" captionStartId="23.[171,183,1573,1592]" captionTargetBox="[156,1139,216,1547]" captionTargetId="figure-14@23.[156,1139,216,1560]" captionTargetPageId="23" captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="6" pageNumber="499">Fig. 10A</figureCitation>
|
||
) [but best viewed from below because setae may be bent posteriorly in some specimens]; pronotum (
|
||
<figureCitation id="0C832A6FEB6EED43FE49FAAC612D8360" box="[428,522,1283,1307]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="6" pageNumber="499">Fig. 14F</figureCitation>
|
||
) relatively narrow (ratio PWM/PL = 2.06 to 2.33) and less broadened basally (ratio PWM/PWA = 1.45 to 1.59), with disc piceous to black and lateral pale band wide and well-defined; free apex of prosternal intercoxal process long (as in
|
||
<figureCitation id="0C832A6FEB6EED43FBC2FAEC67AF8320" box="[1063,1160,1347,1371]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="6" pageNumber="499">Fig. 15C</figureCitation>
|
||
); elytral dark pattern varied (
|
||
<figureCitation id="0C832A6FEB6EED43FE36FACC61458300" box="[467,610,1379,1403]" captionStart="FIGURE" captionStartId="23.[171,183,1573,1592]" captionTargetBox="[156,1139,216,1547]" captionTargetId="figure-14@23.[156,1139,216,1560]" captionTargetPageId="23" captionText="FIGURE 10. Cyclosomus philippinus Heller. A. Holotype female, dorsal habitus (photograph by Olaf Jäger); B. Labels associated with holotype; C. Pale male (Zambales Mountains, Luzon Island, Philippines). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512978" httpUri="https://zenodo.org/record/11512978/files/figure.png" pageId="6" pageNumber="499">Figs. 10A, C</figureCitation>
|
||
), with basal and middle bands of average to greater than average thickness; median lobe of male genitalia with shaft arcuate basally and nearly straight subapically, tapered to a long and pointed apical lamella in lateral aspect (
|
||
<figureCitation id="0C832A6FEB6EED43FBCDFA0B67AE83C7" box="[1064,1161,1444,1468]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="6" pageNumber="499">Fig. 16E</figureCitation>
|
||
), shaft moderately broad basally, gradually tapered to moderately broad and apically blunted, sublaceolate apical lamella in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB6EED43FD69FA4B61FB8387" box="[652,732,1508,1532]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="6" pageNumber="499">Fig. 17</figureCitation>
|
||
); specimen from
|
||
<collectingCountry id="ECAF767AEB6EED43FC7CFA4B677C8387" box="[921,1115,1508,1532]" name="Philippines" pageId="6" pageNumber="499">Philippine Islands</collectingCountry>
|
||
(
|
||
<figureCitation id="0C832A6FEB6EED43FB8FFA4B63E18067" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="6" pageNumber="499">Fig. 21</figureCitation>
|
||
)..............................................
|
||
<taxonomicName id="53B84D69EB6EED43FCA1F9AB67A88067" ID-CoL="6C5XG" authority="Heller" authorityName="Heller" box="[836,1167,1540,1564]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="499" phylum="Arthropoda" rank="species" species="philippinus">
|
||
<emphasis id="A6CCEAF8EB6EED43FCA1F9AB67628067" box="[836,1093,1540,1564]" italics="true" pageId="6" pageNumber="499">Cyclosomus philippinus</emphasis>
|
||
Heller
|
||
</taxonomicName>
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6EED42FF9EF98B67A98748" blockId="6.[123,1174,1219,1725]" lastBlockId="7.[171,1174,219,307]" lastPageId="7" lastPageNumber="500" pageId="6" pageNumber="499">
|
||
<keyLead id="2F4995DFEB6EED42FF9EF98B67A98748" lastPageId="7" lastPageNumber="500" pageId="6" pageNumber="499">
|
||
3’ Elytral epipleuron with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; pronotum (
|
||
<figureCitation id="0C832A6FEB6EED43FDC9F9EB61DF8027" box="[556,760,1604,1628]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="6" pageNumber="499">Figs. 14 C, D, E, I</figureCitation>
|
||
) relatively broader in most specimens (ratio PWM/PL = 2.33 to 2.93) and more broadened basally (ratio PWM/PWA = 1.63 to 1.75), with disc varied in color (pale tan-orange to black) and lateral pale band well-defined or not; free apex of prosternal intercoxal process varied in length, from long to short (
|
||
<figureCitation id="0C832A6FEB6EED43FC0BF90A671A80C6" box="[1006,1085,1701,1725]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="6" pageNumber="499">Fig. 15</figureCitation>
|
||
); elytral dark pattern varied, with basal and middle bands of average to less than average thickness or middle band absent; median lobe of male genitalia not sublanceolate, rounded in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FF56FEB462458748" box="[179,354,283,307]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="7" pageNumber="500">Figs. 17C, D, H</figureCitation>
|
||
); specimen not from
|
||
<collectingCountry id="ECAF767AEB6FED42FDA2FEB4602F8748" box="[583,776,283,307]" name="Philippines" pageId="7" pageNumber="500">Philippine Islands</collectingCountry>
|
||
........................... 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<key id="E02961DBEB6FED4DFF9EFEE967A882CB" lastPageId="8" lastPageNumber="501" pageId="7" pageNumber="500">
|
||
<keyStep id="2F4C2E4FEB6FED42FF9EFEE967A985F9" pageId="7" pageNumber="500">
|
||
<paragraph id="940736EAEB6FED42FF9EFEE967A8849A" blockId="7.[123,1174,326,898]" pageId="7" pageNumber="500">
|
||
<keyLead id="2F4995DFEB6FED42FF9EFEE967A8849A" pageId="7" pageNumber="500">
|
||
4 (3’)Pronotum (
|
||
<figureCitation id="0C832A6FEB6FED42FECAFEE962AA8725" box="[303,397,326,350]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="7" pageNumber="500">Fig. 14C</figureCitation>
|
||
) relatively longer and narrower (ratio PWM/PL less than 2.50), with disc varied in color (rufous to piceous) and lateral pale bands medium-width and well-defined; free apex of prosternal intercoxal process long (as in
|
||
<figureCitation id="0C832A6FEB6FED42FD5DFE28603D87E4" box="[696,794,391,415]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="7" pageNumber="500">Fig. 15C</figureCitation>
|
||
); body form (
|
||
<figureCitation id="0C832A6FEB6FED42FC4AFE28676887E4" box="[943,1103,391,415]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="7" pageNumber="500">Figs. 7A, C, D</figureCitation>
|
||
) more elongate-ovoid (ratio BL/EW = 1.47 to 1.53), with elytra widest at or slightly anterior to mid-length; elytral dark pattern varied in both form and color (
|
||
<figureCitation id="0C832A6FEB6FED42FCFFFE68609C87A4" box="[794,955,455,479]" captionStart="FIGURE" captionStartId="16.[171,183,1579,1598]" captionTargetBox="[182,1114,216,1560]" captionTargetId="figure-9@16.[182,1114,216,1560]" captionTargetPageId="16" captionText="FIGURE 7. Cyclosomus inustus Andrewes. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Pale female (Nanoa Island, Guangdong, China); D. Dark male (“Hat Baoho”, Steong Treng, Cambodia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512970" httpUri="https://zenodo.org/record/11512970/files/figure.png" pageId="7" pageNumber="500">Figs. 7A, C, D</figureCitation>
|
||
), with dark maculae rufous to piceous and with basal and middle bands of average to less than average thickness and continuity or middle band absent; elytral striae very deeply impressed, elytral intervals (especially intervals 3 to 5) slightly but distinctly convex; median lobe of male genitalia long and more slender, shaft of approximately equal thickness throughout, ventral curvature gradually and only slightly arcuate in apical two-thirds in lateral aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FC6FFDC760CB84FB" box="[906,1004,616,640]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="7" pageNumber="500">Fig. 16C</figureCitation>
|
||
), apical lamella longer, rounded apically, and not or only slightly narrowed basally in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FBC2FD2767AF84DB" box="[1063,1160,648,672]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="7" pageNumber="500">Fig. 17C</figureCitation>
|
||
); specimen from southern Asia, from northeastern coastal
|
||
<collectingCountry id="ECAF767AEB6FED42FCF6FD07606B84BB" box="[787,844,680,704]" name="India" pageId="7" pageNumber="500">India</collectingCountry>
|
||
, through Indochina, to coastal southeastern and eastern
|
||
<collectingCountry id="ECAF767AEB6FED42FE5DFD6662DF849A" box="[440,504,713,737]" name="China" pageId="7" pageNumber="500">China</collectingCountry>
|
||
and
|
||
<collectingCountry id="ECAF767AEB6FED42FDC9FD66615D849A" box="[556,634,713,737]" name="Taiwan" pageId="7" pageNumber="500">Taiwan</collectingCountry>
|
||
(
|
||
<figureCitation id="0C832A6FEB6FED42FD6FFD6661FF849A" box="[650,728,713,737]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="7" pageNumber="500">Fig. 20</figureCitation>
|
||
).......
|
||
<taxonomicName id="53B84D69EB6FED42FCACFD6667A8849A" ID-CoL="32X57" authority="Andrewes" authorityName="Andrewes" box="[841,1167,713,737]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="inustus">
|
||
<emphasis id="A6CCEAF8EB6FED42FCACFD66673A849A" box="[841,1053,713,737]" italics="true" pageId="7" pageNumber="500">Cyclosomus inustus</emphasis>
|
||
Andrewes
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6FED42FF9EFD4667A985F9" blockId="7.[123,1174,326,898]" pageId="7" pageNumber="500">
|
||
<keyLead id="2F4995DFEB6FED42FF9EFD4667A985F9" pageId="7" pageNumber="500">
|
||
4’ Pronotum (
|
||
<figureCitation id="0C832A6FEB6FED42FEC0FD4662EA857A" box="[293,461,745,769]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="7" pageNumber="500">Figs. 14D, E, I</figureCitation>
|
||
) shorter and very broad (ratio PWM/PL equal to or greater than 2.65); body form (
|
||
<figureCitation id="0C832A6FEB6FED42FE96FCA662F3855A" box="[371,468,777,801]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="7" pageNumber="500">Figs. 8A</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6FED42FE04FCA66123855A" box="[481,516,777,801]" captionStart="FIGURE" captionStartId="21.[171,183,1155,1174]" captionTargetBox="[120,1176,216,1138]" captionTargetId="figure-233@21.[120,1176,216,1147]" captionTargetPageId="21" captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512976" httpUri="https://zenodo.org/record/11512976/files/figure.png" pageId="7" pageNumber="500">9A</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6FED42FDF5FCA66190855A" box="[528,695,777,801]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="7" pageNumber="500">13A, 13C, 13D</figureCitation>
|
||
) shorter in most specimens (ratio BL/EW = 1.36 to 1.49), more roundly ovoid, broader apically; elytral dark pattern varied in both form and color; elytral striae less deeply impressed in most specimens, elytral intervals flat or nearly so; median lobe of male genitalia varied in form (
|
||
<figureCitation id="0C832A6FEB6FED42FD38FCC5604D85F9" box="[733,874,874,898]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="7" pageNumber="500">Figs. 16D, H</figureCitation>
|
||
)................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2F4C2E4FEB6FED42FF9EFC3A67A98294" pageId="7" pageNumber="500">
|
||
<paragraph id="940736EAEB6FED42FF9EFC3A67A88235" blockId="7.[123,1174,917,1263]" pageId="7" pageNumber="500">
|
||
<keyLead id="2F4995DFEB6FED42FF9EFC3A67A88235" pageId="7" pageNumber="500">
|
||
5 (4’)Dorsal surface (
|
||
<figureCitation id="0C832A6FEB6FED42FE81FC3A629E85D6" box="[356,441,917,941]" captionStart="FIGURE" captionStartId="21.[171,183,1155,1174]" captionTargetBox="[120,1176,216,1138]" captionTargetId="figure-233@21.[120,1176,216,1147]" captionTargetPageId="21" captionText="FIGURE 9. Cyclosomus pallidus Kavanaugh and Cueva-Dabkoski sp. nov. A. Holotype female, dorsal habitus; B. Labels associated with holotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512976" httpUri="https://zenodo.org/record/11512976/files/figure.png" pageId="7" pageNumber="500">Fig. 9A</figureCitation>
|
||
) mainly pale tan-orange, with only pronotal disc, elytral basal dark band and longitudinal dark band on elytral interval 1 slightly darker brown, middle dark band absent; pronotum (
|
||
<figureCitation id="0C832A6FEB6FED42FE97FC7A62E88596" box="[370,463,981,1005]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="7" pageNumber="500">Fig. 14E</figureCitation>
|
||
) very broadly flattened laterally (especially anteriorly), lateral pale bands very broad and only slightly contrasting with color on disc; free apex of prosternal intercoxal process short (as in
|
||
<figureCitation id="0C832A6FEB6FED42FE28FBB961158255" box="[461,562,1046,1070]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="7" pageNumber="500">Fig. 15D</figureCitation>
|
||
); specimen from southeastern
|
||
<collectingCountry id="ECAF767AEB6FED42FC67FBB960FB8255" box="[898,988,1046,1070]" name="Pakistan" pageId="7" pageNumber="500">Pakistan</collectingCountry>
|
||
(
|
||
<figureCitation id="0C832A6FEB6FED42FC0BFBB967678255" box="[1006,1088,1046,1070]" captionStart="FIGURE" captionStartId="39.[171,183,1593,1612]" captionTargetBox="[168,1128,979,1582]" captionTargetId="figure-84@39.[168,1128,979,1582]" captionTargetPageId="39" captionText="FIGURE 21. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow highlighted area with black question mark = type area for C. marginatus Motschulsky; white square = C. pallidus Kavanaugh and Cueva-Dabkoski sp. nov.; white circles = C. philippinus Heller. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513002" httpUri="https://zenodo.org/record/11513002/files/figure.png" pageId="7" pageNumber="500">Fig. 21</figureCitation>
|
||
); males unknown...............
|
||
<emphasis id="A6CCEAF8EB6FED42FE04FB9967A88235" bold="true" box="[481,1167,1078,1102]" pageId="7" pageNumber="500">
|
||
<taxonomicName id="53B84D69EB6FED42FE04FB9967158235" authority="Kavanaugh & Cueva-Dabkoski" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[481,1074,1078,1102]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="pallidus" status="sp. nov.">
|
||
<emphasis id="A6CCEAF8EB6FED42FE04FB9961E48235" bold="true" box="[481,707,1078,1102]" italics="true" pageId="7" pageNumber="500">Cyclosomus pallidus</emphasis>
|
||
Kavanaugh & Cueva-Dabkoski
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDFF5783EB6FED42FBDCFB9867A88235" box="[1081,1167,1079,1102]" pageId="7" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6FED42FF9EFBF967A98294" blockId="7.[123,1174,917,1263]" pageId="7" pageNumber="500">
|
||
<keyLead id="2F4995DFEB6FED42FF9EFBF967A98294" pageId="7" pageNumber="500">
|
||
5’ Dorsal surface (
|
||
<figureCitation id="0C832A6FEB6FED42FEB0FBF962928215" box="[341,437,1110,1134]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="7" pageNumber="500">Figs. 8A</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB6FED42FE24FBF961178215" box="[449,560,1110,1134]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="7" pageNumber="500">13A, C, D</figureCitation>
|
||
) with distinctly contrasting pale and dark areas; pronotal disc darker, rufopiceous to black, with lateral pale bands broad but narrower than above (
|
||
<figureCitation id="0C832A6FEB6FED42FB85FBD9622982D4" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="7" pageNumber="500">Figs. 14D, 14I</figureCitation>
|
||
); elytra pale yellowish-brown with basal, middle and longitudinal dark bands present and dark reddish brown to black; free apex of prosternal intercoxal process short (as in
|
||
<figureCitation id="0C832A6FEB6FED42FB8EFB1863FD8294" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="7" pageNumber="500">Fig. 15D</figureCitation>
|
||
) or medium in length (as in
|
||
<figureCitation id="0C832A6FEB6FED42FDEBFB78614A8294" box="[526,621,1239,1263]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="7" pageNumber="500">Fig. 15B</figureCitation>
|
||
)...................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2F4C2E4FEB6FED42FF9EFAAD67A880C6" pageId="7" pageNumber="500">
|
||
<paragraph id="940736EAEB6FED42FF9EFAAD67A883A0" blockId="7.[123,1174,1282,1725]" pageId="7" pageNumber="500">
|
||
<keyLead id="2F4995DFEB6FED42FF9EFAAD67A883A0" pageId="7" pageNumber="500">
|
||
6 (5’)Body form (
|
||
<figureCitation id="0C832A6FEB6FED42FEA5FAAD62B18361" box="[320,406,1282,1306]" captionStart="FIGURE" captionStartId="19.[171,183,1069,1088]" captionTargetBox="[144,1131,234,1045]" captionTargetId="figure-279@19.[120,1176,216,1058]" captionTargetPageId="19" captionText="FIGURE 8. Cyclosomus marginatus Motschulsky.A. Lectotype female, dorsal habitus (photograph by Rachel Diaz-Bastian); B. Labels associated with lectotype. Scale line = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512974" httpUri="https://zenodo.org/record/11512974/files/figure.png" pageId="7" pageNumber="500">Fig. 8A</figureCitation>
|
||
) roundly ovoid (ratio BL/EW = 1.46 to 1.49; ratio EL/EW = 0.94 to 0.96); free apex of prosternal intercoxal process medium in length (as in
|
||
<figureCitation id="0C832A6FEB6FED42FC4AFA8D67288341" box="[943,1039,1314,1338]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="7" pageNumber="500">Fig. 15B</figureCitation>
|
||
); elytra with middle transverse dark band narrow, extended laterally onto interval 6, portion on intervals 2 to 4 connected to portion on intervals 5 and 6 by in narrow, oblique dark line; median lobe of male genitalia with shaft of approximately equal thickness throughout, ventral curvature straightened subapically in lateral aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FD96FA0C61F083C0" box="[627,727,1443,1467]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="7" pageNumber="500">Fig. 16D</figureCitation>
|
||
), apical lamella slightly shorter in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FF18FA6C627983A0" box="[253,350,1475,1499]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="7" pageNumber="500">Fig. 17D</figureCitation>
|
||
).............................
|
||
<taxonomicName id="53B84D69EB6FED42FD18FA6C67A883A0" ID-CoL="32X59" authority="Motschulsky" authorityName="Motschulsky" box="[765,1167,1475,1500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="marginatus">
|
||
<emphasis id="A6CCEAF8EB6FED42FD18FA6C60D983A7" box="[765,1022,1475,1500]" italics="true" pageId="7" pageNumber="500">Cyclosomus marginatus</emphasis>
|
||
Motschulsky
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB6FED42FF9EFA4B67A880C6" blockId="7.[123,1174,1282,1725]" pageId="7" pageNumber="500">
|
||
<keyLead id="2F4995DFEB6FED42FF9EFA4B67A880C6" pageId="7" pageNumber="500">
|
||
6’ Body form (
|
||
<figureCitation id="0C832A6FEB6FED42FED0FA4B62F88387" box="[309,479,1508,1532]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="7" pageNumber="500">Fig. 13A, C, D</figureCitation>
|
||
) even more roundly ovoid (ratio BL/EW = 1.36 to 1.41; ratio EL/EW = 0.88 to 0.93); free apex of prosternal intercoxal process short (
|
||
<figureCitation id="0C832A6FEB6FED42FC48F9AB67378067" box="[941,1040,1540,1564]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="7" pageNumber="500">Fig. 15D</figureCitation>
|
||
); elytra with middle transverse dark band varied, moderately thick in most specimens (
|
||
<figureCitation id="0C832A6FEB6FED42FC20F98B670F8047" box="[965,1064,1572,1596]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="7" pageNumber="500">Fig. 13A</figureCitation>
|
||
), markedly thickened (
|
||
<figureCitation id="0C832A6FEB6FED42FEA5F9EB62868027" box="[320,417,1604,1628]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="7" pageNumber="500">Fig. 13C</figureCitation>
|
||
) or narrowed (
|
||
<figureCitation id="0C832A6FEB6FED42FDA2F9EB618E8027" box="[583,681,1604,1628]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="7" pageNumber="500">Fig. 13D</figureCitation>
|
||
) (much as in
|
||
<taxonomicName id="53B84D69EB6FED42FCA6F9EB60F88027" ID-CoL="32X59" box="[835,991,1604,1628]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="marginatus">C. marginatus</taxonomicName>
|
||
), in a few specimens; median lobe of male genitalia with shaft distinctly thicker and more arcuate, ventral curvature distinctly curved throughout in lateral aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FD0CF92A606A80E6" box="[745,845,1669,1693]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="7" pageNumber="500">Fig. 16H</figureCitation>
|
||
), apical lamella longer in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB6FED42FEC7F90A62A480C6" box="[290,387,1701,1725]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="7" pageNumber="500">Fig. 17H</figureCitation>
|
||
)....
|
||
<emphasis id="A6CCEAF8EB6FED42FE2DF90A67A880C6" bold="true" box="[456,1167,1701,1725]" pageId="7" pageNumber="500">
|
||
<taxonomicName id="53B84D69EB6FED42FE2DF90A671580C6" authority="Cueva-Dabkoski & Kavanaugh" authorityName="Cueva-Dabkoski & Kavanaugh" authorityYear="2023" box="[456,1074,1701,1725]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="500" phylum="Arthropoda" rank="species" species="vespertilio" status="sp. nov.">
|
||
<emphasis id="A6CCEAF8EB6FED42FE2DF90A61E380C6" bold="true" box="[456,708,1701,1725]" italics="true" pageId="7" pageNumber="500">Cyclosomus vespertilio</emphasis>
|
||
Cueva-Dabkoski & Kavanaugh
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDFF5783EB6FED42FBDCF90967A880C6" box="[1081,1167,1702,1725]" pageId="7" pageNumber="500" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2F4C2E4FEB60ED4DFF9EFF7467A984A7" pageId="8" pageNumber="501">
|
||
<paragraph id="940736EAEB60ED4DFF9EFF7467A98783" blockId="8.[123,1174,219,732]" pageId="8" pageNumber="501">
|
||
<keyLead id="2F4995DFEB60ED4DFF9EFF7467A98783" pageId="8" pageNumber="501">
|
||
7 (2’)Elytral epipleuron with long setae over entire length, visible from dorsal aspect in most specimens [but best viewed from below because setae may be bent posteriorly in some specimens]; pronotum (
|
||
<figureCitation id="0C832A6FEB60ED4DFEC6FEB362A3874F" box="[291,388,284,308]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="8" pageNumber="501">Fig. 14B</figureCitation>
|
||
) light reddish brown to rufopiceous, with lateral pale bands moderately broad but not sharply defined, apical angles broad and broadly rounded apically; elytral dark pattern more thickly developed than average in most specimens (
|
||
<figureCitation id="0C832A6FEB60ED4DFC8AFEF260E9870E" box="[879,974,349,373]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="8" pageNumber="501">Figs. 5C</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB60ED4DFC3EFEF260DA870E" box="[987,1021,349,373]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="8" pageNumber="501">6A</figureCitation>
|
||
) but less than average (
|
||
<figureCitation id="0C832A6FEB60ED4DFEF5FED1625587ED" box="[272,370,382,406]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="8" pageNumber="501">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="0C832A6FEB60ED4DFE64FED1628687ED" box="[385,417,382,406]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="8" pageNumber="501">6B</figureCitation>
|
||
) in a few specimens; male median lobe with apical lamella slightly recurved dorsally in relation to ventral curvature of shaft in lateral aspect (
|
||
<figureCitation id="0C832A6FEB60ED4DFC38FE31676787CD" box="[989,1088,414,438]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="8" pageNumber="501">Fig. 16B</figureCitation>
|
||
), apical lamella broadly triangular with a distinct, more narrowly-rounded projection apicomedially in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB60ED4DFEA0FE4F62838783" box="[325,420,480,504]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="8" pageNumber="501">Fig. 17B</figureCitation>
|
||
)...........................
|
||
<taxonomicName id="53B84D69EB60ED4DFCC2FE4F67A98783" ID-CoL="32X56" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[807,1166,480,504]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="flexuosus">
|
||
<emphasis id="A6CCEAF8EB60ED4DFCC2FE4F67358783" box="[807,1042,480,504]" italics="true" pageId="8" pageNumber="501">Cyclosomus flexuosus</emphasis>
|
||
(Fabricius)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB60ED4DFF9EFDAF67A984A7" blockId="8.[123,1174,219,732]" pageId="8" pageNumber="501">
|
||
<keyLead id="2F4995DFEB60ED4DFF9EFDAF67A984A7" pageId="8" pageNumber="501">
|
||
7’ Elytral epipleuron with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; pronotum rufopiceous to black, with lateral pale bands narrower at least in apical half and better defined, apical angles narrower and less broadly rounded; elytral dark pattern less varied, with middle band of average (
|
||
<figureCitation id="0C832A6FEB60ED4DFD0CFDCD606E8401" box="[745,841,610,634]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="8" pageNumber="501">Fig. 12A</figureCitation>
|
||
) to distinctly greater than average (
|
||
<figureCitation id="0C832A6FEB60ED4DFF05FD2C62AA84E0" box="[224,397,643,667]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="8" pageNumber="501">Figs. 11A, B, C</figureCitation>
|
||
) thickness; male median lobe with apical lamella extended in line with ventral curvature of shaft in lateral aspect (
|
||
<figureCitation id="0C832A6FEB60ED4DFD99FD0C602284C0" box="[636,773,675,699]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="8" pageNumber="501">Figs. 16F, G</figureCitation>
|
||
), apical lamella in dorsal view (
|
||
<figureCitation id="0C832A6FEB60ED4DFB85FD0C63D384A7" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="8" pageNumber="501">Figs. 17F, G</figureCitation>
|
||
) not as above........................................................ 8
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2F4C2E4FEB60ED4DFF9EFD5F67A882CB" pageId="8" pageNumber="501">
|
||
<paragraph id="940736EAEB60ED4DFF9EFD5F67A885B7" blockId="8.[123,1174,752,1200]" pageId="8" pageNumber="501">
|
||
<keyLead id="2F4995DFEB60ED4DFF9EFD5F67A885B7" pageId="8" pageNumber="501">
|
||
8 (7’)Size slightly smaller, BL males =
|
||
<quantity id="53409B0FEB60ED4DFDCCFD5E61E18573" box="[553,710,752,776]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="8.1" metricValueMin="7.7" pageId="8" pageNumber="501" unit="mm" value="7.9" valueMax="8.1" valueMin="7.7">7.7 to 8.1 mm</quantity>
|
||
, females =7.1 to 7.4; elytra (
|
||
<figureCitation id="0C832A6FEB60ED4DFBE4FD5F67AA8573" box="[1025,1165,752,776]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="8" pageNumber="501">Figs. 11A, B</figureCitation>
|
||
) with middle transverse dark band extended from midline at least across interval 8, also onto interval
|
||
<quantity id="53409B0FEB60ED4DFEE1FC9E62088532" box="[260,303,817,841]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="501" unit="in" value="9.0">9 in</quantity>
|
||
some specimens; pronotum (
|
||
<figureCitation id="0C832A6FEB60ED4DFD88FC9E61F78532" box="[621,720,817,841]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="8" pageNumber="501">Fig. 14G</figureCitation>
|
||
) relatively narrow (ratio PWM/PL = 2.25 to 2.43), piceous to black with lateral pale bands slightly narrower, distinctly defined, and slightly narrowed basally; median lobe of male genitalia with apical lamella rounded and of medium length in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB60ED4DFDEDFC3C614185D0" box="[520,614,915,939]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="8" pageNumber="501">Fig. 17F</figureCitation>
|
||
); specimen from West
|
||
<collectingCountry id="ECAF767AEB60ED4DFCBEFC3C609A85D0" box="[859,957,915,939]" name="Malaysia" pageId="8" pageNumber="501">Malaysia</collectingCountry>
|
||
or northern
|
||
<collectingCountry id="ECAF767AEB60ED4DFBA7FC3C63EE85B7" name="Indonesia" pageId="8" pageNumber="501">Indonesia</collectingCountry>
|
||
(
|
||
<figureCitation id="0C832A6FEB60ED4DFF3DFC1B620185B7" box="[216,294,948,972]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="8" pageNumber="501">Fig. 22</figureCitation>
|
||
)....................................
|
||
<taxonomicName id="53B84D69EB60ED4DFCFCFC1B67A885B7" ID-CoL="6C69G" authority="Bouchard" authorityName="Bouchard" box="[793,1167,948,972]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="sumatrensis">
|
||
<emphasis id="A6CCEAF8EB60ED4DFCFCFC1B673885B7" box="[793,1055,948,972]" italics="true" pageId="8" pageNumber="501">Cyclosomus sumatrensis</emphasis>
|
||
Bouchard
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="940736EAEB60ED4DFF9EFC7B67A882CB" blockId="8.[123,1174,752,1200]" pageId="8" pageNumber="501">
|
||
<keyLead id="2F4995DFEB60ED4DFF9EFC7B67A882CB" pageId="8" pageNumber="501">
|
||
8’ Size slightly larger, BL males =
|
||
<quantity id="53409B0FEB60ED4DFDEEFC7B61818597" box="[523,678,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="8.399999999999999" metricValueMax="8.6" metricValueMin="8.2" pageId="8" pageNumber="501" unit="mm" value="8.399999999999999" valueMax="8.6" valueMin="8.2">8.2 to 8.6 mm</quantity>
|
||
, female =
|
||
<quantity id="53409B0FEB60ED4DFCFFFC7B60578597" box="[794,880,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="8" pageNumber="501" unit="mm" value="9.2">9.2 mm</quantity>
|
||
; elytra (
|
||
<figureCitation id="0C832A6FEB60ED4DFC2CFC7B67718597" box="[969,1110,980,1004]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="8" pageNumber="501">Figs. 12A, C</figureCitation>
|
||
) with middle transverse dark band extended from midline to interval
|
||
<quantity id="53409B0FEB60ED4DFCA1FC5A604B8276" box="[836,876,1013,1037]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="8" pageNumber="501" unit="in" value="6.0">6 in</quantity>
|
||
most specimens, onto interval 7 and part of interval
|
||
<quantity id="53409B0FEB60ED4DFE5BFBBA62CF8256" box="[446,488,1045,1069]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="501" unit="in" value="8.0">8 in</quantity>
|
||
some specimens; pronotum (
|
||
<figureCitation id="0C832A6FEB60ED4DFCC0FBBA60A08256" box="[805,903,1045,1069]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="8" pageNumber="501">Fig. 14H</figureCitation>
|
||
) relatively broader (ratio PWM/PL = 2.44 to 2.50), rufopiceous piceous to black with lateral pale bands slightly broader, less well-defined, and distinctly broadened basally; median lobe of male genitalia with apical lamella short and bluntly triangular in dorsal aspect (
|
||
<figureCitation id="0C832A6FEB60ED4DFCC4FBD860A082F4" box="[801,903,1143,1167]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="8" pageNumber="501">Fig. 17G</figureCitation>
|
||
); specimen from eastern
|
||
<collectingCountry id="ECAF767AEB60ED4DFF4EFB3763C582CB" box="[171,226,1176,1200]" name="India" pageId="8" pageNumber="501">India</collectingCountry>
|
||
to westernmost Yunnan (
|
||
<figureCitation id="0C832A6FEB60ED4DFE16FB37616682CB" box="[499,577,1176,1200]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="8" pageNumber="501">Fig. 22</figureCitation>
|
||
)..............
|
||
<taxonomicName id="53B84D69EB60ED4DFCF5FB3767A882CB" ID-CoL="32X5F" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[784,1167,1176,1200]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="suturalis">
|
||
<emphasis id="A6CCEAF8EB60ED4DFCF5FB3760D282CB" box="[784,1013,1176,1200]" italics="true" pageId="8" pageNumber="501">Cyclosomus suturalis</emphasis>
|
||
(Wiedemann)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |