treatments-xml/data/1C/11/87/1C1187FCEB60ED4FFF9EFB7662B28501.xml

<document id="B3563BBDCCA72601580F08AA42CF079C" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB60ED4FFF9EFB7662B28501" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus acutangulus Kavanaugh &amp; Cueva-Dabkoski" docType="treatment" docVersion="12" lastPageNumber="503" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="501" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="982D52110F7ED8ED8E3BBCD8B35708B4">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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<mods:namePart id="3B66AC29E55A001621C2C48B5C52C7CE">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="C2BFE0C8E23477E4192D56DB8721D42A">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:namePart id="D50E39D0FF0A65C971F1C8005C3188C2">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="E9712C7F5B5C36516145C70BD159DCD5">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<treatment id="1C1187FCEB60ED4FFF9EFB7662B28501" ID-DOI="http://doi.org/10.5281/zenodo.12726719" ID-GBIF-Taxon="235039994" ID-Zenodo-Dep="12726719" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501" httpUri="http://treatment.plazi.org/id/1C1187FCEB60ED4FFF9EFB7662B28501" lastPageId="10" lastPageNumber="503" pageId="8" pageNumber="501">
<subSubSection id="DCA26561EB60ED4DFF9EFB7661DB828A" box="[123,764,1241,1265]" pageId="8" pageNumber="501" type="nomenclature">
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<taxonomicName id="53B84D69EB60ED4DFF9EFB7661DB828A" authority="Kavanaugh &amp; Cueva-Dabkoski" authorityName="Kavanaugh &amp; Cueva-Dabkoski" box="[123,764,1241,1265]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="acutangulus">
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<emphasis id="A6CCEAF8EB60ED4DFF9EFB7662A9828A" bold="true" box="[123,398,1241,1265]" italics="true" pageId="8" pageNumber="501">Cyclosomus acutangulus</emphasis>
Kavanaugh &amp; Cueva-Dabkoski
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="DCA26561EB60ED4DFF9EFB556107836B" box="[123,544,1274,1296]" pageId="8" pageNumber="501" type="description">
<paragraph id="940736EAEB60ED4DFF9EFB556107836B" blockId="8.[123,1175,1241,1388]" box="[123,544,1274,1296]" pageId="8" pageNumber="501">
<figureCitation id="0C832A6FEB60ED4DFF9EFB5563F0836B" box="[123,215,1274,1296]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="8" pageNumber="501">Figures 2</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFF07FB5563D7836B" box="[226,240,1274,1296]" captionStart="FIGURE" captionStartId="9.[171,183,1078,1097]" captionTargetBox="[120,1176,236,1061]" captionTargetId="figure-234@9.[120,1176,216,1061]" captionTargetPageId="9" captionText="FIGURE 4. Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski, dorsal habitus. A. Holotype male (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Female (Bhalukpong, Arunachal Pradesh, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512962" httpUri="https://zenodo.org/record/11512962/files/figure.png" pageId="8" pageNumber="501">4</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFF1FFB556201836B" box="[250,294,1274,1296]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="8" pageNumber="501">14A</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFED5FB55627B836B" box="[304,348,1274,1296]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="8" pageNumber="501">15A</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFE83FB5562B6836B" box="[358,401,1274,1296]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="8" pageNumber="501">16A</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFE79FB5562EF836B" box="[412,456,1274,1296]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="8" pageNumber="501">17A</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFE37FB5562D9836B" box="[466,510,1274,1296]" captionStart="FIGURE" captionStartId="37.[171,183,1577,1596]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@37.[120,1176,216,1561]" captionTargetPageId="37" captionText="FIGURE 18. Photographs of habitats for Cyclosomus species. A. C. acutangulus Kavanaugh &amp; Cueva-Dabkoski (Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China; photograph by David H. Kavanaugh); B. C. suturalis (Wiedemann) (Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China; photograph by Hong-Bin Liang)." figureDoi="http://doi.org/10.5281/zenodo.11512996" httpUri="https://zenodo.org/record/11512996/files/figure.png" pageId="8" pageNumber="501">18A</figureCitation>
, 
<figureCitation id="0C832A6FEB60ED4DFDEDFB556107836B" box="[520,544,1274,1296]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="8" pageNumber="501">20</figureCitation>
</paragraph>
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<treatmentCitation id="151910FBEB60ED4DFF9EFAB0602A834E" ID-DOI="http://doi.org/10.5281/zenodo.12726719" ID-Zenodo-Dep="12726719" author="KAVANAUGH, D. H. &amp; M. CUEVA-DABKOSKI &amp; H. B. LIANG" box="[123,781,1311,1333]" httpUri="http://treatment.plazi.org/id/1C1187FCEB60ED4FFF9EFB7662B28501" page="459" pageId="8" pageNumber="501" year="2023">
<taxonomicName id="53B84D69EB60ED4DFF9EFAB0602A834E" authority="Kavanaugh &amp; Cueva-Dabkoski, 2023: 459" authorityName="Kavanaugh &amp; Cueva-Dabkoski" authorityPageNumber="459" authorityYear="2023" box="[123,781,1311,1333]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB60ED4DFF9EFAB0624D834E" box="[123,362,1311,1333]" italics="true" pageId="8" pageNumber="501">Cyclosomus acutangulus</emphasis>
<bibRefCitation id="F0294B1BEB60ED4DFE95FAB0602A834E" author="KAVANAUGH, D. H. &amp; M. CUEVA-DABKOSKI &amp; H. B. LIANG" box="[368,781,1311,1333]" pageId="8" pageNumber="501" pagination="451 - 491" refId="ref23619" refString="KAVANAUGH, D. H., M. CUEVA-DABKOSKI, AND H. B. LIANG. 2023. Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with descriptions of two new species. Proceedings of the California Academy of Sciences (Series 4) 67: 451 - 491." type="journal article" year="2023">Kavanaugh &amp; Cueva-Dabkoski, 2023: 459</bibRefCitation>
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. 
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, a male, deposited in 
<collectionCode id="F2A9AE2FEB60ED4DFB82FAB067B5834E" box="[1127,1170,1311,1333]" pageId="8" pageNumber="501">IOZ</collectionCode>
. Type locality: 
<collectingCountry id="ECAF767AEB60ED4DFEDAFA92625C8328" box="[319,379,1341,1363]" name="China" pageId="8" pageNumber="501">China</collectingCountry>
, 
<collectingRegion id="567CF808EB60ED4DFE6DFA9262F38329" box="[392,468,1341,1362]" country="China" name="Yunnan" pageId="8" pageNumber="501">Yunnan</collectingRegion>
, 
<collectingCounty id="7D664E66EB60ED4DFE07FA9261B88328" box="[482,671,1341,1363]" pageId="8" pageNumber="501">Tengchong County</collectingCounty>
, 
<collectingMunicipality id="7463AC90EB60ED4DFD48FA9260768328" box="[685,849,1341,1363]" pageId="8" pageNumber="501">Wuhe Township</collectingMunicipality>
, 
<location id="91676031EB60ED4DFCBAFA92672C8328" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501:91676031EB60ED4DFCBAFA92672C8328" box="[863,1035,1341,1363]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longchuan River" pageId="8" pageNumber="501" stateProvince="Yunnan">Longchuan River</location>
at 
<location id="91676031EB60ED4DFBD6FA9267B18328" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501:91676031EB60ED4DFBD6FA9267B18328" box="[1075,1174,1341,1363]" country="China" county="Tengchong County" municipality="Wuhe Township" name="Longjiang" pageId="8" pageNumber="501" stateProvince="Yunnan">Longjiang</location>
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Bridge.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB60ED4CFF4EFA2F63CD830C" lastPageId="9" lastPageNumber="502" pageId="8" pageNumber="501" type="diagnosis">
<paragraph id="940736EAEB60ED4CFF4EFA2F63D482CC" blockId="8.[123,1174,1408,1725]" lastBlockId="9.[123,1174,1183,1719]" lastPageId="9" lastPageNumber="502" pageId="8" pageNumber="501">
<emphasis id="A6CCEAF8EB60ED4DFF4EFA2F623883EC" bold="true" box="[171,287,1408,1431]" pageId="8" pageNumber="501">Diagnosis.</emphasis>
Adults of 
<taxonomicName id="53B84D69EB60ED4DFE74FA2F611583EC" box="[401,562,1407,1431]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="501" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB60ED4DFE74FA2F611583EC" box="[401,562,1407,1431]" italics="true" pageId="8" pageNumber="501">C. acutangulus</emphasis>
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can be distinguished from those of all other 
<taxonomicName id="53B84D69EB60ED4DFBF1FAD067B283EC" box="[1044,1173,1407,1431]" pageId="8" pageNumber="501">
<emphasis id="A6CCEAF8EB60ED4DFBF1FAD067B283EC" box="[1044,1173,1407,1431]" italics="true" pageId="8" pageNumber="501">Cyclosomus</emphasis>
</taxonomicName>
species in Asia by the following combination of character states: Body size medium for genus, males = 
<quantity id="53409B0FEB60ED4DFF30FA6E624C83A2" box="[213,363,1473,1497]" metricMagnitude="-3" metricUnit="m" metricValue="7.6499999999999995" metricValueMax="8.2" metricValueMin="7.1" pageId="8" pageNumber="501" unit="mm" value="7.6499999999999995" valueMax="8.2" valueMin="7.1">7.1 to 8.2 mm</quantity>
, females = 
<quantity id="53409B0FEB60ED4DFE00FA6E615B83A2" box="[485,636,1473,1497]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.2" metricValueMin="6.8" pageId="8" pageNumber="501" unit="mm" value="7.5" valueMax="8.2" valueMin="6.8">6.8 to 8.2 mm</quantity>
; pronotum (
<figureCitation id="0C832A6FEB60ED4DFD18FA6E607A83A2" box="[765,861,1473,1497]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="8" pageNumber="501">Fig. 14A</figureCitation>
) with anterior angles narrow, distinctly acute, lateral margins slightly to distinctly sinuate near anterior angles, disk dark piceous to black, with lateral pale bands well-defined and very narrow; free apex of prosternal intercoxal process long (as in 
<figureCitation id="0C832A6FEB60ED4DFEB5F98C62948040" box="[336,435,1571,1595]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="8" pageNumber="501">Fig. 15C</figureCitation>
); elytra with preapical dark spot present or absent; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; median lobe of male genitalia with shaft evenly arcuate and tapered subapically, with apical lamella thick, slightly deflected ventrally in lateral aspect (
<figureCitation id="0C832A6FEB60ED4DFD5CF92B603A80E7" box="[697,797,1668,1692]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="8" pageNumber="501">Fig. 16A</figureCitation>
), shaft slightly broadened for most of its length, then tapered to base of short, narrow, smoothly rounded apical lamella in dorsal aspect (
<figureCitation id="0C832A6FEB61ED4CFF66FB3063C182CC" box="[131,230,1183,1207]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="9" pageNumber="502">Fig. 17A</figureCitation>
).
</paragraph>
<caption id="C0C76662EB61ED4CFF4EFB9962E38207" ID-DOI="http://doi.org/10.5281/zenodo.11512962" ID-Zenodo-Dep="11512962" httpUri="https://zenodo.org/record/11512962/files/figure.png" pageId="9" pageNumber="502" startId="9.[171,183,1078,1097]" targetBox="[120,1176,236,1061]" targetPageId="9" targetType="figure">
<paragraph id="940736EAEB61ED4CFF4EFB9962E38207" blockId="9.[123,1173,1077,1148]" pageId="9" pageNumber="502">
<smallCapsWord id="92E1A036EB61ED4CFF4EFB9963CF8233" baselines="1092,1092" box="[171,232,1078,1097]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="9" pageNumber="502">FIGURE</smallCapsWord>
4. 
<taxonomicName id="53B84D69EB61ED4CFEE0FB9A60278233" authority="Kavanaugh and Cueva-Dabkoski" authorityName="Kavanaugh and Cueva-Dabkoski" box="[261,768,1077,1096]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB61ED4CFEE0FB9A62FD8233" box="[261,474,1077,1096]" italics="true" pageId="9" pageNumber="502">Cyclosomus acutangulus</emphasis>
Kavanaugh and Cueva-Dabkoski
</taxonomicName>
, dorsal habitus. A. Holotype male (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. Female (Bhalukpong, Arunachal Pradesh, India). Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="940736EAEB61ED4CFF4EFB1063CD830C" blockId="9.[123,1174,1183,1719]" pageId="9" pageNumber="502">
Members of 
<taxonomicName id="53B84D69EB61ED4CFED1FB6F62F182AC" box="[308,470,1215,1239]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB61ED4CFED1FB6F62F182AC" box="[308,470,1215,1239]" italics="true" pageId="9" pageNumber="502">C. acutangulus</emphasis>
</taxonomicName>
are most similar to those of 
<taxonomicName id="53B84D69EB61ED4CFCEEFB6F60AF82AC" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[779,904,1215,1239]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="502" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A6CCEAF8EB61ED4CFCEEFB6F60AF82AC" box="[779,904,1215,1239]" italics="true" pageId="9" pageNumber="502">C. suturalis</emphasis>
</taxonomicName>
but differ from the latter in having the pronotum with anterior angles narrow and markedly acute, lateral margins slightly to distinctly sinuate near the anterior angles, and the lateral pale bands well-defined and very narrow. Males of the two species also differ in the curvature of the median lobe in lateral aspect (compare 
<figureCitation id="0C832A6FEB61ED4CFF9EFA9063FF832C" box="[123,216,1343,1367]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="9" pageNumber="502">Fig. 16A</figureCitation>
with 
<figureCitation id="0C832A6FEB61ED4CFEF1FA906253832C" box="[276,372,1343,1367]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="9" pageNumber="502">Fig. 16G</figureCitation>
) and in shape of the apical lamella in dorsal aspect (compare 
<figureCitation id="0C832A6FEB61ED4CFBE7FA906778832C" box="[1026,1119,1343,1367]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="9" pageNumber="502">Fig. 17A</figureCitation>
with 
<figureCitation id="0C832A6FEB61ED4CFF9EFAF063F9830C" box="[123,222,1375,1399]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="9" pageNumber="502">Fig. 17G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB61ED4CFF4EFA2F6226806C" pageId="9" pageNumber="502" type="biology_ecology">
<paragraph id="940736EAEB61ED4CFF4EFA2F6226806C" blockId="9.[123,1174,1183,1719]" pageId="9" pageNumber="502">
<emphasis id="A6CCEAF8EB61ED4CFF4EFA2F62B083EC" bold="true" box="[171,407,1408,1431]" pageId="9" pageNumber="502">Habitat distribution.</emphasis>
In 
<collectingRegion id="567CF808EB61ED4CFE5FFA2F612983EC" box="[442,526,1408,1431]" country="China" name="Yunnan" pageId="9" pageNumber="502">Yunnan</collectingRegion>
, specimens of this species were collected only at night, with the aid of headlamps. They were found active on the surface of dry or only slightly damp sandy substrate, with no or only widely scattered and low vegetation, on the upper edges of open sandy beaches along the Longchuan River (
<figureCitation id="0C832A6FEB61ED4CFDF4FA706154838C" box="[529,627,1503,1527]" captionStart="FIGURE" captionStartId="37.[171,183,1577,1596]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@37.[120,1176,216,1561]" captionTargetPageId="37" captionText="FIGURE 18. Photographs of habitats for Cyclosomus species. A. C. acutangulus Kavanaugh &amp; Cueva-Dabkoski (Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China; photograph by David H. Kavanaugh); B. C. suturalis (Wiedemann) (Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China; photograph by Hong-Bin Liang)." figureDoi="http://doi.org/10.5281/zenodo.11512996" httpUri="https://zenodo.org/record/11512996/files/figure.png" pageId="9" pageNumber="502">Fig. 18A</figureCitation>
) and Nanwanhe at elevations of 1215 and 
<quantity id="53409B0FEB61ED4CFBACFA7067B6838C" box="[1097,1169,1503,1527]" metricMagnitude="2" metricUnit="m" metricValue="9.309999999999999" pageId="9" pageNumber="502" unit="m" value="931.0">931 m</quantity>
, respectively.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB61ED4FFF4EF98F61A5870F" lastPageId="10" lastPageNumber="503" pageId="9" pageNumber="502" type="materials_examined">
<paragraph id="940736EAEB61ED4FFF4EF98F61A5870F" blockId="9.[123,1174,1183,1719]" lastBlockId="10.[123,1174,219,890]" lastPageId="10" lastPageNumber="503" pageId="9" pageNumber="502">
<emphasis id="A6CCEAF8EB61ED4CFF4EF98F62FD804C" bold="true" box="[171,474,1568,1591]" pageId="9" pageNumber="502">Geographical distribution.</emphasis>
<location id="91676031EB61ED4CFE05F98F6124804C" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501:91676031EB61ED4CFE05F98F6124804C" box="[480,515,1568,1591]" name="We" pageId="9" pageNumber="502">We</location>
examined a total of 
<specimenCount id="82BEFD63EB61ED4CFD04F9B06055804C" box="[737,882,1567,1591]" count="30" pageId="9" pageNumber="502" type="generic">30 specimens</specimenCount>
(
<specimenCount id="82BEFD63EB61ED4CFC64F9B060C5804C" box="[897,994,1567,1591]" count="14" pageId="9" pageNumber="502" type="male">14 males</specimenCount>
and 
<specimenCount id="82BEFD63EB61ED4CFBFDF9B067A9804C" box="[1048,1166,1567,1591]" count="16" pageId="9" pageNumber="502" type="female">16 females</specimenCount>
) from the following localities: 
<materialsCitation id="24D03CB7EB61ED4CFE32F9EF67B6800C" ID-GBIF-Occurrence="4905152330" collectingDate="2015-03-25" elevation="931.0" pageId="9" pageNumber="502">
<emphasis id="A6CCEAF8EB61ED4CFE32F9EF6117802C" bold="true" box="[471,560,1600,1623]" pageId="9" pageNumber="502">
<collectingCountry id="ECAF767AEB61ED4CFE32F9EF6117802C" box="[471,560,1600,1623]" name="China" pageId="9" pageNumber="502">CHINA</collectingCountry>
</emphasis>
: 
<collectingRegion id="567CF808EB61ED4CFDA1F9EF61BD802C" box="[580,666,1600,1623]" country="China" name="Yunnan" pageId="9" pageNumber="502">Yunnan</collectingRegion>
: 
<collectingCounty id="7D664E66EB61ED4CFD4EF99060A6802C" box="[683,897,1599,1623]" pageId="9" pageNumber="502">Longchuan County</collectingCounty>
, 
<collectingMunicipality id="7463AC90EB61ED4CFC77F9906721802C" box="[914,1030,1599,1623]" pageId="9" pageNumber="502">Zhangfeng</collectingMunicipality>
(
<location id="91676031EB61ED4CFBFEF99067B6802C" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501:91676031EB61ED4CFBFEF99067B6802C" box="[1051,1169,1599,1623]" country="China" county="Longchuan County" municipality="Zhangfeng" name="Nanwanhe" pageId="9" pageNumber="502" stateProvince="Yunnan">Nanwanhe</location>
, 24.1959°/97.7844°, 
<quantity id="53409B0FEB61ED4CFEB0F9F062B9800C" box="[341,414,1631,1655]" metricMagnitude="2" metricUnit="m" metricValue="9.309999999999999" pageId="9" pageNumber="502" unit="m" value="931.0">
<elevation id="1F95D1D9EB61ED4CFEB0F9F062B9800C" box="[341,414,1631,1655]" metricMagnitude="2" metricUnit="m" metricValue="9.309999999999999" pageId="9" pageNumber="502" unit="m" value="931.0">931 m</elevation>
</quantity>
, 
<date id="E006102AEB61ED4CFE49F9F0617F800C" box="[428,600,1631,1655]" pageId="9" pageNumber="502" value="2015-03-25">
<collectingDate id="F042E9C2EB61ED4CFE49F9F0617F800C" box="[428,600,1631,1655]" pageId="9" pageNumber="502" value="2015-03-25">25 March 2015</collectingDate>
</date>
, 
<collectorName id="394D533CEB61ED4CFD80F9CF618D800C" box="[613,682,1631,1655]" pageId="9" pageNumber="502">Y. Liu</collectorName>
and 
<collectorName id="394D533CEB61ED4CFD01F9CF6065800C" box="[740,834,1631,1655]" pageId="9" pageNumber="502">H.L. Shi</collectorName>
collectors [
<specimenCount id="82BEFD63EB61ED4CFC26F9CF6765800C" box="[963,1090,1631,1655]" count="1" pageId="9" pageNumber="502" type="female">one female</specimenCount>
; 
<collectionCode id="F2A9AE2FEB61ED4CFBB5F9F067A4800C" box="[1104,1155,1631,1655]" pageId="9" pageNumber="502">IOZ</collectionCode>
])
</materialsCitation>
; 
<materialsCitation id="24D03CB7EB61ED4CFF9EF9D067B280CC" ID-GBIF-Occurrence="4905152326" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS, IOZ" collectorName="H. B. Liang &amp; X. C. Shi " country="China" county="Tengchong County" elevation="1215" latitude="24.8941" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="7" longitude="98.675" municipality="Wuhe Township" pageId="9" pageNumber="502" specimenCount="27" specimenCount-female="14" specimenCount-male="13" stateProvince="Yunnan">
<collectingCounty id="7D664E66EB61ED4CFF9EF9D0626080EC" box="[123,327,1663,1687]" pageId="9" pageNumber="502">Tengchong County</collectingCounty>
, 
<collectingMunicipality id="7463AC90EB61ED4CFEB5F9D062DA80EC" box="[336,509,1663,1687]" pageId="9" pageNumber="502">Wuhe Township</collectingMunicipality>
(
<location id="91676031EB61ED4CFDEFF9D0608780EC" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501:91676031EB61ED4CFDEFF9D0608780EC" box="[522,928,1663,1687]" country="China" county="Tengchong County" latitude="24.8941" longLatPrecision="7" longitude="98.675" municipality="Wuhe Township" name="Longjiang Bridge on Longchuanjiang" pageId="9" pageNumber="502" stateProvince="Yunnan">Longjiang Bridge on Longchuanjiang</location>
, 24.89889°/98.66667°, 
<quantity id="53409B0FEB61ED4CFF9EF93063F680CC" box="[123,209,1695,1719]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="9" pageNumber="502" unit="m" value="1215.0">
<elevation id="1F95D1D9EB61ED4CFF9EF93063F680CC" box="[123,209,1695,1719]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="9" pageNumber="502" unit="m" value="1215.0">1215 m</elevation>
</quantity>
, 
<date id="E006102AEB61ED4CFF3AF93062BB80CC" box="[223,412,1695,1719]" pageId="9" pageNumber="502" value="2003-10-30">
<collectingDate id="F042E9C2EB61ED4CFF3AF93062BB80CC" box="[223,412,1695,1719]" pageId="9" pageNumber="502" value="2003-10-30">30 October 2003</collectingDate>
</date>
, 
<collectorName id="394D533CEB61ED4CFE4CF90F610380CC" box="[425,548,1695,1719]" pageId="9" pageNumber="502">H.B. Liang</collectorName>
and 
<collectorName id="394D533CEB61ED4CFDBBF930619980CC" box="[606,702,1695,1719]" pageId="9" pageNumber="502">X.C. Shi</collectorName>
X collectors [
<specimenCount id="82BEFD63EB61ED4CFCB8F93060E080CC" box="[861,967,1695,1719]" count="6" pageId="9" pageNumber="502" type="male">six males</specimenCount>
and 
<specimenCount id="82BEFD63EB61ED4CFBE4F93067A380CC" box="[1025,1156,1695,1719]" count="6" pageId="9" pageNumber="502" type="female">six females</specimenCount>
]),
</materialsCitation>
<materialsCitation id="24D03CB7EB62ED4FFF9EFF7461908768" ID-GBIF-Occurrence="4905152307" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS, IOZ" collectorName="Y. Liu &amp; H. L. Shi " country="China" county="Tengchong County" elevation="1215" latitude="24.8941" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="7" longitude="98.675" municipality="Wuhe Township" pageId="10" pageNumber="503" specimenCount="27" specimenCount-female="14" specimenCount-male="13" stateProvince="Yunnan">
(beach of 
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, 
<geoCoordinate id="F18C502DEB62ED4FFE4AFF7461208688" box="[431,519,219,243]" degrees="24.8941" direction="north" orientation="latitude" pageId="10" pageNumber="503" precision="5" value="24.8941">24.8941</geoCoordinate>
/ 
<geoCoordinate id="F18C502DEB62ED4FFDEAFF74614E8688" box="[527,617,219,243]" degrees="98.6750" direction="east" orientation="longitude" pageId="10" pageNumber="503" precision="5" value="98.675">98.6750</geoCoordinate>
°, 
<quantity id="53409B0FEB62ED4FFD98FF7461F68689" box="[637,721,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="10" pageNumber="503" unit="m" value="1215.0">
<elevation id="1F95D1D9EB62ED4FFD98FF7461F68689" box="[637,721,219,243]" metricMagnitude="3" metricUnit="m" metricValue="1.215" pageId="10" pageNumber="503" unit="m" value="1215.0">1215 m</elevation>
</quantity>
, 
<date id="E006102AEB62ED4FFD38FF7460A28688" box="[733,901,219,243]" pageId="10" pageNumber="503" value="2015-03-22">
<collectingDate id="F042E9C2EB62ED4FFD38FF7460A28688" box="[733,901,219,243]" pageId="10" pageNumber="503" value="2015-03-22">22 March 2015</collectingDate>
</date>
, 
<collectorName id="394D533CEB62ED4FFC6AFF7460F58688" box="[911,978,219,243]" pageId="10" pageNumber="503">Y. Liu</collectorName>
and 
<collectorName id="394D533CEB62ED4FFBEDFF7467428688" box="[1032,1125,219,243]" pageId="10" pageNumber="503">H.L. Shi</collectorName>
collectors [
<specimenCount id="82BEFD63EB62ED4FFF2AFF5462138768" box="[207,308,251,275]" count="6" pageId="10" pageNumber="503" type="male">six males</specimenCount>
and 
<specimenCount id="82BEFD63EB62ED4FFE82FF5361278768" box="[359,512,251,275]" count="7" pageId="10" pageNumber="503" type="female">seven females</specimenCount>
; 
<collectionCode id="F2A9AE2FEB62ED4FFDEEFF5461188768" box="[523,575,251,275]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="10" pageNumber="503" type="Museum">CAS</collectionCode>
and 
<collectionCode id="F2A9AE2FEB62ED4FFD97FF5461838768" box="[626,676,251,275]" pageId="10" pageNumber="503">IOZ</collectionCode>
]),
</materialsCitation>
<materialsCitation id="24D03CB7EB62ED4FFD58FF5467138748" ID-GBIF-Occurrence="4905152318" collectingDate="2003-10-30" collectingDateMax="2015-03-22" collectingDateMin="2003-10-30" collectionCode="CAS, IOZ" collectorName=" D. H. Kavanaugh &amp; R. L. Brett" country="China" county="Tengchong County" elevation="1215" latitude="24.8941" location="Longjiang Bridge on Longchuanjiang" longLatPrecision="7" longitude="98.675" municipality="Wuhe Township" pageId="10" pageNumber="503" specimenCount="27" specimenCount-female="14" specimenCount-male="13" stateProvince="Yunnan">
(
<collectorName id="394D533CEB62ED4FFD21FF5460438768" box="[708,868,251,275]" pageId="10" pageNumber="503">Longjiangqiao</collectorName>
, 24.89176°/98.67551°, 
<quantity id="53409B0FEB62ED4FFB85FF5463B58748" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="503" unit="m" value="1230.0">
<elevation id="1F95D1D9EB62ED4FFB85FF5463B58748" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="10" pageNumber="503" unit="m" value="1230.0">1230 m</elevation>
</quantity>
, 
<date id="E006102AEB62ED4FFF7BFEB462028748" box="[158,293,283,307]" pageId="10" pageNumber="503" value="2006-06-03">
<collectingDate id="F042E9C2EB62ED4FFF7BFEB462028748" box="[158,293,283,307]" pageId="10" pageNumber="503" value="2006-06-03">3 June 2006</collectingDate>
</date>
, 
<collectorName id="394D533CEB62ED4FFED7FEB362C08748" box="[306,487,283,307]" pageId="10" pageNumber="503">D.H. Kavanaugh</collectorName>
and 
<collectorName id="394D533CEB62ED4FFDFAFEB361AC8748" box="[543,651,284,307]" pageId="10" pageNumber="503">R.L. Brett</collectorName>
collectors [
<specimenCount id="82BEFD63EB62ED4FFCEEFEB360578748" box="[779,880,283,307]" count="1" pageId="10" pageNumber="503" type="male">one male</specimenCount>
and 
<specimenCount id="82BEFD63EB62ED4FFC42FEB367018748" box="[935,1062,283,307]" count="1" pageId="10" pageNumber="503" type="female">one female</specimenCount>
])
</materialsCitation>
. 
<materialsCitation id="24D03CB7EB62ED4FFBDAFEB36159870F" ID-GBIF-Occurrence="4905152337" collectingDate="2006-05-26" collectingDateMax="2006-06-03" collectingDateMin="2006-05-26" collectionCode="NHMUK" collectorName="P. Pacholatko" country="India" elevation="150" location="Bhalukpong" pageId="10" pageNumber="503" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arunachal Pradesh">
<emphasis id="A6CCEAF8EB62ED4FFBDAFEB367A98748" bold="true" box="[1087,1166,284,307]" pageId="10" pageNumber="503">
<collectingCountry id="ECAF767AEB62ED4FFBDAFEB367A98748" box="[1087,1166,284,307]" name="India" pageId="10" pageNumber="503">INDIA</collectingCountry>
</emphasis>
: 
<collectingRegion id="567CF808EB62ED4FFF9EFE93626C872F" box="[123,331,316,340]" country="India" name="Arunachal Pradesh" pageId="10" pageNumber="503">Arunachal Pradesh</collectingRegion>
: 
<location id="91676031EB62ED4FFEB3FE9362FF872F" LSID="urn:lsid:plazi:treatment:1C1187FCEB60ED4FFF9EFB7662B28501:91676031EB62ED4FFEB3FE9362FF872F" box="[342,472,316,340]" country="India" name="Bhalukpong" pageId="10" pageNumber="503" stateProvince="Arunachal Pradesh">Bhalukpong</location>
(27.0333°/92.5833°, 
<quantity id="53409B0FEB62ED4FFD27FE93602E872F" box="[706,777,316,340]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="503" unit="m" value="150.0">
<elevation id="1F95D1D9EB62ED4FFD27FE93602E872F" box="[706,777,316,340]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="503" unit="m" value="150.0">150 m</elevation>
</quantity>
, 
<date id="E006102AEB62ED4FFCF0FE9360DF872F" box="[789,1016,316,340]" pageId="10" pageNumber="503" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">
<collectingDate id="F042E9C2EB62ED4FFCF0FE9360DF872F" box="[789,1016,316,340]" pageId="10" pageNumber="503" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">26 May-3 June 2006</collectingDate>
</date>
, 
<collectorName id="394D533CEB62ED4FFBE1FE9267B1872F" box="[1028,1174,316,340]" pageId="10" pageNumber="503">P. Pacholátko</collectorName>
collector [
<specimenCount id="82BEFD63EB62ED4FFF03FEF2626D870F" box="[230,330,348,372]" count="1" pageId="10" pageNumber="503" type="male">one male</specimenCount>
and 
<specimenCount id="82BEFD63EB62ED4FFE9AFEF262DE870F" box="[383,505,348,372]" count="1" pageId="10" pageNumber="503" type="female">one female</specimenCount>
; 
<collectionCode id="F2A9AE2FEB62ED4FFDE1FEF36157870F" box="[516,624,348,372]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="503" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB62ED4FFF4EFED261DE844D" pageId="10" pageNumber="503" type="distribution">
<paragraph id="940736EAEB62ED4FFF4EFED261DE844D" blockId="10.[123,1174,219,890]" pageId="10" pageNumber="503">
At present, this species is known only from one locality in northeastern 
<collectingCountry id="ECAF767AEB62ED4FFC50FED260CB87EE" box="[949,1004,381,405]" name="India" pageId="10" pageNumber="503">India</collectingCountry>
and two localities in western 
<collectingRegion id="567CF808EB62ED4FFEFEFE3162F287CE" box="[283,469,413,437]" country="China" name="Yunnan" pageId="10" pageNumber="503">Yunnan Province</collectingRegion>
, 
<collectingCountry id="ECAF767AEB62ED4FFE3AFE32613887CE" box="[479,543,413,437]" name="China" pageId="10" pageNumber="503">China</collectingCountry>
(
<figureCitation id="0C832A6FEB62ED4FFDC8FE32615D87CE" box="[557,634,413,437]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="10" pageNumber="503">Fig. 20</figureCitation>
<emphasis id="A6CCEAF8EB62ED4FFD9FFE3161A487CE" bold="true" box="[634,643,414,437]" pageId="10" pageNumber="503">)</emphasis>
. The localities in 
<collectingCountry id="ECAF767AEB62ED4FFCA7FE32605E87CE" box="[834,889,413,437]" name="India" pageId="10" pageNumber="503">India</collectingCountry>
and Yunnan are about 
<quantity id="53409B0FEB62ED4FFB8BFE3263BA87AE" metricMagnitude="5" metricUnit="m" metricValue="6.5" pageId="10" pageNumber="503" unit="km" value="650.0">650 km</quantity>
apart, and it is likely that this species occurs also in suitable habitats in the intervening region, including northern 
<collectingCountry id="ECAF767AEB62ED4FFEA3FE716289878D" box="[326,430,478,502]" name="Myanmar" pageId="10" pageNumber="503">Myanmar</collectingCountry>
(
<collectingRegion id="567CF808EB62ED4FFE5EFE71616E878D" box="[443,585,478,502]" country="Myanmar" name="Kachin" pageId="10" pageNumber="503">Kachin State</collectingRegion>
), and northeastern 
<collectingCountry id="ECAF767AEB62ED4FFCF7FE71606E878D" box="[786,841,478,502]" name="India" pageId="10" pageNumber="503">India</collectingCountry>
(
<collectingRegion id="567CF808EB62ED4FFCB3FE716706878D" box="[854,1057,478,502]" country="India" name="Arunachal Pradesh" pageId="10" pageNumber="503">Arunachal Pradesh</collectingRegion>
and 
<collectingRegion id="567CF808EB62ED4FFBB1FE71638E846D" country="India" name="Nagaland" pageId="10" pageNumber="503">Nagaland</collectingRegion>
states), at low elevations (below 
<quantity id="53409B0FEB62ED4FFDC0FE51615B846D" box="[549,636,510,534]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="503" unit="m" value="1500.0">1500 m</quantity>
) along rivers draining the western and southern slopes of the Himalayan ranges in these areas, respectively.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB62ED4FFF4EFDEF62B28501" pageId="10" pageNumber="503" type="description">
<paragraph id="940736EAEB62ED4FFF4EFDEF675B840C" blockId="10.[123,1174,219,890]" pageId="10" pageNumber="503">
<emphasis id="A6CCEAF8EB62ED4FFF4EFDEF62E7842C" bold="true" box="[171,448,576,599]" pageId="10" pageNumber="503">Geographical variation.</emphasis>
The specimens from the locality in northeastern 
<collectingCountry id="ECAF767AEB62ED4FFC1CFD906717842C" box="[1017,1072,575,599]" name="India" pageId="10" pageNumber="503">India</collectingCountry>
lack the preapical dark spot found in specimens from the localities in Yunnan, but otherwise are similar.
</paragraph>
<paragraph id="940736EAEB62ED4FFF4EFD2F62B28501" blockId="10.[123,1174,219,890]" pageId="10" pageNumber="503">
<emphasis id="A6CCEAF8EB62ED4FFF4EFD2F607284EC" bold="true" box="[171,853,640,664]" pageId="10" pageNumber="503">
Geographical relationships with other 
<taxonomicName id="53B84D69EB62ED4FFD94FD2F607284EC" box="[625,853,640,664]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A6CCEAF8EB62ED4FFD94FD2F61D284E3" bold="true" box="[625,757,640,664]" italics="true" pageId="10" pageNumber="503">Cyclosomus</emphasis>
species.
</taxonomicName>
</emphasis>
Members of 
<taxonomicName id="53B84D69EB62ED4FFC15FD2F67B284E3" box="[1008,1173,640,664]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB62ED4FFC15FD2F67B284E3" box="[1008,1173,640,664]" italics="true" pageId="10" pageNumber="503">C. acutangulus</emphasis>
</taxonomicName>
have been found syntopic with those of 
<taxonomicName id="53B84D69EB62ED4FFDDAFD0E61E284C3" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[575,709,672,696]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFDDAFD0E61E284C3" box="[575,709,672,696]" italics="true" pageId="10" pageNumber="503">C. flexuosus</emphasis>
</taxonomicName>
at Bhalukpong, 
<collectingRegion id="567CF808EB62ED4FFC9AFD0F677784C3" box="[895,1104,672,696]" country="India" name="Arunachal Pradesh" pageId="10" pageNumber="503">Arunachal Pradesh</collectingRegion>
, 
<collectingCountry id="ECAF767AEB62ED4FFBBAFD0F67B184C3" box="[1119,1174,672,696]" name="India" pageId="10" pageNumber="503">India</collectingCountry>
[
<collectionCode id="F2A9AE2FEB62ED4FFF64FD6E63CC84A3" box="[129,235,705,728]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="503" type="Museum">NHMUK</collectionCode>
] but nowhere with any other species. The geographical ranges of 
<taxonomicName id="53B84D69EB62ED4FFC27FD6E674284A3" box="[962,1125,704,728]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB62ED4FFC27FD6E674284A3" box="[962,1125,704,728]" italics="true" pageId="10" pageNumber="503">C. acutangulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="53B84D69EB62ED4FFF9EFD4E63D08482" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[123,247,737,761]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A6CCEAF8EB62ED4FFF9EFD4E63D08482" box="[123,247,737,761]" italics="true" pageId="10" pageNumber="503">C. suturalis</emphasis>
</taxonomicName>
may overlap slightly in westernmost Yunnan. In fact, both species have been found on sandy beaches along the Longchuan River (Longchuanjiang) but at different localities more than 
<quantity id="53409B0FEB62ED4FFF9EFC8E63EB8542" box="[123,204,801,825]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="503" unit="km" value="150.0">150 km</quantity>
apart along the river course and at different elevations (the former above 
<quantity id="53409B0FEB62ED4FFC08FC8E67188542" box="[1005,1087,801,825]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="503" unit="m" value="1200.0">1200 m</quantity>
and the latter at 
<quantity id="53409B0FEB62ED4FFF30FCED623B8521" box="[213,284,834,858]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="10" pageNumber="503" unit="m" value="734.0">734 m</quantity>
). Otherwise, the range of 
<taxonomicName id="53B84D69EB62ED4FFDD3FCEC61FE8521" box="[566,729,834,858]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB62ED4FFDD3FCEC61FE8521" box="[566,729,834,858]" italics="true" pageId="10" pageNumber="503">C. acutangulus</emphasis>
</taxonomicName>
is not known to overlap with that of any other 
<taxonomicName id="53B84D69EB62ED4FFF5CFCCD62B28501" box="[185,405,866,890]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A6CCEAF8EB62ED4FFF5CFCCD621D8501" box="[185,314,866,890]" italics="true" pageId="10" pageNumber="503">Cyclosomus</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>