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<document id="75AA5546FC01F969A6291348C14979E3" ID-CLB-Dataset="55892" ID-DOI="10.37520/aemnp.2021.002" ID-GBIF-Dataset="f5f6ab5f-d650-4e1f-b6e3-c21928723245" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037464" ID-ZooBank="0AEECD1F-1C71-4636-89A7-DA31D2DB73D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624896309300" checkinUser="carolina" docAuthor="Yin, Zi-Wei &amp; Li, Ning" docDate="2021" docId="03D13B6DE807513B7F41FEAA3D41F8F9" docLanguage="en" docName="ActaEntMusNatPra.61.1.35-53.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.002" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Pselaphodes banphabat Yin &amp; Li 2021, sp. nov." docType="treatment" docVersion="13" lastPageNumber="42" masterDocId="FFE84315E800513C7E56FFD43F41FFE7" masterDocTitle="Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="53" masterPageNumber="35" pageNumber="42" updateTime="1699021163717" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="6DA4D507108481000C7E1F57A411D727">Eight new species and additional records of the Pselaphodes complex from Laos and Vietnam, with a key to known species (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<mods:namePart id="AEC3A27DA2D2FE4558B8CD5FAFBA6B58">Yin, Zi-Wei</mods:namePart>
<mods:affiliation id="8AA6CD3616563FF64E8FCB6DE80025E5">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China</mods:affiliation>
<mods:nameIdentifier id="0EBB3455EFFE06B26236F7388A399195" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:namePart id="D78CC55A0D54BFE76CC2230D8F1C3BA8">Li, Ning</mods:namePart>
<mods:affiliation id="978EE85BE98F60CF86AD68A0F1E08541">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China.</mods:affiliation>
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<mods:date id="8743FBF544656D74B5C7209630720EFE">2021</mods:date>
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<treatment id="03D13B6DE807513B7F41FEAA3D41F8F9" ID-DOI="http://doi.org/10.5281/zenodo.6474079" ID-GBIF-Taxon="194685196" ID-Zenodo-Dep="6474079" LSID="urn:lsid:plazi:treatment:03D13B6DE807513B7F41FEAA3D41F8F9" httpUri="http://treatment.plazi.org/id/03D13B6DE807513B7F41FEAA3D41F8F9" lastPageNumber="42" pageId="7" pageNumber="42">
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<heading id="D08F3D17E807513B7F41FEAA3D39FE72" box="[279,632,382,405]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="42" reason="3">
<emphasis id="B90C5669E807513B7F41FEAA3D39FE72" bold="true" box="[279,632,382,405]" pageId="7" pageNumber="42">
<taxonomicName id="4C78F1F8E807513B7F41FEAA3D5AFE72" ID-CoL="8Y8PS" authority="Yin &amp; Li, 2021" authorityName="Yin &amp; Li" authorityYear="2021" box="[279,539,382,405]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="banphabat" status="sp. nov.">
<emphasis id="B90C5669E807513B7F41FEAA3D5AFE72" bold="true" box="[279,539,382,405]" italics="true" pageId="7" pageNumber="42">Pselaphodes banphabat</emphasis>
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<taxonomicNameLabel id="A23FEB12E807513B7C74FEAA3D39FE72" box="[546,632,382,405]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C362D9F0E807513B7FC2FE483EBAFE48" box="[404,507,412,431]" pageId="7" pageNumber="42" type="description">
<paragraph id="8BC78A7BE807513B7FC2FE483EBAFE48" blockId="7.[404,507,412,431]" box="[404,507,412,431]" pageId="7" pageNumber="42">
(
<figureCitation id="134396FEE807513B7FCFFE483EA0FE48" box="[409,481,412,431]" captionStart="Fig" captionStartId="5.[151,182,2063,2082]" captionTargetBox="[156,1431,843,2025]" captionTargetId="figure-377@5.[152,1436,835,2033]" captionTargetPageId="5" captionText="Fig. 4. Dorsal habitus of Pselaphodes males. A P. banphabat sp. nov.; B P. dalatensis sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037476" httpUri="https://zenodo.org/record/5037476/files/figure.png" pageId="7" pageNumber="42">Figs 4A</figureCitation>
,
<figureCitation id="134396FEE807513B7FBFFE483EB5FE48" box="[489,500,412,431]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">5</figureCitation>
)
</paragraph>
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<subSubSection id="C362D9F0E807513B7EC1FE123D6DFDE9" pageId="7" pageNumber="42" type="description">
<paragraph id="8BC78A7BE807513B7EC1FE123D6DFDE9" blockId="7.[151,760,451,526]" pageId="7" pageNumber="42">
<materialsCitation id="3B108026E807513B7EC1FE123D69FDE9" ID-GBIF-Occurrence="3898371301" collectingDate="1997-04-27" collectingDateMax="1997-05-01" collectingDateMin="1997-04-27" collectionCode="MHNG" collectorName="Sausa &amp; Jendek'" country="Laos" elevation="150" latitude="18.266666" location="BOLIKHAMSAI" longLatPrecision="1275" longitude="103.183334" pageId="7" pageNumber="42" specimenCount="1" specimenCount-male="1" stateProvince="BOLIKHAMSAI" typeStatus="holotype">
<emphasis id="B90C5669E807513B7EC1FE123E5AFE3E" bold="true" box="[151,283,454,473]" pageId="7" pageNumber="42">Type material</emphasis>
(1 specimen).
<typeStatus id="54C334D9E807513B7FF2FE123D42FE3E" box="[420,515,454,473]" pageId="7" pageNumber="42" type="holotype">HOLOTYPE</typeStatus>
:
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<collectingCountry id="F36FCAEBE807513B7C5BFE123D0AFE3E" box="[525,587,454,473]" name="Laos" pageId="7" pageNumber="42" value="Laos">LAOS</collectingCountry>
:
<collectingRegion id="49BC4499E807513B7C0EFE123D99FE3E" box="[600,728,454,473]" country="Laos" name="Bolikhamxai" pageId="7" pageNumber="42">BOLIKHAMSAI</collectingRegion>
:
</emphasis>
<specimenCount id="9D7E41F2E807513B7CB5FE153DB2FE3D" box="[739,755,449,474]" pageId="7" pageNumber="42" type="male"></specimenCount>
, LAOS:
<collectingDate id="EF825553E807513B7EB0FE353E10FE13" box="[230,337,481,500]" pageId="7" pageNumber="42" value="1997-04-27" valueMax="1997-05-01" valueMin="1997-04-27">
<date id="FFC6ACBBE807513B7EB0FE353E10FE13" box="[230,337,481,500]" pageId="7" pageNumber="42" value="1997-04-27" valueMax="1997-05-01" valueMin="1997-04-27">27.iv-1.v.97</date>
</collectingDate>
,
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<quantity id="4C80279EE807513B7F0BFE353ED9FE13" box="[349,408,481,500]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="42" unit="km" value="70.0">70 km</quantity>
NE Vientiane BAN PHABAT
</locationDeviation>
,
<quantity id="4C80279EE807513B7CEEFE353DB5FE13" box="[696,756,481,500]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="42" unit="m" value="150.0">
<elevation id="00556D48E807513B7CEEFE353DB5FE13" box="[696,756,481,500]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="42" unit="m" value="150.0">150 m</elevation>
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,
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,
<geoCoordinate id="EE4CECBCE807513B7EBFFE2F3E7BFDE9" box="[233,314,507,526]" degrees="103" direction="east" minutes="11" orientation="longitude" pageId="7" pageNumber="42" precision="925" value="103.183334">E103°11</geoCoordinate>
,
<collectorName id="268DEFADE807513B7F14FE2F3E90FDE9" box="[322,465,507,526]" pageId="7" pageNumber="42">Sausa &amp; Jendek</collectorName>
(
<collectionCode id="ED6912BEE807513B7F8AFE2F3D63FDE9" box="[476,546,507,526]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="7" pageNumber="42" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C362D9F0E807513B7EC1FDF33EF0FCB9" pageId="7" pageNumber="42" type="diagnosis">
<paragraph id="8BC78A7BE807513B7EC1FDF33EF0FCB9" blockId="7.[151,761,551,1822]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EC1FDF33E4CFDD9" bold="true" box="[151,269,551,574]" pageId="7" pageNumber="42">Diagnosis.</emphasis>
Male body length
<quantity id="4C80279EE807513B7FB1FDF33D0FFDD9" box="[487,590,551,574]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="7" pageNumber="42" unit="mm" value="3.05">3.05 mm</quantity>
. Antennomeres 911 enlarged and forming club, antennomeres 9 and 10 each with small tubercle at apex and base, respectively. Horn-like metaventral processes long, in lateral view narrowing apically. Protrochanter with tiny ventral spine, mesotrochanter with acute ventral spine; mesofemur with small tooth at middle. Median lobe of aedeagus slightly asymmetrical dorso-ventrally; parameres elongate; endophallus composed of spine-like structures and a single sclerite. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C362D9F0E807513B7EC1FCB33EC7F9B9" pageId="7" pageNumber="42" type="description">
<paragraph id="8BC78A7BE807513B7EC1FCB33DB9F9D9" blockId="7.[151,761,551,1822]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EC1FCB33E2DFC99" bold="true" box="[151,364,871,894]" pageId="7" pageNumber="42">
Description.
<emphasis id="B90C5669E807513B7F64FCB33E2DFC99" bold="true" box="[306,364,871,894]" italics="true" pageId="7" pageNumber="42">Male</emphasis>
</emphasis>
(
<figureCitation id="134396FEE807513B7FD6FCB33E9FFC99" box="[384,478,871,894]" captionStart="Fig" captionStartId="5.[151,182,2063,2082]" captionTargetBox="[156,1431,843,2025]" captionTargetId="figure-377@5.[152,1436,835,2033]" captionTargetPageId="5" captionText="Fig. 4. Dorsal habitus of Pselaphodes males. A P. banphabat sp. nov.; B P. dalatensis sp. nov. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5037476" httpUri="https://zenodo.org/record/5037476/files/figure.png" pageId="7" pageNumber="42">Fig. 4A</figureCitation>
). Body length
<quantity id="4C80279EE807513B7CDDFCB33DB2FC99" box="[651,755,871,894]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="7" pageNumber="42" unit="mm" value="3.05">3.05 mm</quantity>
, Head approximately as long as wide, HL
<quantity id="4C80279EE807513B7C0AFC533DFEFC79" box="[604,703,903,926]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="7" pageNumber="42" unit="mm" value="0.59">0.59 mm</quantity>
, HW
<quantity id="4C80279EE807513B7EC1FC733FBCFC59" box="[151,253,935,958]" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" pageId="7" pageNumber="42" unit="mm" value="0.57">0.57 mm</quantity>
; eyes prominent, each composed of about 44 facets. Antennomeres 911 (
<figureCitation id="134396FEE807513B7F87FC133D69FC39" box="[465,552,967,990]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5A</figureCitation>
) enlarged and forming distinct club, antennomere 9 with a round tubercle at apex, antennomere 10 with similar but larger tubercle at base. Pronotum (
<figureCitation id="134396FEE807513B7F31FBF33EFBFBD9" box="[359,442,1063,1086]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5B</figureCitation>
) as long as wide, PL and PW
<quantity id="4C80279EE807513B7EC1FB933FB8FBB9" box="[151,249,1095,1118]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="7" pageNumber="42" unit="mm" value="0.59">0.59 mm</quantity>
, roundly narrowing apically at apical 2/5. Elytra much broader than long, EL
<quantity id="4C80279EE807513B7FB9FBB33D13FB99" box="[495,594,1127,1150]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="7" pageNumber="42" unit="mm" value="0.83">0.83 mm</quantity>
, EW
<quantity id="4C80279EE807513B7CC6FBB33DB4FB99" box="[656,757,1127,1150]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="7" pageNumber="42" unit="mm" value="1.05">1.05 mm</quantity>
. Horn-like metaventral processes (
<figureCitation id="134396FEE807513B7FA8FB533D0EFB79" box="[510,591,1159,1182]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5C</figureCitation>
) long, in lateral view narrowing apically. Protrochanter (
<figureCitation id="134396FEE807513B7C6AFB733DCFFB59" box="[572,654,1191,1214]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5D</figureCitation>
) with tiny spine at ventral margin, profemur simple; mesotrochanter (
<figureCitation id="134396FEE807513B7EC9FB333FAEFB19" box="[159,239,1255,1278]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5E</figureCitation>
) with acute ventral spine, mesofemur with small tooth at middle; metatrochanter and metafemur (
<figureCitation id="134396FEE807513B7CF6FAD33DB1FAF9" box="[672,752,1287,1310]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5F</figureCitation>
) simple. Abdomen broad at base and narrowed posteriorly, AL
<quantity id="4C80279EE807513B7E97FA933E62FAB9" box="[193,291,1351,1374]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="7" pageNumber="42" unit="mm" value="0.98">0.98 mm</quantity>
, AW
<quantity id="4C80279EE807513B7F08FA933E80FAB9" box="[350,449,1351,1374]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="7" pageNumber="42" unit="mm" value="1.05">1.05 mm</quantity>
; tergite 1 (IV) longest, about 2.5 times as long as tergite V; sternite 7 (IX) (
<figureCitation id="134396FEE807513B7C23FAB33D89FA99" box="[629,712,1383,1406]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5G</figureCitation>
) hemi-membranous, semi-ellipse. Aedeagus (
<figureCitation id="134396FEE807513B7C05FA533DFCFA79" box="[595,701,1415,1438]" captionStart="Fig" captionStartId="6.[151,182,2008,2027]" captionTargetBox="[256,1336,665,1976]" captionTargetId="figure-262@6.[251,1336,665,1980]" captionTargetPageId="6" captionText="Fig. 5. Male diagnostic features of Pselaphodes banphabat sp. nov.A left antennal club, in ventral view; B pronotum; C metaventral process, lateral view; D protrochanter and profemur; E mesotrochanter and mesofemur; F metatrochanter and metafemur; G sternite 7 (IX); HJ aedeagus, in dorsal (H), lateral (I), and ventral (J) view. Scale bars: 0.3 mm in A, B, D, E, F; 0.2 mm in C, H, I, J; 0.1 mm in G." figureDoi="http://doi.org/10.5281/zenodo.5037478" httpUri="https://zenodo.org/record/5037478/files/figure.png" pageId="7" pageNumber="42">Fig. 5HJ</figureCitation>
)
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long; median lobe dorso-ventrally slightly asymmetric; parameres elongate, broadening to near apices, then greatly narrowed at apices, each paramere with one long apical seta; endophallus composed of numerous spine-like structures, one membranous part, and one forked sclerite.
</paragraph>
<paragraph id="8BC78A7BE807513B7EE3F9933EC7F9B9" blockId="7.[151,761,551,1822]" box="[181,390,1607,1630]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EE3F9933E4EF9B9" bold="true" box="[181,271,1607,1630]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EE3F9933E49F9B9" bold="true" box="[181,264,1607,1630]" italics="true" pageId="7" pageNumber="42">Female</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C362D9F0E807513B7EC1F9B33DB8F959" pageId="7" pageNumber="42" type="discussion">
<paragraph id="8BC78A7BE807513B7EC1F9B33DB8F959" blockId="7.[151,761,551,1822]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EC1F9B33E37F999" bold="true" box="[151,374,1639,1662]" pageId="7" pageNumber="42">Comparative notes.</emphasis>
The new species is quite distinctive in the form of the male antennal clubs, which leads to a ready separation of
<taxonomicName id="4C78F1F8E807513B7F34F9733EAEF959" authorityName="Yin &amp; Li" authorityYear="2021" box="[354,495,1703,1726]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="banphabat">
<emphasis id="B90C5669E807513B7F34F9733E33F959" box="[354,370,1703,1726]" italics="true" pageId="7" pageNumber="42">P</emphasis>
.
<emphasis id="B90C5669E807513B7F28F9733EAEF959" box="[382,495,1703,1726]" italics="true" pageId="7" pageNumber="42">banphabat</emphasis>
</taxonomicName>
from all other congeners.
</paragraph>
</subSubSection>
<subSubSection id="C362D9F0E807513B7EC1F9133DA2F919" pageId="7" pageNumber="42" type="etymology">
<paragraph id="8BC78A7BE807513B7EC1F9133DA2F919" blockId="7.[151,761,551,1822]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EC1F9133E5AF939" bold="true" box="[151,283,1735,1758]" pageId="7" pageNumber="42">Etymology.</emphasis>
The new species is named after the
<typeStatus id="54C334D9E807513B7C9EF9133DB6F939" box="[712,759,1735,1758]" pageId="7" pageNumber="42">type</typeStatus>
locality, Ban Phabat. The name is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C362D9F0E807513B7EC1F8D33D41F8F9" box="[151,512,1799,1822]" pageId="7" pageNumber="42" type="distribution">
<paragraph id="8BC78A7BE807513B7EC1F8D33D41F8F9" blockId="7.[151,761,551,1822]" box="[151,512,1799,1822]" pageId="7" pageNumber="42">
<emphasis id="B90C5669E807513B7EC1F8D33E6BF8F9" bold="true" box="[151,298,1799,1822]" pageId="7" pageNumber="42">Distribution.</emphasis>
Laos: Bolikhamsai.
</paragraph>
</subSubSection>
</treatment>
</document>