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<document ID-DOI="10.1590/S1679-62252010000300006" ID-ISSN="1982-0224" ID-Zenodo-Dep="4647452" approvalRequired="26" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="23" approvalRequired_for_treatments="2" checkinTime="1617111332249" checkinUser="carolina" docAuthor="Taphorn, Donald C., Armbruster, Jonathan W., López-Fernández, Hernán &amp; Bernard, Calvin R." docDate="2010" docId="03D12959FFEEFFB4259A82416879FB3B" docLanguage="en" docName="NeotropIchthyol.8.3.615-624.pdf" docOrigin="Neotropical Ichthyology 8 (3)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252010000300006&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.3:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="3" docTitle="Neblinichthys echinasus Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard 2010, new species" docType="treatment" docVersion="2" lastPageId="7" masterDocId="FFE85121FFEAFFB3263E80426D56FFC7" masterDocTitle="Description of Neblinichthys brevibracchium and N. echinasus from the upper Mazaruni River, Guyana (Siluriformes: Loricariidae), and recognition of N. roraima and N. yaravi as distinct species" masterLastPageNumber="615" masterPageNumber="615" pageId="4" pageNumber="615" updateTime="1617113013881" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of Neblinichthys brevibracchium and N. echinasus from the upper Mazaruni River, Guyana (Siluriformes: Loricariidae), and recognition of N. roraima and N. yaravi as distinct species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Taphorn, Donald C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Armbruster, Jonathan W.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>López-Fernández, Hernán</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bernard, Calvin R.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>3</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>615</mods:start>
<mods:end>615</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252010000300006&amp;lng=en&amp;tlng=en</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1590/S1679-62252010000300006</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">4647452</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03D12959FFEEFFB4259A82416879FB3B" httpUri="http://treatment.plazi.org/id/03D12959FFEEFFB4259A82416879FB3B" lastPageId="7" pageId="4">
<subSubSection box="[932,1342,515,539]" pageId="4" type="nomenclature">
<paragraph blockId="4.[932,1342,515,570]" box="[932,1342,515,539]" pageId="4">
<heading box="[932,1342,515,539]" centered="true" fontSize="10" level="2" pageId="4" reason="2">
<emphasis bold="true" box="[932,1342,515,539]" pageId="4">
<taxonomicName authority="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard, 2010" authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[932,1201,515,539]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="echinasus" status="new species">
<emphasis bold="true" box="[932,1201,515,539]" italics="true" pageId="4">Neblinichthys echinasus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1212,1342,515,538]" pageId="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1106,1168,547,570]" pageId="4" type="description">
<paragraph blockId="4.[932,1342,515,570]" box="[1106,1168,547,570]" pageId="4">
<figureCitation box="[1106,1168,547,570]" captionStart="Fig" captionStartId="6.[136,176,1712,1735]" captionTargetBox="[136,1458,257,1701]" captionTargetId="figure-174@6.[136,1459,257,1702]" captionTargetPageId="6" captionText="Fig. 3. Neblinichthys echinasus, holotype, CSBD 1652, 82.9 mm SL, Kukui River, near town of Philipai, upper Mazaruni River drainage." figureDoi="http://doi.org/10.5281/zenodo.4647458" httpUri="https://zenodo.org/record/4647458/files/figure.png" pageId="4">
<emphasis bold="true" box="[1106,1168,547,570]" pageId="4">Fig. 3</emphasis>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" type="materials_examined">
<paragraph blockId="4.[816,1458,611,718]" pageId="4">
<materialsCitation accessionNumber="ROM 83713" collectingDate="2008-04-17" collectionCode="ROM" collectorName="Fernandez, D &amp; Taphorn, E &amp; Liverpool, K &amp; C. Thierens." country="Guyana" county="Philipai" latitude="5.3602777" location="upper Mazaruni River" longLatPrecision="21" longitude="-60.371666" municipality="Kukui River" pageId="4" specimenCode="CSBD 1652" specimenCount="1" typeStatus="holotype">
<emphasis bold="true" box="[816,915,611,632]" pageId="4">
<typeStatus box="[816,911,611,632]" pageId="4">Holotype</typeStatus>
.
</emphasis>
<specimenCode box="[920,1037,611,632]" collectionCode="CSBD" pageId="4">CSBD 1652</specimenCode>
, ex (
<accessionNumber box="[1080,1204,611,632]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ROM83713" pageId="4">
<collectionCode box="[1080,1136,611,632]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" type="Museum">ROM</collectionCode>
83713
</accessionNumber>
),
<quantity box="[1219,1303,611,632]" metricMagnitude="-2" metricUnit="m" metricValue="8.29" pageId="4" unit="mm" value="82.9">82.9 mm</quantity>
SL, presumably male,
<collectingCountry box="[876,954,640,661]" name="Guyana" pageId="4">Guyana</collectingCountry>
,
<location LSID="urn:lsid:plazi:treatment:03D12959FFEEFFB4259A82416879FB3B:8EA7CE94FFEEFFB725FA82C269CDFD52" box="[964,1179,640,661]" country="Guyana" county="Philipai" latitude="5.3602777" longLatPrecision="21" longitude="-60.371666" municipality="Kukui River" name="upper Mazaruni River" pageId="4">upper Mazaruni River</location>
drainage,
<collectingMunicipality box="[1283,1407,640,661]" pageId="4">Kukui River</collectingMunicipality>
, near town of
<collectingCounty box="[893,971,668,689]" pageId="4">Philipai</collectingCounty>
,
<geoCoordinate box="[979,1098,668,689]" degrees="05" direction="north" minutes="21" orientation="latitude" pageId="4" precision="15" seconds="37" value="5.3602777">05°2137”N</geoCoordinate>
<geoCoordinate box="[1102,1225,668,689]" degrees="60" direction="west" minutes="22" orientation="longitude" pageId="4" precision="15" seconds="18" value="-60.371666">60°2218”W</geoCoordinate>
,
<date box="[1234,1354,668,689]" pageId="4" value="2008-04-17">
<collectingDate box="[1234,1354,668,689]" pageId="4" value="2008-04-17">17 Apr 2008</collectingDate>
</date>
, H. López-
<collectorName box="[816,950,697,718]" pageId="4">Fernández, D</collectorName>
.
<collectorName box="[960,1071,697,718]" pageId="4">Taphorn, E</collectorName>
.
<collectorName box="[1081,1210,697,718]" pageId="4">Liverpool, K</collectorName>
. Kramer &amp;
<collectorName box="[1326,1447,697,718]" pageId="4">C. Thierens.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="4.[816,1459,758,894]" pageId="4">
<materialsCitation accessionNumber="ROM 84965" collectingDate="2008-04-17" collectionCode="ROM" collectorName="Fernandez, D &amp; Taphorn, E &amp; Liverpool, K &amp; C. Thierens." country="Guyana" county="Philipai" latitude="5.3602777" location="upper Mazaruni River" longLatPrecision="21" longitude="-60.371666" municipality="Kukui River" pageId="4" specimenCode="1" specimenCount="1" typeStatus="paratype">
<emphasis bold="true" box="[816,926,758,779]" pageId="4">
<typeStatus box="[816,922,758,779]" pageId="4">Paratypes</typeStatus>
.
</emphasis>
<accessionNumber box="[933,1058,758,779]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ROM84965" pageId="4">
<collectionCode box="[933,988,758,779]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" type="Museum">ROM</collectionCode>
84965
</accessionNumber>
,
<specimenCode box="[1068,1080,758,779]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" type="Museum">1</specimenCode>
,
<quantity box="[1093,1181,758,779]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" pageId="4" unit="mm" value="57.5">57.5 mm</quantity>
SL, presumably male, same data as holotype
</materialsCitation>
.
<materialsCitation accessionNumber="AUM 49999, ROM 83692" collectingDate="2008-04-15" collectionCode="AUM, ROM" collectorName="D. Taphorn &amp; E. Liverpool &amp; C. Thierens" country="Guyana" county="Philipai" latitude="5.3602777" location="upper Mazaruni River" longLatPrecision="21" longitude="-60.371666" municipality="Kukui River" pageId="4" specimenCount="1" typeStatus="paratype">
<accessionNumber box="[989,1113,787,808]" httpUri="http://www.ncbi.nlm.nih.gov/protein/AUM49999" pageId="4">
<collectionCode box="[989,1045,787,808]" country="USA" httpUri="http://grbio.org/cool/8wk2-p21y" name="Auburn University Museum of Natural History" pageId="4" type="University or college">AUM</collectionCode>
49999
</accessionNumber>
(ex
<accessionNumber box="[1159,1286,787,808]" httpUri="http://www.ncbi.nlm.nih.gov/protein/ROM83692" pageId="4">
<collectionCode box="[1159,1214,787,808]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" type="Museum">ROM</collectionCode>
83692
</accessionNumber>
), 1, 50.0 mm SL, immature,
<location LSID="urn:lsid:plazi:treatment:03D12959FFEEFFB4259A82416879FB3B:8EA7CE94FFEEFFB725A1836D69D7FC83" box="[927,1153,815,836]" country="Guyana" county="Philipai" latitude="5.3602777" longLatPrecision="21" longitude="-60.371666" municipality="Kukui River" name="upper Mazaruni River" pageId="4">upper Mazaruni River</location>
drainage,
<collectingMunicipality box="[1265,1394,815,836]" pageId="4">Kukui River</collectingMunicipality>
, night collection near
<collectingCounty box="[964,1043,844,865]" pageId="4">Philipai</collectingCounty>
,
<geoCoordinate box="[1051,1171,844,865]" degrees="05" direction="north" minutes="21" orientation="latitude" pageId="4" precision="15" seconds="37" value="5.3602777">05°2137”N</geoCoordinate>
<geoCoordinate box="[1176,1301,844,865]" degrees="60" direction="west" minutes="22" orientation="longitude" pageId="4" precision="15" seconds="18" value="-60.371666">60°2218”W</geoCoordinate>
,
<date box="[1305,1426,844,865]" pageId="4" value="2008-04-15">
<collectingDate box="[1305,1426,844,865]" pageId="4" value="2008-04-15">15 Apr 2008</collectingDate>
</date>
,
<collectorName pageId="4">D. Taphorn</collectorName>
,
<collectorName box="[909,1033,873,894]" pageId="4">E. Liverpool</collectorName>
&amp;
<collectorName box="[1064,1181,873,894]" pageId="4">C. Thierens</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" pageId="4" type="diagnosis">
<paragraph blockId="4.[816,1459,934,1309]" lastBlockId="5.[129,1098,180,205]" lastPageId="5" pageId="4">
<emphasis bold="true" box="[816,934,934,957]" pageId="4">Diagnosis.</emphasis>
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[942,1206,934,957]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="echinasus">
<emphasis box="[942,1206,934,957]" italics="true" pageId="4">Neblinichthys echinasus</emphasis>
</taxonomicName>
can be separated from all other
<taxonomicName authorityName="Ferraris, Isbrucker &amp; Nijssen" authorityYear="1986" box="[910,1058,966,989]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="genus">
<emphasis box="[910,1058,966,989]" italics="true" pageId="4">Neblinichthys</emphasis>
</taxonomicName>
except
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[1140,1341,966,989]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="brevibracchium">
<emphasis box="[1140,1158,966,989]" italics="true" pageId="4">N</emphasis>
.
<emphasis box="[1170,1341,966,989]" italics="true" pageId="4">brevibracchium</emphasis>
</taxonomicName>
, by having a pigmentation pattern on the abdomen consisting of large white spots and vermiculations over a dark gray or black background in adult males (
<emphasis box="[1123,1145,1062,1085]" italics="true" pageId="4">vs</emphasis>
. abdomen with small brown spots and vermiculations or plain); by lacking odontodes on the opercle (rarely one odontode present,
<emphasis box="[1287,1309,1126,1149]" italics="true" pageId="4">vs</emphasis>
. the exposed portion of the opercle completely covered by odontodes), by completely lacking the iris operculum (
<emphasis box="[1216,1238,1190,1213]" italics="true" pageId="4">vs</emphasis>
. iris operculum small or at least the dorsal rim of the pupil straight), and internares width/head length (6.2-6.7%
<emphasis box="[1120,1142,1254,1277]" italics="true" pageId="4">vs</emphasis>
. 7.6-15.5%, see
<tableCitation box="[1313,1424,1254,1277]" captionStart-0="Table 1" captionStart-1="Table2" captionStartId-0="3.[129,191,261,284]" captionStartId-1="5.[129,188,261,284]" captionText-0="Table 1. Morphometric features of Neblinichthys brevibracchium, new species (N = 11) and N. echinasus, new species (N = 3). Landmarks are the numbered landmarks inArmbruster (2003)." captionText-1="Table2. Morphometricfeaturesof Neblinichthyspilosus (N=6),N.roraima(N=4),andN. yaravi(N= 8). Landmarksarethenumbered" pageId="4">Tables 1-2</tableCitation>
). It is distinguished from
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[1051,1254,1286,1309]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="brevibracchium">
<emphasis box="[1051,1069,1286,1309]" italics="true" pageId="4">N</emphasis>
.
<emphasis box="[1083,1254,1286,1309]" italics="true" pageId="4">brevibracchium</emphasis>
</taxonomicName>
by a larger dorsal- 620 Description of
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[634,928,180,201]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="brevibracchium">
<emphasis box="[634,928,180,201]" italics="true" pageId="5">Neblinichthys brevibracchium</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[974,1098,180,201]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="echinasus">
<emphasis box="[974,990,180,201]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[1002,1098,180,201]" italics="true" pageId="5">echinasus</emphasis>
</taxonomicName>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647456" ID-Zenodo-Dep="4647456" httpUri="https://zenodo.org/record/4647456/files/figure.png" pageId="4" startId="4.[816,853,1892,1915]" targetBox="[817,1450,1370,1856]" targetPageId="4">
<paragraph blockId="4.[816,1459,1892,2043]" pageId="4">
<emphasis bold="true" box="[816,884,1892,1915]" pageId="4">Fig. 2.</emphasis>
Distribution of
<taxonomicName authorityName="Ferraris, Isbrucker &amp; Nijssen" authorityYear="1986" box="[1050,1189,1892,1915]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="genus">
<emphasis box="[1050,1189,1892,1915]" italics="true" pageId="4">Neblinichthys</emphasis>
</taxonomicName>
species in the Caroní and Mazaruni basins of Venezuela and Guyana, respectively.Star =
<taxonomicName class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="roraima">
<emphasis box="[1435,1452,1924,1947]" italics="true" pageId="4">N</emphasis>
.
<emphasis box="[816,897,1956,1979]" italics="true" pageId="4">roraima</emphasis>
</taxonomicName>
, black circle =
<taxonomicName box="[1041,1131,1956,1979]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="yaravi">
<emphasis box="[1041,1058,1956,1979]" italics="true" pageId="4">N</emphasis>
.
<emphasis box="[1068,1131,1956,1979]" italics="true" pageId="4">yaravi</emphasis>
</taxonomicName>
,empty circle = type-locality of
<taxonomicName authority="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard, 2010" authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="echinasus" status="new species">
<emphasis box="[1435,1452,1956,1979]" italics="true" pageId="4">N</emphasis>
.
<emphasis box="[816,915,1988,2011]" italics="true" pageId="4">echinasus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[928,1052,1988,2011]" pageId="4" rank="species">new species</taxonomicNameLabel>
, empty square = type-locality and black squares = paratype localities of
<taxonomicName authority="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard, 2010" authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[1130,1320,2020,2043]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="brevibracchium" status="new species">
<emphasis box="[1130,1147,2020,2043]" italics="true" pageId="4">N</emphasis>
.
<emphasis box="[1159,1320,2020,2043]" italics="true" pageId="4">brevibracchium</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[1332,1455,2020,2043]" pageId="4" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</caption>
<caption box="[129,1451,261,284]" pageId="5">
<paragraph blockId="5.[129,1452,261,316]" box="[129,1451,261,284]" pageId="5">
<emphasis bold="true" box="[129,210,261,284]" pageId="5">Table 2.</emphasis>
Morphometric features of
<taxonomicName box="[468,677,261,284]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="pilosus">
<emphasis box="[468,677,261,284]" italics="true" pageId="5">Neblinichthys pilosus</emphasis>
</taxonomicName>
(N = 6),
<taxonomicName box="[761,869,261,284]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="roraima">
<emphasis box="[761,778,261,284]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[788,869,261,284]" italics="true" pageId="5">roraima</emphasis>
</taxonomicName>
(N = 4), and
<taxonomicName box="[991,1083,261,284]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="yaravi">
<emphasis box="[991,1008,261,284]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[1019,1083,261,284]" italics="true" pageId="5">yaravi</emphasis>
</taxonomicName>
(N = 8). Landmarks are the numbered
</paragraph>
</caption>
<paragraph blockId="5.[129,1452,261,316]" lastBlockId="5.[129,771,1348,1787]" pageId="5">
landmarks inArmbruster (2003). Some
<taxonomicName box="[516,625,293,316]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="roraima">
<emphasis box="[516,533,293,316]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[543,625,293,316]" italics="true" pageId="5">roraima</emphasis>
</taxonomicName>
have completely adnate barbels, resulting in a measurement of zero for barbel length. fin base/SL ratio (24.3-27.0%
<emphasis box="[433,454,1348,1371]" italics="true" pageId="5">vs</emphasis>
. 18.1-22.8%), a smaller dorsalanal distance/SL ratio (11.9-12.5%
<emphasis box="[493,515,1380,1403]" italics="true" pageId="5">vs</emphasis>
. 13.7-18.3%), by having the snout tapering shallowly and continuously from eyes to snout tip (
<emphasis box="[242,264,1444,1467]" italics="true" pageId="5">vs</emphasis>
. snout decreasing in steep arc just anterior to eyes and then flattening in area anterior of nares;
<figureCitation box="[650,712,1476,1499]" captionStart="Fig" captionStartId="6.[136,176,1712,1735]" captionTargetBox="[136,1458,257,1701]" captionTargetId="figure-174@6.[136,1459,257,1702]" captionTargetPageId="6" captionText="Fig. 3. Neblinichthys echinasus, holotype, CSBD 1652, 82.9 mm SL, Kukui River, near town of Philipai, upper Mazaruni River drainage." figureDoi="http://doi.org/10.5281/zenodo.4647458" httpUri="https://zenodo.org/record/4647458/files/figure.png" pageId="5">Fig. 3</figureCitation>
), and by having the adpressed dorsal fin reaching the anterior preadipose plate (
<emphasis box="[318,340,1540,1563]" italics="true" pageId="5">vs</emphasis>
. not reaching anterior preadipose plate). From
<taxonomicName box="[196,306,1572,1595]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="pilosus">
<emphasis box="[196,214,1572,1595]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[230,306,1572,1595]" italics="true" pageId="5">pilosus</emphasis>
</taxonomicName>
and
<taxonomicName box="[365,488,1572,1595]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="roraima">
<emphasis box="[365,383,1572,1595]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[400,488,1572,1595]" italics="true" pageId="5">roraima</emphasis>
</taxonomicName>
by having the odontodes along the edge of the snout of nuptial males longer than those on the top of the snout (
<emphasis box="[476,498,1636,1659]" italics="true" pageId="5">vs</emphasis>
. odontodes on the front edge of the snout much longer than those on the sides of the snout; nuptial males not known for
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[509,711,1700,1723]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="brevibracchium">
<emphasis box="[509,527,1700,1723]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[540,711,1700,1723]" italics="true" pageId="5">brevibracchium</emphasis>
</taxonomicName>
or
<taxonomicName class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="yaravi">
<emphasis box="[746,764,1700,1723]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[129,198,1732,1755]" italics="true" pageId="5">yaravi</emphasis>
</taxonomicName>
); from
<taxonomicName box="[283,403,1732,1755]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="roraima">
<emphasis box="[283,301,1732,1755]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[315,403,1732,1755]" italics="true" pageId="5">roraima</emphasis>
</taxonomicName>
by having one to two preadipose plates (
<emphasis box="[206,228,1764,1787]" italics="true" pageId="5">vs</emphasis>
. more than four).
</paragraph>
</subSubSection>
<paragraph pageId="5">
<table box="[156,1433,340,1294]" gridcols="8" gridrows="39" pageId="5">
<tr box="[156,1433,340,359]" gridrow="0" pageId="5" rowspan-0="1" rowspan-1="1">
<td box="[676,910,340,359]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="5">
<taxonomicName box="[758,844,340,359]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="pilosus">
<emphasis box="[758,772,340,359]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[783,844,340,359]" italics="true" pageId="5">pilosus</emphasis>
</taxonomicName>
</td>
<td box="[955,1171,340,359]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="5">
<taxonomicName box="[1015,1110,340,359]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="roraima">
<emphasis box="[1015,1029,340,359]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[1040,1110,340,359]" italics="true" pageId="5">roraima</emphasis>
</taxonomicName>
</td>
<td box="[1216,1433,340,359]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="5">
<taxonomicName box="[1284,1364,340,359]" class="Actinopterygii" family="Loricariidae" genus="Ancistrus" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="species" species="yaravi">
<emphasis box="[1284,1298,340,359]" italics="true" pageId="5">N</emphasis>
.
<emphasis box="[1309,1364,340,359]" italics="true" pageId="5">yaravi</emphasis>
</taxonomicName>
</td>
</tr>
<tr box="[156,1433,366,385]" gridrow="1" pageId="5">
<td box="[156,252,366,385]" gridcol="0" gridrow="1" pageId="5">Landmarks</td>
<td box="[278,533,366,385]" gridcol="1" gridrow="1" pageId="5">Feature</td>
<td box="[676,782,366,385]" gridcol="2" gridrow="1" pageId="5">Mean±SD</td>
<td box="[827,910,366,385]" gridcol="3" gridrow="1" pageId="5">Range</td>
<td box="[955,1043,366,385]" gridcol="4" gridrow="1" pageId="5">Mean±SD</td>
<td box="[1089,1171,366,385]" gridcol="5" gridrow="1" pageId="5">Range</td>
<td box="[1216,1305,366,385]" gridcol="6" gridrow="1" pageId="5">Mean±SD</td>
<td box="[1350,1433,366,385]" gridcol="7" gridrow="1" pageId="5">Range</td>
</tr>
<tr box="[156,1433,392,411]" gridrow="2" pageId="5">
<td box="[156,252,392,411]" gridcol="0" gridrow="2" pageId="5">1-20</td>
<td box="[278,533,392,411]" gridcol="1" gridrow="2" pageId="5">SL (mm)</td>
<td box="[676,782,392,411]" gridcol="2" gridrow="2" pageId="5">55.5</td>
<td box="[827,910,392,411]" gridcol="3" gridrow="2" pageId="5">49.1-61.9</td>
<td box="[955,1043,392,411]" gridcol="4" gridrow="2" pageId="5">43.7</td>
<td box="[1089,1171,392,411]" gridcol="5" gridrow="2" pageId="5">38.7-46.3</td>
<td box="[1216,1305,392,411]" gridcol="6" gridrow="2" pageId="5">48.4</td>
<td box="[1350,1433,392,411]" gridcol="7" gridrow="2" pageId="5">31.0-75.0</td>
</tr>
<tr box="[156,1433,416,435]" gridrow="3" pageId="5" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td box="[676,910,416,435]" colspan="2" colspanRight="1" gridcol="2" gridrow="3" pageId="5">Percents of standard length</td>
</tr>
<tr box="[156,1433,441,460]" gridrow="4" pageId="5">
<td box="[156,252,441,460]" gridcol="0" gridrow="4" pageId="5">1-10</td>
<td box="[278,533,441,460]" gridcol="1" gridrow="4" pageId="5">Predorsal length</td>
<td box="[676,782,441,460]" gridcol="2" gridrow="4" pageId="5">44.4±1.0</td>
<td box="[827,910,441,460]" gridcol="3" gridrow="4" pageId="5">42.8-45.3</td>
<td box="[955,1043,441,460]" gridcol="4" gridrow="4" pageId="5">47.0±2.2</td>
<td box="[1089,1171,441,460]" gridcol="5" gridrow="4" pageId="5">44.9-49.1</td>
<td box="[1216,1305,441,460]" gridcol="6" gridrow="4" pageId="5">43.6±0.9</td>
<td box="[1350,1433,441,460]" gridcol="7" gridrow="4" pageId="5">42.9-45.6</td>
</tr>
<tr box="[156,1433,466,485]" gridrow="5" pageId="5">
<td box="[156,252,466,485]" gridcol="0" gridrow="5" pageId="5">1-7</td>
<td box="[278,533,466,485]" gridcol="1" gridrow="5" pageId="5">Head length (HL)</td>
<td box="[676,782,466,485]" gridcol="2" gridrow="5" pageId="5">34.9±0.9</td>
<td box="[827,910,466,485]" gridcol="3" gridrow="5" pageId="5">33.4-36.1</td>
<td box="[955,1043,466,485]" gridcol="4" gridrow="5" pageId="5">34.4±0.7</td>
<td box="[1089,1171,466,485]" gridcol="5" gridrow="5" pageId="5">33.4-35.0</td>
<td box="[1216,1305,466,485]" gridcol="6" gridrow="5" pageId="5">34.5±1.1</td>
<td box="[1350,1433,466,485]" gridcol="7" gridrow="5" pageId="5">33.4-37.0</td>
</tr>
<tr box="[156,1433,490,509]" gridrow="6" pageId="5">
<td box="[156,252,490,509]" gridcol="0" gridrow="6" pageId="5">7-10</td>
<td box="[278,533,490,509]" gridcol="1" gridrow="6" pageId="5">Head-dorsal length</td>
<td box="[676,782,490,509]" gridcol="2" gridrow="6" pageId="5">9.6±1.3</td>
<td box="[827,910,490,509]" gridcol="3" gridrow="6" pageId="5">7.9-11.2</td>
<td box="[955,1043,490,509]" gridcol="4" gridrow="6" pageId="5">14.1±3.0</td>
<td box="[1089,1171,490,509]" gridcol="5" gridrow="6" pageId="5">11.2-17.7</td>
<td box="[1216,1305,490,509]" gridcol="6" gridrow="6" pageId="5">9.4±0.3</td>
<td box="[1350,1433,490,509]" gridcol="7" gridrow="6" pageId="5">8.9-9.7</td>
</tr>
<tr box="[156,1433,515,534]" gridrow="7" pageId="5">
<td box="[156,252,515,534]" gridcol="0" gridrow="7" pageId="5">8-9</td>
<td box="[278,533,515,534]" gridcol="1" gridrow="7" pageId="5">Cleithral width</td>
<td box="[676,782,515,534]" gridcol="2" gridrow="7" pageId="5">29.1±1.5</td>
<td box="[827,910,515,534]" gridcol="3" gridrow="7" pageId="5">26.7-31.1</td>
<td box="[955,1043,515,534]" gridcol="4" gridrow="7" pageId="5">26.2±0.2</td>
<td box="[1089,1171,515,534]" gridcol="5" gridrow="7" pageId="5">25.9-26.3</td>
<td box="[1216,1305,515,534]" gridcol="6" gridrow="7" pageId="5">31.4±1.3</td>
<td box="[1350,1433,515,534]" gridcol="7" gridrow="7" pageId="5">29.1-33.4</td>
</tr>
<tr box="[156,1433,539,558]" gridrow="8" pageId="5">
<td box="[156,252,539,558]" gridcol="0" gridrow="8" pageId="5">1-12</td>
<td box="[278,533,539,558]" gridcol="1" gridrow="8" pageId="5">Head-pectoral length</td>
<td box="[676,782,539,558]" gridcol="2" gridrow="8" pageId="5">28.5±1.4</td>
<td box="[827,910,539,558]" gridcol="3" gridrow="8" pageId="5">27.3-30.8</td>
<td box="[955,1043,539,558]" gridcol="4" gridrow="8" pageId="5">29.4±0.4</td>
<td box="[1089,1171,539,558]" gridcol="5" gridrow="8" pageId="5">29.2-30.0</td>
<td box="[1216,1305,539,558]" gridcol="6" gridrow="8" pageId="5">26.7±1.6</td>
<td box="[1350,1433,539,558]" gridcol="7" gridrow="8" pageId="5">25.1-30.4</td>
</tr>
<tr box="[156,1433,564,583]" gridrow="9" pageId="5">
<td box="[156,252,564,583]" gridcol="0" gridrow="9" pageId="5">12-13</td>
<td box="[278,533,564,583]" gridcol="1" gridrow="9" pageId="5">Thorax length</td>
<td box="[676,782,564,583]" gridcol="2" gridrow="9" pageId="5">23.0±1.5</td>
<td box="[827,910,564,583]" gridcol="3" gridrow="9" pageId="5">21.3-25.3</td>
<td box="[955,1043,564,583]" gridcol="4" gridrow="9" pageId="5">25.5±0.5</td>
<td box="[1089,1171,564,583]" gridcol="5" gridrow="9" pageId="5">25.0-25.9</td>
<td box="[1216,1305,564,583]" gridcol="6" gridrow="9" pageId="5">23.9± 0.8</td>
<td box="[1350,1433,564,583]" gridcol="7" gridrow="9" pageId="5">22.8-25.0</td>
</tr>
<tr box="[156,1433,588,607]" gridrow="10" pageId="5">
<td box="[156,252,588,607]" gridcol="0" gridrow="10" pageId="5">12-29</td>
<td box="[278,533,588,607]" gridcol="1" gridrow="10" pageId="5">Pectoral-spine length</td>
<td box="[676,782,588,607]" gridcol="2" gridrow="10" pageId="5">24.1±2.2</td>
<td box="[827,910,588,607]" gridcol="3" gridrow="10" pageId="5">22.1-27.6</td>
<td box="[955,1043,588,607]" gridcol="4" gridrow="10" pageId="5">21.9±1.0</td>
<td box="[1089,1171,588,607]" gridcol="5" gridrow="10" pageId="5">20.5-22.8</td>
<td box="[1216,1305,588,607]" gridcol="6" gridrow="10" pageId="5">24.5±1.2</td>
<td box="[1350,1433,588,607]" gridcol="7" gridrow="10" pageId="5">22.4-25.8</td>
</tr>
<tr box="[156,1433,613,632]" gridrow="11" pageId="5">
<td box="[156,252,613,632]" gridcol="0" gridrow="11" pageId="5">13-14</td>
<td box="[278,533,613,632]" gridcol="1" gridrow="11" pageId="5">Abdominal length</td>
<td box="[676,782,613,632]" gridcol="2" gridrow="11" pageId="5">23.7±1.2</td>
<td box="[827,910,613,632]" gridcol="3" gridrow="11" pageId="5">22.6-26.0</td>
<td box="[955,1043,613,632]" gridcol="4" gridrow="11" pageId="5">22.9±1.0</td>
<td box="[1089,1171,613,632]" gridcol="5" gridrow="11" pageId="5">21.8-23.8</td>
<td box="[1216,1305,613,632]" gridcol="6" gridrow="11" pageId="5">23.2±1.5</td>
<td box="[1350,1433,613,632]" gridcol="7" gridrow="11" pageId="5">20.8-25.4</td>
</tr>
<tr box="[156,1433,637,656]" gridrow="12" pageId="5">
<td box="[156,252,637,656]" gridcol="0" gridrow="12" pageId="5">13-30</td>
<td box="[278,533,637,656]" gridcol="1" gridrow="12" pageId="5">Pelvic-spine length</td>
<td box="[676,782,637,656]" gridcol="2" gridrow="12" pageId="5">25.9±2.2</td>
<td box="[827,910,637,656]" gridcol="3" gridrow="12" pageId="5">23.6-29.0</td>
<td box="[955,1043,637,656]" gridcol="4" gridrow="12" pageId="5">21.1±1.5</td>
<td box="[1089,1171,637,656]" gridcol="5" gridrow="12" pageId="5">20.1-23.4</td>
<td box="[1216,1305,637,656]" gridcol="6" gridrow="12" pageId="5">23.9±1.3</td>
<td box="[1350,1433,637,656]" gridcol="7" gridrow="12" pageId="5">22.1-25.5</td>
</tr>
<tr box="[156,1433,662,681]" gridrow="13" pageId="5">
<td box="[156,252,662,681]" gridcol="0" gridrow="13" pageId="5">14-15</td>
<td box="[278,533,662,681]" gridcol="1" gridrow="13" pageId="5">Postanal length</td>
<td box="[676,782,662,681]" gridcol="2" gridrow="13" pageId="5">33.8±0.8</td>
<td box="[827,910,662,681]" gridcol="3" gridrow="13" pageId="5">32.8-34.9</td>
<td box="[955,1043,662,681]" gridcol="4" gridrow="13" pageId="5">31.6±1.2</td>
<td box="[1089,1171,662,681]" gridcol="5" gridrow="13" pageId="5">30.0-32.8</td>
<td box="[1216,1305,662,681]" gridcol="6" gridrow="13" pageId="5">33.3±2.6</td>
<td box="[1350,1433,662,681]" gridcol="7" gridrow="13" pageId="5">30.8-38.0</td>
</tr>
<tr box="[156,1433,687,706]" gridrow="14" pageId="5">
<td box="[156,252,687,706]" gridcol="0" gridrow="14" pageId="5">14-31</td>
<td box="[278,533,687,706]" gridcol="1" gridrow="14" pageId="5">Anal-fin spine length</td>
<td box="[676,782,687,706]" gridcol="2" gridrow="14" pageId="5">12.0±2.7</td>
<td box="[827,910,687,706]" gridcol="3" gridrow="14" pageId="5">9.0-16.0</td>
<td box="[955,1043,687,706]" gridcol="4" gridrow="14" pageId="5">7.2±0.4</td>
<td box="[1089,1171,687,706]" gridcol="5" gridrow="14" pageId="5">7.0-7.5</td>
<td box="[1216,1305,687,706]" gridcol="6" gridrow="14" pageId="5">8.9±1.8</td>
<td box="[1350,1433,687,706]" gridcol="7" gridrow="14" pageId="5">6.3-11.0</td>
</tr>
<tr box="[156,1433,711,730]" gridrow="15" pageId="5">
<td box="[156,252,711,730]" gridcol="0" gridrow="15" pageId="5">10-12</td>
<td box="[278,533,711,730]" gridcol="1" gridrow="15" pageId="5">Dorsal-pectoral distance</td>
<td box="[676,782,711,730]" gridcol="2" gridrow="15" pageId="5">28.6±1.3</td>
<td box="[827,910,711,730]" gridcol="3" gridrow="15" pageId="5">26.5-30.5</td>
<td box="[955,1043,711,730]" gridcol="4" gridrow="15" pageId="5">29.3±1.1</td>
<td box="[1089,1171,711,730]" gridcol="5" gridrow="15" pageId="5">28.5-30.8</td>
<td box="[1216,1305,711,730]" gridcol="6" gridrow="15" pageId="5">27.1±0.6</td>
<td box="[1350,1433,711,730]" gridcol="7" gridrow="15" pageId="5">25.8-27.8</td>
</tr>
<tr box="[156,1433,735,754]" gridrow="16" pageId="5">
<td box="[156,252,735,754]" gridcol="0" gridrow="16" pageId="5">10-11</td>
<td box="[278,533,735,754]" gridcol="1" gridrow="16" pageId="5">Dorsal-spine length</td>
<td box="[676,782,735,754]" gridcol="2" gridrow="16" pageId="5">24.3±2.4</td>
<td box="[827,910,735,754]" gridcol="3" gridrow="16" pageId="5">20.4-27.4</td>
<td box="[955,1043,735,754]" gridcol="4" gridrow="16" pageId="5">22.6±0.7</td>
<td box="[1089,1171,735,754]" gridcol="5" gridrow="16" pageId="5">22.0-23.6</td>
<td box="[1216,1305,735,754]" gridcol="6" gridrow="16" pageId="5">24.1±1.7</td>
<td box="[1350,1433,735,754]" gridcol="7" gridrow="16" pageId="5">20.9-26.8</td>
</tr>
<tr box="[156,1433,760,779]" gridrow="17" pageId="5">
<td box="[156,252,760,779]" gridcol="0" gridrow="17" pageId="5">10-13</td>
<td box="[278,533,760,779]" gridcol="1" gridrow="17" pageId="5">Dorsal-pelvic distance</td>
<td box="[676,782,760,779]" gridcol="2" gridrow="17" pageId="5">19.4±2.1</td>
<td box="[827,910,760,779]" gridcol="3" gridrow="17" pageId="5">15.5-21.0</td>
<td box="[955,1043,760,779]" gridcol="4" gridrow="17" pageId="5">19.0±0.9</td>
<td box="[1089,1171,760,779]" gridcol="5" gridrow="17" pageId="5">17.7-19.9</td>
<td box="[1216,1305,760,779]" gridcol="6" gridrow="17" pageId="5">19.9±1.8</td>
<td box="[1350,1433,760,779]" gridcol="7" gridrow="17" pageId="5">16.4-21.9</td>
</tr>
<tr box="[156,1433,784,803]" gridrow="18" pageId="5">
<td box="[156,252,784,803]" gridcol="0" gridrow="18" pageId="5">10-16</td>
<td box="[278,533,784,803]" gridcol="1" gridrow="18" pageId="5">Dorsal-fin base length</td>
<td box="[676,782,784,803]" gridcol="2" gridrow="18" pageId="5">22.9±1.2</td>
<td box="[827,910,784,803]" gridcol="3" gridrow="18" pageId="5">20.8-24.0</td>
<td box="[955,1043,784,803]" gridcol="4" gridrow="18" pageId="5">19.3±1.1</td>
<td box="[1089,1171,784,803]" gridcol="5" gridrow="18" pageId="5">17.8-20.1</td>
<td box="[1216,1305,784,803]" gridcol="6" gridrow="18" pageId="5">24.3±1.5</td>
<td box="[1350,1433,784,803]" gridcol="7" gridrow="18" pageId="5">22.0-26.6</td>
</tr>
<tr box="[156,1433,809,828]" gridrow="19" pageId="5">
<td box="[156,252,809,828]" gridcol="0" gridrow="19" pageId="5">16-17</td>
<td box="[278,533,809,828]" gridcol="1" gridrow="19" pageId="5">Dorsal-adipose distance</td>
<td box="[676,782,809,828]" gridcol="2" gridrow="19" pageId="5">18.0±1.0</td>
<td box="[827,910,809,828]" gridcol="3" gridrow="19" pageId="5">16.5-19.6</td>
<td box="[955,1043,809,828]" gridcol="4" gridrow="19" pageId="5">19.8±0.8</td>
<td box="[1089,1171,809,828]" gridcol="5" gridrow="19" pageId="5">18.7-20.5</td>
<td box="[1216,1305,809,828]" gridcol="6" gridrow="19" pageId="5">19.4±1.2</td>
<td box="[1350,1433,809,828]" gridcol="7" gridrow="19" pageId="5">17.2-21.0</td>
</tr>
<tr box="[156,1433,834,853]" gridrow="20" pageId="5">
<td box="[156,252,834,853]" gridcol="0" gridrow="20" pageId="5">17-18</td>
<td box="[278,533,834,853]" gridcol="1" gridrow="20" pageId="5">Adipose-spine length</td>
<td box="[676,782,834,853]" gridcol="2" gridrow="20" pageId="5">7.7±1.5</td>
<td box="[827,910,834,853]" gridcol="3" gridrow="20" pageId="5">6.2-9.7</td>
<td box="[955,1043,834,853]" gridcol="4" gridrow="20" pageId="5">8.7±2.3</td>
<td box="[1089,1171,834,853]" gridcol="5" gridrow="20" pageId="5">6.4-11.8</td>
<td box="[1216,1305,834,853]" gridcol="6" gridrow="20" pageId="5">8.4±0.6</td>
<td box="[1350,1433,834,853]" gridcol="7" gridrow="20" pageId="5">7.1-9.0</td>
</tr>
<tr box="[156,1433,858,877]" gridrow="21" pageId="5">
<td box="[156,252,858,877]" gridcol="0" gridrow="21" pageId="5">17-19</td>
<td box="[278,533,858,877]" gridcol="1" gridrow="21" pageId="5">Adipose-up. caudal distance</td>
<td box="[676,782,858,877]" gridcol="2" gridrow="21" pageId="5">14.6±1.2</td>
<td box="[827,910,858,877]" gridcol="3" gridrow="21" pageId="5">13.0-15.9</td>
<td box="[955,1043,858,877]" gridcol="4" gridrow="21" pageId="5">16.8±0.8</td>
<td box="[1089,1171,858,877]" gridcol="5" gridrow="21" pageId="5">15.7-17.5</td>
<td box="[1216,1305,858,877]" gridcol="6" gridrow="21" pageId="5">15.8±1.4</td>
<td box="[1350,1433,858,877]" gridcol="7" gridrow="21" pageId="5">13.4-17.5</td>
</tr>
<tr box="[156,1433,883,902]" gridrow="22" pageId="5">
<td box="[156,252,883,902]" gridcol="0" gridrow="22" pageId="5">15-19</td>
<td box="[278,533,883,902]" gridcol="1" gridrow="22" pageId="5">Caudal peduncle depth</td>
<td box="[676,782,883,902]" gridcol="2" gridrow="22" pageId="5">10.0±0.8</td>
<td box="[827,910,883,902]" gridcol="3" gridrow="22" pageId="5">9.2-11.1</td>
<td box="[955,1043,883,902]" gridcol="4" gridrow="22" pageId="5">10.6±1.5</td>
<td box="[1089,1171,883,902]" gridcol="5" gridrow="22" pageId="5">8.9-12.2</td>
<td box="[1216,1305,883,902]" gridcol="6" gridrow="22" pageId="5">9.0±0.7</td>
<td box="[1350,1433,883,902]" gridcol="7" gridrow="22" pageId="5">8.1 - 10.0</td>
</tr>
<tr box="[156,1433,907,926]" gridrow="23" pageId="5">
<td box="[156,252,907,926]" gridcol="0" gridrow="23" pageId="5">15-17</td>
<td box="[278,533,907,926]" gridcol="1" gridrow="23" pageId="5">Adipose-low. caudal distance</td>
<td box="[676,782,907,926]" gridcol="2" gridrow="23" pageId="5">19.9±1.4</td>
<td box="[827,910,907,926]" gridcol="3" gridrow="23" pageId="5">18.1-21.5</td>
<td box="[955,1043,907,926]" gridcol="4" gridrow="23" pageId="5">18.5±1.2</td>
<td box="[1089,1171,907,926]" gridcol="5" gridrow="23" pageId="5">17.8-20.2</td>
<td box="[1216,1305,907,926]" gridcol="6" gridrow="23" pageId="5">20.7±1.2</td>
<td box="[1350,1433,907,926]" gridcol="7" gridrow="23" pageId="5">18.7-22.0</td>
</tr>
<tr box="[156,1433,932,951]" gridrow="24" pageId="5">
<td box="[156,252,932,951]" gridcol="0" gridrow="24" pageId="5">14-17</td>
<td box="[278,533,932,951]" gridcol="1" gridrow="24" pageId="5">Adipose-anal distance</td>
<td box="[676,782,932,951]" gridcol="2" gridrow="24" pageId="5">20.0±1.5</td>
<td box="[827,910,932,951]" gridcol="3" gridrow="24" pageId="5">17.9-21.4</td>
<td box="[955,1043,932,951]" gridcol="4" gridrow="24" pageId="5">18.0±0.7</td>
<td box="[1089,1171,932,951]" gridcol="5" gridrow="24" pageId="5">17.3-18.7</td>
<td box="[1216,1305,932,951]" gridcol="6" gridrow="24" pageId="5">20.2±0.7</td>
<td box="[1350,1433,932,951]" gridcol="7" gridrow="24" pageId="5">19.0-21.1</td>
</tr>
<tr box="[156,1433,956,975]" gridrow="25" pageId="5">
<td box="[156,252,956,975]" gridcol="0" gridrow="25" pageId="5">14-16</td>
<td box="[278,533,956,975]" gridcol="1" gridrow="25" pageId="5">Dorsal-anal distance</td>
<td box="[676,782,956,975]" gridcol="2" gridrow="25" pageId="5">14.0±0.9</td>
<td box="[827,910,956,975]" gridcol="3" gridrow="25" pageId="5">12.4-15.1</td>
<td box="[955,1043,956,975]" gridcol="4" gridrow="25" pageId="5">13.2±1.9</td>
<td box="[1089,1171,956,975]" gridcol="5" gridrow="25" pageId="5">11.1-15.7</td>
<td box="[1216,1305,956,975]" gridcol="6" gridrow="25" pageId="5">13.3±0.9</td>
<td box="[1350,1433,956,975]" gridcol="7" gridrow="25" pageId="5">12.0-14.7</td>
</tr>
<tr box="[156,1433,981,1000]" gridrow="26" pageId="5">
<td box="[156,252,981,1000]" gridcol="0" gridrow="26" pageId="5">13-16</td>
<td box="[278,533,981,1000]" gridcol="1" gridrow="26" pageId="5">Pelvic-dorsal distance</td>
<td box="[676,782,981,1000]" gridcol="2" gridrow="26" pageId="5">24.7±2.1</td>
<td box="[827,910,981,1000]" gridcol="3" gridrow="26" pageId="5">21.8-27.2</td>
<td box="[955,1043,981,1000]" gridcol="4" gridrow="26" pageId="5">20.8±1.3</td>
<td box="[1089,1171,981,1000]" gridcol="5" gridrow="26" pageId="5">19.0-22.0</td>
<td box="[1216,1305,981,1000]" gridcol="6" gridrow="26" pageId="5">25.7±5.2</td>
<td box="[1350,1433,981,1000]" gridcol="7" gridrow="26" pageId="5">22.1-38.0</td>
</tr>
<tr box="[156,1433,1005,1024]" gridrow="27" pageId="5" rowspan-0="1" rowspan-1="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<td box="[676,910,1005,1024]" colspan="2" colspanRight="1" gridcol="2" gridrow="27" pageId="5">Percents of head length</td>
</tr>
<tr box="[156,1433,1030,1049]" gridrow="28" pageId="5">
<td box="[156,252,1030,1049]" gridcol="0" gridrow="28" pageId="5">5-7</td>
<td box="[278,533,1030,1049]" gridcol="1" gridrow="28" pageId="5">Head-eye length</td>
<td box="[676,782,1030,1049]" gridcol="2" gridrow="28" pageId="5">31.0±1.3</td>
<td box="[827,910,1030,1049]" gridcol="3" gridrow="28" pageId="5">29.8-33.6</td>
<td box="[955,1043,1030,1049]" gridcol="4" gridrow="28" pageId="5">29.2±4.0</td>
<td box="[1089,1171,1030,1049]" gridcol="5" gridrow="28" pageId="5">23.5-32.2</td>
<td box="[1216,1305,1030,1049]" gridcol="6" gridrow="28" pageId="5">29.3±1.0</td>
<td box="[1350,1433,1030,1049]" gridcol="7" gridrow="28" pageId="5">27.3-30.8</td>
</tr>
<tr box="[156,1433,1054,1073]" gridrow="29" pageId="5">
<td box="[156,252,1054,1073]" gridcol="0" gridrow="29" pageId="5">4-5</td>
<td box="[278,533,1054,1073]" gridcol="1" gridrow="29" pageId="5">Orbit diameter</td>
<td box="[676,782,1054,1073]" gridcol="2" gridrow="29" pageId="5">13.2±0.9</td>
<td box="[827,910,1054,1073]" gridcol="3" gridrow="29" pageId="5">12.2-14.3</td>
<td box="[955,1043,1054,1073]" gridcol="4" gridrow="29" pageId="5">11.8±0.7</td>
<td box="[1089,1171,1054,1073]" gridcol="5" gridrow="29" pageId="5">11.1-12.8</td>
<td box="[1216,1305,1054,1073]" gridcol="6" gridrow="29" pageId="5">13.8±0.8</td>
<td box="[1350,1433,1054,1073]" gridcol="7" gridrow="29" pageId="5">12.4-15.0</td>
</tr>
<tr box="[156,1433,1079,1098]" gridrow="30" pageId="5">
<td box="[156,252,1079,1098]" gridcol="0" gridrow="30" pageId="5">1-4</td>
<td box="[278,533,1079,1098]" gridcol="1" gridrow="30" pageId="5">Snout length</td>
<td box="[676,782,1079,1098]" gridcol="2" gridrow="30" pageId="5">49.8±2.9</td>
<td box="[827,910,1079,1098]" gridcol="3" gridrow="30" pageId="5">45.4-52.7</td>
<td box="[955,1043,1079,1098]" gridcol="4" gridrow="30" pageId="5">42.9±1.9</td>
<td box="[1089,1171,1079,1098]" gridcol="5" gridrow="30" pageId="5">41.4-45.7</td>
<td box="[1216,1305,1079,1098]" gridcol="6" gridrow="30" pageId="5">46.8±2.1</td>
<td box="[1350,1433,1079,1098]" gridcol="7" gridrow="30" pageId="5">44.6-50.7</td>
</tr>
<tr box="[156,1433,1103,1122]" gridrow="31" pageId="5">
<td box="[156,252,1103,1122]" gridcol="0" gridrow="31" pageId="5">2-3</td>
<td box="[278,533,1103,1122]" gridcol="1" gridrow="31" pageId="5">Internares width</td>
<td box="[676,782,1103,1122]" gridcol="2" gridrow="31" pageId="5">14.7±0.8</td>
<td box="[827,910,1103,1122]" gridcol="3" gridrow="31" pageId="5">13.4-15.5</td>
<td box="[955,1043,1103,1122]" gridcol="4" gridrow="31" pageId="5">10.1±1.1</td>
<td box="[1089,1171,1103,1122]" gridcol="5" gridrow="31" pageId="5">8.8-11.5</td>
<td box="[1216,1305,1103,1122]" gridcol="6" gridrow="31" pageId="5">8.2±0.6</td>
<td box="[1350,1433,1103,1122]" gridcol="7" gridrow="31" pageId="5">7.6-9.2</td>
</tr>
<tr box="[156,1433,1128,1147]" gridrow="32" pageId="5">
<td box="[156,252,1128,1147]" gridcol="0" gridrow="32" pageId="5">5-6</td>
<td box="[278,533,1128,1147]" gridcol="1" gridrow="32" pageId="5">Interorbital width</td>
<td box="[676,782,1128,1147]" gridcol="2" gridrow="32" pageId="5">32.6±1.9</td>
<td box="[827,910,1128,1147]" gridcol="3" gridrow="32" pageId="5">30.7-35.5</td>
<td box="[955,1043,1128,1147]" gridcol="4" gridrow="32" pageId="5">25.8±2.1</td>
<td box="[1089,1171,1128,1147]" gridcol="5" gridrow="32" pageId="5">23.5-27.9</td>
<td box="[1216,1305,1128,1147]" gridcol="6" gridrow="32" pageId="5">41.7±1.9</td>
<td box="[1350,1433,1128,1147]" gridcol="7" gridrow="32" pageId="5">39.0-44.6</td>
</tr>
<tr box="[156,1433,1152,1171]" gridrow="33" pageId="5">
<td box="[156,252,1152,1171]" gridcol="0" gridrow="33" pageId="5">7-12</td>
<td box="[278,533,1152,1171]" gridcol="1" gridrow="33" pageId="5">Head depth</td>
<td box="[676,782,1152,1171]" gridcol="2" gridrow="33" pageId="5">52.4±2.1</td>
<td box="[827,910,1152,1171]" gridcol="3" gridrow="33" pageId="5">49.5-54.4</td>
<td box="[955,1043,1152,1171]" gridcol="4" gridrow="33" pageId="5">45.9±2.2</td>
<td box="[1089,1171,1152,1171]" gridcol="5" gridrow="33" pageId="5">43.5-48.2</td>
<td box="[1216,1305,1152,1171]" gridcol="6" gridrow="33" pageId="5">50.0±1.1</td>
<td box="[1350,1433,1152,1171]" gridcol="7" gridrow="33" pageId="5">48.9-52.3</td>
</tr>
<tr box="[156,1433,1177,1196]" gridrow="34" pageId="5">
<td box="[156,252,1177,1196]" gridcol="0" gridrow="34" pageId="5">1-24</td>
<td box="[278,533,1177,1196]" gridcol="1" gridrow="34" pageId="5">Mouth length</td>
<td box="[676,782,1177,1196]" gridcol="2" gridrow="34" pageId="5">42.9±4.5</td>
<td box="[827,910,1177,1196]" gridcol="3" gridrow="34" pageId="5">36.2-48.7</td>
<td box="[955,1043,1177,1196]" gridcol="4" gridrow="34" pageId="5">40.2±2.7</td>
<td box="[1089,1171,1177,1196]" gridcol="5" gridrow="34" pageId="5">38.0-43.9</td>
<td box="[1216,1305,1177,1196]" gridcol="6" gridrow="34" pageId="5">42.9±3.3</td>
<td box="[1350,1433,1177,1196]" gridcol="7" gridrow="34" pageId="5">40.2-49.8</td>
</tr>
<tr box="[156,1433,1201,1220]" gridrow="35" pageId="5">
<td box="[156,252,1201,1220]" gridcol="0" gridrow="35" pageId="5">21-22</td>
<td box="[278,533,1201,1220]" gridcol="1" gridrow="35" pageId="5">Mouth width</td>
<td box="[676,782,1201,1220]" gridcol="2" gridrow="35" pageId="5">46.7±3.1</td>
<td box="[827,910,1201,1220]" gridcol="3" gridrow="35" pageId="5">42.8-51.3</td>
<td box="[955,1043,1201,1220]" gridcol="4" gridrow="35" pageId="5">45.4±3.1</td>
<td box="[1089,1171,1201,1220]" gridcol="5" gridrow="35" pageId="5">41.1-48.1</td>
<td box="[1216,1305,1201,1220]" gridcol="6" gridrow="35" pageId="5">47.8±2.4</td>
<td box="[1350,1433,1201,1220]" gridcol="7" gridrow="35" pageId="5">44.1-50.7</td>
</tr>
<tr box="[156,1433,1226,1245]" gridrow="36" pageId="5">
<td box="[156,252,1226,1245]" gridcol="0" gridrow="36" pageId="5">22-23</td>
<td box="[278,533,1226,1245]" gridcol="1" gridrow="36" pageId="5">Barbel length</td>
<td box="[676,782,1226,1245]" gridcol="2" gridrow="36" pageId="5">5.9±1.3</td>
<td box="[827,910,1226,1245]" gridcol="3" gridrow="36" pageId="5">4.3-7.8</td>
<td box="[955,1043,1226,1245]" gridcol="4" gridrow="36" pageId="5">1.3±1.5</td>
<td box="[1089,1171,1226,1245]" gridcol="5" gridrow="36" pageId="5">0.0-2.9</td>
<td box="[1216,1305,1226,1245]" gridcol="6" gridrow="36" pageId="5">2.1±0.5</td>
<td box="[1350,1433,1226,1245]" gridcol="7" gridrow="36" pageId="5">1.3-2.8</td>
</tr>
<tr box="[156,1433,1251,1270]" gridrow="37" pageId="5">
<td box="[156,252,1251,1270]" gridcol="0" gridrow="37" pageId="5">25-26</td>
<td box="[278,533,1251,1270]" gridcol="1" gridrow="37" pageId="5">Dentary tooth cup length</td>
<td box="[676,782,1251,1270]" gridcol="2" gridrow="37" pageId="5">13.6±0.7</td>
<td box="[827,910,1251,1270]" gridcol="3" gridrow="37" pageId="5">12.8-14.3</td>
<td box="[955,1043,1251,1270]" gridcol="4" gridrow="37" pageId="5">15.4±1.6</td>
<td box="[1089,1171,1251,1270]" gridcol="5" gridrow="37" pageId="5">13.6-17.5</td>
<td box="[1216,1305,1251,1270]" gridcol="6" gridrow="37" pageId="5">13.9±0.7</td>
<td box="[1350,1433,1251,1270]" gridcol="7" gridrow="37" pageId="5">13.1-15.3</td>
</tr>
<tr box="[156,1433,1275,1294]" gridrow="38" pageId="5">
<td box="[156,252,1275,1294]" gridcol="0" gridrow="38" pageId="5">27-28</td>
<td box="[278,533,1275,1294]" gridcol="1" gridrow="38" pageId="5">Premaxillary tooth cup length</td>
<td box="[676,782,1275,1294]" gridcol="2" gridrow="38" pageId="5">15.2±1.0</td>
<td box="[827,910,1275,1294]" gridcol="3" gridrow="38" pageId="5">14.0-16.5</td>
<td box="[955,1043,1275,1294]" gridcol="4" gridrow="38" pageId="5">14.9±1.8</td>
<td box="[1089,1171,1275,1294]" gridcol="5" gridrow="38" pageId="5">13.5-17.4</td>
<td box="[1216,1305,1275,1294]" gridcol="6" gridrow="38" pageId="5">13.0±1.3</td>
<td box="[1350,1433,1275,1294]" gridcol="7" gridrow="38" pageId="5">11.9-15.9</td>
</tr>
</table>
</paragraph>
<subSubSection lastPageId="7" pageId="5" type="description">
<paragraph blockId="5.[129,771,1828,2043]" lastBlockId="5.[809,1451,1348,2043]" pageId="5">
<emphasis bold="true" box="[129,269,1828,1851]" pageId="5">Description.</emphasis>
Morphometrics presented in
<tableCitation box="[580,659,1828,1851]" captionStart="Table 1" captionStartId="3.[129,191,261,284]" captionText="Table 1. Morphometric features of Neblinichthys brevibracchium, new species (N = 11) and N. echinasus, new species (N = 3). Landmarks are the numbered landmarks inArmbruster (2003)." pageId="5">Table 1</tableCitation>
. Meristics based on three known specimens, largest
<quantity box="[569,664,1860,1883]" metricMagnitude="-2" metricUnit="m" metricValue="8.29" pageId="5" unit="mm" value="82.9">82.9 mm</quantity>
SL. Body dorsoventrally flattened. Profile from tip of snout sloping posterodorsally to nares, slightly concave between eyes to posterior margin of supraoccipital; gently convex to dorsalfin origin. From deepest point of body (just anterior to dorsalfin origin), profile slopes gently ventrally in straight line to base of caudal-fin spine. Ventral surface flat or slightly concave to anal-fin origin then straight or slightly concave to base of lower caudal-fin spine. Eyes widely separated, almost completely dorsally oriented, not visible from below. In dorsal profile, head margin forms broad
<emphasis box="[1169,1188,1476,1499]" italics="true" pageId="5">U</emphasis>
-shape. Body widest just behind pectoral-fin insertion, not narrowing until reaching pelvic-fin insertions, and then tapering to end of caudal peduncle.
</paragraph>
<paragraph blockId="5.[809,1451,1348,2043]" pageId="5">Cheek plates strongly evertible (to 90° from head), with very numerous elongate as well as even more numerous short and medium sized odontodes, longest reaching base of second or third pectoral-fin ray. Snout with slightly rounded ridges from eye to just anterior to nares. Mesethmoid forming rounded ridge along midline anterior to nares. Dorsal rim of orbit slightly higher than interorbital space. Iris operculum absent, making dorsal rim of pupil round.</paragraph>
<paragraph blockId="5.[809,1451,1348,2043]" pageId="5">Mouth moderate in size with narrow premaxillary and dentary tooth cups forming gentle arcs. Premaxillary teeth 34-59; dentary teeth 33-62. Teeth villiform and bicuspid with very short cusps (medial cusp longer than lateral cusp). Edge of oral disk scalloped, and extending to vertical through anterior margin of eye, not extending beyond lateral margins of head. Enlarged</paragraph>
<paragraph blockId="6.[443,1458,181,205]" box="[443,1458,181,205]" pageId="6">
<emphasis box="[443,1143,181,202]" italics="true" pageId="6">D. C. Taphorn, J. W. Armbruster, H. López-Fernández &amp; C. R. Bernard</emphasis>
621
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647458" ID-Zenodo-Dep="4647458" httpUri="https://zenodo.org/record/4647458/files/figure.png" pageId="6" startId="6.[136,176,1712,1735]" targetBox="[136,1458,257,1701]" targetPageId="6">
<paragraph blockId="6.[136,1458,1712,1767]" pageId="6">
<emphasis bold="true" box="[136,208,1712,1735]" pageId="6">Fig. 3.</emphasis>
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[213,479,1712,1735]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="6" phylum="Chordata" rank="species" species="echinasus">
<emphasis box="[213,479,1712,1735]" italics="true" pageId="6">Neblinichthys echinasus</emphasis>
</taxonomicName>
, holotype, CSBD 1652, 82.9 mm SL, Kukui River, near town of Philipai, upper Mazaruni River drainage.
</paragraph>
</caption>
<paragraph blockId="6.[136,779,1796,2043]" pageId="6">central papilla present in buccal cavity. Maxillary barbel very short, almost completely adnate. Ventral surface of lips papillose. Papillae largest in center of lower lip, decreasing in size radially. No enlarged papillae located behind dentary teeth.</paragraph>
<paragraph blockId="6.[136,779,1796,2043]" lastBlockId="7.[129,1098,180,205]" lastPageId="7" pageId="6">
Dorsal fin II,7; dorsal-fin spinelet visible as flat platelet with short odontodes; dorsal-fin lock functional. Dorsal fin short, just reaching preadipose plate when adpressed. First dorsal-fin ray slightly longer than dorsal-fin spine. Pectoral fin I,6; pectoral-fin spine short (slightly shorter or equal to pelvic-fin spine) not reaching pelvic fin when adpressed. Pectoral-fin spine relatively weak with odontodes numerous and of equal size along entire length. Anterior pectoral-fin rays longer than pectoral-fin spine, decreasing to about half of length of spine posteriorly, covered with odontodes that are largest on dorsal margin. Pelvic fin I,5; pelvic-fin spine thickened and covered with odontodes, reaching middle of 622 Description of
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[634,928,180,201]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="7" phylum="Chordata" rank="species" species="brevibracchium">
<emphasis box="[634,928,180,201]" italics="true" pageId="7">Neblinichthys brevibracchium</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Taphorn &amp; Armbruster &amp; López-Fernández &amp; Bernard" authorityYear="2010" box="[974,1098,180,201]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="7" phylum="Chordata" rank="species" species="echinasus">
<emphasis box="[974,990,180,201]" italics="true" pageId="7">N</emphasis>
.
<emphasis box="[1002,1098,180,201]" italics="true" pageId="7">echinasus</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="7.[809,1452,261,508]" pageId="7">
<typeStatus box="[809,907,261,284]" pageId="7">holotype</typeStatus>
dark brown with bold white to cream colored vermiculations and spots. Ventrum of
<typeStatus box="[1198,1299,293,316]" pageId="7">paratypes</typeStatus>
plain tan. Oral disk with inner papillated surfaces pale tan but brown with white spots on outer anterior margin. All fin spines and rays with alternating wide dark and narrow light bands (pattern most evident in caudal and dorsal fins, least evident in pectorals); fin membranes dark gray in
<typeStatus box="[1119,1214,453,476]" pageId="7">holotype</typeStatus>
, hyaline in
<typeStatus box="[1343,1448,453,476]" pageId="7">paratypes</typeStatus>
. Base of anteriormost dorsal-fin membrane black (
<figureCitation box="[1315,1377,485,508]" captionStart="Fig" captionStartId="6.[136,176,1712,1735]" captionTargetBox="[136,1458,257,1701]" captionTargetId="figure-174@6.[136,1459,257,1702]" captionTargetPageId="6" captionText="Fig. 3. Neblinichthys echinasus, holotype, CSBD 1652, 82.9 mm SL, Kukui River, near town of Philipai, upper Mazaruni River drainage." figureDoi="http://doi.org/10.5281/zenodo.4647458" httpUri="https://zenodo.org/record/4647458/files/figure.png" pageId="7">Fig. 3</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4647460" ID-Zenodo-Dep="4647460" httpUri="https://zenodo.org/record/4647460/files/figure.png" pageId="7" startId="7.[129,168,656,679]" targetBox="[128,770,257,640]" targetPageId="7">
<paragraph blockId="7.[129,771,656,711]" pageId="7">
<emphasis bold="true" box="[129,200,656,679]" pageId="7">Fig. 4.</emphasis>
<taxonomicName authorityName="Ferraris, Isbrucker &amp; Nijssen" authorityYear="1986" box="[209,357,656,679]" class="Actinopterygii" family="Loricariidae" genus="Neblinichthys" kingdom="Animalia" order="Siluriformes" pageId="7" phylum="Chordata" rank="genus">
<emphasis box="[209,357,656,679]" italics="true" pageId="7">Neblinichthys</emphasis>
</taxonomicName>
habitat in the Kukui River at Philipai 05º2137”N 60º2218”W.
</paragraph>
</caption>
<paragraph blockId="7.[129,772,741,1308]" pageId="7">base of anal fin when adpressed; anterior pelvic-fin rays longer than pelvic-fin spine with posterior margin of fin curving out beyond posterior tip of spine. Anal fin I,5; all anal-fin rays longer than weak spine. First anal-fin pterygiophore not exposed to form plate-like structure. Adipose-fin spine straight to slightly curved posteriorly with adipose membrane extending beyond posterior extent of spine. Caudal fin I,14,I; upper caudal-fin spine shorter than first two upper rays, lower spine longer than all rays, and much longer than upper spine. Dorsal procurrent caudal-fin rays five, ventral procurrent caudal-fin rays three. Posterior caudal-fin margin oblique or slightly concave. Rays of all fins supporting small odontodes.</paragraph>
<paragraph blockId="7.[129,772,741,1308]" pageId="7">Median plate series with 21-23 plates. Ventral plates forming gentle arc on caudal peduncle and not forming strong rounded keel. Three rows of plates on caudal peduncle (middorsal and mid-ventral plate series ending at level of adipose fin). Abdomen naked.</paragraph>
<paragraph blockId="7.[129,771,1349,1499]" lastBlockId="7.[809,1452,549,1020]" pageId="7">
<emphasis bold="true" box="[129,366,1349,1372]" pageId="7">Coloration in alcohol.</emphasis>
Head, dorsum of body and sides mostly brown with irregular lighter tan blotches.
<typeStatus box="[610,712,1380,1403]" pageId="7">Holotype</typeStatus>
dark brownish gray with horizontal rows of light spots from behind pectoral-fin insertion to caudal fin on sides. Largest
<typeStatus box="[680,771,1444,1467]" pageId="7">paratype</typeStatus>
with alternating light and dark brown patches. Ventrum of
<emphasis bold="true" box="[809,1057,549,572]" pageId="7">Sexual dimorphism.</emphasis>
Presumed males have extremely hypertrophied odontodes along the entire edge of the snout with the odontodes becoming slightly shorter posteriorly. just prior to the evertible cheek plates (
<figureCitation box="[1219,1283,645,668]" captionStart="Fig" captionStartId="6.[136,176,1712,1735]" captionTargetBox="[136,1458,257,1701]" captionTargetId="figure-174@6.[136,1459,257,1702]" captionTargetPageId="6" captionText="Fig. 3. Neblinichthys echinasus, holotype, CSBD 1652, 82.9 mm SL, Kukui River, near town of Philipai, upper Mazaruni River drainage." figureDoi="http://doi.org/10.5281/zenodo.4647458" httpUri="https://zenodo.org/record/4647458/files/figure.png" pageId="7">Fig. 3</figureCitation>
). Three rows of hypertrophied odontodes on present on the snout, the medial row is relatively short and located on the mesethmoid; the lateral rows are also relatively short, extending from eyes to nares and then intermediate from nares to snout. The areas between the snout edge and the rows of odontodes have some hypertrophied odontodes, but not as densely packed. The remainder of the dorsal and lateral surfaces of the head have slightly elongated odontodes. Odontodes on the lateral plates are very slightly lengthened in nuptial males. No hypertrophied odontodes are present on the pectoral-fin spines. Presumed males have bolder marking on the abdomen.
</paragraph>
</subSubSection>
<subSubSection pageId="7" type="distribution">
<paragraph blockId="7.[809,1451,1061,1116]" pageId="7">
<emphasis bold="true" box="[809,957,1061,1084]" pageId="7">Distribution.</emphasis>
Known from upper Mazaruni River drainage,
<collectingCountry box="[809,891,1093,1116]" name="Guyana" pageId="7">Guyana</collectingCountry>
(
<figureCitation box="[903,966,1093,1116]" captionStart="Fig" captionStartId="4.[816,853,1892,1915]" captionTargetBox="[817,1450,1370,1856]" captionTargetId="figure-745@4.[816,1458,1369,1866]" captionTargetPageId="4" captionText="Fig. 2. Distribution of Neblinichthys species in the Caroní and Mazarunibasins of VenezuelaandGuyana,respectively.Star = N. roraima,black circle=N.yaravi,empty circle=type-localityof N. echinasus, new species, empty square = type-locality and black squares = paratype localities of N. brevibracchium, new species." figureDoi="http://doi.org/10.5281/zenodo.4647456" httpUri="https://zenodo.org/record/4647456/files/figure.png" pageId="7">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" type="etymology">
<paragraph blockId="7.[809,1451,1157,1276]" pageId="7">
<emphasis bold="true" box="[809,934,1157,1180]" pageId="7">Etymology.</emphasis>
From the Latin words
<emphasis box="[1164,1244,1157,1180]" italics="true" pageId="7">echinus</emphasis>
meaning sea-urchin or prickly and
<emphasis box="[985,1048,1189,1212]" italics="true" pageId="7">nasus</emphasis>
meaning nose, in reference to the numerous odontodes that adorn males of this species. Epithet used as a noun in apposition, in masculine form.
</paragraph>
</subSubSection>
</treatment>
</document>