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<document id="1AB3288E1C390EB42AAF6397662AB3FA" ID-CLB-Dataset="20391" ID-DOI="10.11646/zootaxa.5104.3.8" ID-GBIF-Dataset="75c438ea-b4fb-4f34-af3a-5c0872c2062d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332138" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646595153051" checkinUser="plazi" docAuthor="Ahrens, Dirk &amp; Lukic, Daniel" docDate="2022" docId="03D126609828DC2D58EB3D2EFA14BEA0" docLanguage="en" docName="zootaxa.5104.3.8.pdf" docOrigin="Zootaxa 5104 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoserica Ahrens &amp; Lukic, 2022, new species" docType="treatment" docVersion="6" lastPageNumber="443" masterDocId="FFE85E189828DC2F587C3B64FF87BD4B" masterDocTitle="Neoserica (s. str.) phuphami-a further new Neoserica species from Vietnam with highly modified pronotum (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="444" masterPageNumber="441" pageNumber="441" updateTime="1699316923957" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="099BFAF5F6CE7611A9CD0F49D6F50B96" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="915BE3D7EC67D7CA7D13B0EAA4E3FA01">
<mods:title id="2007D488EF23EEB2A25E564049391D45">Neoserica (s. str.) phuphami-a further new Neoserica species from Vietnam with highly modified pronotum (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="6B3175E1166CB4344EB7A32F7E5018ED" type="personal">
<mods:role id="F8A4955FB336577445EBEAE6892DB057">
<mods:roleTerm id="1BEA2C9854A365EC81A632BDA3DAF88E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAC4ED23CEC070033162D4B1CD156F07">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="2B2786BA79CB3ECC56137A5E24943FD7" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:affiliation id="FC1C8FE4BEA6473007878109692610A8">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany. &amp; ahrens. dirk _ col @ gmx. de; https: // orcid. org / 0000 - 0003 - 3524 - 7153</mods:affiliation>
<mods:nameIdentifier id="F798E7E40FF07C9D5DE6900790CFD502" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="6E695D9271B889B56C9C817990EA697B" type="personal">
<mods:role id="0B204569BE736AD4EECCB5B3FEFE0A80">
<mods:roleTerm id="07253056B51DED9649E7908300DDC318">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CFD7CEAAB7BFF1B7AC84DAACF53F2D2A">Lukic, Daniel</mods:namePart>
<mods:nameIdentifier id="686C3DE73C5A8A9DB0F6572110AEBDE1" type="ORCID">0000-0001-9571-3481</mods:nameIdentifier>
<mods:affiliation id="281487A3DBC585AA3D84B2D610CAF401">Zoologisches Forschungsmuseum A. Koenig, Adenauerallee 127, 53113 Bonn, Germany. &amp; Daniel. Lukic @ ruhr-uni-bochum. de; https: // orcid. org / 0000 - 0001 - 9571 - 3481</mods:affiliation>
<mods:nameIdentifier id="65E985741AD93FBB46AC852124684595" type="email">ukic@ruhr-uni-bochum.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7AC17A6DA35165A18F40DEF9525B8496">text</mods:typeOfResource>
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<mods:titleInfo id="07FBFFA37C799E58A592CFF81A3083AC">
<mods:title id="1FB36C6A4D91239A29EA472EC9DFA7E9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="63833EC1805F45C63E94722D3C6B32CF">
<mods:date id="1076CFAAA28376D67C607F21B8425114">2022</mods:date>
<mods:detail id="EFE36762114B184F41BA7B1783DAB029" type="pubDate">
<mods:number id="52FAB3E6B4D57FC652981C85435A4713">2022-02-28</mods:number>
</mods:detail>
<mods:detail id="D7278EE746F0898E54F724C7FF8C496D" type="volume">
<mods:number id="7369D502BC1D0CDA1BF62253B3B9BCFD">5104</mods:number>
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<mods:detail id="E3A0EC4937B3BD645626DEBFD6CEBDD9" type="issue">
<mods:number id="723DFBB80DE3D40735BF95947E6F04B2">3</mods:number>
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<mods:classification id="35F7780614E05BD06C1EBE665E8B6F2C">journal article</mods:classification>
<mods:identifier id="315D9E03F653FDC4E09A899A82B3F673" type="CLB-Dataset">20391</mods:identifier>
<mods:identifier id="7B3AB42E9F057F5437000F84EE3A2C86" type="DOI">10.11646/zootaxa.5104.3.8</mods:identifier>
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<mods:identifier id="5B186FA94D82E9EDC2E0F23286BA573A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0E92359A05052A30370D775BA0EBFCAB" type="ZooBank">600D57B7-5D25-4942-BA4A-753ECF2C5F7D</mods:identifier>
</mods:mods>
<treatment id="03D126609828DC2D58EB3D2EFA14BEA0" ID-DOI="http://doi.org/10.5281/zenodo.6333206" ID-GBIF-Taxon="193876449" ID-Zenodo-Dep="6333206" LSID="urn:lsid:plazi:treatment:03D126609828DC2D58EB3D2EFA14BEA0" httpUri="http://treatment.plazi.org/id/03D126609828DC2D58EB3D2EFA14BEA0" lastPageId="2" lastPageNumber="443" pageId="0" pageNumber="441">
<subSubSection id="C362C4FD9828DC2F58EB3D2EFCD9BB2F" box="[151,862,1609,1636]" pageId="0" pageNumber="441" type="nomenclature">
<paragraph id="8BC797769828DC2F58EB3D2EFCD9BB2F" blockId="0.[151,862,1609,1672]" box="[151,862,1609,1636]" pageId="0" pageNumber="441">
<heading id="D08F201A9828DC2F58EB3D2EFCD9BB2F" bold="true" box="[151,862,1609,1636]" fontSize="11" level="1" pageId="0" pageNumber="441" reason="1">
<emphasis id="B90C4B649828DC2F58EB3D2EFCD9BB2F" bold="true" box="[151,862,1609,1636]" pageId="0" pageNumber="441">
<taxonomicName id="4C78ECF59828DC2F58EB3D2EFE97BB2F" authorityName="Brenske" authorityYear="1894" box="[151,272,1610,1636]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="441" phylum="Arthropoda" rank="genus">
<emphasis id="B90C4B649828DC2F58EB3D2EFE97BB2F" bold="true" box="[151,272,1610,1636]" italics="true" pageId="0" pageNumber="441">Neoserica</emphasis>
</taxonomicName>
(s. str.)
<emphasis id="B90C4B649828DC2F590F3D2DFE68BB2F" bold="true" box="[371,495,1609,1636]" italics="true" pageId="0" pageNumber="441">phuphami</emphasis>
Ahrens &amp; Lukic,
<taxonomicNameLabel id="A23FF61F9828DC2F5AB23D2EFCD9BB2F" box="[718,862,1610,1636]" pageId="0" pageNumber="441" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C362C4FD9828DC2F58EB3D14FE8DBBC3" box="[151,266,1648,1672]" pageId="0" pageNumber="441" type="description">
<paragraph id="8BC797769828DC2F58EB3D14FE8DBBC3" blockId="0.[151,862,1609,1672]" box="[151,266,1648,1672]" pageId="0" pageNumber="441">
<figureCitation id="13438BF39828DC2F58EB3D14FE8DBBC3" box="[151,266,1648,1672]" captionStart="FIGURE 1" captionStartId="1.[151,250,1910,1934]" captionTargetBox="[162,1424,748,1871]" captionTargetId="figure-262@1.[151,1436,739,1885]" captionTargetPageId="1" captionText="FIGURE 1A–G. Neoserica (s. str.) phuphami Ahrens &amp; Lukic, new species (holotype). A, aedeagus, left side lateral view; B, C, parameres, dorsal view; D, aedeagus, right side lateral view; E, habitus, dorsal view; F, habitus, lateral view; G, pronotum and head, dorsal view. Scale (A–D): 0.5 mm, E–G: not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332140" httpUri="https://zenodo.org/record/6332140/files/figure.png" pageId="0" pageNumber="441">Fig. 1A–G</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C362C4FD9828DC2F58EB3DDCFC4BBBBF" pageId="0" pageNumber="441" type="materials_examined">
<paragraph id="8BC797769828DC2F58EB3DDCFC4BBBBF" blockId="0.[151,1436,1719,2032]" pageId="0" pageNumber="441">
<emphasis id="B90C4B649828DC2F58EB3DDCFE33BB84" bold="true" box="[151,436,1719,1744]" inLineHeading="true" pageId="0" pageNumber="441" reason="1">Type material examined.</emphasis>
<materialsCitation id="3B109D2B9828DC2F59C73DDCFC4BBBBF" ID-GBIF-Occurrence="3696833301" collectingDate="2017-04-05" collectingDateMax="2017-04-07" collectingDateMin="2017-04-05" collectionCode="R, ISNB" collectorName="J. Constant &amp; J. Bresseel" country="Vietnam" latitude="20.348057" location="Cuc Phong Nat. Park" longLatPrecision="21" longitude="105.59778" pageId="0" pageNumber="441" specimenCount="1" specimenCount-male="1" stateProvince="Ninh Binh" typeStatus="holotype">
<typeStatus id="54C329D49828DC2F59C73DDCFDA6BB9B" box="[443,545,1720,1744]" pageId="0" pageNumber="441" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D7E5CFF9828DC2F5A513DDCFDC3BB84" box="[557,580,1720,1743]" pageId="0" pageNumber="441" type="male">♂</specimenCount>
: “Coll. I.
<collectionCode id="ED690FB39828DC2F5ADA3DDCFD3BBB9B" box="[678,700,1720,1744]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="0" pageNumber="441">R</collectionCode>
.Sc.N.B.
<collectingCountry id="F36FD7E69828DC2F5B673DDCFCF2BB9B" box="[795,885,1720,1744]" name="Vietnam" pageId="0" pageNumber="441">Vietnam</collectingCountry>
<collectingRegion id="49BC59949828DC2F5B013DDCFBAABB84" box="[893,1069,1720,1744]" country="Vietnam" name="Ninh Binh" pageId="0" pageNumber="441">Ninh Binh prov.</collectingRegion>
<location id="8EA7C1AD9828DC2F5C4F3DDCFA9FBB9B" LSID="urn:lsid:plazi:treatment:03D126609828DC2D58EB3D2EFA14BEA0:8EA7C1AD9828DC2F5C4F3DDCFA9FBB9B" box="[1075,1304,1720,1744]" country="Vietnam" latitude="20.348057" longLatPrecision="21" longitude="105.59778" name="Cuc Phong Nat. Park" pageId="0" pageNumber="441" stateProvince="Ninh Binh">Cuc Phong Nat. Park</location>
<geoCoordinate id="EE4CF1B19828DC2F5D5C3DDCFA1BBB9B" box="[1312,1436,1720,1744]" degrees="20" direction="north" minutes="20" orientation="latitude" pageId="0" pageNumber="441" precision="15" seconds="53" value="20.348057">20°20'53''N</geoCoordinate>
<geoCoordinate id="EE4CF1B19828DC2F58EB3DB8FE99BBBF" box="[151,286,1756,1780]" degrees="105" direction="east" minutes="35" orientation="longitude" pageId="0" pageNumber="441" precision="15" seconds="52" value="105.59778">105°35'52''E</geoCoordinate>
<date id="FFC6B1B69828DC2F59593DB8FE19BBBF" box="[293,414,1756,1780]" pageId="0" pageNumber="441" value="2017-04-05" valueMax="2017-04-07" valueMin="2017-04-05">
<collectingDate id="EF82485E9828DC2F59593DB8FE19BBBF" box="[293,414,1756,1780]" pageId="0" pageNumber="441" value="2017-04-05" valueMax="2017-04-07" valueMin="2017-04-05">5-7.iv.2017</collectingDate>
</date>
leg.
<collectorName id="268DF2A09828DC2F59AE3DB8FDCEBBBF" box="[466,585,1756,1780]" pageId="0" pageNumber="441">J. Constant</collectorName>
&amp;
<collectorName id="268DF2A09828DC2F5A173DB8FD67BBBF" box="[619,736,1756,1780]" pageId="0" pageNumber="441">J. Bresseel</collectorName>
, I.G. 33.447” (
<collectionCode id="ED690FB39828DC2F5B023DB8FC47BBBF" box="[894,960,1756,1780]" pageId="0" pageNumber="441">ISNB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C362C4FD9828DC2D58BB3D9BFCDCBE58" lastPageId="2" lastPageNumber="443" pageId="0" pageNumber="441" type="description">
<paragraph id="8BC797769828DC2F58BB3D9BFB2DBA2B" blockId="0.[151,1436,1719,2032]" pageId="0" pageNumber="441">
<emphasis id="B90C4B649828DC2F58BB3D9BFED5BA5C" bold="true" box="[199,338,1791,1815]" inLineHeading="true" pageId="0" pageNumber="441" reason="1">Description.</emphasis>
Body length:
<quantity id="4C803A939828DC2F59973C64FDB8BA5C" box="[491,575,1792,1816]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="0" pageNumber="441" unit="mm" value="7.3">7.3 mm</quantity>
, length of elytra:
<quantity id="4C803A939828DC2F5A813C64FCD6BA5C" box="[765,849,1792,1816]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="0" pageNumber="441" unit="mm" value="4.8">4.8 mm</quantity>
, width:
<quantity id="4C803A939828DC2F5BDB3C64FC7ABA5C" box="[935,1021,1792,1816]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="0" pageNumber="441" unit="mm" value="4.4">4.4 mm</quantity>
. Body oval, reddish brown, dorsal surface with greenish shine, elytra before apex with a large, round, black spot, with dense, fine, yellowish, adpressed to erect setae, mixed with dense, longer, erect, yellow setae, which are all over the body directed anteriad.
</paragraph>
<paragraph id="8BC797769828DC2E58BB3C08FE72BC74" blockId="0.[151,1436,1719,2032]" lastBlockId="1.[151,1437,151,715]" lastPageId="1" lastPageNumber="442" pageId="0" pageNumber="441">Labroclypeus subrectangular, widest at base, lateral margins moderately convex and convergent to broadly rounded anterior angles, lateral border and ocular canthus producing a distinct obtuse angle; anterior margin concavely emarginated and moderately reflexed; surface weakly convex and shiny, coarsely and finely, densely punctate, with dense long erect setae; frontoclypeal suture finely impressed and moderately bent medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately short and robust, finely and densely punctate, with several short setae. Frons shiny, posterior half dull, finely and coarsely, densely punctate, with dense, long, erect yellow setae, and fine short adpressed ones. Eyes small, ratio of diameter/ interocular width: 0.53. Antenna with ten antennomeres; club with four antennomeres, 1.2 times as long as remaining antennomeres combined, first joint of club subequal club length. Mentum elevated and flattened anteriorly.</paragraph>
<paragraph id="8BC797769829DC2E58BB3A28FE75BF70" blockId="1.[151,1437,151,715]" pageId="1" pageNumber="442">Pronotum moderately wide, widest at base, lateral margins in basal half deeply concavely excavate, strongly curved and convergent in anterior half, anterior angles not produced and blunt, weakly rounded in tip, posterior angles acute; anterior margin straight, with indistinct and widely lacking marginal line; basal margin without marginal line; surface with fine and dense punctures each bearing a short, yellowish, seta interspersed with coarser punctures bearing each a long, erect, yellow seta, along midline punctures less dense; anterior and lateral borders sparsely setose; hypomeron carinate, basal margin of hypomeron strongly produced ventrally. Scutellum triangular, apex sharp, with fine and dense punctures and fine, dense, adpressed setae.</paragraph>
<paragraph id="8BC797769829DC2E58BB3923FB00BF80" blockId="1.[151,1437,151,715]" pageId="1" pageNumber="442">Elytra oval, widest at middle, striae distinctly impressed, with fine and dense punctures; intervals weakly convex, with punctures concentrated along striae, fine punctures bearing fine, short setae mixed with larger ones bearing long erect setae; epipleural edge moderately strong, ending at strongly rounded external apical angle of elytra, epipleura densely setose, apical border narrowly membraneous, with a fine rim of short microtrichomes.</paragraph>
<caption id="DF07C7FE9829DC2E58EB3C12FC9BBA9D" ID-DOI="http://doi.org/10.5281/zenodo.6332140" ID-Zenodo-Dep="6332140" httpUri="https://zenodo.org/record/6332140/files/figure.png" pageId="1" pageNumber="442" startId="1.[151,250,1910,1934]" targetBox="[162,1424,748,1871]" targetPageId="1">
<paragraph id="8BC797769829DC2E58EB3C12FC9BBA9D" blockId="1.[151,1437,1910,2006]" pageId="1" pageNumber="442">
<emphasis id="B90C4B649829DC2E58EB3C12FEC1BAC5" bold="true" box="[151,326,1910,1934]" pageId="1" pageNumber="442">FIGURE 1A–G.</emphasis>
<taxonomicName id="4C78ECF59829DC2E59313C13FE33BAC5" authorityName="Brenske" authorityYear="1894" box="[333,436,1911,1934]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B90C4B649829DC2E59313C13FE33BAC5" box="[333,436,1911,1934]" italics="true" pageId="1" pageNumber="442">Neoserica</emphasis>
</taxonomicName>
(s. str.)
<emphasis id="B90C4B649829DC2E5A743C12FDEABAC5" box="[520,621,1910,1934]" italics="true" pageId="1" pageNumber="442">phuphami</emphasis>
Ahrens &amp; Lukic,
<taxonomicNameLabel id="A23FF61F9829DC2E5B5A3C12FC27BAC5" box="[806,928,1910,1934]" pageId="1" pageNumber="442" rank="species">new species</taxonomicNameLabel>
(holotype).
<emphasis id="B90C4B649829DC2E5C623C12FBB0BAC5" bold="true" box="[1054,1079,1910,1934]" pageId="1" pageNumber="442">A,</emphasis>
aedeagus, left side lateral view;
<emphasis id="B90C4B649829DC2E5DF93C12FF37BAF9" bold="true" pageId="1" pageNumber="442">B, C,</emphasis>
parameres, dorsal view;
<emphasis id="B90C4B649829DC2E59CD3CFEFE4EBAF9" bold="true" box="[433,457,1946,1970]" pageId="1" pageNumber="442">D,</emphasis>
aedeagus, right side lateral view;
<emphasis id="B90C4B649829DC2E5B543CFEFCB8BAF9" bold="true" box="[808,831,1946,1970]" pageId="1" pageNumber="442">E,</emphasis>
habitus, dorsal view;
<emphasis id="B90C4B649829DC2E5C5D3CFEFBB2BAF9" bold="true" box="[1057,1077,1946,1970]" pageId="1" pageNumber="442">F,</emphasis>
habitus, lateral view;
<emphasis id="B90C4B649829DC2E5D653CFEFAB5BAF9" bold="true" box="[1305,1330,1946,1970]" pageId="1" pageNumber="442">G,</emphasis>
pronotum and head, dorsal view. Scale (A
<emphasis id="B90C4B649829DC2E59AB3CDAFE63BA9D" bold="true" box="[471,484,1982,2006]" pageId="1" pageNumber="442">–</emphasis>
D): 0.5 mm, E
<emphasis id="B90C4B649829DC2E5A093CDAFD05BA9D" bold="true" box="[629,642,1982,2006]" pageId="1" pageNumber="442">–</emphasis>
G: not to scale.
</paragraph>
</caption>
<paragraph id="8BC79776982ADC2D58BB3BF3FDB6BC28" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="443">Ventral surface dull, including metacoxa with large, dense punctures and dense, short, light adpressed setae; metacoxa additionally with some robust setae laterally. Abdominal sternites finely and densely punctate and setose, each sternite with an indistinct transverse row of coarse punctures each bearing a short, robust seta. Mesosternum between mesocoxae slightly wider than mesofemur, with numerous strong setae on an indistinct semi-circular carina. Ratio of length of metepisternum/ metacoxa: 1/1.62. Pygidium weakly convex, with coarse, dense punctures and dense longer and shorter setae, without smooth midline.</paragraph>
<paragraph id="8BC79776982ADC2D58BB3A14FB3EBFA4" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="443">Legs moderately slender, femora with fine and dense punctures, densely setose; anterior edge of metafemur acute lacking an adjacent serrated line, posterior margin weakly convex, with a few fine setae medially, ventral posterior margin weakly widened in apical half and finely serrate, dorsal posterior margin serrate. Metatibia moderately long, widest at middle, ratio width/ length: 1/3.41; dorsal margin sharply carinate, with only one groups of spines at three quarters of metatibial length, with a few single robust spines in punctures along entire length of metatibia, beside dorsal margin with a blunt, distinctly serrate carina from base to apex; lateral face longitudinally convex, finely and densely punctate and setose; ventral margin serrate, with five equidistant robust spines; medial face impunctate and glabrous, apex interiorly near tarsal articulation concavely truncate. Tarsomeres glabrous and impunctate dorsally, with sparse, short setae ventrally; metatarsomeres without longitudinal impressions dorsally, with a strongly serrated ridge ventrally and a strong longitudinal carina laterally; first metatarsomere twice as long as dorsal tibial spur and as long as following two tarsomeres combined. Protibia short, bidentate, anterior claws symmetrical, with a sharply pointed basal tooth.</paragraph>
<paragraph id="8BC79776982ADC2D58BB399FFCDCBE58" blockId="2.[151,1437,151,1004]" box="[199,859,763,788]" pageId="2" pageNumber="443">
Aedeagus:
<figureCitation id="13438BF3982ADC2D5942399FFE34BE5F" box="[318,435,763,788]" captionStart="FIGURE 1" captionStartId="1.[151,250,1910,1934]" captionTargetBox="[162,1424,748,1871]" captionTargetId="figure-262@1.[151,1436,739,1885]" captionTargetPageId="1" captionText="FIGURE 1A–G. Neoserica (s. str.) phuphami Ahrens &amp; Lukic, new species (holotype). A, aedeagus, left side lateral view; B, C, parameres, dorsal view; D, aedeagus, right side lateral view; E, habitus, dorsal view; F, habitus, lateral view; G, pronotum and head, dorsal view. Scale (A–D): 0.5 mm, E–G: not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332140" httpUri="https://zenodo.org/record/6332140/files/figure.png" pageId="2" pageNumber="443">Fig. 1A–D</figureCitation>
. Habitus:
<figureCitation id="13438BF3982ADC2D5A63399FFD16BE5F" box="[543,657,763,788]" captionStart="FIGURE 1" captionStartId="1.[151,250,1910,1934]" captionTargetBox="[162,1424,748,1871]" captionTargetId="figure-262@1.[151,1436,739,1885]" captionTargetPageId="1" captionText="FIGURE 1A–G. Neoserica (s. str.) phuphami Ahrens &amp; Lukic, new species (holotype). A, aedeagus, left side lateral view; B, C, parameres, dorsal view; D, aedeagus, right side lateral view; E, habitus, dorsal view; F, habitus, lateral view; G, pronotum and head, dorsal view. Scale (A–D): 0.5 mm, E–G: not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332140" httpUri="https://zenodo.org/record/6332140/files/figure.png" pageId="2" pageNumber="443">Fig. 1E–G</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C362C4FD982ADC2D58BB387BFE74BE8C" pageId="2" pageNumber="443" type="diagnosis">
<paragraph id="8BC79776982ADC2D58BB387BFE74BE8C" blockId="2.[151,1437,151,1004]" pageId="2" pageNumber="443">
<emphasis id="B90C4B64982ADC2D58BB387BFE44BE7C" bold="true" box="[199,451,799,823]" inLineHeading="true" pageId="2" pageNumber="443" reason="1">Differential diagnosis.</emphasis>
<taxonomicName id="4C78ECF5982ADC2D59B03844FDBFBE7C" authorityName="Brenske" authorityYear="1894" box="[460,568,800,823]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="genus">
<emphasis id="B90C4B64982ADC2D59B03844FDBFBE7C" box="[460,568,800,823]" italics="true" pageId="2" pageNumber="443">Neoserica</emphasis>
</taxonomicName>
(s. str.)
<emphasis id="B90C4B64982ADC2D5AE9387BFC87BE7C" box="[661,768,799,823]" italics="true" pageId="2" pageNumber="443">phuphami</emphasis>
Ahrens &amp; Lukic,
<taxonomicNameLabel id="A23FF61F982ADC2D5BBB3844FBCEBE7C" box="[967,1097,799,823]" pageId="2" pageNumber="443" rank="species">new species</taxonomicNameLabel>
differs from
<taxonomicName id="4C78ECF5982ADC2D5CA03844FACFBE7C" authorityName="Brenske" authorityYear="1894" box="[1244,1352,800,823]" class="Insecta" family="Melolonthidae" genus="Neoserica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="genus">
<emphasis id="B90C4B64982ADC2D5CA03844FACFBE7C" box="[1244,1352,800,823]" italics="true" pageId="2" pageNumber="443">Neoserica</emphasis>
</taxonomicName>
(s. str.)
<emphasis id="B90C4B64982ADC2D58EB3827FF7ABE10" box="[151,253,835,859]" italics="true" pageId="2" pageNumber="443">paradoxa</emphasis>
by the slightly longer antennal club as well as by the shape of parameres and apical apophysis of phallobase: the parameres are more or less straight, at apex convexly rounded and with a blunt preapical lobe; the left apical apophysis of phallobase is bluntly truncate at apex; on the right side there is a short, wide but blunt lateral apophysis as well, which is in
<emphasis id="B90C4B64982ADC2D58B738D4FF63BE8C" box="[203,228,944,967]" italics="true" pageId="2" pageNumber="443">N.</emphasis>
(s. str.)
<emphasis id="B90C4B64982ADC2D594638CBFE27BE8C" box="[314,416,943,967]" italics="true" pageId="2" pageNumber="443">paradoxa</emphasis>
absent.
</paragraph>
</subSubSection>
<subSubSection id="C362C4FD982ADC2D58BB38B0FA14BEA0" box="[199,1427,979,1004]" pageId="2" pageNumber="443" type="etymology">
<paragraph id="8BC79776982ADC2D58BB38B0FA14BEA0" blockId="2.[151,1437,151,1004]" box="[199,1427,979,1004]" pageId="2" pageNumber="443">
<emphasis id="B90C4B64982ADC2D58BB38B0FECFBEA7" bold="true" box="[199,328,980,1004]" inLineHeading="true" pageId="2" pageNumber="443" reason="1">Etymology.</emphasis>
The new species (name noun in genitive case) is name after Phu Pham, student of Vietnamese scarabs.
</paragraph>
</subSubSection>
</treatment>
</document>
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