297 lines
36 KiB
XML
297 lines
36 KiB
XML
<document id="1AF635A47940D24A4A365780A1D217F9" ID-CLB-Dataset="8432" ID-DOI="10.11646/zootaxa.4858.1.2" ID-GBIF-Dataset="fa172759-c7a8-4682-b992-8871c155eb3a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4411485" ID-ZooBank="22886812-4F5E-42E4-B8EA-5687BB483AA8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609608283149" checkinUser="plazi" docAuthor="Perkins, Philip D. & Ribera, Ignacio" docDate="2020" docId="03B1ED22FFEF452E9BB05965FB015EB5" docLanguage="en" docName="zootaxa.4858.1.2.pdf" docOrigin="Zootaxa 4858 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adelphydraena spinosa Perkins & Ribera 2020, new species" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FF88955AFFEB45229B275F35FF835D10" masterDocTitle="Three new species and DNA sequence data of the rare South American water beetle genus Adelphydraena Perkins, 1989 (Coleoptera: Hydraenidae)" masterLastPageNumber="52" masterPageNumber="35" pageNumber="39" updateTime="1698851450175" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="814EB170A3BC097F53BAF7CFE56C8570" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="21C4C2F3DCF39C6E4EF9291C5EF0AF5B">
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<mods:title id="77639822BED4802173B078300E9480DE">Three new species and DNA sequence data of the rare South American water beetle genus Adelphydraena Perkins, 1989 (Coleoptera: Hydraenidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="8D73CB6636E88FA78819408E2B95FE47" type="personal">
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<mods:role id="87C7E82D4A30ED9888E96CB3F28B8F97">
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<mods:roleTerm id="208ABF10A7D64251EA33227BE5B2DC7D">Author</mods:roleTerm>
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<mods:namePart id="1B1C53543E322B819F49BF9112500586">Perkins, Philip D.</mods:namePart>
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<mods:nameIdentifier id="52C0E1B71F1771DFF32264135BBF0A73" type="ORCID">0000-0001-8834-9052</mods:nameIdentifier>
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<mods:affiliation id="5B2EBEDD016CA2013DCD2BC7A1D779DC">Department of Entomology, Museum of Comparative Zoology, Harvard University, Cambridge, MA 02138 USA. perkins @ oeb. harvard. edu; https: // orcid. org / 0000 - 0001 - 8834 - 9052</mods:affiliation>
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<mods:nameIdentifier id="A05591D9977775CCAEF52082D5A11A9F" type="email">perkins@oeb.harvard.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="91C6BD8AC27513F0B897E2A6B4A21BC2" type="personal">
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<mods:role id="ADAAA1DE65528B4D2051FED6F02C7BC3">
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<mods:roleTerm id="2E38BC36399B185760F4EA42FB6BC482">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B40394639310B2C6D150A9389839ED39">Ribera, Ignacio</mods:namePart>
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<mods:affiliation id="86A2D2186F5F1A21F84C35C27FDE3037">Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37 - 49, 08003 Barcelona, Spain. ignacio. ribera @ ibe. upf-csic. es</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="EC82CCB188EF180391D0D6963249E648">text</mods:typeOfResource>
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<mods:titleInfo id="EE5804537D1F3E3BA83647AC3BC6B97D">
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<mods:title id="D742173CBAA4972D1890F5A0263998D8">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="352496D3693668D2F9A4586DF2CF7D07">
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<mods:date id="70C8A205B45014F1ACD8CB16DB058993">2020</mods:date>
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<mods:detail id="448B749C81FAF9D9EEA506D8D19F0112" type="pubDate">
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<mods:number id="7D53A51833C05068D1697C84AD4AB3E1">2020-09-29</mods:number>
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</mods:detail>
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<mods:detail id="1E01CDFA6E77BF62194F902CEA7F5D30" type="volume">
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<mods:number id="0F5C84C68B06051079FB3B816E560E71">4858</mods:number>
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</mods:detail>
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<mods:detail id="FDCCC744BB716888E1D30581715BDDBD" type="issue">
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<mods:number id="256AD5BC40D5BC31D225B060DE8367C4">1</mods:number>
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<mods:start id="D94202C424C83476910343BF477D2879">35</mods:start>
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<mods:classification id="F2C71E23508517C2B7C629F67CA7841D">journal article</mods:classification>
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<mods:identifier id="8E639606BFF2F8015CC325A6F1D5CB7D" type="CLB-Dataset">8432</mods:identifier>
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<mods:identifier id="338F0FA2F2B5184D60437174EE85C184" type="DOI">10.11646/zootaxa.4858.1.2</mods:identifier>
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</mods:mods>
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<treatment id="03B1ED22FFEF452E9BB05965FB015EB5" ID-DOI="http://doi.org/10.5281/zenodo.4506847" ID-GBIF-Taxon="177215434" ID-Zenodo-Dep="4506847" LSID="urn:lsid:plazi:treatment:03B1ED22FFEF452E9BB05965FB015EB5" httpUri="http://treatment.plazi.org/id/03B1ED22FFEF452E9BB05965FB015EB5" lastPageId="12" lastPageNumber="47" pageId="4" pageNumber="39">
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<subSubSection id="C3020FBFFFEF45269BB05965FDD35B7A" box="[151,592,1616,1643]" pageId="4" pageNumber="39" type="nomenclature">
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<paragraph id="8BA75C34FFEF45269BB05965FDD35B7A" blockId="4.[151,1160,1616,1679]" box="[151,592,1616,1643]" pageId="4" pageNumber="39">
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<heading id="D0EFEB58FFEF45269BB05965FDD35B7A" bold="true" box="[151,592,1616,1643]" fontSize="11" level="1" pageId="4" pageNumber="39" reason="1">
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<emphasis id="B96C8026FFEF45269BB05965FDD35B7A" bold="true" box="[151,592,1616,1643]" pageId="4" pageNumber="39">
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<taxonomicName id="4C1827B7FFEF45269BB05965FE315B7A" ID-CoL="8XT3H" authority="Perkins & Ribera, 2020" authorityName="Perkins & Ribera" authorityYear="2020" box="[151,434,1616,1643]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
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<emphasis id="B96C8026FFEF45269BB05965FE315B7A" bold="true" box="[151,434,1616,1643]" italics="true" pageId="4" pageNumber="39">Adelphydraena spinosa</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A25F3D5DFFEF45269AE65965FDD35B7A" box="[449,592,1616,1642]" pageId="4" pageNumber="39" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3020FBFFFEF45269BB05941FB0B5B9F" box="[151,1160,1652,1679]" pageId="4" pageNumber="39" type="description">
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<paragraph id="8BA75C34FFEF45269BB05941FB0B5B9F" blockId="4.[151,1160,1616,1679]" box="[151,1160,1652,1679]" pageId="4" pageNumber="39">
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<figureCitation id="132340B1FFEF45269BB05941FF5F5B9E" box="[151,220,1652,1678]" captionStart="FIGURE 1" captionStartId="3.[151,250,2011,2035]" captionTargetBox="[190,1387,333,1965]" captionTargetId="figure-79@3.[158,1428,306,1985]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of Adelphydraena species." figureDoi="http://doi.org/10.5281/zenodo.4411487" httpUri="https://zenodo.org/record/4411487/files/figure.png" pageId="4" pageNumber="39">Figs 1</figureCitation>
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,
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<figureCitation id="132340B1FFEF45269BCD5940FF795B9F" box="[234,250,1653,1679]" captionStart="FIGURE 7" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[182,1403,203,1864]" captionTargetId="figure-30@8.[147,1440,181,1890]" captionTargetPageId="8" captionText="FIGURE 7. Adelphydraena spinosa n. sp., habitus of holotype." figureDoi="http://doi.org/10.5281/zenodo.4411497" httpUri="https://zenodo.org/record/4411497/files/figure.png" pageId="4" pageNumber="39">7</figureCitation>
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,
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<figureCitation id="132340B1FFEF45269A215941FEA15B9E" box="[262,290,1652,1678]" captionStart="FIGURE 13" captionStartId="13.[151,250,1813,1837]" captionTargetBox="[170,1420,215,1741]" captionTargetId="figure-17@13.[156,1432,181,1789]" captionTargetPageId="13" captionText="FIGURE 13. Ventral habitus of Adelphydraena species." figureDoi="http://doi.org/10.5281/zenodo.4411505" httpUri="https://zenodo.org/record/4411505/files/figure.png" pageId="4" pageNumber="39">13</figureCitation>
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(habitus), 9 (aedeagus), 11 (spermatheca and ventrite), 14 (map), 15 (habitat)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3020FBFFFEF45269BB05989FC9A5A76" pageId="4" pageNumber="39" type="materials_examined">
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<paragraph id="8BA75C34FFEF45269BB05989FC9A5A76" blockId="4.[151,1437,1724,2039]" pageId="4" pageNumber="39">
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<emphasis id="B96C8026FFEF45269BB05989FEC65BC6" bold="true" box="[151,325,1724,1750]" pageId="4" pageNumber="39">Type material.</emphasis>
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<materialsCitation id="3B705669FFEF45269A6B5989FEFD5A0E" ID-GBIF-Occurrence="3031207305" collectingDate="2014-03-11" collectionCode="SEMC" collectorName="Short" country="Guyana" elevation="524" latitude="5.0111833" location="Region" longLatPrecision="1" longitude="-59.6367" municipality="Upper Potaro Camp" pageId="4" pageNumber="39" specimenCount="1" typeStatus="holotype">
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<emphasis id="B96C8026FFEF45269A6B5989FE3A5BC6" bold="true" box="[332,441,1724,1750]" pageId="4" pageNumber="39">
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<typeStatus id="54A3E296FFEF45269A6B5989FE3A5BC6" box="[332,441,1724,1750]" pageId="4" pageNumber="39" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male): “
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||
<collectingCountry id="F30F1CA4FFEF452699065989FD1E5BC6" box="[545,669,1724,1750]" name="Guyana" pageId="4" pageNumber="39">GUYANA</collectingCountry>
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,
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||
<location id="8EC70AEFFFEF4526998F5989FD7A5BC6" LSID="urn:lsid:plazi:treatment:03B1ED22FFEF452E9BB05965FB015EB5:8EC70AEFFFEF4526998F5989FD7A5BC6" box="[680,761,1724,1750]" country="Guyana" latitude="5.0111833" longLatPrecision="1" longitude="-59.6367" municipality="Upper Potaro Camp" name="Region" pageId="4" pageNumber="39">Region</location>
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XIII,
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||
<geoCoordinate id="EE2C3AF3FFEF452698195989FC3A5BC6" box="[830,953,1724,1751]" degrees="5" direction="north" minutes="0.671" orientation="latitude" pageId="4" pageNumber="39" precision="1" value="5.0111833">5º 0.671’N</geoCoordinate>
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<geoCoordinate id="EE2C3AF3FFEF452698E75989FBDF5BC6" box="[960,1116,1724,1751]" degrees="59" direction="west" minutes="38.202" orientation="longitude" pageId="4" pageNumber="39" precision="1" value="-59.6367">59º 38.202’W</geoCoordinate>
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,
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<quantity id="4CE0F1D1FFEF45269F405989FB335BC7" box="[1127,1200,1724,1751]" metricMagnitude="2" metricUnit="m" metricValue="5.24" pageId="4" pageNumber="39" unit="m" value="524.0">
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<elevation id="0035BB07FFEF45269F405989FB335BC7" box="[1127,1200,1724,1751]" metricMagnitude="2" metricUnit="m" metricValue="5.24" pageId="4" pageNumber="39" unit="m" value="524.0">524 m</elevation>
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||
</quantity>
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||
,
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<collectingMunicipality id="6BC3C64EFFEF45269F9B5988FA1F5BC6" box="[1212,1436,1724,1751]" pageId="4" pageNumber="39">Upper Potaro Camp</collectingMunicipality>
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||
1 (c.
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||
<locationDeviation id="9A154516FFEF45269BF659D4FE3D5BEB" box="[209,446,1760,1787]" location="Chenapau" pageId="4" pageNumber="39">
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||
<quantity id="4CE0F1D1FFEF45269BF659D4FE885BEA" box="[209,267,1760,1787]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="39" unit="km" value="7.0">7 km</quantity>
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NW Chenapau
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</locationDeviation>
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), Potaro margin trail; leaf packs in rocky stream; leg.
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<collectorName id="26ED39E2FFEF45269F3959D5FBD85BEA" box="[1054,1115,1760,1786]" pageId="4" pageNumber="39">Short</collectorName>
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||
& Baca;
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<date id="FFA67AF4FFEF45269FE059D5FAC15BEA" box="[1223,1346,1760,1786]" pageId="4" pageNumber="39" value="2014-03-11">
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||
<collectingDate id="EFE2831CFFEF45269FE059D5FAC15BEA" box="[1223,1346,1760,1786]" pageId="4" pageNumber="39" value="2014-03-11">11.iii.2014</collectingDate>
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||
</date>
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||
, GY14- 0311-05A” (
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||
<collectionCode id="ED09C4F1FFEF45269A045831FEF65A0E" box="[291,373,1796,1822]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="39" type="Museum">SEMC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B705669FFEF45269AAE5831FC965A76" ID-GBIF-Occurrence="3031207307" collectingDate="2014-03-11" collectionCode="SEMC, MCZ" collectorName="La Cruz, GY" country="Guyana" elevation="524" latitude="5.0112166" location="Same" longLatPrecision="1" longitude="-59.6393" municipality="Upper Potaro Camp" pageId="4" pageNumber="39" specimenCount="5" specimenCount-female="4" specimenCount-male="1" typeStatus="paratype">
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<emphasis id="B96C8026FFEF45269AAE5831FDB25A0E" bold="true" box="[393,561,1796,1822]" pageId="4" pageNumber="39">
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<typeStatus id="54A3E296FFEF45269AAE5831FD825A0E" box="[393,513,1796,1822]" pageId="4" pageNumber="39" type="paratype">Paratypes</typeStatus>
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(5):
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</emphasis>
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<location id="8EC70AEFFFEF4526991E5831FDF45A0E" LSID="urn:lsid:plazi:treatment:03B1ED22FFEF452E9BB05965FB015EB5:8EC70AEFFFEF4526991E5831FDF45A0E" box="[569,631,1796,1822]" latitude="5.0112166" longLatPrecision="1" longitude="-59.6393" municipality="Upper Potaro Camp" name="Same" pageId="4" pageNumber="39">Same</location>
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||
data as holotype (
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||
<specimenCount id="9D1E97BDFFEF452698605831FC175A0E" box="[839,916,1796,1822]" pageId="4" pageNumber="39" type="male">1 male</specimenCount>
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,
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<specimenCount id="9D1E97BDFFEF452698875831FB825A0E" box="[928,1025,1796,1822]" pageId="4" pageNumber="39" type="female">1 female</specimenCount>
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<collectionCode id="ED09C4F1FFEF45269F2F5831FBDA5A0E" box="[1032,1113,1796,1822]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="39" type="Museum">SEMC</collectionCode>
|
||
);
|
||
<collectingMunicipality id="6BC3C64EFFEF45269F4A5830FACD5A0E" box="[1133,1358,1796,1823]" pageId="4" pageNumber="39">Upper Potaro Camp</collectingMunicipality>
|
||
1 (c.
|
||
<locationDeviation id="9A154516FFEF45269EA95830FEED5A53" location="Chenapau" pageId="4" pageNumber="39">
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||
<quantity id="4CE0F1D1FFEF45269EA95830FF385A52" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="39" unit="km" value="7.0">7 km</quantity>
|
||
NW Chenapau
|
||
</locationDeviation>
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||
), stream nr. camp; gravel/sandy stream w/ detritus;
|
||
<geoCoordinate id="EE2C3AF3FFEF45269892581DFBB25A52" box="[949,1073,1832,1859]" degrees="5" direction="north" minutes="0.673" orientation="latitude" pageId="4" pageNumber="39" precision="1" value="5.0112166">5º 0.673’N</geoCoordinate>
|
||
<geoCoordinate id="EE2C3AF3FFEF45269F1F581DFB565A53" box="[1080,1237,1832,1859]" degrees="59" direction="west" minutes="38.358" orientation="longitude" pageId="4" pageNumber="39" precision="1" value="-59.6393">59º 38.358’W</geoCoordinate>
|
||
, Short, Salisbury,
|
||
<collectorName id="26ED39E2FFEF45269BB05878FEA95A76" box="[151,298,1868,1895]" pageId="4" pageNumber="39">La Cruz, GY</collectorName>
|
||
14-0314-01A (
|
||
<specimenCount id="9D1E97BDFFEF45269AE95879FDBE5A76" box="[462,573,1868,1894]" pageId="4" pageNumber="39" type="female">3 females</specimenCount>
|
||
,
|
||
<collectionCode id="ED09C4F1FFEF4526996F5879FD175A76" box="[584,660,1868,1894]" country="USA" name="University of Kansas - Biodiversity Institute" pageId="4" pageNumber="39" type="Museum">SEMC</collectionCode>
|
||
and
|
||
<collectionCode id="ED09C4F1FFEF452699EC5879FC8E5A76" box="[715,781,1868,1894]" country="USA" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="39" type="Museum">MCZ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3020FBFFFEF45269BE05845FB6A5AE6" pageId="4" pageNumber="39" type="diagnosis">
|
||
<paragraph id="8BA75C34FFEF45269BE05845FB6A5AE6" blockId="4.[151,1437,1724,2039]" pageId="4" pageNumber="39">
|
||
<emphasis id="B96C8026FFEF45269BE05845FE4E5A9A" bold="true" box="[199,461,1904,1930]" pageId="4" pageNumber="39">Differential diagnosis.</emphasis>
|
||
Differentiated from other
|
||
<taxonomicName id="4C1827B7FFEF452699D45845FC1D5A9A" authorityName="Perkins" authorityYear="1989" box="[755,926,1904,1930]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B96C8026FFEF452699D45845FC1D5A9A" box="[755,926,1904,1930]" italics="true" pageId="4" pageNumber="39">Adelphydraena</emphasis>
|
||
</taxonomicName>
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by the combination of the chordate pronotum with tumidities almost totally separated by punctures in midline, the elytra with serrate margins, which are posteriorly produced in sharp processes, the labrum with a spinose process on each apicolateral angle, the comparatively large metaventral plaques, which are very narrowly separated anteriorly, and the aedeagus (
|
||
<figureCitation id="132340B1FFEF45269FB158E9FB595AE6" box="[1174,1242,2012,2038]" captionStart="FIGURES9–10" captionStartId="10.[151,264,1989,2013]" captionTargetBox="[325,1261,190,1955]" captionTargetId="figure-30@10.[317,1270,181,1965]" captionTargetPageId="10" captionText="FIGURES9–10.9.Adelphydraenaspinosan.sp.,holotypeaedeagus.10.AdelphydraenaorchymontiPerkins,non-type,aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4411501" httpUri="https://zenodo.org/record/4411501/files/figure.png" pageId="4" pageNumber="39">Fig. 9</figureCitation>
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||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF670CBCFFEE45279BB0590EFCDB5B44" ID-DOI="http://doi.org/10.5281/zenodo.4411491" ID-Zenodo-Dep="4411491" box="[151,856,1595,1620]" httpUri="https://zenodo.org/record/4411491/files/figure.png" pageId="5" pageNumber="40" startId="5.[151,250,1595,1619]" targetBox="[164,1422,181,1572]" targetPageId="5">
|
||
<paragraph id="8BA75C34FFEE45279BB0590EFCDB5B44" blockId="5.[151,856,1595,1620]" box="[151,856,1595,1620]" pageId="5" pageNumber="40">
|
||
<emphasis id="B96C8026FFEE45279BB0590EFE905B43" bold="true" box="[151,275,1595,1619]" pageId="5" pageNumber="40">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C1827B7FFEE45279A3E590EFD015B43" authority="Perkins" authorityName="Perkins" authorityYear="1989" box="[281,642,1595,1619]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="orchymonti">
|
||
<emphasis id="B96C8026FFEE45279A3E590EFDAF5B43" box="[281,556,1595,1619]" italics="true" pageId="5" pageNumber="40">Adelphydraena orchymonti</emphasis>
|
||
Perkins
|
||
</taxonomicName>
|
||
, habitus of non-type.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3020FBFFFEE452E9BE059B4FE915E91" lastPageId="12" lastPageNumber="47" pageId="5" pageNumber="40" type="description">
|
||
<paragraph id="8BA75C34FFEE452E9BE059B4FE915E91" blockId="5.[151,1437,1665,2016]" lastBlockId="12.[151,1437,151,933]" lastPageId="12" lastPageNumber="47" pageId="5" pageNumber="40">
|
||
<emphasis id="B96C8026FFEE45279BE059B4FE145B8B" bold="true" box="[199,407,1665,1691]" pageId="5" pageNumber="40">Description. Size:</emphasis>
|
||
<typeStatus id="54A3E296FFEE45279ABA59B7FE7C5B8C" box="[413,511,1666,1692]" pageId="5" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 1.68/0.88; head width 0.44; pronotum 0.42/0.69, PA 0.52, PB 0.53; elytra 1.04/0.88. Habitus as illustrated (
|
||
<figureCitation id="132340B1FFEE452798F05993FB995BD0" box="[983,1050,1702,1728]" captionStart="FIGURE 7" captionStartId="8.[151,250,1914,1938]" captionTargetBox="[182,1403,203,1864]" captionTargetId="figure-30@8.[147,1440,181,1890]" captionTargetPageId="8" captionText="FIGURE 7. Adelphydraena spinosa n. sp., habitus of holotype." figureDoi="http://doi.org/10.5281/zenodo.4411497" httpUri="https://zenodo.org/record/4411497/files/figure.png" pageId="5" pageNumber="40">Fig. 7</figureCitation>
|
||
).
|
||
<emphasis id="B96C8026FFEE45279F095990FBFF5BAF" bold="true" box="[1070,1148,1701,1727]" pageId="5" pageNumber="40">Color:</emphasis>
|
||
Dorsum brown to reddish brown, legs and maxillary palpi testaceous.
|
||
<emphasis id="B96C8026FFEE4527995959FFFD3D5BF4" bold="true" box="[638,702,1738,1764]" pageId="5" pageNumber="40">Head</emphasis>
|
||
: Frons finely punctate, punctures smaller than eye facets; shiny in middle, microreticulate and dull laterally; posterior line not extended forward at juncture of shiny and microreticulate areas. Clypeus dull, strongly microreticulate. Labrum slightly longer than clypeus; median emargination setose, length nearly 0.66x labrum length; each apicolateral angle produced in minute sharp process. Eyes small in dorsal aspect, width ca. 1/6 interocular distance. Mentum densely micropunctulate, elongate, narrowed anteriorly, lobe on each side of apicomedian emargination curved ventrally and toward midline; apical width 0.5x basal width. Genae without posterior ridge.
|
||
<emphasis id="B96C8026FFEE45279A6C5897FE475AAC" bold="true" box="[331,452,1954,1980]" pageId="5" pageNumber="40">Pronotum</emphasis>
|
||
: Pronotum subcordiform. Posterior 0.5 of disc with raised, rounded relief on each side of midline, reliefs separated anteriorly by ca. 10 large punctures, reliefs narrowly confluent along posterior margin; reliefs very shiny, very finely, sparsely punctate, punctures similar to those of frons. Reliefs bordered laterally and posteriorly by very coarse and dense punctures. Punctures bordering anterior margin smaller, more-or-less forming row on each side; punctures between row and anterior margin of pronotum much smaller, interstices shiny, 0.5–1.0x puncture diameter.All interstices anterior to discal reliefs with extremely sparse, very fine micropunctures similar in size to those of discal reliefs. Lateral depressions coarsely densely punctate. Lateral margin denticulate on anterior 1/2. Anterior margin distinctly trisinuate. Posterior margin straight.
|
||
<emphasis id="B96C8026FFE7452E98B35FEAFC6A5DE9" bold="true" box="[916,1001,223,249]" pageId="12" pageNumber="47">Elytra:</emphasis>
|
||
Each elytron with 9 complete rows of punctures, tenth row represented by few punctures anteriorly; punctures on disc smaller than largest punctures on pronotal disc. Intervals raised slightly, shiny, 2–4x puncture diameter. Punctures in anterior 0.5 of rows 1–5 larger than on remainder of elytron, interstices 1–2x puncture diameter. Raised humeri connected with slightly produced anterior angles. Lateral margin strongly arcuate, serrate; serrations produced as 3–4 short sharp spines in ca. posterior ¼ of each elytron; explanate margin well developed, at midlength ca. 2x distal width of metatibia.
|
||
<emphasis id="B96C8026FFE7452E9E295EA6FA1E5CBD" bold="true" box="[1294,1437,403,429]" pageId="12" pageNumber="47">Prosternum</emphasis>
|
||
with midlongitudinal ridge from anterior margin to between coxae. Prosternal intercoxal process as wide as mesosternal intercoxal process; prosternal midline length ca. 2x prosternal intercoxal process width. Prosternal postcoxal process concave between submedian carinae.
|
||
<emphasis id="B96C8026FFE7452E99FB5ECBFC005F08" bold="true" box="[732,899,510,536]" pageId="12" pageNumber="47">Mesoventrite:</emphasis>
|
||
intercoxal process length 0.5x width, apex very slightly, if at all, arcuate. Submedian carinae attaining thickened anterior border of metaventrite.
|
||
<emphasis id="B96C8026FFE7452E9FD05D17FA1F5F2C" bold="true" box="[1271,1436,546,572]" pageId="12" pageNumber="47">Metaventrite:</emphasis>
|
||
with plaques well developed, slightly raised, together forming nearly complete inverted V-shape, except narrowly separated at anterior extremes (
|
||
<figureCitation id="132340B1FFE7452E9ADA5D5EFDD15F95" box="[509,594,619,645]" captionStart="FIGURE 13" captionStartId="13.[151,250,1813,1837]" captionTargetBox="[170,1420,215,1741]" captionTargetId="figure-17@13.[156,1432,181,1789]" captionTargetPageId="13" captionText="FIGURE 13. Ventral habitus of Adelphydraena species." figureDoi="http://doi.org/10.5281/zenodo.4411505" httpUri="https://zenodo.org/record/4411505/files/figure.png" pageId="12" pageNumber="47">Fig. 13</figureCitation>
|
||
). Submedian and sublateral low carinae extended from anterior margin to near posterior margin of metaventrite.
|
||
<emphasis id="B96C8026FFE7452E996D5DBBFD4B5FB8" bold="true" box="[586,712,654,680]" pageId="12" pageNumber="47">Abdomen:</emphasis>
|
||
Distance separating coxal cavities slightly wider than mesoventral intercoxal process width, ratio ca. 5/4. Submedian carinae of first sternite terminated slightly before posterior border of first sternite, distance separating carinae at posterior extreme ca. equal midline length of first sternite plus fused intercoxal sternite. Sternites 1–5 with hydrofuge pubescence; hydrofuge pubescence distinctly longer between carinae of first sternite than laterally and longer than on sternites 2–5; sternites 6 and 7 shiny, sparsely pubescent.
|
||
<emphasis id="B96C8026FFE7452E9BB05C77FE5E5E4D" bold="true" box="[151,477,834,861]" pageId="12" pageNumber="47">Aedeagus and spermatheca</emphasis>
|
||
as illustrated (
|
||
<figureCitation id="132340B1FFE7452E99A75C76FD4C5E4D" box="[640,719,835,861]" captionStart="FIGURES9–10" captionStartId="10.[151,264,1989,2013]" captionTargetBox="[325,1261,190,1955]" captionTargetId="figure-30@10.[317,1270,181,1965]" captionTargetPageId="10" captionText="FIGURES9–10.9.Adelphydraenaspinosan.sp.,holotypeaedeagus.10.AdelphydraenaorchymontiPerkins,non-type,aedeagus." figureDoi="http://doi.org/10.5281/zenodo.4411501" httpUri="https://zenodo.org/record/4411501/files/figure.png" pageId="12" pageNumber="47">Figs. 9</figureCitation>
|
||
,
|
||
<figureCitation id="132340B1FFE7452E99FD5C76FD745E4D" box="[730,759,835,861]" captionStart="FIGURES 11–12" captionStartId="11.[151,264,1792,1816]" captionTargetBox="[254,1402,218,1729]" captionTargetId="figure-17@11.[185,1402,181,1769]" captionTargetPageId="11" captionText="FIGURES 11–12. 11. Adelphydraena spinosa n. sp., spermatheca and terminal abdominal segments. 12. Adelphydraena surinamensis n. sp., spermatheca and terminal abdominal segments." figureDoi="http://doi.org/10.5281/zenodo.4411503" httpUri="https://zenodo.org/record/4411503/files/figure.png" pageId="12" pageNumber="47">11</figureCitation>
|
||
).
|
||
<emphasis id="B96C8026FFE7452E982B5C76FCCE5E4D" bold="true" box="[780,845,835,861]" pageId="12" pageNumber="47">Legs:</emphasis>
|
||
without apparent modifications.
|
||
<emphasis id="B96C8026FFE7452E9F985C77FAD95E4C" bold="true" box="[1215,1370,834,860]" pageId="12" pageNumber="47">Flight wings:</emphasis>
|
||
Fully developed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF670CBCFFED45249BB058ABFCC55AA7" ID-DOI="http://doi.org/10.5281/zenodo.4411493" ID-Zenodo-Dep="4411493" box="[151,838,1950,1975]" httpUri="https://zenodo.org/record/4411493/files/figure.png" pageId="6" pageNumber="41" startId="6.[151,250,1950,1974]" targetBox="[151,1419,209,1914]" targetPageId="6">
|
||
<paragraph id="8BA75C34FFED45249BB058ABFCC55AA7" blockId="6.[151,838,1950,1975]" box="[151,838,1950,1975]" pageId="6" pageNumber="41">
|
||
<emphasis id="B96C8026FFED45249BB058ABFE905AA6" bold="true" box="[151,275,1950,1974]" pageId="6" pageNumber="41">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C1827B7FFED45249A3E58ABFDF35AA6" authority="Perkins" authorityName="Perkins" authorityYear="1989" box="[281,624,1950,1974]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="B96C8026FFED45249A3E58ABFD995AA6" box="[281,538,1950,1974]" italics="true" pageId="6" pageNumber="41">Adelphydraena spangleri</emphasis>
|
||
Perkins
|
||
</taxonomicName>
|
||
, habitus of non-type.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF670CBCFFEC45259BB058A2FDA35AC4" ID-DOI="http://doi.org/10.5281/zenodo.4411495" ID-Zenodo-Dep="4411495" httpUri="https://zenodo.org/record/4411495/files/figure.png" pageId="7" pageNumber="42" startId="7.[151,264,1943,1967]" targetBox="[287,1304,193,1908]" targetPageId="7">
|
||
<paragraph id="8BA75C34FFEC45259BB058A2FDA35AC4" blockId="7.[151,1436,1943,2004]" pageId="7" pageNumber="42">
|
||
<emphasis id="B96C8026FFEC45259BB058A2FEBF5ABF" bold="true" box="[151,316,1943,1967]" pageId="7" pageNumber="42">FIGURES 5–6.</emphasis>
|
||
5.
|
||
<taxonomicName id="4C1827B7FFEC45259A7858ADFD3A5AA0" authority="Perkins" authorityName="Perkins" authorityYear="1989" box="[351,697,1944,1968]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="spangleri">
|
||
<emphasis id="B96C8026FFEC45259A7858ADFDE15AA0" box="[351,610,1944,1968]" italics="true" pageId="7" pageNumber="42">Adelphydraena spangleri</emphasis>
|
||
Perkins
|
||
</taxonomicName>
|
||
, non-type aedeagus and last tergites. 6.
|
||
<taxonomicName id="4C1827B7FFEC45259F7058ADFA035ABF" authority="Perkins & Ribera, 2020" authorityName="Perkins & Ribera" authorityYear="2020" box="[1111,1408,1944,1968]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="surinamensis" status="sp. nov.">
|
||
<emphasis id="B96C8026FFEC45259F7058ADFA035ABF" box="[1111,1408,1944,1968]" italics="true" pageId="7" pageNumber="42">Adelphydraena surinamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C8026FFEC45259EAF58ADFF355AC4" bold="true" pageId="7" pageNumber="42">
|
||
<taxonomicNameLabel id="A25F3D5DFFEC45259EAF58ADFF355AC4" pageId="7" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype aedeagus and last tergite.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF670CBCFFE3452A9BB0584FFC9C5A82" ID-DOI="http://doi.org/10.5281/zenodo.4411497" ID-Zenodo-Dep="4411497" box="[151,799,1914,1938]" httpUri="https://zenodo.org/record/4411497/files/figure.png" pageId="8" pageNumber="43" startId="8.[151,250,1914,1938]" targetBox="[182,1403,203,1864]" targetPageId="8">
|
||
<paragraph id="8BA75C34FFE3452A9BB0584FFC9C5A82" blockId="8.[151,799,1914,1938]" box="[151,799,1914,1938]" pageId="8" pageNumber="43">
|
||
<emphasis id="B96C8026FFE3452A9BB0584FFE905A82" bold="true" box="[151,275,1914,1938]" pageId="8" pageNumber="43">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C1827B7FFE3452A9A3E584FFD845A82" authority="Perkins & Ribera, 2020" authorityName="Perkins & Ribera" authorityYear="2020" box="[281,519,1914,1938]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
|
||
<emphasis id="B96C8026FFE3452A9A3E584FFD845A82" box="[281,519,1914,1938]" italics="true" pageId="8" pageNumber="43">Adelphydraena spinosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C8026FFE3452A9929584FFDC45A82" bold="true" box="[526,583,1914,1938]" pageId="8" pageNumber="43">
|
||
<taxonomicNameLabel id="A25F3D5DFFE3452A9929584FFDC45A82" box="[526,583,1914,1938]" pageId="8" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF670CBCFFE2452B9BB058AEFCD45AA4" ID-DOI="http://doi.org/10.5281/zenodo.4411499" ID-Zenodo-Dep="4411499" box="[151,855,1947,1972]" httpUri="https://zenodo.org/record/4411499/files/figure.png" pageId="9" pageNumber="44" startId="9.[151,250,1947,1971]" targetBox="[196,1378,239,1902]" targetPageId="9">
|
||
<paragraph id="8BA75C34FFE2452B9BB058AEFCD45AA4" blockId="9.[151,855,1947,1972]" box="[151,855,1947,1972]" pageId="9" pageNumber="44">
|
||
<emphasis id="B96C8026FFE2452B9BB058AEFE905AA4" bold="true" box="[151,275,1947,1972]" pageId="9" pageNumber="44">FIGURE 8.</emphasis>
|
||
<taxonomicName id="4C1827B7FFE2452B9A3E58A9FDC35AA3" authority="Perkins & Ribera, 2020" authorityName="Perkins & Ribera" authorityYear="2020" box="[281,576,1948,1972]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="surinamensis" status="sp. nov.">
|
||
<emphasis id="B96C8026FFE2452B9A3E58A9FDC35AA3" box="[281,576,1948,1972]" italics="true" pageId="9" pageNumber="44">Adelphydraena surinamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C8026FFE2452B996058A9FD035AA4" bold="true" box="[583,640,1948,1972]" pageId="9" pageNumber="44">
|
||
<taxonomicNameLabel id="A25F3D5DFFE2452B996058A9FD035AA4" box="[583,640,1948,1972]" pageId="9" pageNumber="44" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, habitus of holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF670CBCFFE145289BB058F0FA1E5ACD" ID-DOI="http://doi.org/10.5281/zenodo.4411501" ID-Zenodo-Dep="4411501" box="[151,1437,1989,2014]" httpUri="https://zenodo.org/record/4411501/files/figure.png" pageId="10" pageNumber="45" startId="10.[151,264,1989,2013]" targetBox="[325,1261,190,1955]" targetPageId="10">
|
||
<paragraph id="8BA75C34FFE145289BB058F0FA1E5ACD" blockId="10.[151,1437,1989,2014]" box="[151,1437,1989,2014]" pageId="10" pageNumber="45">
|
||
<emphasis id="B96C8026FFE145289BB058F0FEC75ACD" bold="true" box="[151,324,1989,2013]" pageId="10" pageNumber="45">FIGURES 9–10.</emphasis>
|
||
9.
|
||
<emphasis id="B96C8026FFE145289A7B58F0FDC45ACD" box="[348,583,1989,2013]" italics="true" pageId="10" pageNumber="45">Adelphydraenaspinosa</emphasis>
|
||
<emphasis id="B96C8026FFE14528996D58F0FD035ACD" bold="true" box="[586,640,1989,2013]" pageId="10" pageNumber="45">
|
||
<taxonomicNameLabel id="A25F3D5DFFE14528996D58F0FD035ACD" box="[586,640,1989,2013]" pageId="10" pageNumber="45" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotypeaedeagus.10.
|
||
<emphasis id="B96C8026FFE14528984958F0FBFD5ACD" box="[878,1150,1989,2013]" italics="true" pageId="10" pageNumber="45">Adelphydraenaorchymonti</emphasis>
|
||
Perkins, non-type, aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF670CBCFFE045299BB05835FCA05A2D" ID-DOI="http://doi.org/10.5281/zenodo.4411503" ID-Zenodo-Dep="4411503" httpUri="https://zenodo.org/record/4411503/files/figure.png" pageId="11" pageNumber="46" startId="11.[151,264,1792,1816]" targetBox="[254,1402,218,1729]" targetPageId="11">
|
||
<paragraph id="8BA75C34FFE045299BB05835FCA05A2D" blockId="11.[151,1436,1792,1853]" pageId="11" pageNumber="46">
|
||
<emphasis id="B96C8026FFE045299BB05835FED05A08" bold="true" box="[151,339,1792,1816]" pageId="11" pageNumber="46">FIGURES 11–12.</emphasis>
|
||
11.
|
||
<taxonomicName id="4C1827B7FFE045299A585835FDED5A08" authority="Perkins & Ribera, 2020" authorityName="Perkins & Ribera" authorityYear="2020" box="[383,622,1792,1816]" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov.">
|
||
<emphasis id="B96C8026FFE045299A585835FDED5A08" box="[383,622,1792,1816]" italics="true" pageId="11" pageNumber="46">Adelphydraena spinosa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C8026FFE0452999525834FD2D5A09" bold="true" box="[629,686,1793,1817]" pageId="11" pageNumber="46">
|
||
<taxonomicNameLabel id="A25F3D5DFFE0452999525834FD2D5A09" box="[629,686,1793,1817]" pageId="11" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, spermatheca and terminal abdominal segments. 12.
|
||
<taxonomicName id="4C1827B7FFE045299FEC5835FF765A2C" authority="Perkins & Ribera, 2020" authorityName="Perkins & Ribera" authorityYear="2020" class="Insecta" family="Hydraenidae" genus="Adelphydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="surinamensis" status="sp. nov.">
|
||
<emphasis id="B96C8026FFE045299FEC5835FF765A2C" italics="true" pageId="11" pageNumber="46">Adelphydraena surinamensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96C8026FFE045299BDB5810FEB65A2D" bold="true" box="[252,309,1829,1853]" pageId="11" pageNumber="46">
|
||
<taxonomicNameLabel id="A25F3D5DFFE045299BDB5810FEB65A2D" box="[252,309,1829,1853]" pageId="11" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, spermatheca and terminal abdominal segments.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3020FBFFFE7452E9BE05CBEFB015EB5" box="[199,1154,907,933]" pageId="12" pageNumber="47" type="etymology">
|
||
<paragraph id="8BA75C34FFE7452E9BE05CBEFB015EB5" blockId="12.[151,1437,151,933]" box="[199,1154,907,933]" pageId="12" pageNumber="47">
|
||
<emphasis id="B96C8026FFE7452E9BE05CBEFECD5EB5" bold="true" box="[199,334,907,933]" pageId="12" pageNumber="47">Etymology.</emphasis>
|
||
The name is an adjective referring to the spines of the labrum and elytra.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |