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<document id="4FD6CF1B11AB6C204BC0690ED26613ED" ID-DOI="10.5852/ejt.2019.526" ID-GBIF-Dataset="2da65c0a-9c48-43f5-ada3-e088de28dec1" ID-ISSN="2118-9773" ID-Zenodo-Dep="3152217" ID-ZooBank="3FFC527D-E9CA-4D9F-BF59-076FA60171AC" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="existingObjects,operationResults,plazi" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1558535825802" checkinUser="plazi" docAuthor="Nurinsiyah, Ayu Savitri, Neiber, Marco T. &amp; Hausdorf, Bernhard" docDate="2019" docId="03B1DB4C8B314E074C1DFB3F3E98F856" docLanguage="en" docName="ej-526_nurinsiyah_neiber_hausdorf.pdf.imf" docOrigin="European Journal of Taxonomy 526" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Landouria epiplatia" docType="treatment" docVersion="18" lastPageNumber="31" masterDocId="FF88A3348B2D4E194E35FFFC3C0DFFDB" masterDocTitle="Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java" masterLastPageNumber="73" masterPageNumber="1" pageNumber="29" updateTime="1716321918122" updateUser="carolina">
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<mods:title id="1A03D2F1026D47868795F121D820FED6">Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java</mods:title>
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<mods:namePart id="B241500F00D69EC1A582111894053FC4">Nurinsiyah, Ayu Savitri</mods:namePart>
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<mods:affiliation id="655EC64F3566AE764651B40BF4D6D39F">Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia.</mods:affiliation>
<mods:nameIdentifier id="4CEC61B1FAE27300017CBF222E0A3B46" type="email">ayu_nurinsiyah@yahoo.com &amp; ayus002@lipi.go.id</mods:nameIdentifier>
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<mods:namePart id="328E79F060D57B6836A20E0A43A64FC9">Neiber, Marco T.</mods:namePart>
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<mods:affiliation id="9815E285DEDB71D1DDE672BAEFF00F9C">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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<mods:namePart id="65DEEBE79709D7B56F674075F087DE4E">Hausdorf, Bernhard</mods:namePart>
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<mods:affiliation id="EF962ADC04FE49E4FC6E40E30B390EE4">Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<treatment id="03B1DB4C8B314E074C1DFB3F3E98F856" ID-GBIF-Taxon="157239583" LSID="urn:lsid:plazi:treatment:03B1DB4C8B314E074C1DFB3F3E98F856" httpUri="http://treatment.plazi.org/id/03B1DB4C8B314E074C1DFB3F3E98F856" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C30239D18B314E054C1DFB3F3806FB05" box="[552,1035,1219,1246]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BA76A5A8B314E054C1DFB3F3806FB05" blockId="28.[552,1035,1219,1282]" box="[552,1035,1219,1246]" pageId="28" pageNumber="29">
<heading id="D0EFDD368B314E054C1DFB3F3806FB05" box="[552,1035,1219,1246]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="4C1811D98B314E054C1DFB3F3806FB05" ID-CoL="7RTHD" authority="(Mollendorff, 1897)" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[552,1035,1219,1246]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054C1DFB3F3F15FB05" bold="true" box="[552,792,1219,1246]" italics="true" pageId="28" pageNumber="29">Landouria epiplatia</emphasis>
(
<bibRefCitation id="EF8917AB8B314E054D12FB3F380EFB05" author="Mollendorff" box="[807,1027,1219,1246]" firstAuthor="Mollendorff" pageId="28" pageNumber="29" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
)
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</heading>
</paragraph>
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<subSubSection id="C30239D18B314E054C7AFB1B3FE8FADA" box="[591,997,1255,1282]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BA76A5A8B314E054C7AFB1B3FE8FADA" blockId="28.[552,1035,1219,1282]" box="[591,997,1255,1282]" pageId="28" pageNumber="29">
<figureCitation id="132376DF8B314E054C7AFB1B3EA5FADA" box="[591,680,1255,1281]" captionStart="Figs 1524" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 1524. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="28" pageNumber="29">Figs 17</figureCitation>
,
<figureCitation id="132376DF8B314E054C80FB1B3F0CFADA" box="[693,769,1255,1281]" captionStart="Figs 2528" captionStartId="28.[189,241,1858,1884]" captionTargetBox="[221,1366,265,2019]" captionTargetId="figure@27.[221,1366,265,2019]" captionTargetPageId="27" captionText="Figs 2528 (opposite page). Shells of species of Landouria from Java. 25. L. naggsi sp. nov., West Java, Cibitung, near Cikaso Waterfall (holotype, MZB 20424). 26. L. epiplatia (Möllendorff, 1897), West Java, Jampang (lectotype of Plectotropis epiplatia, SMF 8909). 27. L. epiplatia, West Java, Jampang (lectotype of Plectotropis trichotrochium Möllendorff, 1897, SMF 8918). 28. L. epiplatia?, West Java, Bogor, Gunung Gede (ZMH 79946). Scale bar: 5 mm." httpUri="https://zenodo.org/record/3152245/files/figure.png" pageId="28" pageNumber="29">2628</figureCitation>
,
<figureCitation id="132376DF8B314E054D38FB1B3F20FADA" box="[781,813,1255,1281]" captionStart="Figs 2947" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 2947. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="28" pageNumber="29">31</figureCitation>
,
<figureCitation id="132376DF8B314E054D0CFB1B3F54FAD9" box="[825,857,1255,1282]" captionStart="Figs 5255" captionStartId="33.[189,241,1717,1743]" captionTargetBox="[189,1398,265,1993]" captionTargetId="figure@34.[189,1398,265,1993]" captionTargetPageId="34" captionText="Figs 5255 (opposite page). Distribution of species of Landouria in Java (10 grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867)." httpUri="https://zenodo.org/record/3152265/files/figure.png" pageId="28" pageNumber="29">55</figureCitation>
;
<tableCitation id="C69A5FE18B314E054D50FB1B3FE8FADA" box="[869,997,1255,1281]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="28" pageNumber="29" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 34</tableCitation>
</paragraph>
</subSubSection>
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<treatmentCitationGroup id="AB084D748B314E054E88FACC3D50FAB6" pageId="28" pageNumber="29">
<treatmentCitation id="0AB94C4B8B314E054E88FACC3EC1FA91" author="Mollendorff O. F. von" box="[189,716,1328,1354]" page="67" pageId="28" pageNumber="29" year="1897">
<taxonomicName id="4C1811D98B314E054E88FACC3EC1FA91" ID-CoL="7SRFD" authority="Mollendorff, 1897: 67" authorityName="Mollendorff" authorityPageNumber="67" authorityYear="1897" box="[189,716,1328,1354]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054E88FACC3DB1FA91" box="[189,444,1328,1354]" italics="true" pageId="28" pageNumber="29">Plectotropis epiplatia</emphasis>
<bibRefCitation id="EF8917AB8B314E054FF0FACC3EC1FA91" author="Mollendorff O. F. von" box="[453,716,1328,1354]" firstAuthor="Mollendorff" pageId="28" pageNumber="29" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897: 67</bibRefCitation>
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(“
<collectingRegion id="49DCA4B88B314E054CDFFACC3F2BFA91" box="[746,806,1328,1354]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="29">Java</collectingRegion>
”; more exactly (
<bibRefCitation id="EF8917AB8B314E054DDCFACC387CFA90" author="Zilch" box="[1001,1137,1328,1355]" firstAuthor="Zilch" pageId="28" pageNumber="29" pagination="293 - 319" refId="ref35814" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch 1966</bibRefCitation>
): “W-Java: Djampang (2000)”).
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</paragraph>
<paragraph id="8BA76A5A8B314E054E88FA8A3F2AFA6F" blockId="28.[189,1399,1328,1460]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB084D748B314E054E88FA8A3F2AFA6F" pageId="28" pageNumber="29">
<treatmentCitation id="0AB94C4B8B314E054E88FA8A3F21FA4B" author="Mollendorff O. F. von" box="[189,812,1398,1424]" page="67" pageId="28" pageNumber="29" year="1897">
<taxonomicName id="4C1811D98B314E054E88FA8A3F21FA4B" ID-CoL="7SRH4" authority="Mollendorff, 1897: 67" authorityName="Mollendorff" authorityPageNumber="67" authorityYear="1897" box="[189,812,1398,1424]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B314E054E88FA8A3E04FA4B" box="[189,521,1398,1424]" italics="true" pageId="28" pageNumber="29">Plectotropis trichotrochium</emphasis>
<bibRefCitation id="EF8917AB8B314E054C2DFA8A3F21FA4B" author="Mollendorff O. F. von" box="[536,812,1398,1424]" firstAuthor="Mollendorff" pageId="28" pageNumber="29" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897: 67</bibRefCitation>
</taxonomicName>
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(“
<collectingRegion id="49DCA4B88B314E054D7AFA8A3F87FA4B" box="[847,906,1398,1424]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="29">Java</collectingRegion>
”, more exactly (
<bibRefCitation id="EF8917AB8B314E054A6AFA8A38E3FA4A" author="Zilch" box="[1119,1262,1398,1425]" firstAuthor="Zilch" pageId="28" pageNumber="29" pagination="293 - 319" refId="ref35814" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch 1966</bibRefCitation>
): “W-Java: Djampang (2000), s. Susabuni [sic]”).
<emphasis id="B96CB6488B314E054C83FA653F2AFA6F" bold="true" box="[694,807,1433,1460]" pageId="28" pageNumber="29">Syn. nov.</emphasis>
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<paragraph id="8BA76A5A8B314E054E88FA1E3DC1F998" blockId="28.[189,1399,1506,1603]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB084D748B314E054E88FA1E3DC1F998" pageId="28" pageNumber="29">
<taxonomicName id="4C1811D98B314E054E88FA1E3F34FA27" ID-CoL="7RTHD" authority="- van Benthem Jutting 1950: 466" authorityName="- van Benthem Jutting" authorityPageNumber="466" authorityYear="1950" box="[189,825,1506,1533]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054E88FA1E3DA5FA27" box="[189,424,1506,1532]" italics="true" pageId="28" pageNumber="29">Landouria epiplatia</emphasis>
<treatmentCitation id="0AB94C4B8B314E054FF3FA1F3F34FA27" author="van Benthem Jutting W. S. S." box="[454,825,1506,1533]" page="466" pageId="28" pageNumber="29" year="1950">
<bibRefCitation id="EF8917AB8B314E054FF3FA1F3F34FA27" author="van Benthem Jutting W. S. S." box="[454,825,1506,1533]" firstAuthor="van Benthem Jutting" pageId="28" pageNumber="29" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting 1950: 466</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 82. —
<treatmentCitation id="0AB94C4B8B314E054DF6FA1E388FFA27" author="Zilch A." box="[963,1154,1506,1532]" page="293" pageId="28" pageNumber="29" year="1966">
<bibRefCitation id="EF8917AB8B314E054DF6FA1E388FFA27" author="Zilch A." box="[963,1154,1506,1532]" firstAuthor="Zilch" pageId="28" pageNumber="29" pagination="293 - 319" refId="ref35814" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch 1966: 293</bibRefCitation>
</treatmentCitation>
, pl. 7, fig. 2.
<taxonomicName id="4C1811D98B314E054E88F9FA3F23F9C4" ID-CoL="3S6S5" authority="- van Benthem Jutting 1950: 461" authorityName="- van Benthem Jutting" authorityPageNumber="461" authorityYear="1950" box="[189,814,1541,1568]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="rotatoria">
<emphasis id="B96CB6488B314E054E88F9FA3DA5F9C4" box="[189,424,1542,1568]" italics="true" pageId="28" pageNumber="29">Landouria rotatoria</emphasis>
<treatmentCitation id="0AB94C4B8B314E054FF6F9FA3F23F9C4" author="van Benthem Jutting W. S. S." box="[451,814,1541,1568]" page="461" pageId="28" pageNumber="29" year="1950">
<bibRefCitation id="EF8917AB8B314E054FF6F9FA3F23F9C4" author="van Benthem Jutting W. S. S." box="[451,814,1541,1568]" firstAuthor="van Benthem Jutting" pageId="28" pageNumber="29" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting 1950: 461</bibRefCitation>
</treatmentCitation>
</taxonomicName>
(in part). —
<treatmentCitation id="0AB94C4B8B314E054DFCF9F93888F9FB" author="Zilch A." box="[969,1157,1541,1568]" page="294" pageId="28" pageNumber="29" year="1966">
<bibRefCitation id="EF8917AB8B314E054DFCF9F93888F9FB" author="Zilch A." box="[969,1157,1541,1568]" firstAuthor="Zilch" pageId="28" pageNumber="29" pagination="293 - 319" refId="ref35814" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch 1966: 294</bibRefCitation>
</treatmentCitation>
, pl. 7, fig. 6 (in part). (not
<bibRefCitation id="EF8917AB8B314E054F14F9D53DCEF998" author="Pfeiffer" box="[289,451,1577,1603]" firstAuthor="Pfeiffer" pageId="28" pageNumber="29" refId="ref34385" refString="Pfeiffer L. 1842. Symbolae ad Historiam Heliceorum. Sectio alterna. T. Fischer, Cassel [Kassel, Germany]. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B314E054E88F98E3F0EF906" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BA76A5A8B314E054E88F98E3D3BF957" blockId="28.[189,310,1650,1676]" box="[189,310,1650,1676]" pageId="28" pageNumber="29">
<heading id="D0EFDD368B314E054E88F98E3D3BF957" bold="true" box="[189,310,1650,1676]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B96CB6488B314E054E88F98E3D3BF957" bold="true" box="[189,310,1650,1676]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B314E054E88F95C3F0EF906" blockId="28.[189,1398,1696,1758]" pageId="28" pageNumber="29">
<taxonomicName id="4C1811D98B314E054E88F95C3DA4F961" authorityName="- van Benthem Jutting" authorityYear="1950" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[189,425,1696,1722]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054E88F95C3DA4F961" box="[189,425,1696,1722]" italics="true" pageId="28" pageNumber="29">Landouria epiplatia</emphasis>
</taxonomicName>
is characterized by a brownish, sharply keeled, almost disc-like shell with an only slightly elevated spire and a very wide umbilicus.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B314E054E88F8BE3F37F831" pageId="28" pageNumber="29" type="description">
<caption id="DF673AD28B314E054E88F8BE3F37F831" httpUri="https://zenodo.org/record/3152245/files/figure.png" pageId="28" pageNumber="29" targetBox="[221,1366,265,2019]" targetPageId="27">
<paragraph id="8BA76A5A8B314E054E88F8BE3F37F831" blockId="28.[189,1399,1858,2026]" pageId="28" pageNumber="29">
<emphasis id="B96CB6488B314E054E88F8BE3D4CF887" bold="true" box="[189,321,1858,1884]" pageId="28" pageNumber="29">Figs 2528</emphasis>
(opposite page). Shells of species of
<taxonomicName id="4C1811D98B314E054CC7F8BF3F63F886" authorityName="Godwin-Austen" authorityYear="1918" box="[754,878,1859,1885]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus">
<emphasis id="B96CB6488B314E054CC7F8BF3F63F886" box="[754,878,1859,1885]" italics="true" pageId="28" pageNumber="29">Landouria</emphasis>
</taxonomicName>
from
<collectingRegion id="49DCA4B88B314E054D81F8BF3FE1F886" box="[948,1004,1859,1885]" country="Indonesia" name="Jawa Timur" pageId="28" pageNumber="29">Java</collectingRegion>
.
<emphasis id="B96CB6488B314E054DC3F8BE381EF887" bold="true" box="[1014,1043,1858,1884]" pageId="28" pageNumber="29">25</emphasis>
.
<taxonomicName id="4C1811D98B314E054A14F8BF3880F887" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1057,1165,1859,1884]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="naggsi" status="sp. nov.">
<emphasis id="B96CB6488B314E054A14F8BF3880F887" box="[1057,1165,1859,1884]" italics="true" pageId="28" pageNumber="29">L. naggsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B314E054AA1F8BF38FDF886" box="[1172,1264,1859,1885]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B314E054ACEF8BF397FF886" box="[1275,1394,1859,1885]" country="Indonesia" name="Jawa Barat" pageId="28" pageNumber="29">West Java</collectingRegion>
, Cibitung, near Cikaso Waterfall (
<typeStatus id="54A3D4F88B314E054C7BF89A3EB4F85B" box="[590,697,1894,1920]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
, MZB 20424).
<emphasis id="B96CB6488B314E054D42F89A3F99F85B" bold="true" box="[887,916,1894,1920]" pageId="28" pageNumber="29">26</emphasis>
.
<taxonomicName id="4C1811D98B314E054D92F89B3923F85B" authority="(Mollendorff, 1897)" authorityName="Mollendorff" authorityYear="1897" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[935,1326,1894,1920]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054D92F89B383CF85B" box="[935,1073,1894,1920]" italics="true" pageId="28" pageNumber="29">L. epiplatia</emphasis>
(
<bibRefCitation id="EF8917AB8B314E054A71F89A3928F85B" author="Mollendorff" box="[1092,1317,1894,1920]" firstAuthor="Mollendorff" pageId="28" pageNumber="29" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B314E054B08F89A3CF9F878" country="Indonesia" name="Jawa Barat" pageId="28" pageNumber="29">West Java</collectingRegion>
, Jampang (
<typeStatus id="54A3D4F88B314E054F4EF8753DE5F878" box="[379,488,1929,1955]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C1811D98B314E054C21F8753F19F878" authorityName="Mollendorff" authorityYear="1897" box="[532,788,1929,1955]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054C21F8753F19F878" box="[532,788,1929,1955]" italics="true" pageId="28" pageNumber="29">Plectotropis epiplatia</emphasis>
</taxonomicName>
, SMF 8909).
<emphasis id="B96CB6488B314E054D8BF8753FD6F878" bold="true" box="[958,987,1929,1955]" pageId="28" pageNumber="29">27</emphasis>
.
<taxonomicName id="4C1811D98B314E054DD9F8763878F878" authorityName="Mollendorff" authorityYear="1897" box="[1004,1141,1929,1955]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054DD9F8763878F878" box="[1004,1141,1929,1955]" italics="true" pageId="28" pageNumber="29">L. epiplatia</emphasis>
</taxonomicName>
,
<collectingRegion id="49DCA4B88B314E054AB3F875390DF878" box="[1158,1280,1929,1955]" country="Indonesia" name="Jawa Barat" pageId="28" pageNumber="29">West Java</collectingRegion>
, Jampang (
<typeStatus id="54A3D4F88B314E054EF0F8503D3FF81D" box="[197,306,1964,1990]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C1811D98B314E054F6EF8503F89F81D" authority="Mollendorff, 1897" authorityName="Mollendorff" authorityYear="1897" box="[347,900,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B314E054F6EF8503EACF81D" box="[347,673,1964,1990]" italics="true" pageId="28" pageNumber="29">Plectotropis trichotrochium</emphasis>
<bibRefCitation id="EF8917AB8B314E054C9CF8503F89F81D" author="Mollendorff" box="[681,900,1964,1990]" firstAuthor="Mollendorff" pageId="28" pageNumber="29" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
</taxonomicName>
, SMF 8918).
<emphasis id="B96CB6488B314E054A1DF8503848F81D" bold="true" box="[1064,1093,1964,1990]" pageId="28" pageNumber="29">28</emphasis>
.
<taxonomicName id="4C1811D98B314E054A60F85138D0F81D" authorityName="Mollendorff" authorityYear="1897" box="[1109,1245,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B314E054A60F85138D0F81D" box="[1109,1245,1964,1990]" italics="true" pageId="28" pageNumber="29">L. epiplatia</emphasis>
</taxonomicName>
?,
<collectingRegion id="49DCA4B88B314E054ACCF851397FF81C" box="[1273,1394,1965,1991]" country="Indonesia" name="Jawa Barat" pageId="28" pageNumber="29">West Java</collectingRegion>
, Bogor, Gunung Gede (ZMH 79946). Scale bar:
<quantity id="4CE0C7BF8B314E054CDAF8333F3BF831" box="[751,822,1999,2026]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="28" pageNumber="29" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C30239D18B304E074E88FEF7387FFE93" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BA76A5A8B304E044E88FEF73DA7FEFE" blockId="29.[189,1399,267,477]" box="[189,426,267,293]" pageId="29" pageNumber="30">
<heading id="D0EFDD368B304E044E88FEF73DA7FEFE" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B96CB6488B304E044E88FEF73DA7FEFE" bold="true" box="[189,426,267,293]" pageId="29" pageNumber="30">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B304E044EDFFECA3D94FE94" blockId="29.[189,1399,267,477]" box="[234,409,309,336]" pageId="29" pageNumber="30">
<emphasis id="B96CB6488B304E044EDFFECA3D94FE94" bold="true" box="[234,409,309,336]" pageId="29" pageNumber="30">
<typeStatus id="54A3D4F88B304E044EDFFECA3D2AFE8B" box="[234,295,310,336]" pageId="29" pageNumber="30">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="8BA76A5A8B304E044E88FEA53CF2FE06" blockId="29.[189,1399,267,477]" pageId="29" pageNumber="30">
<materialsCitation id="3B7060078B304E044E88FEA53ED3FE4D" ID-GBIF-Occurrence="2243137320" collectionCode="SMF" country="Indonesia" elevation="600" latitude="-7.05" location="Jampang" longLatPrecision="1303" longitude="106.8" pageId="29" pageNumber="30" specimenCode="SMF 8909" specimenCount="1" stateProvince="West Java" typeStatus="lectotype">
<collectingCountry id="F30F2ACA8B304E044E88FEA53D56FEA8" box="[189,347,345,371]" name="Indonesia" pageId="29" pageNumber="30">INDONESIA</collectingCountry>
<collectingRegion id="49DCA4B88B304E044F4EFEA53DF1FEA8" box="[379,508,345,371]" country="Indonesia" name="Jawa Barat" pageId="29" pageNumber="30">
<emphasis id="B96CB6488B304E044F4EFEA53DF1FEA8" bold="true" box="[379,508,345,371]" pageId="29" pageNumber="30">West Java</emphasis>
</collectingRegion>
<typeStatus id="54A3D4F88B304E044C2DFEA53E89FEA8" box="[536,644,345,371]" pageId="29" pageNumber="30" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C1811D98B304E044C98FEA53FA1FEA8" authorityName="Mollendorff" authorityYear="1897" box="[685,940,345,371]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B304E044C98FEA53FA1FEA8" box="[685,940,345,371]" italics="true" pageId="29" pageNumber="30">
<originalDetermination id="A806D2A68B304E044C98FEA53FA1FEA8" box="[685,940,345,371]" pageId="29" pageNumber="30">Plectotropis epiplatia</originalDetermination>
</emphasis>
</taxonomicName>
(designated by
<bibRefCitation id="EF8917AB8B304E044A5FFEA538FDFEAF" author="Zilch" box="[1130,1264,345,372]" firstAuthor="Zilch" pageId="29" pageNumber="30" pagination="293 - 319" refId="ref35814" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch 1966</bibRefCitation>
);
<location id="8EC73C818B304E044B33FEA5397FFEA8" LSID="urn:lsid:plazi:treatment:03B1DB4C8B314E074C1DFB3F3E98F856:8EC73C818B304E044B33FEA5397FFEA8" box="[1286,1394,345,371]" country="Indonesia" latitude="-7.05" longLatPrecision="1303" longitude="106.8" name="Jampang" pageId="29" pageNumber="30" stateProvince="West Java">Jampang</location>
;
<geoCoordinate id="EE2C0C9D8B304E044E88FE863D18FE4D" box="[189,277,378,406]" degrees="7" direction="south" minutes="03" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-7.05">7°03 S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B304E044F17FE863D97FE4C" box="[290,410,378,407]" degrees="106" direction="east" minutes="48" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="106.8">106°48 E</geoCoordinate>
; ca
<elevation id="00358D698B304E044FFCFE803E5EFE4D" box="[457,595,380,407]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="m" value="600.0">
<quantity id="4CE0C7BF8B304E044FFCFE803E1EFE4C" box="[457,531,380,407]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
;
<specimenCode id="DBBEC2218B304E044C54FE803ED3FE4D" box="[609,734,380,406]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="29" pageNumber="30">SMF 8909</specimenCode>
</materialsCitation>
<materialsCitation id="3B7060078B304E044CCDFE803DE7FE62" ID-GBIF-Occurrence="2243137413" collectionCode="SMF" country="Indonesia" elevation="600" latitude="-7.05" location="Jampang" longLatPrecision="1303" longitude="106.8" pageId="29" pageNumber="30" specimenCode="SMF 8910" specimenCount="1" stateProvince="Jawa Barat" typeStatus="holotype">
<specimenCount id="9D1EA1D38B304E044CCDFE803FA1FE4D" box="[760,940,380,406]" count="1" pageId="29" pageNumber="30" type="generic" typeStatus="paralectotype">1 paralectotype</specimenCount>
of
<taxonomicName id="4C1811D98B304E044DE1FE8038DFFE4D" authorityName="Mollendorff" authorityYear="1897" box="[980,1234,380,406]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B304E044DE1FE8038DFFE4D" box="[980,1234,380,406]" italics="true" pageId="29" pageNumber="30">
<originalDetermination id="A806D2A68B304E044DE1FE8038DFFE4D" box="[980,1234,380,406]" pageId="29" pageNumber="30">Plectotropis epiplatia</originalDetermination>
</emphasis>
</taxonomicName>
; same data as for preceding;
<specimenCode id="DBBEC2218B304E044F59FE633DE7FE62" box="[364,490,415,441]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="29" pageNumber="30">SMF 8910</specimenCode>
</materialsCitation>
<materialsCitation id="3B7060078B304E044C31FE633CF6FE06" ID-GBIF-Occurrence="2243137331" collectionCode="SMF" country="Indonesia" elevation="600" latitude="-7.05" location="Jampang" longLatPrecision="1303" longitude="106.8" pageId="29" pageNumber="30" specimenCode="SMF 8918" specimenCount="1" stateProvince="Jawa Barat" typeStatus="lectotype">
<typeStatus id="54A3D4F88B304E044C31FE633E7DFE62" box="[516,624,415,441]" pageId="29" pageNumber="30" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C1811D98B304E044CACFE633FD3FE62" authorityName="Mollendorff" authorityYear="1897" box="[665,990,415,441]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B304E044CACFE633FD3FE62" box="[665,990,415,441]" italics="true" pageId="29" pageNumber="30">
<originalDetermination id="A806D2A68B304E044CACFE633FD3FE62" box="[665,990,415,441]" pageId="29" pageNumber="30">Plectotropis trichotrochium</originalDetermination>
</emphasis>
</taxonomicName>
; same data as for preceding;
<specimenCode id="DBBEC2218B304E044B09FE633CF6FE06" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="29" pageNumber="30">SMF 8918</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA76A5A8B304E074EDFFE00387FFE93" blockId="29.[189,1399,508,605]" lastBlockId="30.[189,1399,265,329]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B96CB6488B304E044EDFFE003DABFDCD" bold="true" box="[234,422,508,534]" pageId="29" pageNumber="30">Other material</emphasis>
<materialsCitation id="3B7060078B304E044E88FDDC3EC4FDE2" ID-GBIF-Occurrence="2243140019" box="[189,713,543,570]" collectionCode="ZMB" country="Indonesia" location="Java" pageId="29" pageNumber="30" specimenCode="47682" specimenCount="1" stateProvince="Java">
<collectingCountry id="F30F2ACA8B304E044E88FDDC3D56FDE1" box="[189,347,544,570]" name="Indonesia" pageId="29" pageNumber="30">INDONESIA</collectingCountry>
<specimenCount id="9D1EA1D38B304E044F4CFDDC3DDDFDE1" box="[377,464,544,570]" count="1" pageId="29" pageNumber="30" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="49DCA4B88B304E044FDDFDDC3E2DFDE1" box="[488,544,544,570]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">Java</collectingRegion>
;
<collectionCode id="ED09F29F8B304E044C1BFDDC3E63FDE1" box="[558,622,544,570]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="29" pageNumber="30" type="Museum">ZMB</collectionCode>
<specimenCode id="DBBEC2218B304E044C42FDE33EC9FDE2" box="[631,708,543,569]" pageId="29" pageNumber="30">47682</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B7060078B304E044CE7FDDC3E37FD86" ID-GBIF-Occurrence="2243140019" collectionCode="SMF" country="Indonesia" latitude="-7.0" location="West Java" longLatPrecision="1303" longitude="106.583336" pageId="29" pageNumber="30" specimenCode="96542" specimenCount="4" stateProvince="West Java">
<collectingRegion id="49DCA4B88B304E044CDFFDDC3F60FDE1" box="[746,877,544,570]" country="Indonesia" name="Jawa Barat" pageId="29" pageNumber="30">
<emphasis id="B96CB6488B304E044CDFFDDC3F60FDE1" bold="true" box="[746,877,544,570]" pageId="29" pageNumber="30">West Java</emphasis>
</collectingRegion>
<specimenCount id="9D1EA1D38B304E044DBFFDDC3FECFDE1" box="[906,993,544,570]" count="4" pageId="29" pageNumber="30" type="generic">4 spec.</specimenCount>
;
<locationDeviation id="9A1573788B304E044DC5FDE3397FFDE1" box="[1008,1394,543,570]" pageId="29" pageNumber="30">
<quantity id="4CE0C7BF8B304E044DC5FDE33820FDE2" box="[1008,1069,543,570]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="km" value="5.0">5 km</quantity>
from Palabuhan Ratu Bay
</locationDeviation>
;
<geoCoordinate id="EE2C0C9D8B304E044E88FDBD3D1BFD86" box="[189,278,577,605]" degrees="7" direction="south" minutes="00" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-7.0">7°00 S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B304E044F10FDBD3D92FD86" box="[293,415,577,605]" degrees="106" direction="east" minutes="35" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="106.583336">106°35 E</geoCoordinate>
;
<collectionCode id="ED09F29F8B304E044F98FDBF3DE5FD86" box="[429,488,579,605]" collectionName="Germany, Frankfurt-am-Main, Forschungsinstitut und Naturmuseum Senckenberg" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="29" pageNumber="30">SMF</collectionCode>
<specimenCode id="DBBEC2218B304E044FC4FDBE3E37FD86" box="[497,570,578,605]" pageId="29" pageNumber="30">96542</specimenCode>
</materialsCitation>
<materialsCitation id="3B7060078B304E044C62FDBF396EFD86" ID-GBIF-Occurrence="2243140018" box="[599,1379,577,605]" collectionCode="SMF" country="Indonesia" elevation="600" latitude="-7.05" location="Jampang" longLatPrecision="1303" longitude="106.8" pageId="29" pageNumber="30" specimenCode="SMF 8911" specimenCount="2" stateProvince="Jawa Barat">
<specimenCount id="9D1EA1D38B304E044C62FDBF3EA3FD86" box="[599,686,579,605]" count="2" pageId="29" pageNumber="30" type="generic">2 spec.</specimenCount>
;
<location id="8EC73C818B304E044C89FDBF3F25FD86" LSID="urn:lsid:plazi:treatment:03B1DB4C8B314E074C1DFB3F3E98F856:8EC73C818B304E044C89FDBF3F25FD86" box="[700,808,579,605]" country="Indonesia" latitude="-7.05" longLatPrecision="1303" longitude="106.8" name="Jampang" pageId="29" pageNumber="30" stateProvince="Jawa Barat">Jampang</location>
;
<geoCoordinate id="EE2C0C9D8B304E044D00FDBD3F83FD86" box="[821,910,577,605]" degrees="7" direction="south" minutes="03" orientation="latitude" pageId="29" pageNumber="30" precision="925" value="-7.05">7°03 S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B304E044DA8FDBD381AFD86" box="[925,1047,577,605]" degrees="106" direction="east" minutes="48" orientation="longitude" pageId="29" pageNumber="30" precision="925" value="106.8">106°48 E</geoCoordinate>
; ca
<elevation id="00358D698B304E044A7DFDBF38DBFD86" box="[1096,1238,579,605]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="m" value="600.0">
<quantity id="4CE0C7BF8B304E044A7DFDBF3898FD86" box="[1096,1173,579,605]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="29" pageNumber="30" unit="m" value="600.0">600 m</quantity>
a.s.l.
</elevation>
;
<specimenCode id="DBBEC2218B304E044AD3FDBF396EFD86" box="[1254,1379,579,605]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="29" pageNumber="30">SMF 8911</specimenCode>
</materialsCitation>
<materialsCitation id="3B7060078B334E074E88FEF73903FEFE" ID-GBIF-Occurrence="2243140021" box="[189,1294,265,294]" collectionCode="ZMA" country="Indonesia" latitude="-6.9166665" location="Sukabumi" longLatPrecision="1303" longitude="106.916664" pageId="30" pageNumber="31" specimenCode="ZMA 407610" specimenCount="2" stateProvince="Jawa Barat">
<specimenCount id="9D1EA1D38B334E074E88FEF73D03FEFD" box="[189,270,267,294]" count="2" pageId="30" pageNumber="31" type="generic">2 spec.</specimenCount>
(vouchers for
<bibRefCitation id="EF8917AB8B334E074F8DFEF03EE2FEFD" author="van Benthem Jutting" box="[440,751,267,294]" firstAuthor="van Benthem Jutting" pageId="30" pageNumber="31" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting 1950</bibRefCitation>
);
<location id="8EC73C818B334E074D36FEF73F73FEFE" LSID="urn:lsid:plazi:treatment:03B1DB4C8B314E074C1DFB3F3E98F856:8EC73C818B334E074D36FEF73F73FEFE" box="[771,894,267,293]" country="Indonesia" latitude="-6.9166665" longLatPrecision="1303" longitude="106.916664" name="Sukabumi" pageId="30" pageNumber="31" stateProvince="Jawa Barat">Sukabumi</location>
;
<geoCoordinate id="EE2C0C9D8B334E074DBCFEF53FEDFEFE" box="[905,992,265,293]" degrees="6" direction="south" minutes="55" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="-6.9166665">6°55 S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B334E074DD8FEF53869FEFD" box="[1005,1124,265,294]" degrees="106" direction="east" minutes="55" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="106.916664">106°55 E</geoCoordinate>
;
<specimenCode id="DBBEC2218B334E074A45FEF03903FEFE" box="[1136,1294,267,294]" collectionCode="ZMA" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="30" pageNumber="31">ZMA 407610</specimenCode>
</materialsCitation>
<materialsCitation id="3B7060078B334E074B13FEF73863FE93" ID-GBIF-Occurrence="2243137407" collectionCode="ZMH" country="Indonesia" latitude="-6.7666664" location="Gunung Gede" longLatPrecision="1304" longitude="106.98333" municipality="Bogor" pageId="30" pageNumber="31" specimenCode="79946" specimenCount="1" stateProvince="Jawa Barat">
<specimenCount id="9D1EA1D38B334E074B13FEF7397AFEFD" box="[1318,1399,267,294]" count="1" pageId="30" pageNumber="31" type="generic">1 spec.</specimenCount>
(det. anat.; see remarks);
<collectingMunicipality id="6BC3F0208B334E074FD0FED33E3DFE92" box="[485,560,303,329]" pageId="30" pageNumber="31">Bogor</collectingMunicipality>
,
<location id="8EC73C818B334E074C09FED33EE9FE93" LSID="urn:lsid:plazi:treatment:03B1DB4C8B314E074C1DFB3F3E98F856:8EC73C818B334E074C09FED33EE9FE93" box="[572,740,302,329]" country="Indonesia" latitude="-6.7666664" longLatPrecision="1304" longitude="106.98333" municipality="Bogor" name="Gunung Gede" pageId="30" pageNumber="31" stateProvince="Jawa Barat">Gunung Gede</location>
;
<geoCoordinate id="EE2C0C9D8B334E074CDAFED13F4AFE92" box="[751,839,301,329]" degrees="6" direction="south" minutes="46" orientation="latitude" pageId="30" pageNumber="31" precision="925" value="-6.7666664">6°46 S</geoCoordinate>
,
<geoCoordinate id="EE2C0C9D8B334E074D61FED13FC1FE92" box="[852,972,301,329]" degrees="106" direction="east" minutes="59" orientation="longitude" pageId="30" pageNumber="31" precision="925" value="106.98333">106°59 E</geoCoordinate>
;
<collectionCode id="ED09F29F8B334E074DEDFED33814FE92" box="[984,1049,303,329]" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="30" pageNumber="31" type="Museum">ZMH</collectionCode>
<specimenCode id="DBBEC2218B334E074A14FED23863FE93" box="[1057,1134,302,328]" pageId="30" pageNumber="31">79946</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF673AD28B304E044E88FAAA3843F831" httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="29" pageNumber="30" startId="29.[189,241,1366,1392]" targetBox="[189,1399,649,1337]" targetPageId="29">
<paragraph id="8BA76A5A8B304E044E88FAAA3843F831" blockId="29.[189,1399,1366,2026]" pageId="29" pageNumber="30">
<emphasis id="B96CB6488B304E044E88FAAA3D47FAAB" bold="true" box="[189,330,1366,1392]" pageId="29" pageNumber="30">Figs 2947.</emphasis>
Flagella of species of
<taxonomicName id="4C1811D98B304E044C62FAAA3EDEFAAB" authorityName="Godwin-Austen" authorityYear="1918" box="[599,723,1366,1392]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="genus">
<emphasis id="B96CB6488B304E044C62FAAA3EDEFAAB" box="[599,723,1366,1392]" italics="true" pageId="29" pageNumber="30">Landouria</emphasis>
</taxonomicName>
from
<collectingRegion id="49DCA4B88B304E044D29FAAA3F59FAAB" box="[796,852,1366,1392]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">Java</collectingRegion>
.
<emphasis id="B96CB6488B304E044D55FAAA3F70FAAB" bold="true" box="[864,893,1366,1392]" pageId="29" pageNumber="30">29</emphasis>
.
<taxonomicName id="4C1811D98B304E044DB8FAAB38E0FAAB" authority="(Pfeiffer, 1842)" authorityName="Pfeiffer" authorityYear="1842" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[909,1261,1366,1392]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="winteriana">
<emphasis id="B96CB6488B304E044DB8FAAB3821FAAB" box="[909,1068,1367,1392]" italics="true" pageId="29" pageNumber="30">L. winteriana</emphasis>
(
<bibRefCitation id="EF8917AB8B304E044A09FAAA38E9FAAB" author="Pfeiffer" box="[1084,1252,1366,1392]" firstAuthor="Pfeiffer" pageId="29" pageNumber="30" refId="ref34385" refString="Pfeiffer L. 1842. Symbolae ad Historiam Heliceorum. Sectio alterna. T. Fischer, Cassel [Kassel, Germany]. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B304E044ACCFAAA397FFAAB" box="[1273,1394,1366,1392]" country="Indonesia" name="Jawa Barat" pageId="29" pageNumber="30">West Java</collectingRegion>
, Padalarang, Gunung Masigit (
<typeStatus id="54A3D4F88B304E044C1DFA853E87FA48" box="[552,650,1401,1427]" pageId="29" pageNumber="30" type="neotype">neotype</typeStatus>
, ZMA 376193a).
<emphasis id="B96CB6488B304E044D50FA853F8FFA48" bold="true" box="[869,898,1401,1427]" pageId="29" pageNumber="30">30</emphasis>
.
<taxonomicName id="4C1811D98B304E044DA0FA8638EAFA48" authority="(Pfeiffer, 1842)" authorityName="Pfeiffer" authorityYear="1842" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[917,1255,1401,1427]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="rotatoria">
<emphasis id="B96CB6488B304E044DA0FA86382DFA48" box="[917,1056,1402,1427]" italics="true" pageId="29" pageNumber="30">L. rotatoria</emphasis>
(
<bibRefCitation id="EF8917AB8B304E044A06FA8538D3FA48" author="Pfeiffer" box="[1075,1246,1401,1427]" firstAuthor="Pfeiffer" pageId="29" pageNumber="30" refId="ref34385" refString="Pfeiffer L. 1842. Symbolae ad Historiam Heliceorum. Sectio alterna. T. Fischer, Cassel [Kassel, Germany]. https: // doi. org / 10.5962 / bhl. title. 11903" type="book" year="1842">Pfeiffer, 1842</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B304E044AC3FA86397FFA4F" box="[1270,1394,1402,1428]" country="Indonesia" name="Jawa Barat" pageId="29" pageNumber="30">West Java</collectingRegion>
, Gunung Salak,
<quantity id="4CE0C7BF8B304E044F41FA603DC0FA6C" box="[372,461,1436,1463]" metricMagnitude="3" metricUnit="m" metricValue="1.166" pageId="29" pageNumber="30" unit="m" value="1166.0">1166 m</quantity>
a.s.l. (
<typeStatus id="54A3D4F88B304E044C2EFA603E70FA6D" box="[539,637,1436,1462]" pageId="29" pageNumber="30" type="neotype">neotype</typeStatus>
, MZB 19205).
<emphasis id="B96CB6488B304E044D01FA603F5CFA6D" bold="true" box="[820,849,1436,1462]" pageId="29" pageNumber="30">31</emphasis>
.
<taxonomicName id="4C1811D98B304E044D55FA6138ECFA6D" authority="(Mollendorff, 1897)" authorityName="Mollendorff" authorityYear="1897" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[864,1249,1436,1462]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B304E044D55FA613FE5FA6D" box="[864,1000,1436,1462]" italics="true" pageId="29" pageNumber="30">L. epiplatia</emphasis>
(
<bibRefCitation id="EF8917AB8B304E044DCDFA6038DAFA6D" author="Mollendorff" box="[1016,1239,1436,1462]" firstAuthor="Mollendorff" pageId="29" pageNumber="30" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
)
</taxonomicName>
?,
<collectingRegion id="49DCA4B88B304E044ACCFA61397FFA6C" box="[1273,1394,1437,1463]" country="Indonesia" name="Jawa Barat" pageId="29" pageNumber="30">West Java</collectingRegion>
, Bogor, Gunung Gede (ZMH 79946).
<emphasis id="B96CB6488B304E044C45FA433E80FA02" bold="true" box="[624,653,1471,1497]" pageId="29" pageNumber="30">32</emphasis>
.
<taxonomicName id="4C1811D98B304E044CA9FA3C3F95FA01" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[668,920,1472,1498]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="nusakambangensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044CA9FA3C3F95FA01" box="[668,920,1472,1498]" italics="true" pageId="29" pageNumber="30">L. nusakambangensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044DABFA3C3FF6FA01" box="[926,1019,1472,1498]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044A33FA433897FA01" box="[1030,1178,1471,1498]" country="Indonesia" name="Jawa Tengah" pageId="29" pageNumber="30">Central Java</collectingRegion>
, Nusa Kambangan Island, nature monument Gligir (
<typeStatus id="54A3D4F88B304E044C7CFA1F3EB9FA26" box="[585,692,1507,1533]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, ZMA 412298a).
<emphasis id="B96CB6488B304E044DB9FA1F3FA4FA26" bold="true" box="[908,937,1507,1533]" pageId="29" pageNumber="30">33</emphasis>
.
<taxonomicName id="4C1811D98B304E044D8EFA18386EFA26" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[955,1123,1507,1533]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="petrukensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044D8EFA18386EFA26" box="[955,1123,1507,1533]" italics="true" pageId="29" pageNumber="30">L. petrukensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044A59FA1F38C1FA26" box="[1132,1228,1507,1533]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044AEFFA1F397FFA26" box="[1242,1394,1507,1533]" country="Indonesia" name="Jawa Tengah" pageId="29" pageNumber="30">Central Java</collectingRegion>
, Karangbolong, way to Petruk Cave (
<typeStatus id="54A3D4F88B304E044C4DF9FA3EEEF9FB" box="[632,739,1542,1568]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 19203).
<emphasis id="B96CB6488B304E044D95F9FA3FB0F9FB" bold="true" box="[928,957,1542,1568]" pageId="29" pageNumber="30">34</emphasis>
.
<taxonomicName id="4C1811D98B304E044DFAF9FB387FF9FB" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[975,1138,1542,1568]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="tholiformis" status="sp. nov.">
<emphasis id="B96CB6488B304E044DFAF9FB387FF9FB" box="[975,1138,1542,1568]" italics="true" pageId="29" pageNumber="30">L. tholiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044A49F9FA38D6F9FB" box="[1148,1243,1542,1568]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044ADCF9FA397FF9FB" box="[1257,1394,1542,1568]" country="Indonesia" name="Yogyakarta" pageId="29" pageNumber="30">Yogyakarta</collectingRegion>
, Gunung Kidul, Tepus, near Poktunggal Beach (
<typeStatus id="54A3D4F88B304E044D30F9D53F7DF998" box="[773,880,1577,1603]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 17999).
<emphasis id="B96CB6488B304E044A06F9D5385DF998" bold="true" box="[1075,1104,1577,1603]" pageId="29" pageNumber="30">35</emphasis>
.
<taxonomicName id="4C1811D98B304E044A51F9D63909F998" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1124,1284,1577,1603]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="abdidalem" status="sp. nov.">
<emphasis id="B96CB6488B304E044A51F9D63909F998" box="[1124,1284,1577,1603]" italics="true" pageId="29" pageNumber="30">L. abdidalem</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044B25F9D6397FF99F" box="[1296,1394,1578,1604]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044E88F9B03D4BF9BD" box="[189,326,1612,1638]" country="Indonesia" name="Yogyakarta" pageId="29" pageNumber="30">Yogyakarta</collectingRegion>
, Kulonprogo, Sabrang Kidul (
<typeStatus id="54A3D4F88B304E044C90F9B03F1DF9BD" box="[677,784,1612,1638]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 17664).
<emphasis id="B96CB6488B304E044DF6F9B03FEDF9BD" bold="true" box="[963,992,1612,1638]" pageId="29" pageNumber="30">36</emphasis>
.
<taxonomicName id="4C1811D98B304E044DDAF9B1389EF9BD" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1007,1171,1612,1638]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="madurensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044DDAF9B1389EF9BD" box="[1007,1171,1612,1638]" italics="true" pageId="29" pageNumber="30">L. madurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044AAFF9B138FBF9BC" box="[1178,1270,1613,1639]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044B37F9B1397FF9BC" box="[1282,1394,1613,1639]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">East Java</collectingRegion>
, Madura Island, Sampang, Nepa Monkey lowland rainforest (
<typeStatus id="54A3D4F88B304E044DA5F98C3FF7F951" box="[912,1018,1648,1674]" pageId="29" pageNumber="30" type="paratype">paratype</typeStatus>
, MZB 19209).
<emphasis id="B96CB6488B304E044A86F99338DDF952" bold="true" box="[1203,1232,1647,1673]" pageId="29" pageNumber="30">37</emphasis>
.
<taxonomicName id="4C1811D98B304E044AD5F98C397BF952" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1248,1398,1648,1673]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="sewuensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044AD5F98C397BF952" box="[1248,1398,1648,1673]" italics="true" pageId="29" pageNumber="30">L. sewuensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044E88F96F3D11F976" box="[189,284,1683,1709]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044F1CF96F3DBFF976" box="[297,434,1683,1709]" country="Indonesia" name="Yogyakarta" pageId="29" pageNumber="30">Yogyakarta</collectingRegion>
, Gunung Kidul, Tepus, Mbajo Hill (
<typeStatus id="54A3D4F88B304E044D5DF96F3FDEF976" box="[872,979,1683,1709]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 19255).
<emphasis id="B96CB6488B304E044ABAF96F38A1F976" bold="true" box="[1167,1196,1683,1709]" pageId="29" pageNumber="30">38</emphasis>
.
<taxonomicName id="4C1811D98B304E044A88F968397AF976" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1213,1399,1683,1709]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="sukoliloensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044A88F968397AF976" box="[1213,1399,1683,1709]" italics="true" pageId="29" pageNumber="30">L. sukoliloensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044E88F94A3D16F90B" box="[189,283,1718,1744]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044F12F94A3DB0F90B" box="[295,445,1718,1744]" country="Indonesia" name="Jawa Tengah" pageId="29" pageNumber="30">Central Java</collectingRegion>
, Kayen, near Watu Payung (
<typeStatus id="54A3D4F88B304E044D38F94A3F75F90B" box="[781,888,1718,1744]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 19219).
<emphasis id="B96CB6488B304E044A1BF94A3846F90B" bold="true" box="[1070,1099,1718,1744]" pageId="29" pageNumber="30">39</emphasis>
.
<taxonomicName id="4C1811D98B304E044A6EF94B3901F90B" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1115,1292,1718,1744]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="tonywhitteni" status="sp. nov.">
<emphasis id="B96CB6488B304E044A6EF94B3901F90B" box="[1115,1292,1718,1744]" italics="true" pageId="29" pageNumber="30">L. tonywhitteni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044B21F94A397FF90B" box="[1300,1394,1718,1744]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044E88F9253D5FF92F" box="[189,338,1753,1780]" country="Indonesia" name="Jawa Tengah" pageId="29" pageNumber="30">Central Java</collectingRegion>
, Taruman Village, near Sendang Sedayu (
<typeStatus id="54A3D4F88B304E044D08F9253FA5F928" box="[829,936,1753,1779]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 19214).
<emphasis id="B96CB6488B304E044A6BF9253876F928" bold="true" box="[1118,1147,1753,1779]" pageId="29" pageNumber="30">40</emphasis>
.
<taxonomicName id="4C1811D98B304E044ABFF9263901F928" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1162,1292,1753,1779]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="nodifera" status="sp. nov.">
<emphasis id="B96CB6488B304E044ABFF9263901F928" box="[1162,1292,1753,1779]" italics="true" pageId="29" pageNumber="30">L. nodifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044B21F926397FF92F" box="[1300,1394,1754,1780]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044E88F9003D42F8CC" box="[189,335,1788,1815]" country="Indonesia" name="Jawa Tengah" pageId="29" pageNumber="30">Central Java</collectingRegion>
, Larangan Village, Lanang Spring (
<typeStatus id="54A3D4F88B304E044CD2F9003F5FF8CD" box="[743,850,1788,1814]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 19227).
<emphasis id="B96CB6488B304E044A35F9003810F8CD" bold="true" box="[1024,1053,1788,1814]" pageId="29" pageNumber="30">41</emphasis>
.
<taxonomicName id="4C1811D98B304E044A1FF90138D7F8CD" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1066,1242,1789,1814]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="pacitanensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044A1FF90138D7F8CD" box="[1066,1242,1789,1814]" italics="true" pageId="29" pageNumber="30">L. pacitanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044AEAF9013937F8CC" box="[1247,1338,1789,1815]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044B71F9013CF9F8E1" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">East Java</collectingRegion>
, Pacitan, near Srau Beach (MZB 19231).
<emphasis id="B96CB6488B304E044CE0F8E33EFFF8E2" bold="true" box="[725,754,1823,1849]" pageId="29" pageNumber="30">42</emphasis>
.
<taxonomicName id="4C1811D98B304E044D35F8DC387CF8E1" authority="(Martens, 1867)" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" baseAuthorityName="Martens" baseAuthorityYear="1867" box="[768,1137,1823,1850]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="intumescens">
<emphasis id="B96CB6488B304E044D35F8DC3FA1F8E2" box="[768,940,1824,1849]" italics="true" pageId="29" pageNumber="30">L. intumescens</emphasis>
(
<bibRefCitation id="EF8917AB8B304E044D8EF8DC3865F8E1" author="Martens" box="[955,1128,1823,1850]" firstAuthor="Martens" pageId="29" pageNumber="30" refId="ref33696" refString="Martens E. von. 1867. Die Landschnecken. In: Die Preussische Expedition nach Ost-Asien. Nach amtlichen Quellen. Zoologischer Theil. Vol. 2. Decker, Berlin. https: // doi. org / 10.5962 / bhl. title. 12726" type="book" year="1867">Martens, 1867</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B304E044A49F8DC38E6F8E1" box="[1148,1259,1824,1850]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">East Java</collectingRegion>
, Guwoterus (MZB 19207).
<emphasis id="B96CB6488B304E044F43F8BF3D9EF886" bold="true" box="[374,403,1859,1885]" pageId="29" pageNumber="30">43</emphasis>
.
<taxonomicName id="4C1811D98B304E044F90F8B83F14F886" authority="(Mousson, 1848)" authorityName="Mousson" authorityYear="1848" baseAuthorityName="Mousson" baseAuthorityYear="1848" box="[421,793,1859,1885]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="smimensis">
<emphasis id="B96CB6488B304E044F90F8B83E4DF886" box="[421,576,1860,1885]" italics="true" pageId="29" pageNumber="30">L. smimensis</emphasis>
(
<bibRefCitation id="EF8917AB8B304E044C67F8BF3F02F886" author="Mousson" box="[594,783,1859,1885]" firstAuthor="Mousson" pageId="29" pageNumber="30" pagination="264 - 273" refId="ref33832" refString="Mousson A. 1848. Ueber die Land- und Susswassermollusken von Java. Mittheilungen der naturforschenden Gesellschaft in Zurich 1: 264 - 273." type="journal article" year="1848">Mousson, 1848</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B304E044D12F8BF3F91F886" box="[807,924,1859,1885]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">East Java</collectingRegion>
, Tosari (ZMA 390881).
<emphasis id="B96CB6488B304E044AF0F8BF38EFF886" bold="true" box="[1221,1250,1859,1885]" pageId="29" pageNumber="30">44</emphasis>
.
<taxonomicName id="4C1811D98B304E044AC1F8B8397BF886" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1268,1398,1859,1885]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="zonifera" status="sp. nov.">
<emphasis id="B96CB6488B304E044AC1F8B8397BF886" box="[1268,1398,1859,1885]" italics="true" pageId="29" pageNumber="30">L. zonifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044E88F89A3D11F85B" box="[189,284,1894,1920]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044F1CF89A3DBFF85B" box="[297,434,1894,1920]" country="Indonesia" name="Yogyakarta" pageId="29" pageNumber="30">Yogyakarta</collectingRegion>
, Kulonprogo, Sibolong (
<typeStatus id="54A3D4F88B304E044CEEF89A3F4BF85B" box="[731,838,1894,1920]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 17727).
<emphasis id="B96CB6488B304E044A34F89A3813F85B" bold="true" box="[1025,1054,1894,1920]" pageId="29" pageNumber="30">45</emphasis>
.
<taxonomicName id="4C1811D98B304E044A05F89B3D09F878" authority="(Mollendorff, 1897)" authorityName="Mollendorff" authorityYear="1897" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="ciliocincta">
<emphasis id="B96CB6488B304E044A05F89B38C0F85B" box="[1072,1229,1894,1920]" italics="true" pageId="29" pageNumber="30">L. ciliocincta</emphasis>
(
<bibRefCitation id="EF8917AB8B304E044AEBF89A3CF6F878" author="Mollendorff" firstAuthor="Mollendorff" pageId="29" pageNumber="30" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
)
</taxonomicName>
,
<collectingRegion id="49DCA4B88B304E044F25F8763D8FF87F" box="[272,386,1930,1956]" country="Indonesia" name="Jawa Timur" pageId="29" pageNumber="30">East Java</collectingRegion>
, Malang, Kondangmerak (MZB 19211).
<emphasis id="B96CB6488B304E044D53F8753F8EF878" bold="true" box="[870,899,1929,1955]" pageId="29" pageNumber="30">46</emphasis>
.
<taxonomicName id="4C1811D98B304E044DA7F876382AF878" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[914,1063,1929,1955]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="pakidulan" status="sp. nov.">
<emphasis id="B96CB6488B304E044DA7F876382AF878" box="[914,1063,1929,1955]" italics="true" pageId="29" pageNumber="30">L. pakidulan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044A1AF8763881F87F" box="[1071,1164,1930,1956]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044AACF8753923F87F" box="[1177,1326,1929,1956]" country="Indonesia" name="Jawa Tengah" pageId="29" pageNumber="30">Central Java</collectingRegion>
, Nusa Kambangan Island, nature monument Gligir (
<typeStatus id="54A3D4F88B304E044CE9F8503F4AF81D" box="[732,839,1964,1990]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, ZMA 417011a).
<emphasis id="B96CB6488B304E044A2EF8503835F81D" bold="true" box="[1051,1080,1964,1990]" pageId="29" pageNumber="30">47</emphasis>
.
<taxonomicName id="4C1811D98B304E044A7CF8513907F81D" authority="Nurinsiyah &amp; Neiber &amp; Hausdorf, 2019" authorityName="Nurinsiyah &amp; Neiber &amp; Hausdorf" authorityYear="2019" box="[1097,1290,1964,1990]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="menorehensis" status="sp. nov.">
<emphasis id="B96CB6488B304E044A7CF8513907F81D" box="[1097,1290,1964,1990]" italics="true" pageId="29" pageNumber="30">L. menorehensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F0B338B304E044B26F851397FF81C" box="[1299,1394,1965,1991]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
,
<collectingRegion id="49DCA4B88B304E044E88F8333D4BF832" box="[189,326,1999,2025]" country="Indonesia" name="Yogyakarta" pageId="29" pageNumber="30">Yogyakarta</collectingRegion>
, Kulonprogo, Sibolong (
<typeStatus id="54A3D4F88B304E044C52F8333EDFF832" box="[615,722,1999,2025]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
, MZB 19201). Scale bar:
<quantity id="4CE0C7BF8B304E044A36F82C3847F831" box="[1027,1098,2000,2026]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<subSubSection id="C30239D18B334E074E88FE8B3F9FFC65" pageId="30" pageNumber="31" type="description">
<paragraph id="8BA76A5A8B334E074E88FE8B3D43FE4A" blockId="30.[189,334,375,401]" box="[189,334,375,401]" pageId="30" pageNumber="31">
<heading id="D0EFDD368B334E074E88FE8B3D43FE4A" bold="true" box="[189,334,375,401]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B96CB6488B334E074E88FE8B3D43FE4A" bold="true" box="[189,334,375,401]" pageId="30" pageNumber="31">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B334E074E88FE573D8CFD46" blockId="30.[189,1399,426,669]" pageId="30" pageNumber="31">
SHELL (Figs 2628;
<tableCitation id="C69A5FE18B334E074F98FE573E3EFE1E" box="[429,563,427,453]" captionStart-0="Table 3" captionStart-1="Table 4" captionStartId-0="13.[189,258,268,294]" captionStartId-1="22.[188,257,551,579]" captionTargetBox-0="[201,1370,481,1908]" captionTargetBox-1="[205,1929,661,1732]" captionTargetPageId-0="13" captionTargetPageId-1="22" captionText-0="Table 3 (continued on next 3 pages). Shell measurements for species of Landouria Godwin-Austen, 1918 (in mm). Abbreviations: D = shell diameter; D3W = diameter of the third whorl; da = diameter of the aperture; H = shell height; ha = height of the aperture; max = maximum; min = minimum; n = number of measured spec.; std = standard deviation; U = diameter of the umbilicus; W = number of whorls." captionText-1="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri-0="http://table.plazi.org/id/DF673AD28B204E144E88FEF03D19FE68" httpUri-1="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="30" pageNumber="31" tableUuid-0="DF673AD28B204E144E88FEF03D19FE68" tableUuid-1="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Tables 34</tableCitation>
). Almost disc-like, with 5.256.25 hardly convex whorls; protoconch almost smooth; teleoconch with irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, slightly reflexed and slightly thickened; umbilicus almost concentric, very wide, comprising 3043% of shell diameter, hardly obscured by columellar edge.
</paragraph>
<paragraph id="8BA76A5A8B334E074E88FD313F9FFC65" blockId="30.[189,1399,716,959]" pageId="30" pageNumber="31">
GENITALIA (
<figureCitation id="132376DF8B334E074F76FD303DADFD3C" box="[323,416,716,743]" captionStart="Figs 1524" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 1524. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="30" pageNumber="31">Figs 17</figureCitation>
,
<figureCitation id="132376DF8B334E074F84FD313DDCFD3C" box="[433,465,717,743]" captionStart="Figs 2947" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 2947. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="30" pageNumber="31">31</figureCitation>
;
<tableCitation id="C69A5FE18B334E074FD5FD303E30FD3C" box="[480,573,716,743]" captionStart="Table 4" captionStartId="22.[188,257,551,579]" captionTargetBox="[205,1929,661,1732]" captionTargetPageId="22" captionText="Table 4 (continued on next 3 pages). Diagnostic characteristics of the species of Landouria Godwin-Austen, 1918 from Java. Characteristics most important for distinguishing similar taxa are printed in bold." httpUri="http://table.plazi.org/id/DF673AD28B3B4E0F4E89FDDB3F73FDBC" pageId="30" pageNumber="31" tableUuid="DF673AD28B3B4E0F4E89FDDB3F73FDBC">Table 4</tableCitation>
). Atrium short; penis very long, with a small bulge at proximal end containing a short penis papilla; without penis coecum; membranaceous sheath around distal part of penis connected with epiphallus by tissue; penial retractor runs from diaphragm to proximal part of cylindrical epiphallus; flagellum very long, broadest in its middle part, without node, tapering towards its proximal end; vas deferens narrow and long; vagina long; oviduct short, broad; peduncle of bursa copulatrix subdivided into a broader, abruptly tapering distal part and a narrower, longer proximal part. Right ommatophoral retractor runs between penis and vagina.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B334E074E88FC123EB5F930" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BA76A5A8B334E074E88FC123D22FBD3" blockId="30.[189,303,1006,1032]" box="[189,303,1006,1032]" pageId="30" pageNumber="31">
<heading id="D0EFDD368B334E074E88FC123D22FBD3" bold="true" box="[189,303,1006,1032]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B96CB6488B334E074E88FC123D22FBD3" bold="true" box="[189,303,1006,1032]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B334E074E88FBDD3E6AFB10" blockId="30.[189,1399,1057,1228]" pageId="30" pageNumber="31">
<taxonomicName id="4C1811D98B334E074E88FBDD3DAAFBE0" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[189,423,1057,1083]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074E88FBDD3DAAFBE0" box="[189,423,1057,1083]" italics="true" pageId="30" pageNumber="31">Landouria epiplatia</emphasis>
</taxonomicName>
was described from Jampang in
<collectingRegion id="49DCA4B88B334E074D16FBDE3F96FBE7" box="[803,923,1058,1084]" country="Indonesia" name="Jawa Barat" pageId="30" pageNumber="31">West Java</collectingRegion>
. However, there are several places called Jampang in
<collectingRegion id="49DCA4B88B334E074F67FBBA3DC3FBBB" box="[338,462,1094,1120]" country="Indonesia" name="Jawa Barat" pageId="30" pageNumber="31">West Java</collectingRegion>
, two among them being more important ones: Jampang Kulon and Jampang Tengah in Sukabumi Regency. Jampang Kulon is at ca
<quantity id="4CE0C7BF8B334E074D6FFB953FAAFB5F" box="[858,935,1129,1156]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="30" pageNumber="31" unit="m" value="300.0">300 m</quantity>
a.s.l., Jampang Tengah is at ca
<quantity id="4CE0C7BF8B334E074B1FFB95397AFB5F" box="[1322,1399,1129,1156]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="30" pageNumber="31" unit="m" value="500.0">500 m</quantity>
a.s.l. and closer to Sukabumi, the locality given by
<bibRefCitation id="EF8917AB8B334E074D2CFB72386BFB73" author="van Benthem Jutting" box="[793,1126,1165,1192]" firstAuthor="van Benthem Jutting" pageId="30" pageNumber="31" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting (1950)</bibRefCitation>
. Thus, we suppose that Jampang Tengah is the
<typeStatus id="54A3D4F88B334E074FFAFB4E3E0FFB17" box="[463,514,1202,1228]" pageId="30" pageNumber="31">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BA76A5A8B334E074E88FB073EE4F9EE" blockId="30.[189,1398,1275,1589]" pageId="30" pageNumber="31">
<taxonomicName id="4C1811D98B334E074E88FB073EE0FACE" authority="Mollendorff, 1897" authorityName="Mollendorff" authorityYear="1897" box="[189,749,1275,1301]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B334E074E88FB073E08FACE" box="[189,517,1275,1301]" italics="true" pageId="30" pageNumber="31">Plectotropis trichotrochium</emphasis>
<bibRefCitation id="EF8917AB8B334E074C25FB073EE0FACE" author="Mollendorff" box="[528,749,1275,1301]" firstAuthor="Mollendorff" pageId="30" pageNumber="31" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
</taxonomicName>
(Fig. 27) was synonymized with
<taxonomicName id="4C1811D98B334E074AB8FB003914FACE" ID-CoL="3S6S5" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[1165,1305,1276,1301]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="rotatoria">
<emphasis id="B96CB6488B334E074AB8FB003914FACE" box="[1165,1305,1276,1301]" italics="true" pageId="30" pageNumber="31">L. rotatoria</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8917AB8B334E074B79FB003DDAFAE1" author="van Benthem Jutting" firstAuthor="van Benthem Jutting" pageId="30" pageNumber="31" pagination="381 - 505" refId="ref35442" refString="van Benthem Jutting W. S. S. 1950. Systematic studies on the non-marine Mollusca of the Indo-Australian archipelago. II. Critical revision of the Javanese pulmonate land-shells of the families Helicarionidae, Pleurodontidae, Fruticicolidae and Streptaxidae. Treubia 20: 381 - 505." type="journal article" year="1950">van Benthem Jutting (1950)</bibRefCitation>
and
<bibRefCitation id="EF8917AB8B334E074C26FAE33EA1FAE1" author="Zilch" box="[531,684,1311,1338]" firstAuthor="Zilch" pageId="30" pageNumber="31" pagination="293 - 319" refId="ref35814" refString="Zilch A. 1966. Die Typen und Typoide des Natur-Museums Senckenberg, 35: Mollusca, Camaenidae (5). Archiv fur Molluskenkunde 95: 293 - 319." type="journal article" year="1966">Zilch (1966)</bibRefCitation>
. However, in contrast to
<taxonomicName id="4C1811D98B334E074DECFADC386EFAE2" ID-CoL="3S6S5" baseAuthorityName="Pfeiffer" baseAuthorityYear="1842" box="[985,1123,1312,1337]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="rotatoria">
<emphasis id="B96CB6488B334E074DECFADC386EFAE2" box="[985,1123,1312,1337]" italics="true" pageId="30" pageNumber="31">L. rotatoria</emphasis>
</taxonomicName>
, the spire of the single specimen on which
<taxonomicName id="4C1811D98B334E074F9AFAB83E70FA86" authorityName="Mollendorff" authorityYear="1897" box="[431,637,1347,1373]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B334E074F9AFAB83E70FA86" box="[431,637,1347,1373]" italics="true" pageId="30" pageNumber="31">P. trichotrochium</emphasis>
</taxonomicName>
was based is hardly elevated. We suppose that the
<typeStatus id="54A3D4F88B334E074AD8FABF3959FA86" box="[1261,1364,1347,1373]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1811D98B334E074E88FA943D86FA5A" authorityName="Mollendorff" authorityYear="1897" box="[189,395,1383,1409]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B334E074E88FA943D86FA5A" box="[189,395,1383,1409]" italics="true" pageId="30" pageNumber="31">P. trichotrochium</emphasis>
</taxonomicName>
is a subadult specimen of
<taxonomicName id="4C1811D98B334E074CF8FA943F5BFA5A" ID-CoL="7RTHD" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[717,854,1383,1409]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074CF8FA943F5BFA5A" box="[717,854,1383,1409]" italics="true" pageId="30" pageNumber="31">L. epiplatia</emphasis>
</taxonomicName>
that formed a slightly expanded peristome at a premature stage. The specimen is furthermore characterized by stronger radial wrinkles and more distinct incised spiral lines than is usual in
<taxonomicName id="4C1811D98B334E074C80FA4C3F31FA12" ID-CoL="7RTHD" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[693,828,1455,1481]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074C80FA4C3F31FA12" box="[693,828,1455,1481]" italics="true" pageId="30" pageNumber="31">L. epiplatia</emphasis>
</taxonomicName>
. Nevertheless, its occurrence at the type locality of
<taxonomicName id="4C1811D98B334E074ED5FA283D66FA36" ID-CoL="7RTHD" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[224,363,1491,1517]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074ED5FA283D66FA36" box="[224,363,1491,1517]" italics="true" pageId="30" pageNumber="31">L. epiplatia</emphasis>
</taxonomicName>
also indicates that it is an aberrant specimen of that species. As first revisers (ICZN 1999: Art. 24), we determine precedence of the name
<taxonomicName id="4C1811D98B334E074D64FA0B3935F9CA" authority="Mollendorff, 1897" authorityName="Mollendorff" authorityYear="1897" box="[849,1336,1527,1553]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074D64FA0B385FF9CA" box="[849,1106,1527,1553]" italics="true" pageId="30" pageNumber="31">Plectotropis epiplatia</emphasis>
<bibRefCitation id="EF8917AB8B334E074A69FA0B3935F9CA" author="Mollendorff" box="[1116,1336,1527,1553]" firstAuthor="Mollendorff" pageId="30" pageNumber="31" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
</taxonomicName>
over
<taxonomicName id="4C1811D98B334E074E88F9E73EE9F9EE" authority="Mollendorff, 1897" authorityName="Mollendorff" authorityYear="1897" box="[189,740,1563,1589]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="trichotrochium">
<emphasis id="B96CB6488B334E074E88F9E73E0FF9EE" box="[189,514,1563,1589]" italics="true" pageId="30" pageNumber="31">Plectotropis trichotrochium</emphasis>
<bibRefCitation id="EF8917AB8B334E074C3CF9E73EE9F9EE" author="Mollendorff" box="[521,740,1563,1589]" firstAuthor="Mollendorff" pageId="30" pageNumber="31" pagination="89 - 97" refId="ref33803" refString="Mollendorff O. F. von. 1897. Neue Landschnecken von Java. Nachrichtsblatt der deutschen malakozoologischen Gesellschaft 29: 57 - 72, 89 - 97." type="journal article" year="1897">Möllendorff, 1897</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA76A5A8B334E074E88F9993EB5F930" blockId="30.[189,1399,1637,1771]" pageId="30" pageNumber="31">
The description of the genitalia is based on a single subadult specimen from Gunung Gede (ZMH 79946;
<figureCitation id="132376DF8B334E074F23F9753D62F978" box="[278,367,1673,1699]" captionStart="Figs 1524" captionStartId="21.[189,241,1646,1672]" captionTargetBox="[206,1377,265,1616]" captionTargetId="figure@21.[204,1383,242,1625]" captionTargetPageId="21" captionText="Figs 1524. Genitalia of species of Landouria from Java. 15. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 16. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 17. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 18. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 19. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 20. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 21. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 22. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 23. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 24. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). Scale bars: 1 mm." httpUri="https://zenodo.org/record/3152243/files/figure.png" pageId="30" pageNumber="31">Figs 17</figureCitation>
,
<figureCitation id="132376DF8B334E074F49F9753D91F978" box="[380,412,1673,1699]" captionStart="Figs 2947" captionStartId="29.[189,241,1366,1392]" captionTargetBox="[189,1399,649,1337]" captionTargetId="figure@29.[189,1399,649,1337]" captionTargetPageId="29" captionText="Figs 2947. Flagella of species of Landouria from Java. 29. L. winteriana (Pfeiffer, 1842), West Java, Padalarang, Gunung Masigit (neotype, ZMA 376193a). 30. L. rotatoria (Pfeiffer, 1842), West Java, Gunung Salak, 1166 m a.s.l. (neotype, MZB 19205). 31. L. epiplatia (Möllendorff, 1897)?, West Java, Bogor, Gunung Gede (ZMH 79946). 32. L. nusakambangensis sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 412298a). 33. L. petrukensis sp. nov., Central Java, Karangbolong, way to Petruk Cave (holotype, MZB 19203). 34. L. tholiformis sp. nov., Yogyakarta, Gunung Kidul, Tepus, near Poktunggal Beach (holotype, MZB 17999). 35. L. abdidalem sp. nov., Yogyakarta, Kulonprogo, Sabrang Kidul (holotype, MZB 17664). 36. L. madurensis sp. nov., East Java, Madura Island, Sampang, Nepa Monkey lowland rainforest (paratype, MZB 19209). 37. L. sewuensis sp. nov., Yogyakarta, Gunung Kidul, Tepus, Mbajo Hill (holotype, MZB 19255). 38. L. sukoliloensis sp. nov., Central Java, Kayen, near Watu Payung (holotype, MZB 19219). 39. L. tonywhitteni sp. nov., Central Java, Taruman Village, near Sendang Sedayu (holotype, MZB 19214). 40. L. nodifera sp. nov., Central Java, Larangan Village, Lanang Spring (holotype, MZB 19227). 41. L. pacitanensis sp. nov., East Java, Pacitan, near Srau Beach (MZB 19231). 42. L. intumescens (Martens, 1867), East Java, Guwoterus (MZB 19207). 43. L. smimensis (Mousson, 1848), East Java, Tosari (ZMA 390881). 44. L. zonifera sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 17727). 45. L. ciliocincta (Möllendorff, 1897), East Java, Malang, Kondangmerak (MZB 19211). 46. L. pakidulan sp. nov., Central Java, Nusa Kambangan Island, nature monument Gligir (holotype, ZMA 417011a). 47. L. menorehensis sp. nov., Yogyakarta, Kulonprogo, Sibolong (holotype, MZB 19201). Scale bar: 1 mm." httpUri="https://zenodo.org/record/3152255/files/figure.png" pageId="30" pageNumber="31">31</figureCitation>
), which differs from typical
<taxonomicName id="4C1811D98B334E074CD8F9763F78F978" authorityName="Mollendorff" authorityYear="1897" box="[749,885,1673,1699]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074CD8F9763F78F978" box="[749,885,1673,1699]" italics="true" pageId="30" pageNumber="31">L. epiplatia</emphasis>
</taxonomicName>
in the slightly tighter coiled, more elevated spire. The available material is insufficient to decide whether this specimen falls within the variability of
<taxonomicName id="4C1811D98B334E074E88F92E3D49F930" authorityName="Mollendorff" authorityYear="1897" box="[189,324,1745,1771]" class="Gastropoda" family="Camaenidae" genus="Plectotropis" higherTaxonomySource="CoL" kingdom="Plantae" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074E88F92E3D49F930" box="[189,324,1745,1771]" italics="true" pageId="30" pageNumber="31">L. epiplatia</emphasis>
</taxonomicName>
or represents a distinct species.
</paragraph>
</subSubSection>
<subSubSection id="C30239D18B334E074E88F8E73E98F856" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BA76A5A8B334E074E88F8E73D5BF8EE" blockId="30.[189,342,1819,1845]" box="[189,342,1819,1845]" pageId="30" pageNumber="31">
<heading id="D0EFDD368B334E074E88F8E73D5BF8EE" bold="true" box="[189,342,1819,1845]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="B96CB6488B334E074E88F8E73D5BF8EE" bold="true" box="[189,342,1819,1845]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA76A5A8B334E074E88F8B23E98F856" blockId="30.[189,1398,1870,1933]" pageId="30" pageNumber="31">
<taxonomicName id="4C1811D98B334E074E88F8B23DA4F8B3" baseAuthorityName="Mollendorff" baseAuthorityYear="1897" box="[189,425,1870,1896]" class="Gastropoda" family="Camaenidae" genus="Landouria" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="epiplatia">
<emphasis id="B96CB6488B334E074E88F8B23DA4F8B3" box="[189,425,1870,1896]" italics="true" pageId="30" pageNumber="31">Landouria epiplatia</emphasis>
</taxonomicName>
is known from the northern part of Sukabumi Regency in
<collectingRegion id="49DCA4B88B334E074A54F8B238DAF8B3" box="[1121,1239,1870,1896]" country="Indonesia" name="Jawa Barat" pageId="30" pageNumber="31">West Java</collectingRegion>
and its range might extend to Gunung Gede (Fig. 55).
</paragraph>
</subSubSection>
</treatment>
</document>