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<document id="A2A02E40B0B1CFC69F0F9A1DC7425188" ID-CLB-Dataset="55668" ID-DOI="10.5252/geodiversitas2022v44a17" ID-GBIF-Dataset="fbd49218-183d-4aac-9368-0dd19bb340d2" ID-ISSN="1638-9395" ID-Zenodo-Dep="6518503" ID-ZooBank="urn:lsid:zoobank.org:pub:E4F744B3-E6DA-4ACC-836C-974ED25724D7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651669834440" checkinUser="felipe" docAuthor="Olive, Sébastien, Taverne, Louis &amp; Brito, Paulo M." docDate="2022" docId="03B187F7D45DD566FECFFC5AFE88FA1A" docLanguage="en" docName="Geodiversitas.44.17.505-514.pdf.imf" docOrigin="Geodiversitas 44 (17)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Pleuropholis germinalis Olive &amp; Taverne &amp; Brito 2022, n. sp." docType="treatment" docUuid="0E3ABDB0-1F72-4ADF-A1D4-68AFE8CF1B0C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="510" masterDocId="FF88FF8FD458D56EFFE9FFEEFF90FFB7" masterDocTitle="Pleuropholis germinalis n. sp., a new Pleuropholidae (Neopterygii, Teleostei) from the Early Cretaceous of Bernissart, Belgium" masterLastPageNumber="514" masterPageNumber="505" pageNumber="508" updateTime="1699329486868" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="C1D2533A79A15B802E488C9B59AC2E99">Pleuropholis germinalis n. sp., a new Pleuropholidae (Neopterygii, Teleostei) from the Early Cretaceous of Bernissart, Belgium</mods:title>
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<mods:namePart id="59E3A80889182750AE0E305F1AE14F3E">Olive, Sébastien</mods:namePart>
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<mods:namePart id="8A9A1DF2FED28AD1E3DC14E2DC357826">Taverne, Louis</mods:namePart>
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<mods:namePart id="44E30E284C455B60E87F0A8D57690441">Brito, Paulo M.</mods:namePart>
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<paragraph id="8BA736E1D45DD56BFECFFC5AFDF6FC78" blockId="5.[294,614,948,975]" box="[294,614,948,975]" pageId="5" pageNumber="508">
<heading id="D0EF818DD45DD56BFECFFC5AFDF6FC78" box="[294,614,948,975]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="508" reason="2">
<taxonomicName id="4C184D62D45DD56BFECFFC5AFDB4FC78" authority="Olive &amp; Taverne &amp; Brito, 2022" authorityName="Olive &amp; Taverne &amp; Brito" authorityYear="2022" box="[294,548,948,975]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="species" species="germinalis" status="sp. nov.">
<emphasis id="B96CEAF3D45DD56BFECFFC5AFDB4FC78" bold="true" box="[294,548,948,975]" italics="true" pageId="5" pageNumber="508">Pleuropholis germinalis</emphasis>
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<taxonomicNameLabel id="A25F5788D45DD56BFDC2FC5BFDF6FC78" box="[555,614,949,975]" pageId="5" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="8BA736E1D45DD56BFE78FC3BFE6AFC58" blockId="5.[401,506,980,1007]" box="[401,506,980,1007]" pageId="5" pageNumber="508">
(
<figureCitation id="13232A64D45DD56BFE73FC3AFE61FC58" box="[410,497,980,1007]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="6.[132,143,1519,1536]" captionStartId-1="7.[132,143,1476,1493]" captionStartId-2="8.[135,146,974,991]" captionStartId-3="9.[132,143,833,850]" captionTargetBox-0="[155,1432,215,1475]" captionTargetBox-1="[207,1381,214,1432]" captionTargetBox-2="[696,1380,658,927]" captionTargetBox-3="[134,1453,215,791]" captionTargetId-0="figure-267@6.[784,1416,754,1296]" captionTargetId-1="figure-270@7.[205,839,496,1037]" captionTargetId-2="graphics-579@8.[700,1381,665,919]" captionTargetId-3="figure-667@9.[698,1265,133,661]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." captionText-1="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." captionText-2="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." captionText-3="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6518509" figureDoi-1="http://doi.org/10.5281/zenodo.6518511" figureDoi-2="http://doi.org/10.5281/zenodo.6518513" figureDoi-3="http://doi.org/10.5281/zenodo.6518515" httpUri-0="https://zenodo.org/record/6518509/files/figure.png" httpUri-1="https://zenodo.org/record/6518511/files/figure.png" httpUri-2="https://zenodo.org/record/6518513/files/figure.png" httpUri-3="https://zenodo.org/record/6518515/files/figure.png" pageId="5" pageNumber="508">Figs 2-5</figureCitation>
)
</paragraph>
<paragraph id="8BA736E1D45DD56BFF6DFBF9FC97FB9F" blockId="5.[132,775,1047,1064]" box="[132,775,1047,1064]" pageId="5" pageNumber="508">
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urn:lsid:zoobank.org:act:
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<paragraph id="8BA736E1D45DD56BFF6DFBB6FD51FBC7" blockId="5.[132,705,1112,1136]" box="[132,705,1112,1136]" pageId="5" pageNumber="508">
<taxonomicName id="4C184D62D45DD56BFF6DFBB6FE87FBC7" box="[132,279,1112,1136]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B96CEAF3D45DD56BFF6DFBB6FF66FBC7" box="[132,246,1112,1136]" italics="true" pageId="5" pageNumber="508">Pleuropholis</emphasis>
sp.
</taxonomicName>
indet.
<bibRefCitation id="EF894B10D45DD56BFE86FBB6FDB4FBC7" author="TRAQUAIR R. H." box="[367,548,1112,1136]" journalOrPublisher="Memoires du Musee royal d'Histoire naturelle de Belgique" pageId="5" pageNumber="508" pagination="1 - 65" part="6" publicationUrl="https://www.biodiversitylibrary.org/page/42309609" refId="ref7555" refString="TRAQUAIR R. H. 1911. - Les poissons wealdiens de Bernissart. Memoires du Musee royal d'Histoire naturelle de Belgique 6: 1 - 65. https: // www. biodiversitylibrary. org / page / 42309609" title="Les poissons wealdiens de Bernissart" type="journal article" year="1911">Traquair 1911: 45</bibRefCitation>
, pl. IX, figs 1-3.
</paragraph>
</subSubSection>
<subSubSection id="C302656AD45DD56BFF6DFB66FEF2FA1F" pageId="5" pageNumber="508" type="diagnosis">
<paragraph id="8BA736E1D45DD56BFF6DFB66FEF2FA1F" blockId="5.[130,775,1160,1448]" pageId="5" pageNumber="508">
DIAGNOSIS. — Small sized actinopterygian (SL of roughly
<quantity id="4CE09B04D45DD56BFD51FB66FC90FB17" box="[696,768,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="508" unit="mm" value="64.0">64 mm</quantity>
) with the dorsal fin located posteriorly and at the same level as the anal fin. Species of
<taxonomicName id="4C184D62D45DD56BFEE2FB53FEEFFB62" authorityName="Egerton" authorityYear="1858" box="[267,383,1213,1237]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B96CEAF3D45DD56BFEE2FB53FEEFFB62" box="[267,383,1213,1237]" italics="true" pageId="5" pageNumber="508">Pleuropholis</emphasis>
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with the following unique combination of characters: robust preopercle with the horizontal limb as wide and high as the vertical limb; preopercle with the posterior edge entirely smooth; preopercle canal placed at equal distance from the dorsal and ventral borders of the horizontal branch (autapomorphy of
<taxonomicName id="4C184D62D45DD56BFF77FAACFE88FAEE" authority="Olive &amp; Taverne &amp; Brito, 2022" authorityName="Olive &amp; Taverne &amp; Brito" authorityYear="2022" box="[158,280,1345,1369]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="species" species="germinalis" status="sp. nov.">
<emphasis id="B96CEAF3D45DD56BFF77FAACFE88FAEE" box="[158,280,1345,1369]" italics="true" pageId="5" pageNumber="508">P. germinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F5788D45DD56BFEF5FAACFEC4FAEE" box="[284,340,1345,1369]" pageId="5" pageNumber="508" rank="species">n. sp.</taxonomicNameLabel>
); short and upturned maxilla; dentary with leptolepid notch in its median-dorsal part; quadrato-mandibular articulation lying anteriorly to the orbital margin; smooth posterior border of flank scales.
</paragraph>
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<paragraph id="8BA736E1D45DD56BFF6DFA2FFF75FA44" blockId="5.[132,775,1472,1523]" pageId="5" pageNumber="508">
<materialsCitation id="3B703CBCD45DD56BFF6DFA2FFF75FA44" ID-GBIF-Occurrence="3763236302" collectionCode="IRSNB" pageId="5" pageNumber="508" specimenCount="1" typeStatus="holotype">
<typeStatus id="54A38843D45DD56BFF6DFA2FFF60FA6F" box="[132,240,1473,1496]" pageId="5" pageNumber="508" type="holotype">HOLOTYPE</typeStatus>
. —
<collectionCode id="ED09AE24D45DD56BFEC9FA2EFEF9FA6F" box="[288,361,1472,1496]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="5" pageNumber="508">IRSNB</collectionCode>
P.1236 and 1235 (part and counterpart;
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,
<figureCitation id="13232A64D45DD56BFF22FA35FF48FA44" box="[203,216,1499,1523]" captionStart="FIG" captionStartId="7.[132,143,1476,1493]" captionTargetBox="[207,1381,214,1432]" captionTargetId="figure-270@7.[205,839,496,1037]" captionTargetPageId="7" captionText="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." figureDoi="http://doi.org/10.5281/zenodo.6518511" httpUri="https://zenodo.org/record/6518511/files/figure.png" pageId="5" pageNumber="508">3</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA736E1D45DD56BFF6DF9E5FE65F995" blockId="5.[132,501,1546,1570]" box="[132,501,1546,1570]" pageId="5" pageNumber="508">
<materialsCitation id="3B703CBCD45DD56BFF6DF9E5FE65F995" ID-GBIF-Occurrence="3763236301" box="[132,501,1546,1570]" pageId="5" pageNumber="508" typeStatus="paratype">
<typeStatus id="54A38843D45DD56BFF6DF9E5FF73F995" box="[132,227,1547,1570]" pageId="5" pageNumber="508" type="paratype">PARATYPE</typeStatus>
. —
<collectionCode id="ED09AE24D45DD56BFEE6F9E4FEC7F995" box="[271,343,1546,1570]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="5" pageNumber="508">IRSNB</collectionCode>
P.1234 (
<figureCitation id="13232A64D45DD56BFE46F9E4FE77F995" box="[431,487,1546,1570]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="5" pageNumber="508">Fig. 4</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA736E1D45DD56BFF6DF9D4FD50F9E5" blockId="5.[132,704,1594,1618]" box="[132,704,1594,1618]" pageId="5" pageNumber="508">
OTHER SPECIMENS. —
<materialsCitation id="3B703CBCD45DD56BFE8CF9D4FD50F9E5" ID-GBIF-Occurrence="3763236303" box="[357,704,1594,1618]" pageId="5" pageNumber="508">
IRSNB Vert-01680-00651 (
<figureCitation id="13232A64D45DD56BFD93F9D4FD22F9E5" box="[634,690,1594,1618]" captionStart="FIG" captionStartId="9.[132,143,833,850]" captionTargetBox="[134,1453,215,791]" captionTargetId="figure-667@9.[698,1265,133,661]" captionTargetPageId="9" captionText="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6518515" httpUri="https://zenodo.org/record/6518515/files/figure.png" pageId="5" pageNumber="508">Fig. 5</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302656AD45DD56BFF6DF984FE1FF95C" pageId="5" pageNumber="508" type="etymology">
<paragraph id="8BA736E1D45DD56BFF6DF984FE1FF95C" blockId="5.[132,777,1641,1771]" pageId="5" pageNumber="508">ETYMOLOGY. — The specific name refers to the novel of Emile Zola, Germinal. The novel focuses on the coalmine work and the harsh conditions of miners. Without miners, fossils from Bernissart would have never been extracted and this specific assignment consequently honors them.</paragraph>
</subSubSection>
<subSubSection id="C302656AD45DD56BFF6DF8EDFD66F8E7" pageId="5" pageNumber="508" type="materials_examined">
<paragraph id="8BA736E1D45DD56BFF6DF8EDFD66F8E7" blockId="5.[131,775,1795,1872]" pageId="5" pageNumber="508">
<collectionCode id="ED09AE24D45DD56BFF6DF8EDFE35F8AE" box="[132,421,1795,1818]" pageId="5" pageNumber="508">TYPE LOCALITY AND HORIZON</collectionCode>
. — Bernissart, Province of Hainaut,
<collectingCountry id="F30F7671D45DD56BFF6AF8F3FF4BF882" box="[131,219,1821,1845]" name="Belgium" pageId="5" pageNumber="508">Belgium</collectingCountry>
. Sainte-Barbe Clays Formation, uppermost Barremian to lowermost Aptian (
<bibRefCitation id="EF894B10D45DD56BFEB2F8D6FE66F8E7" author="YANS J. &amp; DEJAX J. &amp; PONS D. &amp; TAVERNE L. &amp; BULTYNCK P." box="[347,502,1848,1872]" journalOrPublisher="Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre" pageId="5" pageNumber="508" pagination="91 - 95" part="76" refId="ref7884" refString="YANS J., DEJAX J., PONS D., TAVERNE L. &amp; BULTYNCK P. 2006. - The iguanodons of Bernissart are middle Barremian to early Aptian in age. Bulletin de l'Institut royal des Sciences naturelles de Belgique, Sciences de la Terre 76: 91 - 95." title="The iguanodons of Bernissart are middle Barremian to early Aptian in age" type="journal article" year="2006">
Yans
<emphasis id="B96CEAF3D45DD56BFE67F8D6FE2DF8E7" box="[398,445,1848,1872]" italics="true" pageId="5" pageNumber="508">et al.</emphasis>
2006
</bibRefCitation>
,
<bibRefCitation id="EF894B10D45DD56BFDEBF8D6FDA5F8E7" author="SPAGNA P. &amp; YANS J. &amp; SCHNYDER J. &amp; DUPUIS C." box="[514,565,1848,1872]" editor="GODEFROIT P." journalOrPublisher="Indiana University Press, Indiana" pageId="5" pageNumber="508" pagination="87 - 96" publicationUrl="http://www.jstor.org/stable/j.ctt16gzfhx.14" refId="ref7249" refString="SPAGNA P., YANS J., SCHNYDER J. &amp; DUPUIS C. 2012. - The Paleoenvironment of the Bernissart Iguanodons: Sedimentological Analysis of the Lower Cretaceous Wealden Facies in the Bernissart Area, in GODEFROIT P. (ed.), Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Indiana University Press, Indiana: 87 - 96. http: // www. jstor. org / stable / j. ctt 16 gzfhx. 14" title="The Paleoenvironment of the Bernissart Iguanodons: Sedimentological Analysis of the Lower Cretaceous Wealden Facies in the Bernissart Area" type="book chapter" volumeTitle="Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems" year="2012">2012</bibRefCitation>
;
<bibRefCitation id="EF894B10D45DD56BFDA8F8D6FD78F8E7" author="Dejax" box="[577,744,1848,1872]" etAl="et al." firstAuthor="Dejax" pageId="5" pageNumber="508" pagination="25 - 38" refId="ref5964" refString="DEJAX J., PONS D. &amp; YANS J. 2007. - Palynology of the dinosaurbearing Wealden facies in the natural pit of Bernissart (Belgium). Review of Palaeobotany and Palynology 144: 25 - 38. https: // doi. org / 10.1016 / j. revpalbo. 2005.10.004" type="journal article" year="2007">
Dejax
<emphasis id="B96CEAF3D45DD56BFD68F8D6FD20F8E7" box="[641,688,1848,1872]" italics="true" pageId="5" pageNumber="508">et al.</emphasis>
2007
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302656AD45DD566FF6DF89CFE88FA1A" lastPageId="8" lastPageNumber="511" pageId="5" pageNumber="508" type="description">
<paragraph id="8BA736E1D45DD56BFF6DF89CFE85F83B" blockId="5.[131,775,1906,2028]" box="[132,277,1906,1932]" pageId="5" pageNumber="508">DESCRIPTION</paragraph>
<paragraph id="8BA736E1D45DD56BFF6DF87FFC97F85C" blockId="5.[131,775,1906,2028]" pageId="5" pageNumber="508">
With a standard length of approximately
<quantity id="4CE09B04D45DD56BFDAAF87FFD01F81B" box="[579,657,1937,1964]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="508" unit="mm" value="64.0">64 mm</quantity>
(specimen IRSNB P.1234,
<figureCitation id="13232A64D45DD56BFED6F85FFEEFF87C" box="[319,383,1969,1995]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="5" pageNumber="508">Fig. 4</figureCitation>
), this is a small sized actinopterygian, although one of the largest within
<taxonomicName id="4C184D62D45DD56BFDBCF83FFC93F85C" authorityName="Saint-Seine" authorityYear="1949" box="[597,771,2001,2027]" class="Actinopterygii" family="Pleuropholidae" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="family">Pleuropholidae</taxonomicName>
.
</paragraph>
<paragraph id="8BA736E1D45DD56BFCC6FF39FABEFE26" blockId="5.[811,1457,215,401]" pageId="5" pageNumber="508">
The highest part of the fish is located in the mid-length of the body (
<figureCitation id="13232A64D45DD56BFC48FF19FC71FEA6" box="[929,993,247,273]" captionStart="FIG" captionStartId="7.[132,143,1476,1493]" captionTargetBox="[207,1381,214,1432]" captionTargetId="figure-270@7.[205,839,496,1037]" captionTargetPageId="7" captionText="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." figureDoi="http://doi.org/10.5281/zenodo.6518511" httpUri="https://zenodo.org/record/6518511/files/figure.png" pageId="5" pageNumber="508">Fig. 3</figureCitation>
). The head, from the anterior part of the rostral to the posterior part of the opercle, is roughly one fourth of the standard length. The dorsal fin is situated well posteriorly, at the same level as the anal fin. The dermal bones and the scales are covered with ganoin.
</paragraph>
<paragraph id="8BA736E1D45DD56BFCC4FE58FCF4FE67" blockId="5.[813,868,438,464]" box="[813,868,438,464]" pageId="5" pageNumber="508">
<heading id="D0EF818DD45DD56BFCC4FE58FCF4FE67" box="[813,868,438,464]" fontSize="11" level="2" pageId="5" pageNumber="508" reason="8">
<emphasis id="B96CEAF3D45DD56BFCC4FE58FCF4FE67" box="[813,868,438,464]" italics="true" pageId="5" pageNumber="508">Skull</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736E1D45DD56BFCC4FE38FA81FD67" blockId="5.[811,1457,470,2028]" pageId="5" pageNumber="508">
The skull is fragmented in all specimens exhibiting it. The skull roof is preserved in right lateral view on specimen IRSNB P.1236 (
<figureCitation id="13232A64D45DD56BFC36FDF8FB8DFD87" box="[991,1053,534,560]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
) and is partially preserved in left lateral view, where it is somewhat distorted, in IRSNB P.1234 (
<figureCitation id="13232A64D45DD56BFCDFFDB8FCE3FDC7" box="[822,883,598,624]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="5" pageNumber="508">Fig. 4</figureCitation>
). Regarding IRSNB vert-01680-00651 (
<figureCitation id="13232A64D45DD56BFAC5FDB8FAF9FDC6" box="[1324,1385,598,625]" captionStart="FIG" captionStartId="9.[132,143,833,850]" captionTargetBox="[134,1453,215,791]" captionTargetId="figure-667@9.[698,1265,133,661]" captionTargetPageId="9" captionText="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6518515" httpUri="https://zenodo.org/record/6518515/files/figure.png" pageId="5" pageNumber="508">Fig. 5</figureCitation>
), only the posterior part of the skull, in lateral view, is preserved. The following description of the skull is therefore mainly based on specimen IRSNB P.1236 (
<figureCitation id="13232A64D45DD56BFB29FD5BFA90FD67" box="[1216,1280,693,720]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BA736E1D45DD56BFCAAFD3BFA21FB59" blockId="5.[811,1457,470,2028]" pageId="5" pageNumber="508">
The rostral is not well preserved in any of the specimens and its posterior suture with the nasals is not clearly discernible. The nasals are longer than wide, located anteriorly to the frontals. Frontals are the longest bones of the skull roof. Like in other species of teleosts (e.g.
<taxonomicName id="4C184D62D45DD56BFAC8FCBBFAB2FC38" authority=", Ichthyokentema, Siemensichthys Arratia, 2000" authorityName=", Ichthyokentema, Siemensichthys Arratia" authorityYear="2000" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B96CEAF3D45DD56BFAC8FCBBFA37FCD8" box="[1313,1447,853,879]" italics="true" pageId="5" pageNumber="508">Pleuropholis</emphasis>
,
<emphasis id="B96CEAF3D45DD56BFCC4FC9BFB12FC38" box="[813,1154,885,911]" italics="true" pageId="5" pageNumber="508">Ichthyokentema, Siemensichthys</emphasis>
<bibRefCitation id="EF894B10D45DD56BFB62FC9BFAB2FC38" author="Arratia" box="[1163,1314,885,911]" firstAuthor="Arratia" pageId="5" pageNumber="508" pagination="113 - 143" refId="ref5073" refString="ARRATIA G. 2000. - New teleostean fishes from the Jurassic of southern Germany and the systematic problems concerning the ' pholidophoriforms'. Palaontologische Zeitschrift 74 (1 - 2): 113 - 143. https: // doi. org / 10.1007 / BF 02987957" type="journal article" year="2000">Arratia, 2000</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C184D62D45DD56BFAD8FC98FC5DFC18" authority="Wagner, 1863" authorityName="Wagner" authorityYear="1863" class="Actinopterygii" family="Eurycormidae" genus="Eurycormus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pholidophoriformes" pageId="5" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B96CEAF3D45DD56BFAD8FC98FA3FFC38" box="[1329,1455,886,911]" italics="true" pageId="5" pageNumber="508">Eurycormus</emphasis>
Wagner, 1863
</taxonomicName>
, and
<taxonomicName id="4C184D62D45DD56BFBE3FC7BFCF7FC78" authority="Brito &amp; Gallo, 2002" authorityName="Brito &amp; Gallo" authorityYear="2002" class="Actinopterygii" family="Pleuropholidae" genus="Gondwanapleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B96CEAF3D45DD56BFBE3FC7BFA92FC18" box="[1034,1282,917,943]" italics="true" pageId="5" pageNumber="508">Gondwanapleuropholis</emphasis>
<bibRefCitation id="EF894B10D45DD56BFAE3FC7BFCF7FC78" author="Brito &amp; Gallo" firstAuthor="Brito" pageId="5" pageNumber="508" pagination="697 - 703" refId="ref5632" refString="BRITO P. M. &amp; GALLO V. 2002. - A new pleuropholid, Gondwanapleuropholis longimaxillaris n. g., n. sp. (Actinopterygii: Teleostei) from the Jurassic of north east Brazil. Comptes Rendus Palevol 1 (8): 697 - 703. https: // doi. org / 10.1016 / S 1631 - 0683 (02) 00080 - 5" type="journal article" year="2002">Brito &amp; Gallo, 2002</bibRefCitation>
</taxonomicName>
) the frontals are broad, and do not display the anterior narrowing found in more advanced teleosts (
<bibRefCitation id="EF894B10D45DD56BFAF3FC3AFA38FC58" author="Arratia" box="[1306,1448,980,1007]" firstAuthor="Arratia" pageId="5" pageNumber="508" pagination="113 - 143" refId="ref5073" refString="ARRATIA G. 2000. - New teleostean fishes from the Jurassic of southern Germany and the systematic problems concerning the ' pholidophoriforms'. Palaontologische Zeitschrift 74 (1 - 2): 113 - 143. https: // doi. org / 10.1007 / BF 02987957" type="journal article" year="2000">Arratia 2000</bibRefCitation>
;
<bibRefCitation id="EF894B10D45DD56BFCC4FC1AFB9BFBB8" author="Brito &amp; Gallo" box="[813,1035,1012,1039]" firstAuthor="Brito" pageId="5" pageNumber="508" pagination="697 - 703" refId="ref5632" refString="BRITO P. M. &amp; GALLO V. 2002. - A new pleuropholid, Gondwanapleuropholis longimaxillaris n. g., n. sp. (Actinopterygii: Teleostei) from the Jurassic of north east Brazil. Comptes Rendus Palevol 1 (8): 697 - 703. https: // doi. org / 10.1016 / S 1631 - 0683 (02) 00080 - 5" type="journal article" year="2002">Brito &amp; Gallo 2002</bibRefCitation>
). The parietals (
<figureCitation id="13232A64D45DD56BFB2BFC1AFA91FBB8" box="[1218,1281,1012,1039]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
) are subrectangular, slightly longer than wide and their length represents approximately two thirds of the length of the frontals. The extrascapulars are located posteriorly to the parietals and are barely distinguishable in the available material. Only the impression of the right dermopterotic is present in the specimen IRSNB P.1236 (
<figureCitation id="13232A64D45DD56BFB6BFB5AFB53FB79" box="[1154,1219,1204,1230]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
). The dermopterotic is a subrectangular bone, that widens in its posterior part.
</paragraph>
<paragraph id="8BA736E1D45DD56BFCAAFB1DFB93F99B" blockId="5.[811,1457,470,2028]" pageId="5" pageNumber="508">
The circumorbital series, as well as the posterior cheek region, are not well preserved in any of our specimens. The circumorbital series is formed by at least four infraorbitals, the dermosphenotic, and at least two supraorbitals. The infraorbital series is very fragmented in the specimen IRSNB P.1236 (
<figureCitation id="13232A64D45DD56BFC0AFA7DFBB2FA1A" box="[995,1058,1427,1453]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
). The dermosphenotic is in contact with the dorsal edge of the most posterior infraorbital; it is a somewhat rectangular bone that participates in the posterodorsal corner of the orbit, below the frontal and the dermopterotic.
</paragraph>
<paragraph id="8BA736E1D45DD56BFCAAF9DCFB87F8DB" blockId="5.[811,1457,470,2028]" pageId="5" pageNumber="508">
The preopercle is a “L-shaped” robust bone in which the horizontal limb is as wide and high as the vertical limb. These limbs are placed perpendicular to each other (
<figureCitation id="13232A64D45DD56BFCDFF97CFCEAF91B" box="[822,890,1682,1708]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
). The posterior edge of the preopercle is totally smooth (
<figureCitation id="13232A64D45DD56BFC7FF95CFC4CF97B" box="[918,988,1714,1740]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Figs 2</figureCitation>
;
<figureCitation id="13232A64D45DD56BFC07F95CFC6CF97A" box="[1006,1020,1714,1741]" captionStart="FIG" captionStartId="9.[132,143,833,850]" captionTargetBox="[134,1453,215,791]" captionTargetId="figure-667@9.[698,1265,133,661]" captionTargetPageId="9" captionText="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6518515" httpUri="https://zenodo.org/record/6518515/files/figure.png" pageId="5" pageNumber="508">5</figureCitation>
). The preopercular canal is placed at equal distance from the dorsal and ventral borders of the horizontal limb of the preopercle (
<figureCitation id="13232A64D45DD56BFB4CF91CFB75F8BB" box="[1189,1253,1778,1804]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Figs 2</figureCitation>
;
<figureCitation id="13232A64D45DD56BFB1AF91CFA91F8BA" box="[1267,1281,1778,1805]" captionStart="FIG" captionStartId="9.[132,143,833,850]" captionTargetBox="[134,1453,215,791]" captionTargetId="figure-667@9.[698,1265,133,661]" captionTargetPageId="9" captionText="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6518515" httpUri="https://zenodo.org/record/6518515/files/figure.png" pageId="5" pageNumber="508">5</figureCitation>
). The opercular series is formed by a slightly deeper than wide opercle, a subtriangular subopercle, and an interopercle (incomplete on IRSNB P.01236).
</paragraph>
<paragraph id="8BA736E1D45DD568FCAAF89FFDB2F93B" blockId="5.[811,1457,470,2028]" lastBlockId="6.[130,776,1618,2028]" lastPageId="6" lastPageNumber="509" pageId="5" pageNumber="508">
The maxilla is upturned, rather narrow and short, but badly preserved (
<figureCitation id="13232A64D45DD56BFC05F87FFBBBF81C" box="[1004,1067,1937,1963]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="5" pageNumber="508">Fig. 2</figureCitation>
). The oral margin of the maxilla is not preserved here. The single supramaxilla is a moderately large bone placed posteriorly to the maxilla and along the anterior margin of the orbit (
<figureCitation id="13232A64D45ED568FE0DF9BCFDB8F9DB" box="[484,552,1618,1644]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="6" pageNumber="509">Fig. 2</figureCitation>
). No premaxilla is preserved in the available specimens.
</paragraph>
<caption id="DF676669D45ED568FF6DFA01FDFBF9A1" ID-DOI="http://doi.org/10.5281/zenodo.6518509" ID-Zenodo-Dep="6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="6" pageNumber="509" startId="6.[132,143,1519,1536]" targetBox="[155,1432,215,1475]" targetPageId="6">
<paragraph id="8BA736E1D45ED568FF6DFA01FDFBF9A1" blockId="6.[132,1455,1519,1558]" pageId="6" pageNumber="509">
FIG. 2. — Skeleton of
<taxonomicName id="4C184D62D45ED568FEDDFA01FE63F9B7" authority="Olive &amp; Taverne &amp; Brito, 2022" authorityName="Olive &amp; Taverne &amp; Brito" authorityYear="2022" box="[308,499,1519,1536]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="509" phylum="Chordata" rank="species" species="germinalis" status="sp. nov.">
<emphasis id="B96CEAF3D45ED568FEDDFA01FE63F9B7" box="[308,499,1519,1536]" italics="true" pageId="6" pageNumber="509">Pleuropholis germinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F5788D45ED568FE1EFA01FDB8F9B7" box="[503,552,1519,1536]" pageId="6" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
, IRSNB P.01236, holotype:
<emphasis id="B96CEAF3D45ED568FCE5FA01FC89F9B7" bold="true" box="[780,793,1519,1536]" pageId="6" pageNumber="509">A</emphasis>
, photograph;
<emphasis id="B96CEAF3D45ED568FC64FA01FC0AF9B7" bold="true" box="[909,922,1519,1536]" pageId="6" pageNumber="509">B</emphasis>
, close-up on the head whitened with ammonium chloride;
<emphasis id="B96CEAF3D45ED568FA92FA01FA19F9B7" bold="true" box="[1403,1417,1519,1536]" pageId="6" pageNumber="509">C</emphasis>
, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm.
</paragraph>
</caption>
<paragraph id="8BA736E1D45ED568FF72F97DFC98F85C" blockId="6.[130,776,1618,2028]" pageId="6" pageNumber="509">
Dentary and angular are exposed laterally. The dentary is a short stout subsquare bone that increases in height posteriorly. The dentary presents, in its median-dorsal part, a leptolepid notch, like in
<taxonomicName id="4C184D62D45ED568FEF2F91DFD84F8BB" authority="Bravi, 1988" authorityName="Bravi" authorityYear="1988" box="[283,532,1778,1804]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="509" phylum="Chordata" rank="species" species="decastroi">
<emphasis id="B96CEAF3D45ED568FEF2F91DFE1CF8BB" box="[283,396,1778,1804]" italics="true" pageId="6" pageNumber="509">P. decastroi</emphasis>
<bibRefCitation id="EF894B10D45ED568FE7AF91CFD84F8BB" author="Bravi" box="[403,532,1778,1804]" firstAuthor="Bravi" pageId="6" pageNumber="509" pagination="575 - 586" refId="ref5461" refString="BRAVI S. 1988. - Contributo allo studio del giacimento ad ittioliti di Pietraroja (Benevento). I: Pleuropholis decastroi n. sp. (Pisces, Actinopterygii, Pholidophoriformes). Memorie della Societa Geologica Italiana 41: 575 - 586." type="journal article" year="1988">Bravi, 1988</bibRefCitation>
</taxonomicName>
, from the Early Cretaceous of
<collectingCountry id="F30F7671D45ED568FF08F8FCFE81F89B" box="[225,273,1810,1836]" name="Italy" pageId="6" pageNumber="509">Italy</collectingCountry>
(
<bibRefCitation id="EF894B10D45ED568FEC9F8FCFDBEF89B" author="TAVERNE L. &amp; CAPASSO L." box="[288,558,1810,1836]" journalOrPublisher="Geo-Eco-Trop" pageId="6" pageNumber="509" pagination="35 - 51" part="43" refId="ref7510" refString="TAVERNE L. &amp; CAPASSO L. 2019. - New data on Pleuropholis decastroi (Teleostei, Pleuropholidae), a &quot; pholidophoriform &quot; fish from the Lower Cretaceous of the Eurafrican Mesogea. Geo-Eco-Trop 43 (1): 35 - 51." title="New data on Pleuropholis decastroi (Teleostei, Pleuropholidae), a &quot; pholidophoriform &quot; fish from the Lower Cretaceous of the Eurafrican Mesogea" type="journal article" year="2019">Taverne &amp; Capasso 2019</bibRefCitation>
). The dentary is traversed by the mandibular sensory canal, which extends along its ventral border. Posteriorly, the dentary sutures with the angular that is partially exposed in IRSNB P.1236 and forms the posterior corner of the lower jaw (
<figureCitation id="13232A64D45ED568FDCEF87FFDF6F81C" box="[551,614,1937,1963]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="6" pageNumber="509">Fig. 2</figureCitation>
). Other bones of the lower jaw are not exposed in the specimens studied here. No teeth have been observed in the available material.
</paragraph>
<paragraph id="8BA736E1D45ED568FCC4F9BCFC4DF9DB" blockId="6.[813,989,1618,1644]" box="[813,989,1618,1644]" pageId="6" pageNumber="509">
<heading id="D0EF818DD45ED568FCC4F9BCFC4DF9DB" box="[813,989,1618,1644]" fontSize="11" level="2" pageId="6" pageNumber="509" reason="8">
<emphasis id="B96CEAF3D45ED568FCC4F9BCFC4DF9DB" box="[813,989,1618,1644]" italics="true" pageId="6" pageNumber="509">Vertebral column</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736E1D45ED568FCC4F99CFA3FF91A" blockId="6.[812,1455,1650,1709]" pageId="6" pageNumber="509">
Only few vertebral elements of the abdominal region are exposed where some scales are displaced or lost (
<figureCitation id="13232A64D45ED568FAB5F97CFA0DF91B" box="[1372,1437,1682,1708]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="6" pageNumber="509">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BA736E1D45ED568FCC4F93CFBB2F95B" blockId="6.[813,1058,1746,1772]" box="[813,1058,1746,1772]" pageId="6" pageNumber="509">
<heading id="D0EF818DD45ED568FCC4F93CFBB2F95B" box="[813,1058,1746,1772]" fontSize="11" level="2" pageId="6" pageNumber="509" reason="8">
<emphasis id="B96CEAF3D45ED568FCC4F93CFBB2F95B" box="[813,1058,1746,1772]" italics="true" pageId="6" pageNumber="509">Paired girdles and fins</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736E1D45ED569FCC4F91CFD14F8BB" blockId="6.[810,1456,1778,2028]" lastBlockId="7.[131,774,1618,1804]" lastPageId="7" lastPageNumber="510" pageId="6" pageNumber="509">
Some elements of the pectoral girdle are preserved as fragmented elements, impressions in the matrix, or are hidden by the presence of displaced scales (
<figureCitation id="13232A64D45ED568FAF8F8DFFAFBF8FB" box="[1297,1387,1841,1868]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="6.[132,143,1519,1536]" captionStartId-1="7.[132,143,1476,1493]" captionStartId-2="8.[135,146,974,991]" captionTargetBox-0="[155,1432,215,1475]" captionTargetBox-1="[207,1381,214,1432]" captionTargetBox-2="[696,1380,658,927]" captionTargetId-0="figure-267@6.[784,1416,754,1296]" captionTargetId-1="figure-270@7.[205,839,496,1037]" captionTargetId-2="graphics-579@8.[700,1381,665,919]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." captionText-1="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." captionText-2="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi-0="http://doi.org/10.5281/zenodo.6518509" figureDoi-1="http://doi.org/10.5281/zenodo.6518511" figureDoi-2="http://doi.org/10.5281/zenodo.6518513" httpUri-0="https://zenodo.org/record/6518509/files/figure.png" httpUri-1="https://zenodo.org/record/6518511/files/figure.png" httpUri-2="https://zenodo.org/record/6518513/files/figure.png" pageId="6" pageNumber="509">Figs 2-4</figureCitation>
). The cleithrum is a robust bone located ventrally to the short and small supracleithrum. It is smooth, and anteriorly underlies the subopercle. Fragmented and displaced elements of the putative ventral branch of the cleithrum are found in IRSNB P.1236 (
<figureCitation id="13232A64D45ED568FB67F83FFB42F85C" box="[1166,1234,2001,2027]" captionStart="FIG" captionStartId="6.[132,143,1519,1536]" captionTargetBox="[155,1432,215,1475]" captionTargetId="figure-267@6.[784,1416,754,1296]" captionTargetPageId="6" captionText="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." figureDoi="http://doi.org/10.5281/zenodo.6518509" httpUri="https://zenodo.org/record/6518509/files/figure.png" pageId="6" pageNumber="509">Fig. 2</figureCitation>
). The pectoral fin is positioned close to the ventral margin of the body and below the large scales of the body flank in IRSNB P.1234 (
<figureCitation id="13232A64D45FD569FF09F97CFE8FF91B" box="[224,287,1682,1708]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="7" pageNumber="510">Fig. 4</figureCitation>
). At least nine rays are preserved. In IRSNB P.1235, a pelvic fin is preserved, in which roughly seven rays are preserved (
<figureCitation id="13232A64D45FD569FE89F93CFE31F95B" box="[352,417,1746,1772]" captionStart="FIG" captionStartId="7.[132,143,1476,1493]" captionTargetBox="[207,1381,214,1432]" captionTargetId="figure-270@7.[205,839,496,1037]" captionTargetPageId="7" captionText="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." figureDoi="http://doi.org/10.5281/zenodo.6518511" httpUri="https://zenodo.org/record/6518511/files/figure.png" pageId="7" pageNumber="510">Fig. 3</figureCitation>
). No bones of the pelvic girdle are exposed in any of the studied specimens.
</paragraph>
<caption id="DF676669D45FD569FF6DFA2AFCFFFA5C" ID-DOI="http://doi.org/10.5281/zenodo.6518511" ID-Zenodo-Dep="6518511" httpUri="https://zenodo.org/record/6518511/files/figure.png" pageId="7" pageNumber="510" startId="7.[132,143,1476,1493]" targetBox="[207,1381,214,1432]" targetPageId="7">
<paragraph id="8BA736E1D45FD569FF6DFA2AFCFFFA5C" blockId="7.[132,1455,1476,1515]" pageId="7" pageNumber="510">
FIG. 3. — Skeleton of
<taxonomicName id="4C184D62D45FD569FED3FA2AFE6AFA62" authority="Olive &amp; Taverne &amp; Brito, 2022" authorityName="Olive &amp; Taverne &amp; Brito" authorityYear="2022" box="[314,506,1476,1493]" class="Actinopterygii" family="Pleuropholidae" genus="Pleuropholis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="7" pageNumber="510" phylum="Chordata" rank="species" species="germinalis" status="sp. nov.">
<emphasis id="B96CEAF3D45FD569FED3FA2AFE6AFA62" box="[314,506,1476,1493]" italics="true" pageId="7" pageNumber="510">Pleuropholis germinalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F5788D45FD569FDE8FA2AFDA4FA62" box="[513,564,1476,1493]" pageId="7" pageNumber="510" rank="species">n. sp.</taxonomicNameLabel>
, IRSNB P.01235, holotype, counterpart of IRSNB P.01236:
<emphasis id="B96CEAF3D45FD569FBCCFA2AFBA2FA62" bold="true" box="[1061,1074,1476,1493]" pageId="7" pageNumber="510">A</emphasis>
, photograph;
<emphasis id="B96CEAF3D45FD569FB40FA2AFB26FA62" bold="true" box="[1193,1206,1476,1493]" pageId="7" pageNumber="510">B</emphasis>
, line drawing close-up on the dorsal fin;
<emphasis id="B96CEAF3D45FD569FF32FA34FF79FA5C" bold="true" box="[219,233,1498,1515]" pageId="7" pageNumber="510">C</emphasis>
, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm.
</paragraph>
</caption>
<paragraph id="8BA736E1D45FD569FF6DF8DFFEC6F8FB" blockId="7.[132,342,1841,1868]" box="[132,342,1841,1868]" pageId="7" pageNumber="510">
<heading id="D0EF818DD45FD569FF6DF8DFFEC6F8FB" box="[132,342,1841,1868]" fontSize="11" level="2" pageId="7" pageNumber="510" reason="8">
<emphasis id="B96CEAF3D45FD569FF6DF8DFFEC6F8FB" box="[132,342,1841,1868]" italics="true" pageId="7" pageNumber="510">Dorsal and anal fins</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736E1D45FD569FF6DF8BFFDBFF85C" blockId="7.[130,775,1873,2028]" pageId="7" pageNumber="510">
The dorsal fin (
<figureCitation id="13232A64D45FD569FEC3F8BFFEFAF8DB" box="[298,362,1873,1900]" captionStart="FIG" captionStartId="7.[132,143,1476,1493]" captionTargetBox="[207,1381,214,1432]" captionTargetId="figure-270@7.[205,839,496,1037]" captionTargetPageId="7" captionText="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." figureDoi="http://doi.org/10.5281/zenodo.6518511" httpUri="https://zenodo.org/record/6518511/files/figure.png" pageId="7" pageNumber="510">Figs 3</figureCitation>
;
<figureCitation id="13232A64D45FD569FE90F8BCFE17F8DB" box="[377,391,1874,1900]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="7" pageNumber="510">4</figureCitation>
) counts at least 9 dorsal fin rays on IRSNB P.1235 (
<figureCitation id="13232A64D45FD569FEDDF89FFEE2F83C" box="[308,370,1905,1931]" captionStart="FIG" captionStartId="7.[132,143,1476,1493]" captionTargetBox="[207,1381,214,1432]" captionTargetId="figure-270@7.[205,839,496,1037]" captionTargetPageId="7" captionText="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." figureDoi="http://doi.org/10.5281/zenodo.6518511" httpUri="https://zenodo.org/record/6518511/files/figure.png" pageId="7" pageNumber="510">Fig. 3</figureCitation>
) whereas the anal fin (
<figureCitation id="13232A64D45FD569FD8CF89FFD33F83C" box="[613,675,1905,1931]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="7" pageNumber="510">Fig. 4</figureCitation>
) displays at least 11 fin rays. The anterior margin of the anal fin displays at least 7 fringing fulcra. Elongated pterygiophores, supporting the anal fin, are observed anteriorly.
</paragraph>
<paragraph id="8BA736E1D45FD569FCC4F9BCFC0CF9DB" blockId="7.[813,924,1618,1644]" box="[813,924,1618,1644]" pageId="7" pageNumber="510">
<heading id="D0EF818DD45FD569FCC4F9BCFC0CF9DB" box="[813,924,1618,1644]" fontSize="11" level="2" pageId="7" pageNumber="510" reason="8">
<emphasis id="B96CEAF3D45FD569FCC4F9BCFC0CF9DB" box="[813,924,1618,1644]" italics="true" pageId="7" pageNumber="510">Caudal fin</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736E1D45FD569FCC4F99CFA20F8DC" blockId="7.[810,1456,1650,1900]" pageId="7" pageNumber="510">
The caudal pedicle is slender and covered with rhombic caudal scales(
<figureCitation id="13232A64D45FD569FC86F97CFC38F91B" box="[879,936,1682,1708]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="7" pageNumber="510">Figs 4</figureCitation>
;
<figureCitation id="13232A64D45FD569FC5AF97CFC50F91A" box="[947,960,1682,1709]" captionStart="FIG" captionStartId="9.[132,143,833,850]" captionTargetBox="[134,1453,215,791]" captionTargetId="figure-667@9.[698,1265,133,661]" captionTargetPageId="9" captionText="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6518515" httpUri="https://zenodo.org/record/6518515/files/figure.png" pageId="7" pageNumber="510">5</figureCitation>
).The caudal endoskeleton is covered by scales and not visible for analysis. Although the posterior tip of the caudal fin is not preserved, the caudal fin is almost complete in IRSNB P.1234 (
<figureCitation id="13232A64D45FD569FC68F91CFC2AF8BB" box="[897,954,1778,1804]" captionStart="FIG" captionStartId="8.[135,146,974,991]" captionTargetBox="[696,1380,658,927]" captionTargetId="graphics-579@8.[700,1381,665,919]" captionTargetPageId="8" captionText="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." figureDoi="http://doi.org/10.5281/zenodo.6518513" httpUri="https://zenodo.org/record/6518513/files/figure.png" pageId="7" pageNumber="510">Fig. 4</figureCitation>
). The caudal fin is hemiheterocercal and includes at least 19 evenly segmented rays. On the dorsal margin of the caudal fin, there are at least 4 dorsal caudal fulcra. The ventral margin of the caudal fin is preserved in none of the specimens.
</paragraph>
<paragraph id="8BA736E1D45FD569FCC4F87FFCF8F81C" blockId="7.[813,872,1937,1963]" box="[813,872,1937,1963]" pageId="7" pageNumber="510">
<heading id="D0EF818DD45FD569FCC4F87FFCF8F81C" box="[813,872,1937,1963]" fontSize="11" level="2" pageId="7" pageNumber="510" reason="8">
<emphasis id="B96CEAF3D45FD569FCC4F87FFCF8F81C" box="[813,872,1937,1963]" italics="true" pageId="7" pageNumber="510">Scales</emphasis>
</heading>
</paragraph>
<paragraph id="8BA736E1D45FD566FCC4F85FFE88FA1A" blockId="7.[813,1456,1969,2028]" lastBlockId="8.[131,777,1044,1453]" lastPageId="8" lastPageNumber="511" pageId="7" pageNumber="510">
The scales are thick and of ganoid
<typeStatus id="54A38843D45FD569FB5EF85CFB77F87B" box="[1207,1255,1970,1996]" pageId="7" pageNumber="510">type</typeStatus>
(
<figureCitation id="13232A64D45FD569FB10F85FFAC4F87B" box="[1273,1364,1969,1996]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStart-3="FIG" captionStartId-0="6.[132,143,1519,1536]" captionStartId-1="7.[132,143,1476,1493]" captionStartId-2="8.[135,146,974,991]" captionStartId-3="9.[132,143,833,850]" captionTargetBox-0="[155,1432,215,1475]" captionTargetBox-1="[207,1381,214,1432]" captionTargetBox-2="[696,1380,658,927]" captionTargetBox-3="[134,1453,215,791]" captionTargetId-0="figure-267@6.[784,1416,754,1296]" captionTargetId-1="figure-270@7.[205,839,496,1037]" captionTargetId-2="graphics-579@8.[700,1381,665,919]" captionTargetId-3="figure-667@9.[698,1265,133,661]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="FIG. 2. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01236, holotype: A, photograph; B, close-up on the head whitened with ammonium chloride;C, line drawing reconstruction. Scale bars: A, 1 cm; B, C, 0.5 cm." captionText-1="FIG. 3. — Skeleton of Pleuropholis germinalis n. sp., IRSNB P.01235, holotype, counterpart of IRSNB P.01236: A, photograph; B, line drawing close-up on the dorsal fin; C, line drawing close-up on the pelvic fin. Scale bars: A, 0.5 cm; B, C, 0.25 cm." captionText-2="FIG. 4. — Skeleton of Pleuropholis germinalis n.sp., IRSNB P.01234,paratype:A, photograph;B, line drawing of the vertebral skeleton and of fins.Scale bars:1 cm." captionText-3="FIG. 5. — Photograph of the skeleton of Pleuropholis germinalis n. sp. with a close-up on the preopercle, IRSNB vert.01680-0065. Scale bars: A, 1 cm; B, 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6518509" figureDoi-1="http://doi.org/10.5281/zenodo.6518511" figureDoi-2="http://doi.org/10.5281/zenodo.6518513" figureDoi-3="http://doi.org/10.5281/zenodo.6518515" httpUri-0="https://zenodo.org/record/6518509/files/figure.png" httpUri-1="https://zenodo.org/record/6518511/files/figure.png" httpUri-2="https://zenodo.org/record/6518513/files/figure.png" httpUri-3="https://zenodo.org/record/6518515/files/figure.png" pageId="7" pageNumber="510">Figs 2-5</figureCitation>
). Those along the flank are rectangular in shape, 5-6 times deeper B Cl than long, with anterior and posterior borders being slightly curved. The entire posterior border of all flank scales is smooth. Dorsally to the flank scale row, two rows of rhomboidal scales are observed. The most dorsal scales, anterior to the dorsal fin, could potentially be pair scute-like elements. However, the preservation of this feature is not good enough to be conclusive. Ventrally to the flank scale row, at least two scale rows (probably three) are present. Scales are relatively small and rhomboidal. The lateral line is not observable through the body flank scales; pores or thickenings revealing the course of branch of the lateral line system are not observable probably due to a lack of preservation.
</paragraph>
</subSubSection>
</treatment>
</document>