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<document id="52ECD8B21F794951C52036CF23748748" ID-DOI="10.12651/JSR.2024.13.3.340" ID-ISSN="2713-8615" ID-Zenodo-Dep="14659397" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1736971599052" checkinUser="felipe" docAuthor="Chullasorn, Supawadee, Kangtia, Pawana, Klangsin, Pradchek &amp; Song, Sung Joon" docDate="2024" docId="03B187F2FFBAFFD06BDDC1D7FE1EFB4D" docLanguage="en" docName="JSpeciesRes.13.3.340-351.pdf" docOrigin="Journal of Species Research 13 (3)" docSource="http://zoobank.org/0b0dffeb-709a-403d-800e-a925b06265c9" docStyle="DocumentStyle:4D398BFF45FF8CA9E08F546BB06AE372.2:JSpeciesRes.2012-.journal_article" docStyleId="4D398BFF45FF8CA9E08F546BB06AE372" docStyleName="JSpeciesRes.2012-.journal_article" docStyleVersion="2" docTitle="Brianola rawaiensis Chullasorn, Kangtia, Klangsin &amp; Song, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="350" masterDocId="FF88FF8AFFB8FFDA6B4DC011FFD7FFAC" masterDocTitle="A new species of Brianola Monard, 1926 (Copepoda: Harpacticoida: Canuellidae) from Rawai Beach, Phuket Island, Thailand" masterLastPageNumber="351" masterPageNumber="340" pageNumber="342" updateTime="1736973211098" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="840E6F60588469B1CED2EE259F1C5929">A new species of Brianola Monard, 1926 (Copepoda: Harpacticoida: Canuellidae) from Rawai Beach, Phuket Island, Thailand</mods:title>
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<mods:namePart id="8C073DF0166E2802FA72B37107B4983F">Chullasorn, Supawadee</mods:namePart>
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<mods:namePart id="26046C99C79F705E0495875FF12D0911">Kangtia, Pawana</mods:namePart>
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<mods:namePart id="7E0A3CFF3C9D7C5254DB1FDF772CA822">Klangsin, Pradchek</mods:namePart>
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<mods:namePart id="5D8C86A8BBA471DD20980DA377E22CD3">Song, Sung Joon</mods:namePart>
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<mods:date id="9D9433DAFF872BDBAB3C9B881AB0A431">2024</mods:date>
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<treatment id="03B187F2FFBAFFD06BDDC1D7FE1EFB4D" LSID="urn:lsid:plazi:treatment:03B187F2FFBAFFD06BDDC1D7FE1EFB4D" httpUri="http://treatment.plazi.org/id/03B187F2FFBAFFD06BDDC1D7FE1EFB4D" lastPageId="10" lastPageNumber="350" pageId="2" pageNumber="342">
<subSubSection id="C302656FFFBAFFD86BDDC1D7FD92FE72" box="[144,581,454,479]" pageId="2" pageNumber="342" type="nomenclature">
<paragraph id="8BA736E4FFBAFFD86BDDC1D7FD92FE72" blockId="2.[144,581,454,479]" box="[144,581,454,479]" pageId="2" pageNumber="342">
<heading id="D0EF8188FFBAFFD86BDDC1D7FD92FE72" bold="true" box="[144,581,454,479]" fontSize="10" level="2" pageId="2" pageNumber="342" reason="4">
<emphasis id="B96CEAF6FFBAFFD86BDDC1D7FD92FE72" bold="true" box="[144,581,454,479]" pageId="2" pageNumber="342">
<taxonomicName id="4C184D67FFBAFFD86BDDC1D7FEBAFE73" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[144,365,454,479]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="2" pageNumber="342" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBAFFD86BDDC1D7FEBAFE73" bold="true" box="[144,365,454,479]" italics="true" pageId="2" pageNumber="342">Brianola rawaiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F578DFFBAFFD86A39C1D6FE1EFE73" box="[372,457,455,479]" pageId="2" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13232A61FFBAFFD86A98C1D7FDEAFE72" box="[469,573,454,479]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStartId-0="1.[424,457,1938,1960]" captionStartId-1="2.[268,301,1975,1997]" captionStartId-2="3.[468,501,1689,1711]" captionStartId-3="4.[254,287,1722,1744]" captionStartId-4="5.[447,480,1529,1551]" captionStartId-5="6.[412,445,1530,1552]" captionStartId-6="7.[435,468,1463,1485]" captionStartId-7="8.[276,309,1374,1396]" captionStartId-8="9.[475,508,1635,1657]" captionTargetBox-0="[357,1260,716,1906]" captionTargetBox-1="[341,1176,817,1938]" captionTargetBox-2="[244,1374,257,1653]" captionTargetBox-3="[232,1320,264,1680]" captionTargetBox-4="[289,1320,258,1494]" captionTargetBox-5="[300,1294,258,1498]" captionTargetBox-6="[385,1251,261,1430]" captionTargetBox-7="[366,1159,257,1339]" captionTargetBox-8="[381,1303,262,1602]" captionTargetId-0="figure-229@1.[355,1262,713,1906]" captionTargetId-1="figure-211@2.[341,1216,812,1941]" captionTargetId-2="figure-118@3.[243,1375,257,1655]" captionTargetId-3="figure-114@4.[231,1326,257,1688]" captionTargetId-4="figure-207@5.[296,1322,257,1497]" captionTargetId-5="figure-198@6.[260,1298,257,1499]" captionTargetId-6="figure-254@7.[365,1252,257,1431]" captionTargetId-7="figure-253@8.[346,1212,257,1342]" captionTargetId-8="figure-14@9.[376,1306,257,1602]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="7" captionTargetPageId-7="8" captionTargetPageId-8="9" captionText-0="Fig. 1. Brianola rawaiensis sp. nov., female: (A) habitus, dorsal; (B) habitus, lateral." captionText-1="Fig. 2. Brianola rawaiensis sp. nov., female: (A) urosome dorsal; (B); urosome ventral; (C) genital area; (D) P5." captionText-2="Fig. 3. Brianola rawaiensis sp. nov., female: (A) antennule(disarticulated)." captionText-3="Fig. 4. Brianola rawaiensis sp. nov., female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped." captionText-4="Fig. 5. Brianola rawaiensis sp. nov., female: (A) P1, anterior; (B) P2, posterior." captionText-5="Fig. 6. Brianola rawaiensis sp. nov., female: (A) P3, posterior; (B) P4, posterior." captionText-6="Fig. 7.Brianola rawaiensis sp. nov., male: (A) habitus, dorsal; (B) habitus, lateral." captionText-7="Fig. 8. Brianola rawaiensis sp. nov., male: (A) urosome, dorsal; (B) urosome, ventral;(C) genital area;(D) P5." captionText-8="Fig. 9. Brianola rawaiensis sp. nov., male: (A) antennule (disarticulated)." figureDoi-0="http://doi.org/10.5281/zenodo.14659399" figureDoi-1="http://doi.org/10.5281/zenodo.14659401" figureDoi-2="http://doi.org/10.5281/zenodo.14659403" figureDoi-3="http://doi.org/10.5281/zenodo.14659406" figureDoi-4="http://doi.org/10.5281/zenodo.14659408" figureDoi-5="http://doi.org/10.5281/zenodo.14659410" figureDoi-6="http://doi.org/10.5281/zenodo.14659412" figureDoi-7="http://doi.org/10.5281/zenodo.14659414" figureDoi-8="http://doi.org/10.5281/zenodo.14659416" httpUri-0="https://zenodo.org/record/14659399/files/figure.png" httpUri-1="https://zenodo.org/record/14659401/files/figure.png" httpUri-2="https://zenodo.org/record/14659403/files/figure.png" httpUri-3="https://zenodo.org/record/14659406/files/figure.png" httpUri-4="https://zenodo.org/record/14659408/files/figure.png" httpUri-5="https://zenodo.org/record/14659410/files/figure.png" httpUri-6="https://zenodo.org/record/14659412/files/figure.png" httpUri-7="https://zenodo.org/record/14659414/files/figure.png" httpUri-8="https://zenodo.org/record/14659416/files/figure.png" pageId="2" pageNumber="342">Figs. 1-9</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302656FFFBAFFD86BDDC216FF34FD4D" pageId="2" pageNumber="342" type="materials_examined">
<paragraph id="8BA736E4FFBAFFD86BDDC216FD81FD93" blockId="2.[144,748,519,737]" pageId="2" pageNumber="342">
<materialsCitation id="3B703CB9FFBAFFD86BDDC216FD85FD93" country="Thailand" county="Island" latitude="7.7713885" location="Intertidal" longLatPrecision="15" longitude="98.32103" municipality="Rawai Beach" pageId="2" pageNumber="342" specimenCount="1" stateProvince="Phuket" typeStatus="holotype">
<emphasis id="B96CEAF6FFBAFFD86BDDC216FEFFFDB3" bold="true" box="[144,296,519,543]" pageId="2" pageNumber="342">
<typeStatus id="54A38846FFBAFFD86BDDC216FF1FFDB3" box="[144,200,519,543]" pageId="2" pageNumber="342">Type</typeStatus>
locality.
</emphasis>
<location id="8EC7603FFFBAFFD86A62C216FE46FDB3" LSID="urn:lsid:plazi:treatment:03B187F2FFBAFFD06BDDC1D7FE1EFB4D:8EC7603FFFBAFFD86A62C216FE46FDB3" box="[303,401,519,543]" country="Thailand" county="Island" latitude="7.7713885" longLatPrecision="15" longitude="98.32103" municipality="Rawai Beach" name="Intertidal" pageId="2" pageNumber="342" stateProvince="Phuket">Intertidal</location>
zone at
<collectingMunicipality id="6BC3AC9EFFBAFFD86AA3C216FDAAFDB3" box="[494,637,519,543]" pageId="2" pageNumber="342">Rawai Beach</collectingMunicipality>
of
<collectingRegion id="49DCF806FFBAFFD869EFC216FD3BFDB3" box="[674,748,519,543]" country="Thailand" name="Phuket" pageId="2" pageNumber="342">Phuket</collectingRegion>
<collectingCounty id="62C64E68FFBAFFD86BDDC236FF03FD93" box="[144,212,551,575]" pageId="2" pageNumber="342">Island</collectingCounty>
,
<collectingCountry id="F30F7674FFBAFFD86B93C236FE97FD93" box="[222,320,551,575]" name="Thailand" pageId="2" pageNumber="342">Thailand</collectingCountry>
,
<geoCoordinate id="EE2C5023FFBAFFD86A07C236FE6DFD93" box="[330,442,550,576]" degrees="7" direction="north" minutes="46" orientation="latitude" pageId="2" pageNumber="342" precision="15" seconds="17" value="7.7713885">
7°46
<emphasis id="B96CEAF6FFBAFFD86A31C237FE55FDEC" bold="true" box="[380,386,550,576]" italics="true" pageId="2" pageNumber="342">ʹ</emphasis>
17
<emphasis id="B96CEAF6FFBAFFD86AD0C237FE70FDEC" bold="true" box="[413,423,550,576]" italics="true" pageId="2" pageNumber="342">ʺ</emphasis>
N
</geoCoordinate>
<geoCoordinate id="EE2C5023FFBAFFD86A8CC239FD85FD93" box="[449,594,550,576]" degrees="98" direction="east" minutes="19" orientation="longitude" pageId="2" pageNumber="342" precision="1" seconds="15.7" value="98.32103">
98°19
<emphasis id="B96CEAF6FFBAFFD8694CC237FDD0FDEC" bold="true" box="[513,519,550,576]" italics="true" pageId="2" pageNumber="342">ʹ</emphasis>
15.7
<emphasis id="B96CEAF6FFBAFFD86978C237FDE8FDEC" bold="true" box="[565,575,550,576]" italics="true" pageId="2" pageNumber="342">ʺ</emphasis>
E
</geoCoordinate>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA736E4FFBAFFD86BDDC259FF34FD4D" blockId="2.[144,748,519,737]" pageId="2" pageNumber="342">
<emphasis id="B96CEAF6FFBAFFD86BDDC259FEA7FDCC" bold="true" box="[144,368,584,608]" pageId="2" pageNumber="342">Material examined.</emphasis>
<materialsCitation id="3B703CB9FFBAFFD86A34C259FF34FD4D" collectionCode="NIBR" pageId="2" pageNumber="342" specimenCode="IV0000910337, NIBRIV0000910338, NIBRIV0000910339" specimenCount="6" specimenCount-female="3" specimenCount-male="3" typeStatus="holotype">
<typeStatus id="54A38846FFBAFFD86A34C259FE09FDCC" box="[377,478,584,608]" pageId="2" pageNumber="342" type="holotype">Holotype</typeStatus>
: ovigerous female (
<collectionCode id="ED09AE21FFBAFFD869E1C259FD3BFDCC" box="[684,748,584,608]" country="South Korea" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="2" pageNumber="342">NIBR</collectionCode>
<specimenCode id="DBBE9E9FFFBAFFD86BDDC279FEF9FD2C" box="[144,302,616,640]" collectionCode="IV" pageId="2" pageNumber="342">IV0000910337</specimenCode>
) dissected on 8 slides; adult male
<typeStatus id="54A38846FFBAFFD869D4C279FD3BFD2C" box="[665,748,616,640]" pageId="2" pageNumber="342" type="allotype">allotype</typeStatus>
(
<specimenCode id="DBBE9E9FFFBAFFD86BDBC299FEA4FD0C" box="[150,371,648,672]" collectionCode="NIBRIV" pageId="2" pageNumber="342">NIBRIV0000910338</specimenCode>
) dissected on 7 slides,
<specimenCount id="9D1EFD6DFFBAFFD86913C299FD16FD0C" box="[606,705,648,672]" count="3" pageId="2" pageNumber="342" type="female">3 females</specimenCount>
and
<specimenCount id="9D1EFD6DFFBAFFD86BDDC2B8FE97FD6C" box="[144,320,680,705]" count="3" pageId="2" pageNumber="342" type="male" typeStatus="paratype">3 paratype males</specimenCount>
(
<specimenCode id="DBBE9E9FFFBAFFD86A06C2B8FDFEFD6D" box="[331,553,681,705]" collectionCode="NIBRIV" pageId="2" pageNumber="342">NIBRIV0000910339</specimenCode>
) preserved in 70% ethanol.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302656FFFBAFFD86864C114FAC2FE91" pageId="2" pageNumber="342" type="etymology">
<paragraph id="8BA736E4FFBAFFD86864C114FAC2FE91" blockId="2.[809,1414,261,737]" pageId="2" pageNumber="342">
<emphasis id="B96CEAF6FFBAFFD86864C114FC7EFEB1" bold="true" box="[809,937,261,285]" pageId="2" pageNumber="342">Etymology.</emphasis>
The species name refers to the
<typeStatus id="54A38846FFBAFFD86E4DC114FAF9FEB1" box="[1280,1326,261,285]" pageId="2" pageNumber="342">type</typeStatus>
locality at Rawai muddy sand Beach of
<collectingRegion id="49DCF806FFBAFFD86F31C134FB11FE91" box="[1148,1222,293,317]" country="Thailand" name="Phuket" pageId="2" pageNumber="342">Phuket</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C302656FFFBAFFD06864C154FE1EFB4D" lastPageId="10" lastPageNumber="350" pageId="2" pageNumber="342" type="description">
<paragraph id="8BA736E4FFBAFFD86864C154FC1CFD2C" blockId="2.[809,1414,261,737]" pageId="2" pageNumber="342">
<emphasis id="B96CEAF6FFBAFFD86864C154FC62FEF1" bold="true" box="[809,949,325,349]" pageId="2" pageNumber="342">Description.</emphasis>
Female. Habitus (
<figureCitation id="13232A61FFBAFFD86F35C154FB39FEF2" box="[1144,1262,325,350]" captionStart="Fig" captionStartId="1.[424,457,1938,1960]" captionTargetBox="[357,1260,716,1906]" captionTargetId="figure-229@1.[355,1262,713,1906]" captionTargetPageId="1" captionText="Fig. 1. Brianola rawaiensis sp. nov., female: (A) habitus, dorsal; (B) habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14659399" httpUri="https://zenodo.org/record/14659399/files/figure.png" pageId="2" pageNumber="342">Fig. 1A, B</figureCitation>
). Body length 992 μm. Body length of
<typeStatus id="54A38846FFBAFFD86F79C177FB4EFED2" box="[1076,1177,358,382]" pageId="2" pageNumber="342" type="paratype">paratypes</typeStatus>
821
<emphasis id="B96CEAF6FFBAFFD86F84C17BFB0FFED2" bold="true" box="[1225,1240,362,382]" pageId="2" pageNumber="342">-</emphasis>
1,026 μm (
<emphasis id="B96CEAF6FFBAFFD86E07C177FA80FED1" box="[1354,1367,358,381]" italics="true" pageId="2" pageNumber="342">n</emphasis>
= 6, mean = 919 μm). Body fusiform, the entire body surface ornamented with minutely punctate on the cephalothorax and thoracic segments, otherwise unornamented and with an entire hyaline frill; prosome gradually tapering anteriorly, few sensilla on the body surface. Nauplius eye not visible. Prosome four-segmented, comprising cephalosome fused to first pedigerous somite, and three free pedigerous somites.
</paragraph>
<paragraph id="8BA736E4FFBAFFD86872C299FA51FD4C" blockId="2.[809,1414,261,737]" pageId="2" pageNumber="342">Cephalothorax longer than three prosomites combined, posterior margin of second to fourth prosomites ornamented with rows of hyaline frills. Posterior margin of uroso-</paragraph>
<paragraph id="8BA736E4FFBAFFD86FE5C3B0FB68FC13" blockId="2.[1192,1215,929,959]" box="[1192,1215,929,959]" pageId="2" pageNumber="342">
<emphasis id="B96CEAF6FFBAFFD86FE5C3B0FB68FC13" bold="true" box="[1192,1215,929,959]" pageId="2" pageNumber="342">B</emphasis>
</paragraph>
<caption id="DF67666CFFBAFFD86A41C7A6FADDF861" ID-DOI="http://doi.org/10.5281/zenodo.14659401" ID-Zenodo-Dep="14659401" box="[268,1290,1975,1997]" httpUri="https://zenodo.org/record/14659401/files/figure.png" pageId="2" pageNumber="342" startId="2.[268,301,1975,1997]" targetBox="[341,1176,817,1938]" targetPageId="2" targetType="figure">
<paragraph id="8BA736E4FFBAFFD86A41C7A6FADDF861" blockId="2.[268,1290,1975,1997]" box="[268,1290,1975,1997]" pageId="2" pageNumber="342">
<emphasis id="B96CEAF6FFBAFFD86A41C7A6FE9FF860" bold="true" box="[268,328,1975,1997]" pageId="2" pageNumber="342">Fig. 2.</emphasis>
<taxonomicName id="4C184D67FFBAFFD86A00C7A6FDD2F861" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[333,517,1975,1997]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="2" pageNumber="342" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBAFFD86A00C7A6FDD2F861" box="[333,517,1975,1997]" italics="true" pageId="2" pageNumber="342">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFBAFFD86946C7A9FD83F861" bold="true" box="[523,596,1976,1997]" pageId="2" pageNumber="342">
<taxonomicNameLabel id="A25F578DFFBAFFD86946C7A9FD83F861" box="[523,596,1976,1997]" pageId="2" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) urosome dorsal; (B); urosome ventral; (C) genital area; (D) P5.
</paragraph>
</caption>
<caption id="DF67666CFFBBFFD96A99C688FB56F903" ID-DOI="http://doi.org/10.5281/zenodo.14659403" ID-Zenodo-Dep="14659403" box="[468,1153,1689,1711]" httpUri="https://zenodo.org/record/14659403/files/figure.png" pageId="3" pageNumber="343" startId="3.[468,501,1689,1711]" targetBox="[244,1374,257,1653]" targetPageId="3" targetType="figure">
<paragraph id="8BA736E4FFBBFFD96A99C688FB56F903" blockId="3.[468,1153,1689,1711]" box="[468,1153,1689,1711]" pageId="3" pageNumber="343">
<emphasis id="B96CEAF6FFBBFFD96A99C688FDC7F902" bold="true" box="[468,528,1689,1711]" pageId="3" pageNumber="343">Fig. 3.</emphasis>
<taxonomicName id="4C184D67FFBBFFD96959C688FD1BF903" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[532,716,1689,1711]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="3" pageNumber="343" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBBFFD96959C688FD1BF903" box="[532,716,1689,1711]" italics="true" pageId="3" pageNumber="343">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFBBFFD9699FC688FCCCF902" bold="true" box="[722,795,1689,1710]" pageId="3" pageNumber="343">
<taxonomicNameLabel id="A25F578DFFBBFFD9699FC688FCCCF902" box="[722,795,1689,1710]" pageId="3" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) antennule (disarticulated).
</paragraph>
</caption>
<paragraph id="8BA736E4FFBBFFD96BE3C6E4FCDCF8E2" blockId="3.[174,779,1781,1967]" pageId="3" pageNumber="343">mites ornamented with fine spinules and hyaline frills. Rostrum very large, bell-shaped, slightly exceeding the length of antennules, with two sensilla (but could not see).</paragraph>
<paragraph id="8BA736E4FFBBFFDE6B88C746FB73F8E2" blockId="3.[174,779,1781,1967]" lastBlockId="4.[809,1413,1814,1967]" lastPageId="4" lastPageNumber="344" pageId="3" pageNumber="343">
Urosome (
<figureCitation id="13232A61FFBBFFD96A63C747FE4BF8C2" box="[302,412,1878,1902]" captionStart="Fig" captionStartId="2.[268,301,1975,1997]" captionTargetBox="[341,1176,817,1938]" captionTargetId="figure-211@2.[341,1216,812,1941]" captionTargetPageId="2" captionText="Fig. 2. Brianola rawaiensis sp. nov., female: (A) urosome dorsal; (B); urosome ventral; (C) genital area; (D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659401" httpUri="https://zenodo.org/record/14659401/files/figure.png" pageId="3" pageNumber="343">Fig. 2A, B</figureCitation>
) five-segmented, consisting of rectangular somites, ornamented with rows of fine spinules and hyaline frills dorsally and ventrally, genital doublesomite fused, two abdominal somites, and anal somite. Genital area indistinct, ventrally with one copulatory pore, and two genital apertures (
<figureCitation id="13232A61FFBBFFD96F10C727FB66F8E2" box="[1117,1201,1846,1870]" captionStart="Fig" captionStartId="2.[268,301,1975,1997]" captionTargetBox="[341,1176,817,1938]" captionTargetId="figure-211@2.[341,1216,812,1941]" captionTargetPageId="2" captionText="Fig. 2. Brianola rawaiensis sp. nov., female: (A) urosome dorsal; (B); urosome ventral; (C) genital area; (D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659401" httpUri="https://zenodo.org/record/14659401/files/figure.png" pageId="3" pageNumber="343">Fig. 2C</figureCitation>
). Third and fourth urosomites with rows of spinules on the surface and are ornamented with hyaline frills along the outer margin dorsally and ventrally. Anal operculum semicircular and smooth. Caudal rami unsegmented, twice as long as wide, ornamented with spinules on the outer margin. Each ramus (
<figureCitation id="13232A61FFBCFFDE6BDAC747FF22F8C3" box="[151,245,1878,1903]" captionStart="Fig" captionStartId="1.[424,457,1938,1960]" captionTargetBox="[357,1260,716,1906]" captionTargetId="figure-229@1.[355,1262,713,1906]" captionTargetPageId="1" captionText="Fig. 1. Brianola rawaiensis sp. nov., female: (A) habitus, dorsal; (B) habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14659399" httpUri="https://zenodo.org/record/14659399/files/figure.png" pageId="4" pageNumber="344">Figs. 1A</figureCitation>
;
<figureCitation id="13232A61FFBCFFDE6A4DC746FEE8F8C3" box="[256,319,1879,1903]" captionStart="Fig" captionStartId="2.[268,301,1975,1997]" captionTargetBox="[341,1176,817,1938]" captionTargetId="figure-211@2.[341,1216,812,1941]" captionTargetPageId="2" captionText="Fig. 2. Brianola rawaiensis sp. nov., female: (A) urosome dorsal; (B); urosome ventral; (C) genital area; (D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659401" httpUri="https://zenodo.org/record/14659401/files/figure.png" pageId="4" pageNumber="344">2A, B</figureCitation>
) armed with seven setae: seta I short and located at the outer margin close to seta IV; seta II longer than seta I and III; seta IV shorter than seta V; seta V longest; seta VI short close to seta V; seta VII shortest located close to the insertion point of seta V.
</paragraph>
<caption id="DF67666CFFBCFFDE6BB3C6ABFACDF97C" ID-DOI="http://doi.org/10.5281/zenodo.14659406" ID-Zenodo-Dep="14659406" box="[254,1306,1722,1744]" httpUri="https://zenodo.org/record/14659406/files/figure.png" pageId="4" pageNumber="344" startId="4.[254,287,1722,1744]" targetBox="[232,1320,264,1680]" targetPageId="4" targetType="figure">
<paragraph id="8BA736E4FFBCFFDE6BB3C6ABFACDF97C" blockId="4.[254,1306,1722,1744]" box="[254,1306,1722,1744]" pageId="4" pageNumber="344">
<emphasis id="B96CEAF6FFBCFFDE6BB3C6ABFEEDF963" bold="true" box="[254,314,1722,1744]" pageId="4" pageNumber="344">Fig. 4.</emphasis>
<taxonomicName id="4C184D67FFBCFFDE6A73C6ABFE21F963" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[318,502,1722,1743]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="4" pageNumber="344" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBCFFDE6A73C6ABFE21F963" box="[318,502,1722,1743]" italics="true" pageId="4" pageNumber="344">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFBCFFDE6AB1C6ABFD93F963" bold="true" box="[508,580,1722,1743]" pageId="4" pageNumber="344">
<taxonomicNameLabel id="A25F578DFFBCFFDE6AB1C6ABFD93F963" box="[508,580,1722,1743]" pageId="4" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped.
</paragraph>
</caption>
<paragraph id="8BA736E4FFBCFFDF6872C747FD4EF943" blockId="4.[809,1413,1814,1967]" lastBlockId="5.[174,779,1622,1968]" lastPageId="5" lastPageNumber="345" pageId="4" pageNumber="344">
Antennule (
<figureCitation id="13232A61FFBCFFDE68F7C747FBD9F8C3" box="[954,1038,1878,1903]" captionStart="Fig" captionStartId="3.[468,501,1689,1711]" captionTargetBox="[244,1374,257,1653]" captionTargetId="figure-118@3.[243,1375,257,1655]" captionTargetPageId="3" captionText="Fig. 3. Brianola rawaiensis sp. nov., female: (A) antennule(disarticulated)." figureDoi="http://doi.org/10.5281/zenodo.14659403" httpUri="https://zenodo.org/record/14659403/files/figure.png" pageId="4" pageNumber="344">Fig. 3A</figureCitation>
) four-segmented indistinctly, without spinules on the dorsal surface. First segment with five spinules at the inner margin and six pinnate setae at the inner corner. Second segment with seven spinulose and twelve smooth setae plus two aesthetascs. Third segment bearing two spinulate setae and one smooth seta. Distal segment with seven spinulate and six smooth setae. Armature formula: 1-(6), 2-(19 + 2 ae), 3-(3), 4-(13).
</paragraph>
<caption id="DF67666CFFBDFFDF6AF2C5E8FB43F9A3" ID-DOI="http://doi.org/10.5281/zenodo.14659408" ID-Zenodo-Dep="14659408" box="[447,1172,1529,1551]" httpUri="https://zenodo.org/record/14659408/files/figure.png" pageId="5" pageNumber="345" startId="5.[447,480,1529,1551]" targetBox="[289,1320,258,1494]" targetPageId="5" targetType="figure">
<paragraph id="8BA736E4FFBDFFDF6AF2C5E8FB43F9A3" blockId="5.[447,1172,1529,1551]" box="[447,1172,1529,1551]" pageId="5" pageNumber="345">
<emphasis id="B96CEAF6FFBDFFDF6AF2C5E8FE2CF9A3" bold="true" box="[447,507,1529,1551]" pageId="5" pageNumber="345">Fig. 5.</emphasis>
<taxonomicName id="4C184D67FFBDFFDF694DC5E8FD6EF9A3" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[512,697,1529,1551]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="5" pageNumber="345" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBDFFDF694DC5E8FD6EF9A3" box="[512,697,1529,1551]" italics="true" pageId="5" pageNumber="345">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFBDFFDF69F3C5EBFCD0F9A3" bold="true" box="[702,775,1530,1551]" pageId="5" pageNumber="345">
<taxonomicNameLabel id="A25F578DFFBDFFDF69F3C5EBFCD0F9A3" box="[702,775,1530,1551]" pageId="5" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) P1, anterior; (B) P2, posterior.
</paragraph>
</caption>
<paragraph id="8BA736E4FFBDFFDF6B88C6E6FB78F921" blockId="5.[174,779,1622,1968]" lastBlockId="5.[839,1445,1622,1968]" pageId="5" pageNumber="345">
Antenna (
<figureCitation id="13232A61FFBDFFDF6A64C6E6FEA8F8A3" box="[297,383,1783,1807]" captionStart="Fig" captionStartId="4.[254,287,1722,1744]" captionTargetBox="[232,1320,264,1680]" captionTargetId="figure-114@4.[231,1326,257,1688]" captionTargetPageId="4" captionText="Fig. 4. Brianola rawaiensis sp. nov., female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14659406" httpUri="https://zenodo.org/record/14659406/files/figure.png" pageId="5" pageNumber="345">Fig. 4A</figureCitation>
). Basis with small spinules on middle and outer margin. Exopod consist of eight segments; fourth segment with one long naked seta; ultimate segment with four pinnate setae. Endopod three-segmented; enp-1 with one long setule and spinular row along outer margin and anterior surface, enp-2 with one naked seta and three spinulose setae, enp-3 with six spinulose setae, one pinnate seta and one naked short seta.
</paragraph>
<paragraph id="8BA736E4FFBDFFDF6813C687FA8AF8C3" blockId="5.[839,1445,1622,1968]" pageId="5" pageNumber="345">
Mandible (
<figureCitation id="13232A61FFBDFFDF6880C687FBCBF902" box="[973,1052,1686,1710]" captionStart="Fig" captionStartId="4.[254,287,1722,1744]" captionTargetBox="[232,1320,264,1680]" captionTargetId="figure-114@4.[231,1326,257,1688]" captionTargetPageId="4" captionText="Fig. 4. Brianola rawaiensis sp. nov., female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14659406" httpUri="https://zenodo.org/record/14659406/files/figure.png" pageId="5" pageNumber="345">Fig. 4B</figureCitation>
) with large and elongated coxa, without surface ornamentation. Gnathobase well-developed, cutting edge with two rows of strong teeth in different sizes. Basis bearing two naked setae. Endopod two-segmented, enp-1 with two pinnate and one spinulose setae; enp-2 with three plumose setae fused with segment. Exopod one-segmented, with seven naked and three plumose setae.
</paragraph>
<paragraph id="8BA736E4FFBDFFDF6813C769FA73F81C" blockId="5.[839,1445,1622,1968]" pageId="5" pageNumber="345">
Maxillule (
<figureCitation id="13232A61FFBDFFDF689EC769FBFEF83C" box="[979,1065,1912,1936]" captionStart="Fig" captionStartId="4.[254,287,1722,1744]" captionTargetBox="[232,1320,264,1680]" captionTargetId="figure-114@4.[231,1326,257,1688]" captionTargetPageId="4" captionText="Fig. 4. Brianola rawaiensis sp. nov., female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14659406" httpUri="https://zenodo.org/record/14659406/files/figure.png" pageId="5" pageNumber="345">Fig. 4C</figureCitation>
). Praecoxa arthrite with two juxtaposed slender setae on anterior surface and with eight dis-
</paragraph>
<paragraph id="8BA736E4FFBEFFDC6A55C229FD52F8E3" blockId="6.[280,302,568,598]" lastBlockId="6.[144,749,1622,1969]" pageId="6" pageNumber="346">
<emphasis id="B96CEAF6FFBEFFDC6A55C229FEF9FDFA" bold="true" box="[280,302,568,598]" pageId="6" pageNumber="346">A</emphasis>
tal spines. Coxal endite cylindrical, bearing three spinulose setae; epipodite represented by three plumose setae. Basis ornamented with rows of spinules on surface, basal endite bearing six spinulose and two smooth setae. Endopod two-segmented, end-1 bearing three spinulose and two smooth setae; enp-2 bearing four spinulose and two plumose setae. Exopod one-segmented, with six plumose setae fused with segment and one pectinate seta.
</paragraph>
<caption id="DF67666CFFBEFFDC6AD1C5EBFBACF9BC" ID-DOI="http://doi.org/10.5281/zenodo.14659410" ID-Zenodo-Dep="14659410" box="[412,1147,1530,1552]" httpUri="https://zenodo.org/record/14659410/files/figure.png" pageId="6" pageNumber="346" startId="6.[412,445,1530,1552]" targetBox="[300,1294,258,1498]" targetPageId="6" targetType="figure">
<paragraph id="8BA736E4FFBEFFDC6AD1C5EBFBACF9BC" blockId="6.[412,1147,1530,1552]" box="[412,1147,1530,1552]" pageId="6" pageNumber="346">
<emphasis id="B96CEAF6FFBEFFDC6AD1C5EBFE0FF9A3" bold="true" box="[412,472,1530,1552]" pageId="6" pageNumber="346">Fig. 6.</emphasis>
<taxonomicName id="4C184D67FFBEFFDC6A90C5EBFD42F9A3" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[477,661,1530,1551]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="6" pageNumber="346" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBEFFDC6A90C5EBFD42F9A3" box="[477,661,1530,1551]" italics="true" pageId="6" pageNumber="346">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFBEFFDC69D6C5EBFD33F9A3" bold="true" box="[667,740,1530,1551]" pageId="6" pageNumber="346">
<taxonomicNameLabel id="A25F578DFFBEFFDC69D6C5EBFD33F9A3" box="[667,740,1530,1551]" pageId="6" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: (A) P3, posterior; (B) P4, posterior.
</paragraph>
</caption>
<paragraph id="8BA736E4FFBEFFDC6BEBC749FAA1F943" blockId="6.[144,749,1622,1969]" lastBlockId="6.[809,1414,1622,1969]" pageId="6" pageNumber="346">
Maxilla (
<figureCitation id="13232A61FFBEFFDC6A49C749FE81F8DC" box="[260,342,1880,1904]" captionStart="Fig" captionStartId="4.[254,287,1722,1744]" captionTargetBox="[232,1320,264,1680]" captionTargetId="figure-114@4.[231,1326,257,1688]" captionTargetPageId="4" captionText="Fig. 4. Brianola rawaiensis sp. nov., female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14659406" httpUri="https://zenodo.org/record/14659406/files/figure.png" pageId="6" pageNumber="346">Fig. 4D</figureCitation>
) comprising praecoxa, coxa, allobasis, and one-segmented endopod. Praecoxa bearing one row of spinules, two praecoxal endites with two pinnate setae and two spinulose setae; coxa ornamented with different sizes of spinules on surface, bearing two coxal endites with three spinulose setae each; basis extended into a strong claw and one spinulose spine, basal endite with two plumose setae. Endopod one-segmented, with seven smooth setae.
</paragraph>
<paragraph id="8BA736E4FFBEFFDC6872C6E6FA54F81C" blockId="6.[809,1414,1622,1969]" pageId="6" pageNumber="346">
Maxilliped (
<figureCitation id="13232A61FFBEFFDC68F2C6E6FBC7F8A3" box="[959,1040,1783,1807]" captionStart="Fig" captionStartId="4.[254,287,1722,1744]" captionTargetBox="[232,1320,264,1680]" captionTargetId="figure-114@4.[231,1326,257,1688]" captionTargetPageId="4" captionText="Fig. 4. Brianola rawaiensis sp. nov., female: (A) antenna; (B) mandible; (C) maxillule; (D) maxilla; (E) maxilliped." figureDoi="http://doi.org/10.5281/zenodo.14659406" httpUri="https://zenodo.org/record/14659406/files/figure.png" pageId="6" pageNumber="346">Fig. 4E</figureCitation>
) phyllopodial, comprising syncoxa (praecox and coxa fused); basis, and one-segmented endopod. Syncoxa bearing one long plumose seta at base and five spinulose and one naked seta. Basis bearing four spinulose setae, and one row of long setules along outer margin. Endopod with three spinulose and four plumose setae.
</paragraph>
<caption id="DF67666CFFBFFFDD6AFEC5A6FB75FA61" ID-DOI="http://doi.org/10.5281/zenodo.14659412" ID-Zenodo-Dep="14659412" box="[435,1186,1463,1485]" httpUri="https://zenodo.org/record/14659412/files/figure.png" pageId="7" pageNumber="347" startId="7.[435,468,1463,1485]" targetBox="[385,1251,261,1430]" targetPageId="7" targetType="figure">
<paragraph id="8BA736E4FFBFFFDD6AFEC5A6FB75FA61" blockId="7.[435,1186,1463,1485]" box="[435,1186,1463,1485]" pageId="7" pageNumber="347">
<emphasis id="B96CEAF6FFBFFFDD6AFEC5A6FE38FA60" bold="true" box="[435,495,1463,1485]" pageId="7" pageNumber="347">Fig. 7.</emphasis>
<taxonomicName id="4C184D67FFBFFFDD6ABFC5A6FD7CFA60" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[498,683,1463,1484]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="7" pageNumber="347" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFBFFFDD6ABFC5A6FD7CFA60" box="[498,683,1463,1484]" italics="true" pageId="7" pageNumber="347">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFBFFFDD69FCC5A6FD2EFA60" bold="true" box="[689,761,1463,1484]" pageId="7" pageNumber="347">
<taxonomicNameLabel id="A25F578DFFBFFFDD69FCC5A6FD2EFA60" box="[689,761,1463,1484]" pageId="7" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: (A) habitus, dorsal; (B) habitus, lateral.
</paragraph>
</caption>
<paragraph id="8BA736E4FFBFFFDD6B88C602FED7F920" blockId="7.[174,779,1555,1966]" pageId="7" pageNumber="347">
All swimming legs (
<figureCitation id="13232A61FFBFFFDD6AD4C602FDC1F986" box="[409,534,1555,1579]" captionStart="Fig" captionStartId="5.[447,480,1529,1551]" captionTargetBox="[289,1320,258,1494]" captionTargetId="figure-207@5.[296,1322,257,1497]" captionTargetPageId="5" captionText="Fig. 5. Brianola rawaiensis sp. nov., female: (A) P1, anterior; (B) P2, posterior." figureDoi="http://doi.org/10.5281/zenodo.14659408" httpUri="https://zenodo.org/record/14659408/files/figure.png" pageId="7" pageNumber="347">Figs. 5A, B</figureCitation>
;
<figureCitation id="13232A61FFBFFFDD696FC602FDB5F987" box="[546,610,1555,1579]" captionStart="Fig" captionStartId="6.[412,445,1530,1552]" captionTargetBox="[300,1294,258,1498]" captionTargetId="figure-198@6.[260,1298,257,1499]" captionTargetPageId="6" captionText="Fig. 6. Brianola rawaiensis sp. nov., female: (A) P3, posterior; (B) P4, posterior." figureDoi="http://doi.org/10.5281/zenodo.14659410" httpUri="https://zenodo.org/record/14659410/files/figure.png" pageId="7" pageNumber="347">6A, B</figureCitation>
) biramous; P1
<emphasis id="B96CEAF6FFBFFFDD69B1C606FCDCF987" bold="true" box="[764,779,1559,1579]" pageId="7" pageNumber="347">-</emphasis>
P4 with three-segmented exopods and endopods. Members of each leg pair are connected by simple intercoxal sclerite.
</paragraph>
<paragraph id="8BA736E4FFBFFFDD6B88C685FA74F920" blockId="7.[174,779,1555,1966]" lastBlockId="7.[839,1444,1555,1966]" pageId="7" pageNumber="347">
P1 (
<figureCitation id="13232A61FFBFFFDD6BA0C685FE94F900" box="[237,323,1684,1708]" captionStart="Fig" captionStartId="5.[447,480,1529,1551]" captionTargetBox="[289,1320,258,1494]" captionTargetId="figure-207@5.[296,1322,257,1497]" captionTargetPageId="5" captionText="Fig. 5. Brianola rawaiensis sp. nov., female: (A) P1, anterior; (B) P2, posterior." figureDoi="http://doi.org/10.5281/zenodo.14659408" httpUri="https://zenodo.org/record/14659408/files/figure.png" pageId="7" pageNumber="347">Fig. 5A</figureCitation>
). Coxa with one inner pinnate spine, ornamented with rows of spinules on the surface, outer margin with two setules and five spinules. Basis ornamented with setules distally, with one inner pinnate spine and one outer spinulose seta. Exopod shorter than endopod. Exp-1 with one outer pinnate spine; exp-2 with one outer pectinate spine and one inner plumose seta; exp-3 with one outer pectinate and one outer pinnate spine, one distal pinnate spine and one distal spinulose spine, and one inner plumose seta. Enp-1 and enp-2 with plumose inner seta each; enp-3 with one naked and one pinnate outer spine, one spinulose spine and one plumose seta distally, and one short bare seta and one plumose seta on inner margin.
</paragraph>
<paragraph id="8BA736E4FFBFFFDD6813C685FA73F802" blockId="7.[839,1444,1555,1966]" pageId="7" pageNumber="347">
P2 (
<figureCitation id="13232A61FFBFFFDD68C5C685FC08F900" box="[904,991,1684,1708]" captionStart="Fig" captionStartId="5.[447,480,1529,1551]" captionTargetBox="[289,1320,258,1494]" captionTargetId="figure-207@5.[296,1322,257,1497]" captionTargetPageId="5" captionText="Fig. 5. Brianola rawaiensis sp. nov., female: (A) P1, anterior; (B) P2, posterior." figureDoi="http://doi.org/10.5281/zenodo.14659408" httpUri="https://zenodo.org/record/14659408/files/figure.png" pageId="7" pageNumber="347">Fig. 5B</figureCitation>
). Coxa ornamented with several spinular rows of unequal length on the anterior surface, and a row of spinules along the outer margin. Basis ornamented with small and large spinules along the outer margin and one outer small seta. Exopod shorter than endopod. Exp-1 with one outer pinnate spine; exp-2 with one outer pinnate spine and one inner plumose seta; exp-3 with one outer pinnate spine, one distal spinulose spine, and one inner plumose seta. Enp-1 and enp-2 with one inner plumose
</paragraph>
<paragraph id="8BA736E4FFB0FFD26A10C1BDFB16FE66" blockId="8.[349,371,428,458]" box="[349,1217,428,458]" lastBlockId="8.[1194,1217,428,458]" pageId="8" pageNumber="348">
<emphasis id="B96CEAF6FFB0FFD26A10C1BDFB16FE66" bold="true" box="[349,1217,428,458]" pageId="8" pageNumber="348">A B</emphasis>
</paragraph>
<caption id="DF67666CFFB0FFD26A59C54FFAD5FAD8" ID-DOI="http://doi.org/10.5281/zenodo.14659414" ID-Zenodo-Dep="14659414" box="[276,1282,1374,1396]" httpUri="https://zenodo.org/record/14659414/files/figure.png" pageId="8" pageNumber="348" startId="8.[276,309,1374,1396]" targetBox="[366,1159,257,1339]" targetPageId="8" targetType="figure">
<paragraph id="8BA736E4FFB0FFD26A59C54FFAD5FAD8" blockId="8.[276,1282,1374,1396]" box="[276,1282,1374,1396]" pageId="8" pageNumber="348">
<emphasis id="B96CEAF6FFB0FFD26A59C54FFE87FADF" bold="true" box="[276,336,1374,1396]" pageId="8" pageNumber="348">Fig. 8.</emphasis>
<taxonomicName id="4C184D67FFB0FFD26A18C54FFDD9FAD8" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[341,526,1374,1396]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="8" pageNumber="348" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFB0FFD26A18C54FFDD9FAD8" box="[341,526,1374,1396]" italics="true" pageId="8" pageNumber="348">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFB0FFD26959C54EFD8BFAD8" bold="true" box="[532,604,1375,1396]" pageId="8" pageNumber="348">
<taxonomicNameLabel id="A25F578DFFB0FFD26959C54EFD8BFAD8" box="[532,604,1375,1396]" pageId="8" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: (A) urosome, dorsal; (B) urosome, ventral; (C) genital area; (D) P5.
</paragraph>
</caption>
<paragraph id="8BA736E4FFB0FFD26BDDC5AAFEFEF9BF" blockId="8.[144,749,1467,1975]" pageId="8" pageNumber="348">seta each; enp-3 with one outer pinnate spine, one distal pinnate spine and one distal spinulose spine, and two inner plumose setae.</paragraph>
<paragraph id="8BA736E4FFB0FFD26BEBC60DFE71F938" blockId="8.[144,749,1467,1975]" pageId="8" pageNumber="348">
P3 (
<figureCitation id="13232A61FFB0FFD26B9DC60AFEF0F99F" box="[208,295,1563,1587]" captionStart="Fig" captionStartId="6.[412,445,1530,1552]" captionTargetBox="[300,1294,258,1498]" captionTargetId="figure-198@6.[260,1298,257,1499]" captionTargetPageId="6" captionText="Fig. 6. Brianola rawaiensis sp. nov., female: (A) P3, posterior; (B) P4, posterior." figureDoi="http://doi.org/10.5281/zenodo.14659410" httpUri="https://zenodo.org/record/14659410/files/figure.png" pageId="8" pageNumber="348">Fig. 6A</figureCitation>
) rather like P2 except for setal formulae. Exp-3 and enp-3 with one outer pinnate spine, one distal pinnate spine and one distal spinulose spine, and one inner plumose seta, respectively.
</paragraph>
<paragraph id="8BA736E4FFB0FFD26BEBC68CFE4DF83A" blockId="8.[144,749,1467,1975]" pageId="8" pageNumber="348">
P4 (
<figureCitation id="13232A61FFB0FFD26B82C68DFEF4F918" box="[207,291,1692,1716]" captionStart="Fig" captionStartId="6.[412,445,1530,1552]" captionTargetBox="[300,1294,258,1498]" captionTargetId="figure-198@6.[260,1298,257,1499]" captionTargetPageId="6" captionText="Fig. 6. Brianola rawaiensis sp. nov., female: (A) P3, posterior; (B) P4, posterior." figureDoi="http://doi.org/10.5281/zenodo.14659410" httpUri="https://zenodo.org/record/14659410/files/figure.png" pageId="8" pageNumber="348">Fig. 6B</figureCitation>
). Coxa and basis ornamentation are almost the same as P2 and P3. Exp-1 and exp-2 with outer pinnate spine each; exp-3 with one outer pinnate spine, one distal pinnate spine and one distal spinulose spine, and one inner pectinate spine. Enp-1with very short smooth inner seta; enp-2 bare; enp-3 with outer bare spine, one distal pinnate spine and one distal spinulose spine, and one inner pectinate spine.
</paragraph>
<paragraph id="8BA736E4FFB0FFD26BEBC78EFD3BF81B" blockId="8.[144,749,1467,1975]" box="[166,748,1951,1975]" pageId="8" pageNumber="348">The spine and setal formula of the swimming legs of</paragraph>
<paragraph id="8BA736E4FFB0FFD26864C5ABFAD9FA7F" blockId="8.[809,1294,1466,1491]" box="[809,1294,1466,1491]" pageId="8" pageNumber="348">
<taxonomicName id="4C184D67FFB0FFD26864C5ABFBC7FA7E" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[809,1040,1466,1490]" class="Copepoda" family="Canuellidae" genus="Brianoloa" kingdom="Animalia" order="Polyarthra" pageId="8" pageNumber="348" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFB0FFD26864C5ABFBC7FA7E" box="[809,1040,1466,1490]" italics="true" pageId="8" pageNumber="348">Brianoloa rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFB0FFD26F5BC5AAFBBCFA7F" bold="true" box="[1046,1131,1467,1491]" pageId="8" pageNumber="348">
<taxonomicNameLabel id="A25F578DFFB0FFD26F5BC5AAFBBCFA7F" box="[1046,1131,1467,1491]" pageId="8" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are as follows:
</paragraph>
<paragraph id="8BA736E4FFB0FFD26884C5FBFB31F9AE" blockId="8.[831,1254,1514,1667]" box="[969,1254,1514,1538]" pageId="8" pageNumber="348">Exopod Endopod</paragraph>
<paragraph id="8BA736E4FFB0FFD26872C61BFB08F98E" blockId="8.[831,1254,1514,1667]" box="[831,1247,1546,1570]" pageId="8" pageNumber="348">P1 0.1.122 1.1.222</paragraph>
<paragraph id="8BA736E4FFB0FFD26872C63AFB08F9EF" blockId="8.[831,1254,1514,1667]" box="[831,1247,1579,1603]" pageId="8" pageNumber="348">P2 0.1.111 1.1.221</paragraph>
<paragraph id="8BA736E4FFB0FFD26872C65AFB09F9CF" blockId="8.[831,1254,1514,1667]" box="[831,1246,1611,1635]" pageId="8" pageNumber="348">P3 0.1.121 1.1.121</paragraph>
<paragraph id="8BA736E4FFB0FFD26872C67AFB09F92F" blockId="8.[831,1254,1514,1667]" box="[831,1246,1643,1667]" pageId="8" pageNumber="348">P4 0.0.121 1.0.121</paragraph>
<paragraph id="8BA736E4FFB0FFD26872C68BFAE6F95F" blockId="8.[809,1415,1690,1973]" pageId="8" pageNumber="348">
P5 (
<figureCitation id="13232A61FFB0FFD2682AC68BFC6EF91F" box="[871,953,1690,1715]" captionStart="Fig" captionStartId="2.[268,301,1975,1997]" captionTargetBox="[341,1176,817,1938]" captionTargetId="figure-211@2.[341,1216,812,1941]" captionTargetPageId="2" captionText="Fig. 2. Brianola rawaiensis sp. nov., female: (A) urosome dorsal; (B); urosome ventral; (C) genital area; (D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659401" httpUri="https://zenodo.org/record/14659401/files/figure.png" pageId="8" pageNumber="348">Fig. 2D</figureCitation>
) reduced, incorporated into somite; connected by deeply incised hyaline frills; with four pinnate setae: outer seta longest, two inner setae equal in length.
</paragraph>
<paragraph id="8BA736E4FFB0FFD26864C6EAFAD1F819" blockId="8.[809,1415,1690,1973]" pageId="8" pageNumber="348">
<emphasis id="B96CEAF6FFB0FFD26864C6EAFB57F8B8" bold="true" box="[809,1152,1787,1812]" pageId="8" pageNumber="348">
Description of
<typeStatus id="54A38846FFB0FFD26897C6EDFBE0F8B8" box="[986,1079,1788,1812]" pageId="8" pageNumber="348" type="allotype">allotype</typeStatus>
male.
</emphasis>
Habitus is almost as in female (
<figureCitation id="13232A61FFB0FFD26830C70DFC25F898" box="[893,1010,1820,1844]" captionStart="Fig" captionStartId="7.[435,468,1463,1485]" captionTargetBox="[385,1251,261,1430]" captionTargetId="figure-254@7.[365,1252,257,1431]" captionTargetPageId="7" captionText="Fig. 7.Brianola rawaiensis sp. nov., male: (A) habitus, dorsal; (B) habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.14659412" httpUri="https://zenodo.org/record/14659412/files/figure.png" pageId="8" pageNumber="348">Fig. 7A, B</figureCitation>
). The body length of the
<typeStatus id="54A38846FFB0FFD26E4FC70DFA81F898" box="[1282,1366,1820,1844]" pageId="8" pageNumber="348" type="allotype">allotype</typeStatus>
902 μm. Body length of
<typeStatus id="54A38846FFB0FFD268B0C72DFBB6F8F8" box="[1021,1121,1852,1876]" pageId="8" pageNumber="348" type="paratype">paratypes</typeStatus>
males 796
<emphasis id="B96CEAF6FFB0FFD26F9EC751FB35F8F8" bold="true" box="[1235,1250,1856,1876]" pageId="8" pageNumber="348">-</emphasis>
929 μm (
<emphasis id="B96CEAF6FFB0FFD26E70C72DFA9DF8FF" box="[1341,1354,1852,1875]" italics="true" pageId="8" pageNumber="348">n</emphasis>
= 12, mean = 874 μm). Antennae, mouthparts, swimming legs, and caudal rami as in female. Sexual dimorphisms are expressed in antennules and abdominal somites.
</paragraph>
<caption id="DF67666CFFB1FFD36A96C672FBAFF9D5" ID-DOI="http://doi.org/10.5281/zenodo.14659416" ID-Zenodo-Dep="14659416" box="[475,1144,1635,1657]" httpUri="https://zenodo.org/record/14659416/files/figure.png" pageId="9" pageNumber="349" startId="9.[475,508,1635,1657]" targetBox="[381,1303,262,1602]" targetPageId="9" targetType="figure">
<paragraph id="8BA736E4FFB1FFD36A96C672FBAFF9D5" blockId="9.[475,1144,1635,1657]" box="[475,1144,1635,1657]" pageId="9" pageNumber="349">
<emphasis id="B96CEAF6FFB1FFD36A96C672FDC0F9D4" bold="true" box="[475,535,1635,1657]" pageId="9" pageNumber="349">Fig. 9.</emphasis>
<taxonomicName id="4C184D67FFB1FFD36950C672FD02F9D5" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[541,725,1635,1657]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="9" pageNumber="349" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFB1FFD36950C672FD02F9D5" box="[541,725,1635,1657]" italics="true" pageId="9" pageNumber="349">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFB1FFD36996C675FCF4F9D5" bold="true" box="[731,803,1636,1657]" pageId="9" pageNumber="349">
<taxonomicNameLabel id="A25F578DFFB1FFD36996C675FCF4F9D5" box="[731,803,1636,1657]" pageId="9" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: (A) antennule (disarticulated).
</paragraph>
</caption>
<caption id="DF67666CFFB1FFD36BE3C6D0FC6FF97A" box="[174,952,1728,1750]" pageId="9" pageNumber="349" startId="9.[174,227,1729,1750]" targetType="table">
<paragraph id="8BA736E4FFB1FFD36BE3C6D0FC6FF97A" blockId="9.[174,952,1728,1750]" box="[174,952,1728,1750]" pageId="9" pageNumber="349">
<emphasis id="B96CEAF6FFB1FFD36BE3C6D0FED7F979" bold="true" box="[174,256,1728,1750]" pageId="9" pageNumber="349">Table 1.</emphasis>
Character difference between
<taxonomicName id="4C184D67FFB1FFD3695BC6D1FD19F97A" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[534,718,1728,1750]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="9" pageNumber="349" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFB1FFD3695BC6D1FD19F97A" box="[534,718,1728,1750]" italics="true" pageId="9" pageNumber="349">Brianola rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFB1FFD36999C6D0FCCBF97A" bold="true" box="[724,796,1729,1750]" pageId="9" pageNumber="349">
<taxonomicNameLabel id="A25F578DFFB1FFD36999C6D0FCCBF97A" box="[724,796,1729,1750]" pageId="9" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C184D67FFB1FFD36805C6D0FC64F979" authorityName="Nazari, Mirshamsi, Sari, Aliabadian &amp; Martinez Arbizu" authorityYear="2018" box="[840,947,1728,1750]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="9" pageNumber="349" phylum="Arthropoda" rank="species" species="haliensis">
<emphasis id="B96CEAF6FFB1FFD36805C6D0FC64F979" box="[840,947,1728,1750]" italics="true" pageId="9" pageNumber="349">B. haliensis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA736E4FFB1FFD3693BC6E7FAAAF809" pageId="9" pageNumber="349">
<table id="F918C444FFB100256B99C6E7FAAAF809" box="[212,1405,1782,1957]" gridcols="3" gridrows="5" pageId="9" pageNumber="349">
<tr id="352834A6FFB100256B99C6E7FAAAF8A7" box="[212,1405,1782,1803]" gridrow="0" pageId="9" pageNumber="349" rowspan-0="1">
<th id="76F95DDAFFB100256922C6E7FC90F8A7" box="[623,839,1782,1803]" gridcol="1" gridrow="0" pageId="9" pageNumber="349">
<taxonomicName id="4C184D67FFB1FFD3693BC6E7FD26F8A7" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[630,753,1782,1803]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="9" pageNumber="349" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFB1FFD3693BC6E7FD26F8A7" box="[630,753,1782,1803]" italics="true" pageId="9" pageNumber="349">B. rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFB1FFD369BAC6E7FCE8F8A7" bold="true" box="[759,831,1782,1803]" pageId="9" pageNumber="349">
<taxonomicNameLabel id="A25F578DFFB1FFD369BAC6E7FCE8F8A7" box="[759,831,1782,1803]" pageId="9" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76F95DDAFFB100256FC3C6E7FAAAF8A7" box="[1166,1405,1782,1803]" gridcol="2" gridrow="0" pageId="9" pageNumber="349">
<taxonomicName id="4C184D67FFB1FFD36F9CC6E7FAECF8A7" authorityName="Nazari, Mirshamsi, Sari, Aliabadian &amp; Martinez Arbizu" authorityYear="2018" box="[1233,1339,1782,1803]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="9" pageNumber="349" phylum="Arthropoda" rank="species" species="haliensis">
<emphasis id="B96CEAF6FFB1FFD36F9CC6E7FAECF8A7" box="[1233,1339,1782,1803]" italics="true" pageId="9" pageNumber="349">B. haliensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="352834A6FFB100256B99C73AFAAAF8ED" box="[212,1405,1835,1857]" gridrow="1" pageId="9" pageNumber="349">
<th id="76F95DDAFFB100256B99C73AFEF2F8ED" box="[212,293,1835,1857]" gridcol="0" gridrow="1" pageId="9" pageNumber="349">P1 exp-1</th>
<td id="76F95DDAFFB100256922C73AFC90F8ED" box="[623,839,1835,1857]" gridcol="1" gridrow="1" pageId="9" pageNumber="349">One outer pinnate spine</td>
<td id="76F95DDAFFB100256FC3C73AFAAAF8ED" box="[1166,1405,1835,1857]" gridcol="2" gridrow="1" pageId="9" pageNumber="349">One outer pectinate spine</td>
</tr>
<tr id="352834A6FFB100256B99C75DFAAAF8CE" box="[212,1405,1868,1890]" gridrow="2" pageId="9" pageNumber="349">
<th id="76F95DDAFFB100256B99C75DFEF2F8CE" box="[212,293,1868,1890]" gridcol="0" gridrow="2" pageId="9" pageNumber="349">P1 exp-2</th>
<td id="76F95DDAFFB100256922C75DFC90F8CE" box="[623,839,1868,1890]" gridcol="1" gridrow="2" pageId="9" pageNumber="349">One plumose inner seta</td>
<td id="76F95DDAFFB100256FC3C75DFAAAF8CE" box="[1166,1405,1868,1890]" gridcol="2" gridrow="2" pageId="9" pageNumber="349">Without inner seta</td>
</tr>
<tr id="352834A6FFB100256B99C77CFAAAF82F" box="[212,1405,1901,1923]" gridrow="3" pageId="9" pageNumber="349">
<th id="76F95DDAFFB100256B99C77CFEF2F82F" box="[212,293,1901,1923]" gridcol="0" gridrow="3" pageId="9" pageNumber="349">P2 exp-3</th>
<td id="76F95DDAFFB100256922C77CFC90F82F" box="[623,839,1901,1923]" gridcol="1" gridrow="3" pageId="9" pageNumber="349">2 spines and 1 seta</td>
<td id="76F95DDAFFB100256FC3C77CFAAAF82F" box="[1166,1405,1901,1923]" gridcol="2" gridrow="3" pageId="9" pageNumber="349">3 spines and 1 seta</td>
</tr>
<tr id="352834A6FFB100256B99C79EFAAAF809" box="[212,1405,1935,1957]" gridrow="4" pageId="9" pageNumber="349">
<th id="76F95DDAFFB100256B99C79EFEF2F809" box="[212,293,1935,1957]" gridcol="0" gridrow="4" pageId="9" pageNumber="349">P4 enp-1</th>
<td id="76F95DDAFFB100256922C79EFC90F809" box="[623,839,1935,1957]" gridcol="1" gridrow="4" pageId="9" pageNumber="349">Without inner seta/spine</td>
<td id="76F95DDAFFB100256FC3C79EFAAAF809" box="[1166,1405,1935,1957]" gridcol="2" gridrow="4" pageId="9" pageNumber="349">One short stout inner spine</td>
</tr>
</table>
</paragraph>
<caption id="DF67666CFFB2FFD06BDDC115FC4DFEB6" box="[144,922,260,282]" pageId="10" pageNumber="350" startId="10.[144,197,260,281]" targetType="table">
<paragraph id="8BA736E4FFB2FFD06BDDC115FC4DFEB6" blockId="10.[144,922,260,282]" box="[144,922,260,282]" pageId="10" pageNumber="350">
<emphasis id="B96CEAF6FFB2FFD06BDDC115FF35FEB5" bold="true" box="[144,226,260,281]" pageId="10" pageNumber="350">Table 2.</emphasis>
Occurrence information about species belonging to
<taxonomicName id="4C184D67FFB2FFD069F0C115FC41FEB6" authority="Monard, 1926" authorityName="Monard" authorityYear="1926" box="[701,918,260,282]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="genus">
<emphasis id="B96CEAF6FFB2FFD069F0C115FCD9FEB5" box="[701,782,260,281]" italics="true" pageId="10" pageNumber="350">Brianola</emphasis>
<bibRefCitation id="EF894B15FFB2FFD0685EC115FC41FEB6" author="Monard, A." box="[787,918,260,282]" pageId="10" pageNumber="350" pagination="419 - 434" refId="ref4556" refString="Monard, A. 1926. Sur les Harpacticus de Banyuls. Bulletin de la Societe Zoologique de France 51: 419 - 434." type="journal article" year="1926">Monard, 1926</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA736E4FFB2FFD06A0AC12BFAEDFD23" pageId="10" pageNumber="350">
<table id="F918C444FFB200256BDDC12BFA52FD23" box="[144,1413,314,655]" gridcols="4" gridrows="10" pageId="10" pageNumber="350">
<tr id="352834A6FFB200256BDDC12BFA52FEFC" box="[144,1413,314,336]" gridrow="0" pageId="10" pageNumber="350" rowspan-0="1">
<th id="76F95DDAFFB200256B90C12BFE20FEFC" box="[221,503,314,336]" gridcol="1" gridrow="0" pageId="10" pageNumber="350">Species</th>
<th id="76F95DDAFFB200256994C12BFC0AFEFC" box="[729,989,314,336]" gridcol="2" gridrow="0" pageId="10" pageNumber="350">Locality</th>
<th id="76F95DDAFFB200256F8CC12BFA52FEFC" box="[1217,1413,314,336]" gridcol="3" gridrow="0" pageId="10" pageNumber="350">Source</th>
</tr>
<tr id="352834A6FFB200256BDDC17FFA52FE29" box="[144,1413,366,389]" gridrow="1" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC17FFF76FE29" box="[144,161,366,389]" gridcol="0" gridrow="1" pageId="10" pageNumber="350">1.</th>
<td id="76F95DDAFFB200256B90C17FFE20FE29" box="[221,503,366,389]" gridcol="1" gridrow="1" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C17EFE17FE29" authority="Monard, 1926" authorityName="Monard" authorityYear="1926" box="[221,448,366,389]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="stebleri">
<emphasis id="B96CEAF6FFB2FFD06B90C17EFEEDFE2F" box="[221,314,366,388]" italics="true" pageId="10" pageNumber="350">B. stebleri</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A0DC17EFE17FE29" author="Monard, A." box="[320,448,367,389]" pageId="10" pageNumber="350" pagination="419 - 434" refId="ref4556" refString="Monard, A. 1926. Sur les Harpacticus de Banyuls. Bulletin de la Societe Zoologique de France 51: 419 - 434." type="journal article" year="1926">Monard, 1926</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C17FFC0AFE29" box="[729,989,366,389]" gridcol="2" gridrow="1" pageId="10" pageNumber="350">France</td>
<td id="76F95DDAFFB200256F8CC17FFA52FE29" box="[1217,1413,366,389]" gridcol="3" gridrow="1" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC17EFA9EFE29" author="Monard, A." box="[1217,1353,367,389]" pageId="10" pageNumber="350" pagination="419 - 434" refId="ref4556" refString="Monard, A. 1926. Sur les Harpacticus de Banyuls. Bulletin de la Societe Zoologique de France 51: 419 - 434." type="journal article" year="1926">Monard (1926)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC181FA52FE0A" box="[144,1413,400,422]" gridrow="2" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC181FF76FE0A" box="[144,161,400,422]" gridcol="0" gridrow="2" pageId="10" pageNumber="350">2.</th>
<td id="76F95DDAFFB200256B90C181FE20FE0A" box="[221,503,400,422]" gridcol="1" gridrow="2" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C181FE58FE0A" authority="Por, 1967" authorityName="Por" authorityYear="1967" box="[221,399,400,422]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="exigua">
<emphasis id="B96CEAF6FFB2FFD06B90C181FEE5FE09" box="[221,306,400,421]" italics="true" pageId="10" pageNumber="350">B. exigua</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A75C181FE58FE0A" author="Por, F. D." box="[312,399,400,422]" pageId="10" pageNumber="350" pagination="101 - 165" refId="ref4626" refString="Por, F. D. 1967. Level bottom Harpacticoida (Crustacea, Copepoda) from Elat (Red Sea), part I. Israel Journal of Zoology 16: 101 - 165." type="journal article" year="1967">Por, 1967</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C181FC0AFE0A" box="[729,989,400,422]" gridcol="2" gridrow="2" pageId="10" pageNumber="350">Gulf of Eilat in Israel</td>
<td id="76F95DDAFFB200256F8CC181FA52FE0A" box="[1217,1413,400,422]" gridcol="3" gridrow="2" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC181FAF6FE0A" author="Por, F. D." box="[1217,1313,400,422]" pageId="10" pageNumber="350" pagination="101 - 165" refId="ref4626" refString="Por, F. D. 1967. Level bottom Harpacticoida (Crustacea, Copepoda) from Elat (Red Sea), part I. Israel Journal of Zoology 16: 101 - 165." type="journal article" year="1967">Por (1967)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC1A0FA52FE6B" box="[144,1413,433,455]" gridrow="3" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC1A0FF76FE6B" box="[144,161,433,455]" gridcol="0" gridrow="3" pageId="10" pageNumber="350">3.</th>
<td id="76F95DDAFFB200256B90C1A0FE20FE6B" box="[221,503,433,455]" gridcol="1" gridrow="3" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C1A3FE05FE6B" authority="Bozic, 1968" authorityName="Bozic" authorityYear="1968" box="[221,466,433,455]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="curvirostris">
<emphasis id="B96CEAF6FFB2FFD06B90C1A3FE88FE6B" box="[221,351,434,455]" italics="true" pageId="10" pageNumber="350">B. curvirostris</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A28C1A0FE05FE6B" author="Bozic, B." box="[357,466,433,455]" pageId="10" pageNumber="350" pagination="570 - 573" refId="ref4264" refString="Bozic, B. 1968. Copepodes de La reunion III. Brianola curvirostris n. sp. Bulletin Du Museum National D'History Naturelle 40: 570 - 573." type="journal article" year="1968">Bozic, 1968</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C1A0FC0AFE6B" box="[729,989,433,455]" gridcol="2" gridrow="3" pageId="10" pageNumber="350">Mediterranean Sea</td>
<td id="76F95DDAFFB200256F8CC1A0FA52FE6B" box="[1217,1413,433,455]" gridcol="3" gridrow="3" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC1A0FAE1FE6B" author="Bozic, B." box="[1217,1334,433,455]" pageId="10" pageNumber="350" pagination="570 - 573" refId="ref4264" refString="Bozic, B. 1968. Copepodes de La reunion III. Brianola curvirostris n. sp. Bulletin Du Museum National D'History Naturelle 40: 570 - 573." type="journal article" year="1968">Bozic (1968)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC1C3FA52FE44" box="[144,1413,466,488]" gridrow="4" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC1C3FF76FE44" box="[144,161,466,488]" gridcol="0" gridrow="4" pageId="10" pageNumber="350">4.</th>
<td id="76F95DDAFFB200256B90C1C3FE20FE44" box="[221,503,466,488]" gridcol="1" gridrow="4" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C1C2FE1FFE44" authority="Hamond, 1973" authorityName="Hamond" authorityYear="1973" box="[221,456,466,488]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="B96CEAF6FFB2FFD06B90C1C2FEECFE4B" box="[221,315,466,488]" italics="true" pageId="10" pageNumber="350">B. elegans</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A0CC1C3FE1FFE44" author="Hamond, R." box="[321,456,466,488]" pageId="10" pageNumber="350" pagination="165 - 201" refId="ref4336" refString="Hamond, R. 1973. Four new copepods (Crustacea: Harpacticoida, Canuellidae) simultaneously occurring with Diogenes senex (Crustacea: Paguridae) near Sydney. Proceedings of the Linnean Society of New South Wales 97 (3): 165 - 201." type="journal article" year="1973">Hamond, 1973</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C1C3FC0AFE44" box="[729,989,466,488]" gridcol="2" gridrow="4" pageId="10" pageNumber="350">Metropolitan Sydney</td>
<td id="76F95DDAFFB200256F8CC1C3FA52FE44" box="[1217,1413,466,488]" gridcol="3" gridrow="4" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC1C3FA87FE44" author="Hamond, R." box="[1217,1360,466,488]" pageId="10" pageNumber="350" pagination="165 - 201" refId="ref4336" refString="Hamond, R. 1973. Four new copepods (Crustacea: Harpacticoida, Canuellidae) simultaneously occurring with Diogenes senex (Crustacea: Paguridae) near Sydney. Proceedings of the Linnean Society of New South Wales 97 (3): 165 - 201." type="journal article" year="1973">Hamond (1973)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC1E2FA52FDA6" box="[144,1413,499,522]" gridrow="5" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC1E2FF76FDA6" box="[144,161,499,522]" gridcol="0" gridrow="5" pageId="10" pageNumber="350">5.</th>
<td id="76F95DDAFFB200256B90C1E2FE20FDA6" box="[221,503,499,522]" gridcol="1" gridrow="5" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C1E5FE3AFDA5" authority="Hamond, 1973" authorityName="Hamond" authorityYear="1973" box="[221,493,499,522]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="sydneyensis">
<emphasis id="B96CEAF6FFB2FFD06B90C1E5FEB6FDA4" box="[221,353,499,521]" italics="true" pageId="10" pageNumber="350">B. sydneyensis</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A2AC1E5FE3AFDA5" author="Hamond, R." box="[359,493,500,522]" pageId="10" pageNumber="350" pagination="165 - 201" refId="ref4336" refString="Hamond, R. 1973. Four new copepods (Crustacea: Harpacticoida, Canuellidae) simultaneously occurring with Diogenes senex (Crustacea: Paguridae) near Sydney. Proceedings of the Linnean Society of New South Wales 97 (3): 165 - 201." type="journal article" year="1973">Hamond, 1973</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C1E2FC0AFDA6" box="[729,989,499,522]" gridcol="2" gridrow="5" pageId="10" pageNumber="350">Metropolitan Sydney</td>
<td id="76F95DDAFFB200256F8CC1E2FA52FDA6" box="[1217,1413,499,522]" gridcol="3" gridrow="5" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC1E5FA87FDA5" author="Hamond, R." box="[1217,1360,500,522]" pageId="10" pageNumber="350" pagination="165 - 201" refId="ref4336" refString="Hamond, R. 1973. Four new copepods (Crustacea: Harpacticoida, Canuellidae) simultaneously occurring with Diogenes senex (Crustacea: Paguridae) near Sydney. Proceedings of the Linnean Society of New South Wales 97 (3): 165 - 201." type="journal article" year="1973">Hamond (1973)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC204FA52FD87" box="[144,1413,533,555]" gridrow="6" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC204FF76FD87" box="[144,161,533,555]" gridcol="0" gridrow="6" pageId="10" pageNumber="350">6.</th>
<td id="76F95DDAFFB200256B90C204FE20FD87" box="[221,503,533,555]" gridcol="1" gridrow="6" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C204FE02FD86" authority="Fiers, 1982" authorityName="Fiers" authorityYear="1982" box="[221,469,533,555]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="vangoethemi">
<emphasis id="B96CEAF6FFB2FFD06B90C204FEBDFD86" box="[221,362,533,554]" italics="true" pageId="10" pageNumber="350">B. vangoethemi</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A3DC204FE02FD86" author="Fiers, F." box="[368,469,533,555]" pageId="10" pageNumber="350" pagination="1 - 32" refId="ref4294" refString="Fiers, F. 1982. New Canuellidae from the northern coast of Papua New Guinea (Copepoda: Harpacticoida). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie 54 (4): 1 - 32." type="journal article" year="1982">Fiers, 1982</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C204FC0AFD87" box="[729,989,533,555]" gridcol="2" gridrow="6" pageId="10" pageNumber="350">Papua New Guinea</td>
<td id="76F95DDAFFB200256F8CC204FA52FD87" box="[1217,1413,533,555]" gridcol="3" gridrow="6" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC204FAF9FD86" author="Fiers, F." box="[1217,1326,533,555]" pageId="10" pageNumber="350" pagination="1 - 32" refId="ref4294" refString="Fiers, F. 1982. New Canuellidae from the northern coast of Papua New Guinea (Copepoda: Harpacticoida). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Biologie 54 (4): 1 - 32." type="journal article" year="1982">Fiers (1982)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC227FA52FDE0" box="[144,1413,566,588]" gridrow="7" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC227FF76FDE0" box="[144,161,566,588]" gridcol="0" gridrow="7" pageId="10" pageNumber="350">7.</th>
<td id="76F95DDAFFB200256B90C227FE20FDE0" box="[221,503,566,588]" gridcol="1" gridrow="7" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C226FE20FDE7" authority="Wells &amp; Rao, 1987" authorityName="Wells &amp; Rao" authorityYear="1987" box="[221,503,566,588]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="hamondi">
<emphasis id="B96CEAF6FFB2FFD06B90C226FE92FDE7" box="[221,325,566,588]" italics="true" pageId="10" pageNumber="350">B. hamondi</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A07C227FE20FDE7" author="Wells, J. B. J. &amp; G. C. Rao" box="[330,503,566,588]" pageId="10" pageNumber="350" pagination="1 - 385" refId="ref4694" refString="Wells, J. B. J. and G. C. Rao. 1987. Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India 16 (4): 1 - 385." type="journal article" year="1987">Wells &amp; Rao, 1987</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C227FC0AFDE0" box="[729,989,566,588]" gridcol="2" gridrow="7" pageId="10" pageNumber="350">Andaman &amp; Nicobar Islands</td>
<td id="76F95DDAFFB200256F8CC227FA52FDE0" box="[1217,1413,566,588]" gridcol="3" gridrow="7" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC227FA52FDE7" author="Wells, J. B. J. &amp; G. C. Rao" box="[1217,1413,566,588]" pageId="10" pageNumber="350" pagination="1 - 385" refId="ref4694" refString="Wells, J. B. J. and G. C. Rao. 1987. Littoral Harpacticoida (Crustacea: Copepoda) from Andaman and Nicobar Islands. Memoirs of the Zoological Survey of India 16 (4): 1 - 385." type="journal article" year="1987">Wells and Rao (1987)</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC246FA52FDC1" box="[144,1413,599,621]" gridrow="8" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC246FF76FDC1" box="[144,161,599,621]" gridcol="0" gridrow="8" pageId="10" pageNumber="350">8.</th>
<td id="76F95DDAFFB200256B90C246FE20FDC1" box="[221,503,599,621]" gridcol="1" gridrow="8" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C249FE23FDC1" authority="Nazari et al., 2018" box="[221,500,599,621]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="haliensis">
<emphasis id="B96CEAF6FFB2FFD06B90C249FE90FDC0" box="[221,327,599,621]" italics="true" pageId="10" pageNumber="350">B. haliensis</emphasis>
<bibRefCitation id="EF894B15FFB2FFD06A00C246FE23FDC1" author="Nazari, F. &amp; O. Mirshami &amp; A. Sari &amp; M. Aliabadian &amp; P. Martinez Arbizu" box="[333,500,599,621]" pageId="10" pageNumber="350" pagination="401 - 441" refId="ref4581" refString="Nazari, F., O. Mirshami, A. Sari, M. Aliabadian and P. Martinez Arbizu. 2018. Three new Canuellidae (Copepoda: Canuelloida) from Iran. Zootaxa 4446 (4): 401 - 441." type="journal article" year="2018">
Nazari
<emphasis id="B96CEAF6FFB2FFD06AC2C249FE60FDC0" box="[399,439,599,621]" italics="true" pageId="10" pageNumber="350">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
</td>
<td id="76F95DDAFFB200256994C246FC0AFDC1" box="[729,989,599,621]" gridcol="2" gridrow="8" pageId="10" pageNumber="350">Nayband, Iran</td>
<td id="76F95DDAFFB200256F8CC246FA52FDC1" box="[1217,1413,599,621]" gridcol="3" gridrow="8" pageId="10" pageNumber="350">
<bibRefCitation id="EF894B15FFB2FFD06F8CC246FAA7FDC1" author="Nazari, F. &amp; O. Mirshami &amp; A. Sari &amp; M. Aliabadian &amp; P. Martinez Arbizu" box="[1217,1392,599,621]" pageId="10" pageNumber="350" pagination="401 - 441" refId="ref4581" refString="Nazari, F., O. Mirshami, A. Sari, M. Aliabadian and P. Martinez Arbizu. 2018. Three new Canuellidae (Copepoda: Canuelloida) from Iran. Zootaxa 4446 (4): 401 - 441." type="journal article" year="2018">
Nazari
<emphasis id="B96CEAF6FFB2FFD06E4EC249FAFCFDC0" box="[1283,1323,599,621]" italics="true" pageId="10" pageNumber="350">et al</emphasis>
. (2018)
</bibRefCitation>
</td>
</tr>
<tr id="352834A6FFB200256BDDC268FA52FD23" box="[144,1413,633,655]" gridrow="9" pageId="10" pageNumber="350">
<th id="76F95DDAFFB200256BDDC268FF76FD23" box="[144,161,633,655]" gridcol="0" gridrow="9" pageId="10" pageNumber="350">9</th>
<td id="76F95DDAFFB200256B90C268FE20FD23" box="[221,503,633,655]" gridcol="1" gridrow="9" pageId="10" pageNumber="350">
<taxonomicName id="4C184D67FFB2FFD06B90C268FE8FFD22" authorityName="Chullasorn &amp; Kangtia &amp; Klangsin &amp; Song" authorityYear="2024" box="[221,344,633,654]" class="Copepoda" family="Canuellidae" genus="Brianola" kingdom="Animalia" order="Polyarthra" pageId="10" pageNumber="350" phylum="Arthropoda" rank="species" species="rawaiensis" status="sp. nov.">
<emphasis id="B96CEAF6FFB2FFD06B90C268FE8FFD22" box="[221,344,633,654]" italics="true" pageId="10" pageNumber="350">B. rawaiensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAF6FFB2FFD06A13C268FE70FD22" bold="true" box="[350,423,633,654]" pageId="10" pageNumber="350">
<taxonomicNameLabel id="A25F578DFFB2FFD06A13C268FE70FD22" box="[350,423,633,654]" pageId="10" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</td>
<td id="76F95DDAFFB200256994C268FC0AFD23" box="[729,989,633,655]" gridcol="2" gridrow="9" pageId="10" pageNumber="350">Phuket, Thailand</td>
<td id="76F95DDAFFB200256F8CC268FA52FD23" box="[1217,1413,633,655]" gridcol="3" gridrow="9" pageId="10" pageNumber="350">Present study</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA736E4FFB2FFD06BEBC2F4FE9CFBEC" blockId="10.[144,749,741,1249]" pageId="10" pageNumber="350">
Antennule (
<figureCitation id="13232A61FFB2FFD06A53C2F4FEB8FD51" box="[286,367,741,765]" captionStart="Fig" captionStartId="9.[475,508,1635,1657]" captionTargetBox="[381,1303,262,1602]" captionTargetId="figure-14@9.[376,1306,257,1602]" captionTargetPageId="9" captionText="Fig. 9. Brianola rawaiensis sp. nov., male: (A) antennule (disarticulated)." figureDoi="http://doi.org/10.5281/zenodo.14659416" httpUri="https://zenodo.org/record/14659416/files/figure.png" pageId="10" pageNumber="350">Fig. 9A</figureCitation>
) five-segmented. First segment larg- est, with two smooth and one unipinnate seta, and three small spinules close to the base of the unipinnate seta. Second segment large, with three pinnate, four spinulose, and seven smooth setae/spines and two aesthetascs. Third segment small, with one spinulose and one small smooth seta. Fourth segment large, with two plumose, one spinulose, three pinnate, and one smooth seta and one modified element. Distal segment small, with one pinnate and five smooth setae. Armature formula: 1-(6), 2-(14 +2 ae), 3-(2), 4-(7), 5-(6).
</paragraph>
<paragraph id="8BA736E4FFB2FFD06BEBC458FE1EFB4D" blockId="10.[144,749,741,1249]" pageId="10" pageNumber="350">
Urosome (
<figureCitation id="13232A61FFB2FFD06A5EC459FE54FBCC" box="[275,387,1096,1120]" captionStart="Fig" captionStartId="8.[276,309,1374,1396]" captionTargetBox="[366,1159,257,1339]" captionTargetId="figure-253@8.[346,1212,257,1342]" captionTargetPageId="8" captionText="Fig. 8. Brianola rawaiensis sp. nov., male: (A) urosome, dorsal; (B) urosome, ventral;(C) genital area;(D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659414" httpUri="https://zenodo.org/record/14659414/files/figure.png" pageId="10" pageNumber="350">Fig. 8A, B</figureCitation>
) five-segmented and consisting of P5-bearing somite, genital somite, two urosomites, and anal somite. Genital area and P6 oval-shaped ornamented with spinules as
<figureCitation id="13232A61FFB2FFD06A72C4B8FE47FB6D" box="[319,400,1193,1217]" captionStart="Fig" captionStartId="8.[276,309,1374,1396]" captionTargetBox="[366,1159,257,1339]" captionTargetId="figure-253@8.[346,1212,257,1342]" captionTargetPageId="8" captionText="Fig. 8. Brianola rawaiensis sp. nov., male: (A) urosome, dorsal; (B) urosome, ventral;(C) genital area;(D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659414" httpUri="https://zenodo.org/record/14659414/files/figure.png" pageId="10" pageNumber="350">Fig. 8C</figureCitation>
. Spermatophores paired. P5 with four bipinnate setae (
<figureCitation id="13232A61FFB2FFD06A24C4D8FE6BFB4D" box="[361,444,1225,1249]" captionStart="Fig" captionStartId="8.[276,309,1374,1396]" captionTargetBox="[366,1159,257,1339]" captionTargetId="figure-253@8.[346,1212,257,1342]" captionTargetPageId="8" captionText="Fig. 8. Brianola rawaiensis sp. nov., male: (A) urosome, dorsal; (B) urosome, ventral;(C) genital area;(D) P5." figureDoi="http://doi.org/10.5281/zenodo.14659414" httpUri="https://zenodo.org/record/14659414/files/figure.png" pageId="10" pageNumber="350">Fig. 8D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>