245 lines
31 KiB
XML
245 lines
31 KiB
XML
<document id="A4C88A5C6E80B78EC7538484A2FA9F60" ID-DOI="10.5281/zenodo.7909894" ID-ISSN="2305-2562" ID-Zenodo-Dep="7909894" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683577476113" checkinUser="felipe" docAuthor="Bursey, M. L. & Herbert, D. G." docDate="2004" docId="03B187DCFFFBFFCEFD8BFA8F0FC54629" docLanguage="en" docName="AfrInvertebr.45.1.249–262.pdf" docOrigin="African Invertebrates 45" docStyle="DocumentStyle:C20B10C002986E51083406A4EACF003A.3:AfrInvertebr.2001-2007.journal_article" docStyleId="C20B10C002986E51083406A4EACF003A" docStyleName="AfrInvertebr.2001-2007.journal_article" docStyleVersion="3" docTitle="Gulella hamerae Bursey & Herbert 2004, sp. n." docType="treatment" docVersion="6" lastPageNumber="252" masterDocId="FF88FFA4FFFAFFCDFF9AFFD90C4F434E" masterDocTitle="Four new narrow-range endemic species of Gulella from Eastern Cape, South Africa (Mollusca: Pulmonata: Streptaxidae)" masterLastPageNumber="262" masterPageNumber="249" pageNumber="250" updateTime="1699498715153" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:mods id="C2B954979150818B221D998F64583961" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="781BCEADDE0FF52FF156AFE1A94D8015">
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<mods:title id="79500B87E52B1E1C66C0CAA31574D328">Four new narrow-range endemic species of Gulella from Eastern Cape, South Africa (Mollusca: Pulmonata: Streptaxidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="0FDC5C68BB5DC376E7472DC2C52A7577" type="personal">
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<mods:role id="86401B12BAE84B12A1E85EAD4E53B400">
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<mods:roleTerm id="8ED2FB177C2EBE8C0176F1A042689722">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E9E43E7698A9BCE9FF9FE742A405A97C">Bursey, M. L.</mods:namePart>
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</mods:name>
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<mods:name id="15B3D2CD5B990EB009D45B7C167BBEED" type="personal">
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<mods:role id="E0E1EDCEDBF3A5A3DAFAABD629FEAD85">
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<mods:roleTerm id="4E8038BF1B730C8B66FE08CC0B70996A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="58EDFC2EAECFAEB0B7B79AD20AFCCC0B">Herbert, D. G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C6FF06272370E9FA64AE0104E0E90E11">text</mods:typeOfResource>
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<mods:titleInfo id="2B69B67FF70D6453BA914AA7EC215429">
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<mods:title id="53CDE09F41ABF612B3885E9A9D25F13B">African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part id="B22ACA113B44C510B27D35F8FFF8495E">
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<mods:date id="83B12C4A48547555C46E4808712CEA3B">2004</mods:date>
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<mods:detail id="49D029DAC4F9B5FF8F14EA39A5F9DF0C" type="pubDate">
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<mods:number id="DEDD09333706F0577FD6D2E1AA454240">2004-12-31</mods:number>
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</mods:detail>
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<mods:detail id="5A1B89BF6932ADD613BE027DF16DB345" type="volume">
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<mods:number id="B6F0E8F1FDD195406643A89D1F952937">45</mods:number>
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</mods:detail>
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<mods:extent id="2931061A8814FA474F154710756C2A7D" unit="page">
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<mods:start id="86E9552582B93E57794F6AE8548CA8A4">249</mods:start>
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<mods:end id="35F779968B9FAC8108A7D07526F89EFB">262</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="9A6C745E23B70D9E9386AC4B105E19C1">journal article</mods:classification>
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<mods:identifier id="18894865D332ED29732920F334157AED" type="DOI">10.5281/zenodo.7909894</mods:identifier>
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<mods:identifier id="5BB5DADEF9FB6A64AD279DE10704A8D4" type="ISSN">2305-2562</mods:identifier>
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<mods:identifier id="B9E19DC6C4CF7CC4040701BC18EB5240" type="Zenodo-Dep">7909894</mods:identifier>
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<treatment id="03B187DCFFFBFFCEFD8BFA8F0FC54629" ID-DOI="http://doi.org/10.5281/zenodo.7910318" ID-Zenodo-Dep="7910318" LSID="urn:lsid:plazi:treatment:03B187DCFFFBFFCEFD8BFA8F0FC54629" httpUri="http://treatment.plazi.org/id/03B187DCFFFBFFCEFD8BFA8F0FC54629" lastPageId="3" lastPageNumber="252" pageId="1" pageNumber="250">
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<subSubSection id="C3026541FFFBFFCCFD8BFA8F0F554621" box="[529,794,1366,1391]" pageId="1" pageNumber="250" type="nomenclature">
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<paragraph id="8BA736CAFFFBFFCCFD8BFA8F0F554621" blockId="1.[181,1142,1292,1671]" box="[529,794,1366,1391]" pageId="1" pageNumber="250">
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<heading id="D0EF81A6FFFBFFCCFD8BFA8F0F554621" box="[529,794,1366,1391]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="250" reason="2">
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<taxonomicName id="4C184D49FFFBFFCCFD8BFA8F0E984621" ID-CoL="3HLRM" authority="Bursey & Herbert, 2004" authorityName="Bursey & Herbert" authorityYear="2004" box="[529,727,1366,1391]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="1" pageNumber="250" phylum="Mollusca" rank="species" species="hamerae" status="sp. nov.">
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<emphasis id="B96CEAD8FFFBFFCCFD8BFA8F0E984621" bold="true" box="[529,727,1366,1391]" pageId="1" pageNumber="250">Gulella hamerae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57A3FFFBFFCCFD44FA8F0F554621" box="[734,794,1366,1391]" pageId="1" pageNumber="250" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026541FFFBFFCEFDCAFAA70FC54629" lastPageId="3" lastPageNumber="252" pageId="1" pageNumber="250" type="description">
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<paragraph id="8BA736CAFFFBFFCCFDCAFAA70E9446D9" blockId="1.[181,1142,1292,1671]" box="[592,731,1406,1431]" pageId="1" pageNumber="250">
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<figureCitation id="13232A4FFFFBFFCCFDCAFAA70EFC46D9" box="[592,691,1406,1431]" captionStart="Figs 1–5" captionStartId="2.[143,182,1171,1191]" captionTargetBox="[140,1105,201,1149]" captionTargetId="figure-148@2.[127,1116,190,1165]" captionTargetPageId="2" captionText="Figs 1–5. Gulella hamerae sp. n., holotype. 1, 2, apertural and lateral views of shell, length = 2.7 mm; 3, oblique view into aperture, bar = 0.25 mm; 4, detail of sculpture (start of last whorl), bar = 100 µm; 5, microsculpture on embryonic shell, bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7917361" httpUri="https://zenodo.org/record/7917361/files/figure.png" pageId="1" pageNumber="250">Figs 1–5</figureCitation>
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,
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<figureCitation id="13232A4FFFFBFFCCFD25FAA70E9446D9" box="[703,731,1406,1431]" captionStart="Fig" captionStartId="10.[280,313,1647,1667]" captionTargetBox="[184,1064,733,1622]" captionTargetId="figure-237@10.[185,1062,734,1621]" captionTargetPageId="10" captionText="Fig. 24. Map of northern Eastern Cape showing distribution of new species." figureDoi="http://doi.org/10.5281/zenodo.7917365" httpUri="https://zenodo.org/record/7917365/files/figure.png" pageId="1" pageNumber="250">24</figureCitation>
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</paragraph>
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<paragraph id="8BA736CAFFFBFFCCFF2FFA7F0D0146B1" blockId="1.[181,1142,1292,1671]" pageId="1" pageNumber="250">
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Etymology: Named for Dr Michelle Hamer, University of
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<collectingRegion id="49DCF828FFFBFFCCFCCDFA7F084546F1" box="[855,1034,1446,1471]" country="South Africa" name="KwaZulu-Natal" pageId="1" pageNumber="250">KwaZulu-Natal</collectingRegion>
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, who has devoted much time and effort to the study and conservation of southern African invertebrates.
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</paragraph>
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<paragraph id="8BA736CAFFFBFFCFFF2FF9D70D2E4670" blockId="1.[181,1142,1292,1671]" lastBlockId="2.[144,1104,1285,1670]" lastPageId="2" lastPageNumber="251" pageId="1" pageNumber="250">
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||
Diagnosis: Shell minute, oval-cylindrical; sculptured with strong axial ribs; peristome entire, but fused to base of preceding whorl in parietal region; aperture y-shaped, columella strongly indented; dentition five-fold: a parietal lamella, two labral denticles, a basal denticle and a large curved superficial tooth on indented columella; embryonic shell sculptured with close-set, spiral lirae; umbilicus open, relatively wide (width
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<quantity id="4CE09B2FFFF8FFCFFB85FADC0CA84670" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="251" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4– 1.6 mm</quantity>
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) and deep.
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</paragraph>
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<caption id="DF676642FFF8FFCFFF15FB4A0F1D4799" ID-DOI="http://doi.org/10.5281/zenodo.7917361" ID-Zenodo-Dep="7917361" httpUri="https://zenodo.org/record/7917361/files/figure.png" pageId="2" pageNumber="251" startId="2.[143,182,1171,1191]" targetBox="[140,1105,201,1149]" targetPageId="2" targetType="figure">
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<paragraph id="8BA736CAFFF8FFCFFF15FB4A0F1D4799" blockId="2.[143,1103,1171,1239]" pageId="2" pageNumber="251">
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Figs 1–5.
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<taxonomicName id="4C184D49FFF8FFCFFF77FB4A0DC847E9" authority="Bursey & Herbert, 2004" authorityName="Bursey & Herbert" authorityYear="2004" box="[237,391,1171,1191]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="251" phylum="Mollusca" rank="species" species="hamerae" status="sp. nov.">
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<emphasis id="B96CEAD8FFF8FFCFFF77FB4A0DC847E9" box="[237,391,1171,1191]" italics="true" pageId="2" pageNumber="251">Gulella hamerae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57A3FFF8FFCFFE15FB4A0D8C47E9" box="[399,451,1171,1191]" pageId="2" pageNumber="251" rank="species">sp. n.</taxonomicNameLabel>
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, holotype. 1, 2, apertural and lateral views of shell, length = 2.7 mm; 3, oblique view into aperture, bar = 0.25 mm; 4, detail of sculpture (start of last whorl), bar = 100 µm; 5, microsculpture on embryonic shell, bar = 100 µm.
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</paragraph>
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||
</caption>
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||
<paragraph id="8BA736CAFFF8FFCEFF0AFA940E874109" blockId="2.[144,1104,1285,1670]" lastBlockId="3.[181,1142,206,583]" lastPageId="3" lastPageNumber="252" pageId="2" pageNumber="251">
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Description: Shell minute, oval-cylindrical, length
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<quantity id="4CE09B2FFFF8FFCFFD48FA940F134628" box="[722,860,1357,1382]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.1" metricValueMin="2.7" pageId="2" pageNumber="251" unit="mm" value="2.9" valueMax="3.1" valueMin="2.7">2.7–3.1 mm</quantity>
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, width
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<quantity id="4CE09B2FFFF8FFCFFC28FA9408044628" box="[946,1099,1357,1382]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="2" pageNumber="251" unit="mm" value="1.575" valueMax="1.65" valueMin="1.5">1.5–1.65 mm</quantity>
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; length:width 1.70–1.96. Embryonic shell approx.
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<quantity id="4CE09B2FFFF8FFCFFD54FAB40F7A46C8" box="[718,821,1389,1414]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="251" unit="mm" value="0.88">0.88 mm</quantity>
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in diameter, comprising approx. 2.25 whorls; sculptured more or less throughout with close-set, microscopic raised spiral threads (
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<figureCitation id="13232A4FFFF8FFCFFEEEFA740DFA4688" box="[372,437,1453,1478]" captionStart="Figs 1–5" captionStartId="2.[143,182,1171,1191]" captionTargetBox="[140,1105,201,1149]" captionTargetId="figure-148@2.[127,1116,190,1165]" captionTargetPageId="2" captionText="Figs 1–5. Gulella hamerae sp. n., holotype. 1, 2, apertural and lateral views of shell, length = 2.7 mm; 3, oblique view into aperture, bar = 0.25 mm; 4, detail of sculpture (start of last whorl), bar = 100 µm; 5, microsculpture on embryonic shell, bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7917361" httpUri="https://zenodo.org/record/7917361/files/figure.png" pageId="2" pageNumber="251">Fig. 5</figureCitation>
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||
), last half whorl also with fine, close-set axial riblets radiating from suture; junction between embryonic shell and teleoconch distinct. Teleoconch comprising approx. 4.5 whorls; first whorl convex, but subsequent ones rather more flat-sided; suture narrowly indented; sculpture of strong, sigmoid axial ribs, extending from suture to suture (47–49 on penultimate whorl); prosocline on spire whorls and almost orthocline on last whorl; rib intervals lacking obvious microsculpture (
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<figureCitation id="13232A4FFFF8FFCFFB99F99408084528" box="[1027,1095,1613,1638]" captionStart="Figs 1–5" captionStartId="2.[143,182,1171,1191]" captionTargetBox="[140,1105,201,1149]" captionTargetId="figure-148@2.[127,1116,190,1165]" captionTargetPageId="2" captionText="Figs 1–5. Gulella hamerae sp. n., holotype. 1, 2, apertural and lateral views of shell, length = 2.7 mm; 3, oblique view into aperture, bar = 0.25 mm; 4, detail of sculpture (start of last whorl), bar = 100 µm; 5, microsculpture on embryonic shell, bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7917361" httpUri="https://zenodo.org/record/7917361/files/figure.png" pageId="2" pageNumber="251">Fig. 4</figureCitation>
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) and with only occasional traces of spiral threads on sides of ribs. Peristome entire and fused with base of penultimate whorl in parietal region, flaring outward elsewhere. Aperture constricted, y-shaped with lower limb curving toward shell axis, columella strongly indented; apertural dentition five-fold (
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<figureCitation id="13232A4FFFF9FFCEFD40FED70F6F4269" box="[730,800,270,295]" captionStart="Figs 1–5" captionStartId="2.[143,182,1171,1191]" captionTargetBox="[140,1105,201,1149]" captionTargetId="figure-148@2.[127,1116,190,1165]" captionTargetPageId="2" captionText="Figs 1–5. Gulella hamerae sp. n., holotype. 1, 2, apertural and lateral views of shell, length = 2.7 mm; 3, oblique view into aperture, bar = 0.25 mm; 4, detail of sculpture (start of last whorl), bar = 100 µm; 5, microsculpture on embryonic shell, bar = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7917361" httpUri="https://zenodo.org/record/7917361/files/figure.png" pageId="3" pageNumber="252">Fig. 3</figureCitation>
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): 1) a strong parietal lamella, outer portion oblique and then curving inward so that remainder runs into the aperture more or less at right angles, a distinct ridge-like swelling at point of curvature; 2-3) two broad labral teeth, upper one flat-topped and level with apertural margin, lower one flat-topped or somewhat rounded and inset well behind upper one; 4) a deep-set, small, transverse basal tooth; 5) a large curved columella tooth level with aperture margin, but angled inward basally; a lamella on inner part of columella is not evident. Labral complex corresponds with an external pit behind flaring outer lip. Umbilicus relatively wide (width
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<quantity id="4CE09B2FFFF9FFCEFE9EFDD70DC44169" box="[260,395,526,551]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="3" pageNumber="252" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
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) and deep, with a conspicuous pit underlying columella tooth. Shell translucent, uniformly milky-white when fresh.
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</paragraph>
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<paragraph id="8BA736CAFFF9FFCEFF2FFD80083941DC" blockId="3.[181,1142,601,882]" pageId="3" pageNumber="252">
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Type material:
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<materialsCitation id="3B703C97FFF9FFCEFEC1FD80083941DC" collectionCode="NMSA" collectorName="H. J. Puzey" country="South Africa" location="Umtata" municipality="Nqadu Forest" pageId="3" pageNumber="252" specimenCode="W1217" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
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||
<typeStatus id="54A38868FFF9FFCEFEC1FD800D87413C" box="[347,456,601,626]" pageId="3" pageNumber="252" type="holotype">Holotype</typeStatus>
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||
:
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||
<collectionCode id="ED09AE0FFFF9FFCEFE4BFD800E6F413C" box="[465,544,601,626]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="252" type="Museum">NMSA</collectionCode>
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||
<specimenCode id="DBBE9EB1FFF9FFCEFDBFFD800E36413C" box="[549,633,601,626]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="252" type="Museum">W1217</specimenCode>
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||
/T1999, length
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<quantity id="4CE09B2FFFF9FFCEFCBAFD800F38413C" box="[800,887,601,626]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="252" unit="mm" value="2.7">2.7 mm</quantity>
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||
, width
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||
<quantity id="4CE09B2FFFF9FFCEFC5CFD800864413C" box="[966,1067,601,626]" metricMagnitude="-3" metricUnit="m" metricValue="1.52" pageId="3" pageNumber="252" unit="mm" value="1.52">1.52 mm</quantity>
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||
.
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||
<collectingCountry id="F30F765AFFF9FFCEFBAEFD800CB141DC" name="South Africa" pageId="3" pageNumber="252">South Africa</collectingCountry>
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||
,
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||
<collectingRegion id="49DCF828FFF9FFCEFE9DFDA00DD641DC" box="[263,409,633,658]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="252">Eastern Cape</collectingRegion>
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||
, Transkei,
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||
<collectingMunicipality id="6BC3ACB0FFF9FFCEFD96FDA00EEE41DC" box="[524,673,633,658]" pageId="3" pageNumber="252">Nqadu Forest</collectingMunicipality>
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||
, north of
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||
<location id="8EC76011FFF9FFCEFC9EFDA00F1741DC" LSID="urn:lsid:plazi:treatment:03B187DCFFFBFFCEFD8BFA8F0FC54629:8EC76011FFF9FFCEFC9EFDA00F1741DC" box="[772,856,633,658]" country="South Africa" municipality="Nqadu Forest" name="Umtata" pageId="3" pageNumber="252" stateProvince="Eastern Cape">Umtata</location>
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||
, leg.
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||
<collectorName id="26ED531CFFF9FFCEFC17FDA0084E41DC" box="[909,1025,633,658]" pageId="3" pageNumber="252">H.J. Puzey</collectorName>
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||
(undated).
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||
</materialsCitation>
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||
</paragraph>
|
||
<paragraph id="8BA736CAFFF9FFCEFF4AFD400F0B419C" blockId="3.[181,1142,601,882]" pageId="3" pageNumber="252">
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||
<materialsCitation id="3B703C97FFF9FFCEFF4AFD400F0F419C" collectingDate="2001-05-12" collectionCode="NMSA" collectorName="D. Herbert" county="Transkei" latitude="-31.42775" location="Umtata" longLatPrecision="1" longitude="28.751966" municipality="Nqadu Forest" pageId="3" pageNumber="252" specimenCode="V9127" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
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||
<typeStatus id="54A38868FFF9FFCEFF4AFD400D7F41FC" box="[208,304,665,690]" pageId="3" pageNumber="252" type="paratype">Paratype</typeStatus>
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||
1,
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<collectionCode id="ED09AE0FFFF9FFCEFED4FD400DD341FC" box="[334,412,665,690]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="252" type="Museum">NMSA</collectionCode>
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||
<specimenCode id="DBBE9EB1FFF9FFCEFE3AFD400DA141FC" box="[416,494,665,690]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="252" type="Museum">V9127</specimenCode>
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||
/T1998,
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||
<collectingRegion id="49DCF828FFF9FFCEFDDCFD400E9741FC" box="[582,728,665,690]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="252">Eastern Cape</collectingRegion>
|
||
,
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||
<collectingCounty id="62C64E46FFF9FFCEFD7AFD400F0D41FC" box="[736,834,665,690]" pageId="3" pageNumber="252">Transkei</collectingCounty>
|
||
,
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||
<collectingMunicipality id="6BC3ACB0FFF9FFCEFCD0FD400F9241FC" box="[842,989,665,690]" pageId="3" pageNumber="252">Nqadu Forest</collectingMunicipality>
|
||
(
|
||
<geoCoordinate id="EE2C500DFFF9FFCEFC70FD40083D41FC" box="[1002,1138,665,690]" degrees="31" direction="south" minutes="25.665" orientation="latitude" pageId="3" pageNumber="252" precision="1" value="-31.42775">31º25.665'S</geoCoordinate>
|
||
:
|
||
<geoCoordinate id="EE2C500DFFF9FFCEFF2FFD600D0E419C" box="[181,321,697,722]" degrees="28" direction="east" minutes="45.118" orientation="longitude" pageId="3" pageNumber="252" precision="1" value="28.751966">28º45.118'E</geoCoordinate>
|
||
), north of
|
||
<location id="8EC76011FFF9FFCEFE2EFD600E46419C" LSID="urn:lsid:plazi:treatment:03B187DCFFFBFFCEFD8BFA8F0FC54629:8EC76011FFF9FFCEFE2EFD600E46419C" box="[436,521,697,722]" county="Transkei" latitude="-31.42775" longLatPrecision="1" longitude="28.751966" municipality="Nqadu Forest" name="Umtata" pageId="3" pageNumber="252" stateProvince="Eastern Cape">Umtata</location>
|
||
, leg.
|
||
<collectorName id="26ED531CFFF9FFCEFDDFFD600EF0419C" box="[581,703,697,722]" pageId="3" pageNumber="252">D. Herbert</collectorName>
|
||
,
|
||
<date id="FFA6100AFFF9FFCEFD50FD600F0F419C" box="[714,832,697,722]" pageId="3" pageNumber="252" value="2001-05-12">
|
||
<collectingDate id="EFE2E9E2FFF9FFCEFD50FD600F0F419C" box="[714,832,697,722]" pageId="3" pageNumber="252" value="2001-05-12">12/v/2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA736CAFFF9FFCEFF4AFD000DC7407C" blockId="3.[181,1142,601,882]" pageId="3" pageNumber="252">
|
||
<materialsCitation id="3B703C97FFF9FFCEFF4AFD00083941BC" box="[208,1142,729,754]" collectionCode="NMSA, NMW" pageId="3" pageNumber="252" specimenCode="W223" specimenCount="2" typeStatus="paratype">
|
||
<typeStatus id="54A38868FFF9FFCEFF4AFD000D7041BC" box="[208,319,729,754]" pageId="3" pageNumber="252" type="paratype">Paratypes</typeStatus>
|
||
2, 3,
|
||
<collectionCode id="ED09AE0FFFF9FFCEFE11FD000D9441BC" box="[395,475,729,754]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="252" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBBE9EB1FFF9FFCEFE7FFD000E6441BC" box="[485,555,729,754]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="3" pageNumber="252" type="Museum">W223</specimenCode>
|
||
/T2009,
|
||
<typeStatus id="54A38868FFF9FFCEFD15FD000EBE41BC" box="[655,753,729,754]" pageId="3" pageNumber="252" type="paratype">paratype</typeStatus>
|
||
4,
|
||
<collectionCode id="ED09AE0FFFF9FFCEFC87FD000F2A41BC" box="[797,869,729,754]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="252">NMW</collectionCode>
|
||
Z.2004.030.00001 and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B703C97FFF9FFCEFF2FFD200DCB407C" collectionCode="ELM" collectorName="W. G. Rump" location="Umtata" municipality="Nqadu Forest" pageId="3" pageNumber="252" specimenCode="ELM 13661" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
|
||
<typeStatus id="54A38868FFF9FFCEFF2FFD200D5A405C" box="[181,277,761,786]" pageId="3" pageNumber="252" type="paratype">paratype</typeStatus>
|
||
5,
|
||
<specimenCode id="DBBE9EB1FFF9FFCEFEA0FD200D8A405C" box="[314,453,761,786]" collectionCode="ELM" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="3" pageNumber="252">ELM 13661</specimenCode>
|
||
,
|
||
<collectingRegion id="49DCF828FFF9FFCEFE4BFD200E27405C" box="[465,616,761,786]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="252">Eastern Cape</collectingRegion>
|
||
, Transkei,
|
||
<collectingMunicipality id="6BC3ACB0FFF9FFCEFD79FD200F30405C" box="[739,895,761,786]" pageId="3" pageNumber="252">Nqadu Forest</collectingMunicipality>
|
||
, north of
|
||
<location id="8EC76011FFF9FFCEFC71FD20080F405C" LSID="urn:lsid:plazi:treatment:03B187DCFFFBFFCEFD8BFA8F0FC54629:8EC76011FFF9FFCEFC71FD20080F405C" box="[1003,1088,761,786]" municipality="Nqadu Forest" name="Umtata" pageId="3" pageNumber="252" stateProvince="Eastern Cape">Umtata</location>
|
||
, leg.
|
||
<collectorName id="26ED531CFFF9FFCEFF2FFCC00D71407C" box="[181,318,793,818]" pageId="3" pageNumber="252">W.G. Rump</collectorName>
|
||
, 1935
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA736CAFFF9FFCEFF4AFCE00811403C" blockId="3.[181,1142,601,882]" pageId="3" pageNumber="252">
|
||
<materialsCitation id="3B703C97FFF9FFCEFF4AFCE00815403C" collectionCode="RMNH" collectorName="W. G. Rump & W. Falcon & H. Boswell & A. C. van Bruggen" location="Umtata" municipality="Nqadu Forest" pageId="3" pageNumber="252" specimenCode="RMNH 98152" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
|
||
<typeStatus id="54A38868FFF9FFCEFF4AFCE00D73401C" box="[208,316,825,850]" pageId="3" pageNumber="252" type="paratype">Paratypes</typeStatus>
|
||
6,7,
|
||
<specimenCode id="DBBE9EB1FFF9FFCEFEF5FCE00E5F401C" box="[367,528,825,850]" collectionCode="RMNH" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="3" pageNumber="252" type="Museum">RMNH 98152</specimenCode>
|
||
,
|
||
<collectingRegion id="49DCF828FFF9FFCEFD83FCE00EE2401C" box="[537,685,825,850]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="252">Eastern Cape</collectingRegion>
|
||
, Transkei,
|
||
<collectingMunicipality id="6BC3ACB0FFF9FFCEFCBBFCE00FF7401C" box="[801,952,825,850]" pageId="3" pageNumber="252">Nqadu Forest</collectingMunicipality>
|
||
, north of
|
||
<location id="8EC76011FFF9FFCEFB87FCE0083D401C" LSID="urn:lsid:plazi:treatment:03B187DCFFFBFFCEFD8BFA8F0FC54629:8EC76011FFF9FFCEFB87FCE0083D401C" box="[1053,1138,825,850]" municipality="Nqadu Forest" name="Umtata" pageId="3" pageNumber="252" stateProvince="Eastern Cape">Umtata</location>
|
||
, leg.
|
||
<collectorName id="26ED531CFFF9FFCEFF7FFC800D22403C" box="[229,365,857,882]" pageId="3" pageNumber="252">W.G. Rump</collectorName>
|
||
, 1935,
|
||
<emphasis id="B96CEAD8FFF9FFCEFE25FC800D97403F" box="[447,472,857,881]" italics="true" pageId="3" pageNumber="252">ex</emphasis>
|
||
coll’n
|
||
<collectorName id="26ED531CFFF9FFCEFDBCFC800ED4403C" box="[550,667,857,882]" pageId="3" pageNumber="252">W. Falcon</collectorName>
|
||
, don.
|
||
<collectorName id="26ED531CFFF9FFCEFD44FC800F12403C" box="[734,861,857,882]" pageId="3" pageNumber="252">H. Boswell</collectorName>
|
||
via
|
||
<collectorName id="26ED531CFFF9FFCEFC11FC800815403C" box="[907,1114,857,882]" pageId="3" pageNumber="252">A.C. van Bruggen</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA736CAFFF9FFCEFF2FFC5A0F4B40F2" blockId="3.[181,1141,899,956]" pageId="3" pageNumber="252">
|
||
Distribution and habitat (
|
||
<figureCitation id="13232A4FFFF9FFCEFE49FC5A0E6740D2" box="[467,552,899,924]" captionStart="Fig" captionStartId="10.[280,313,1647,1667]" captionTargetBox="[184,1064,733,1622]" captionTargetId="figure-237@10.[185,1062,734,1621]" captionTargetPageId="10" captionText="Fig. 24. Map of northern Eastern Cape showing distribution of new species." figureDoi="http://doi.org/10.5281/zenodo.7917365" httpUri="https://zenodo.org/record/7917365/files/figure.png" pageId="3" pageNumber="252">Fig. 24</figureCitation>
|
||
): Known only from Nqadu Forest, north of Umtata,
|
||
<collectingRegion id="49DCF828FFF9FFCEFF2FFC7A0D0240F2" box="[181,333,931,956]" country="South Africa" name="Eastern Cape" pageId="3" pageNumber="252">Eastern Cape</collectingRegion>
|
||
; Transkei mist-belt forest, in leaf-litter.
|
||
</paragraph>
|
||
<paragraph id="8BA736CAFFF9FFCEFF2FFC170FC54629" blockId="3.[181,1142,974,1383]" pageId="3" pageNumber="252">
|
||
Remarks:
|
||
<taxonomicName id="4C184D49FFF9FFCEFEB3FC170DE140A8" authorityName="Bursey & Herbert" authorityYear="2004" box="[297,430,974,998]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="252" phylum="Mollusca" rank="species" species="hamerae">
|
||
<emphasis id="B96CEAD8FFF9FFCEFEB3FC170DE140A8" box="[297,430,974,998]" italics="true" pageId="3" pageNumber="252">G. hamerae</emphasis>
|
||
</taxonomicName>
|
||
is evidently related to
|
||
<taxonomicName id="4C184D49FFF9FFCEFD29FC17084B40A9" authority="Connolly, 1939" authorityName="Connolly" authorityYear="1939" box="[691,1028,974,999]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="252" phylum="Mollusca" rank="species" species="claustralis">
|
||
<emphasis id="B96CEAD8FFF9FFCEFD29FC170F0340A8" box="[691,844,974,998]" italics="true" pageId="3" pageNumber="252">G. claustralis</emphasis>
|
||
Connolly, 1939
|
||
</taxonomicName>
|
||
, a species known only from two mist-belt forest localities in northern Pondoland and southern
|
||
<collectingRegion id="49DCF828FFF9FFCEFF2FFBD70D2D4769" box="[181,354,1038,1063]" country="South Africa" name="KwaZulu-Natal" pageId="3" pageNumber="252">KwaZulu-Natal</collectingRegion>
|
||
(
|
||
<bibRefCitation id="EF894B3BFFF9FFCEFEF4FBD70E354769" author="HERBERT, D. G. & KILBURN, R. N." box="[366,634,1038,1063]" pageId="3" pageNumber="252" refId="ref6374" refString="HERBERT, D. G. & KILBURN, R. N. 2004. Field guide to the land snails and slugs of eastern South Africa. Pietermaritzburg: Natal Museum." type="book" year="2004">Herbert & Kilburn 2004</bibRefCitation>
|
||
). The strong axial sculpture, y-shaped aperture, entire peristome, indented columella lip, and spirally striate embryonic shell are evidence of this. It differs, however, in the form of the labral teeth (lower cusp absent) and in its much more strongly indented columella, the inner margin of which possesses a single strong, smoothly curved tooth, instead of two distinct teeth. In terms of its columella morphology,
|
||
<taxonomicName id="4C184D49FFF9FFCEFED3FB770D834788" authorityName="Bursey & Herbert" authorityYear="2004" box="[329,460,1198,1222]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="252" phylum="Mollusca" rank="species" species="hamerae">
|
||
<emphasis id="B96CEAD8FFF9FFCEFED3FB770D834788" box="[329,460,1198,1222]" italics="true" pageId="3" pageNumber="252">G. hamerae</emphasis>
|
||
</taxonomicName>
|
||
somewhat resembles
|
||
<taxonomicName id="4C184D49FFF9FFCEFD27FB7708414789" authority="Bruggen, 1972" authorityName="Bruggen" authorityYear="1972" box="[701,1038,1198,1223]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="252" phylum="Mollusca" rank="species" species="incurvidens">
|
||
<emphasis id="B96CEAD8FFF9FFCEFD27FB770F2E4788" box="[701,865,1198,1222]" italics="true" pageId="3" pageNumber="252">G. incurvidens</emphasis>
|
||
Bruggen, 1972
|
||
</taxonomicName>
|
||
, from the northern Drakensberg and Soutpansberg (
|
||
<collectingRegion id="49DCF828FFF9FFCEFD17FB170F6C47A9" box="[653,803,1230,1255]" country="South Africa" name="Mpumalanga" pageId="3" pageNumber="252">Mpumalanga</collectingRegion>
|
||
and
|
||
<collectingRegion id="49DCF828FFF9FFCEFCC7FB170F8B47A9" box="[861,964,1230,1255]" country="South Africa" name="Limpopo" pageId="3" pageNumber="252">Limpopo</collectingRegion>
|
||
provinces), but that species evidently lacks spiral sculpture on the embryonic shell, has a single in-running labral tooth and the basal part of its aperture is twisted obliquely upward. It seems probable that
|
||
<taxonomicName id="4C184D49FFF9FFCEFE0DFAF70E554608" authorityName="Bursey & Herbert" authorityYear="2004" box="[407,538,1326,1350]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="252" phylum="Mollusca" rank="species" species="hamerae">
|
||
<emphasis id="B96CEAD8FFF9FFCEFE0DFAF70E554608" box="[407,538,1326,1350]" italics="true" pageId="3" pageNumber="252">G. hamerae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D49FFF9FFCEFDD4FAF70EAA4608" authorityName="Connolly" authorityYear="1939" box="[590,741,1326,1350]" class="Gastropoda" family="Streptaxidae" genus="Gulella" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="252" phylum="Mollusca" rank="species" species="claustralis">
|
||
<emphasis id="B96CEAD8FFF9FFCEFDD4FAF70EAA4608" box="[590,741,1326,1350]" italics="true" pageId="3" pageNumber="252">G. claustralis</emphasis>
|
||
</taxonomicName>
|
||
are sister taxa, having evolved from fragmented populations of a single, more widespread progenitor.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |