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<document id="030D9AF442F08859CE276F3353258321" ID-DOI="10.5281/zenodo.277016" ID-GBIF-Dataset="c678007e-60bb-4123-acdb-3588c6b6c7fe" ID-ISSN="1175-5326" ID-Zenodo-Dep="277016" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460261786154" checkinUser="plazi" docAuthor="Rowley, Jodi J. L., Le, Duong Thi Thuy, Tran, Dao Thi Anh &amp; Hoang, Huy Duc" docDate="2011" docId="03B187D6FFD4FFB22393FE15FBD54A05" docLanguage="en" docName="zt02796p028.pdf" docOrigin="Zootaxa 2796" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptolalax bidoupensis Rowley, Le, Tran &amp; Hoang, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="26" masterDocId="FF88FFAEFFD6FFB92304FFEAFF9E4B19" masterDocTitle="A new species of Leptolalax (Anura: Megophryidae) from southern Vietnam" masterLastPageNumber="28" masterPageNumber="15" pageNumber="17" updateTime="1698244594168" updateUser="plazi">
<mods:mods id="D2E6CE12E0DA874D049AEBC3BE824A6B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="F70C85C28C78816FEA2625D21D7C5EBC">A new species of Leptolalax (Anura: Megophryidae) from southern Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="F29A6D1B2803B3D997F0A172BF220003" type="personal">
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<mods:namePart id="C566B5E5D5DD4C6B24F505D7BB581552">Rowley, Jodi J. L.</mods:namePart>
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<mods:namePart id="E574F3768E1519B2CBD45C85422460D6">Le, Duong Thi Thuy</mods:namePart>
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<mods:namePart id="64C68F4F4D9665F7895106E97AED16E9">Tran, Dao Thi Anh</mods:namePart>
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<mods:namePart id="515267A3E635FE02CDFF71BCDBD673FD">Hoang, Huy Duc</mods:namePart>
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<mods:title id="33495FEE86F59076651325139A172C9F">Zootaxa</mods:title>
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<mods:part id="C5BFCCDBFBA662F3D0CEABDE2BBAEB20">
<mods:date id="3C56B223DAD054D8E80E040B9DE489DA">2011</mods:date>
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<treatment id="03B187D6FFD4FFB22393FE15FBD54A05" ID-DOI="http://doi.org/10.5281/zenodo.5612780" ID-GBIF-Taxon="119393492" ID-Zenodo-Dep="5612780" LSID="urn:lsid:plazi:treatment:03B187D6FFD4FFB22393FE15FBD54A05" httpUri="http://treatment.plazi.org/id/03B187D6FFD4FFB22393FE15FBD54A05" lastPageId="11" lastPageNumber="26" pageId="2" pageNumber="17">
<subSubSection id="C302654BFFD4FFBB2393FE15FD8C4900" box="[151,530,511,537]" pageId="2" pageNumber="17" type="nomenclature">
<paragraph id="8BA736C0FFD4FFBB2393FE15FD8C4900" blockId="2.[151,530,511,537]" box="[151,530,511,537]" pageId="2" pageNumber="17">
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<emphasis id="B96CEAD2FFD4FFBB2393FE15FD8C4900" bold="true" box="[151,530,511,537]" pageId="2" pageNumber="17">
<taxonomicName id="4C184D43FFD4FFBB2393FE15FE304900" ID-CoL="3TQPG" box="[151,430,511,537]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD4FFBB2393FE15FE304900" bold="true" box="[151,430,511,537]" italics="true" pageId="2" pageNumber="17">Leptolalax bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD4FFBB22B1FE15FD8C4900" box="[437,530,511,537]" pageId="2" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302654BFFD4FFBB2393FDADFEB24E6F" pageId="2" pageNumber="17" type="materials_examined">
<paragraph id="8BA736C0FFD4FFBB2393FDADFE1B49D3" blockId="2.[151,1437,582,2011]" pageId="2" pageNumber="17">
<emphasis id="B96CEAD2FFD4FFBB2393FDADFE934946" bold="true" box="[151,269,583,607]" pageId="2" pageNumber="17">
<typeStatus id="54A38862FFD4FFBB2393FDADFE964946" box="[151,264,583,607]" pageId="2" pageNumber="17" type="holotype">Holotype</typeStatus>
:
</emphasis>
AMS R 173133, adult male, calling on clay bank
<quantity id="4CE09B25FFD4FFBB203BFDACFCE14947" box="[831,895,582,606]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="17" unit="m" value="0.2">0.2 m</quantity>
from
<quantity id="4CE09B25FFD4FFBB20C7FDACFB944947" box="[963,1034,582,606]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="2" pageNumber="17" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">14 m</quantity>
wide, medium-high gradient, rocky stream in montane evergreen forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB279CFD80FB69499B" box="[1176,1271,618,642]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB2602FD80FA06499B" box="[1286,1432,618,642]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.19225">12.19225º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB2393FD64FEAC49BF" box="[151,306,654,678]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71494">108.71494º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB223AFD64FE0B49BF" box="[318,405,654,678]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="2" pageNumber="17" unit="m" value="1730.0">1730 m</quantity>
,
<figureCitation id="13232A45FFD4FFBB22A6FD64FD9E49BF" box="[418,512,654,678]" captionStart="FIGURE 1" captionStartId="4.[151,250,1412,1435]" captionTargetBox="[151,1436,298,1391]" captionTargetId="figure@4.[151,1436,298,1391]" captionTargetPageId="4" captionText="FIGURE 1. Collection site of Leptolalax bidoupensis sp. nov. from Bidoup-Nui Ba National Park (black star) and type localities of Leptolalax applebyi (white circle) and Leptolalax melicus (black circles)." httpUri="https://zenodo.org/record/277017/files/figure.png" pageId="2" pageNumber="17">Figure 1</figureCitation>
). Collected at 23:55 h on
<date id="FFA61000FFD4FFBB202DFD64FC5E49BF" box="[809,960,654,679]" pageId="2" pageNumber="17" value="2008-05-19">19 May 2008</date>
by J. J. L. Rowley, Hoang D. H., Le T. T. D., and Tran T. A. D.
</paragraph>
<paragraph id="8BA736C0FFD4FFBB23C1FD3DFEB24E6F" blockId="2.[151,1437,582,2011]" pageId="2" pageNumber="17">
<emphasis id="B96CEAD2FFD4FFBB23C1FD3DFED849F6" bold="true" box="[197,326,727,751]" pageId="2" pageNumber="17">
<typeStatus id="54A38862FFD4FFBB23C1FD3DFEDF49F6" box="[197,321,727,751]" pageId="2" pageNumber="17" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectionCode id="ED09AE05FFD4FFBB224AFD3CFE1849F7" box="[334,390,726,750]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="17">UNS</collectionCode>
00101/AMS R 173135, adult male, calling on tree root
<quantity id="4CE09B25FFD4FFBB270EFD3CFBD249F7" box="[1034,1100,726,750]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="17" unit="m" value="0.2">0.2 m</quantity>
above
<quantity id="4CE09B25FFD4FFBB27A4FD3CFB7649F7" box="[1184,1256,726,750]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="17" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">25 m</quantity>
wide, mediumhigh gradient, rocky stream in montane evergreen forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB2393FCF4FF68482F" box="[151,246,798,822]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB2200FCF4FE0B482F" box="[260,405,798,822]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.19106">12.19106º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB22A4FCF4FDA7482F" box="[416,569,798,822]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71703">108.71703º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB2141FCF4FD03482F" box="[581,669,798,822]" metricMagnitude="3" metricUnit="m" metricValue="1.641" pageId="2" pageNumber="17" unit="m" value="1641.0">1641 m</quantity>
), collected at 20:25 h on
<date id="FFA61000FFD4FFBB20B1FCF4FBD0482F" box="[949,1102,798,823]" pageId="2" pageNumber="17" value="2008-05-20">20 May 2008</date>
.
<collectionCode id="ED09AE05FFD4FFBB275CFCF4FB0E482F" box="[1112,1168,798,822]" httpUri="http://grbio.org/cool/0nv9-p2k7" name="University of Science, Ho Chi Minh City, Vietnam" pageId="2" pageNumber="17">UNS</collectionCode>
00102/AMS R 173137, metamorph, in water of swampy area adjacent to a swift, rocky stream in cloud forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB22CAFC8CFDB34867" box="[462,557,870,894]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB2138FC8CFD514867" box="[572,719,870,894]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.18644">12.18644º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB21D8FC8CFCE84867" box="[732,886,870,894]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71486">108.71486º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB2087FC8CFC424867" box="[899,988,870,894]" metricMagnitude="3" metricUnit="m" metricValue="1.627" pageId="2" pageNumber="17" unit="m" value="1627.0">1627 m</quantity>
), collected at 19:45 h on
<date id="FFA61000FFD4FFBB27FAFC8CFA074867" box="[1278,1433,870,895]" pageId="2" pageNumber="17" value="2008-05-18">18 May 2008</date>
. AMS R 173134, adult female, on clay bank
<quantity id="4CE09B25FFD4FFBB218DFC60FD5748BB" box="[649,713,906,930]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="17" unit="m" value="0.2">0.2 m</quantity>
from
<quantity id="4CE09B25FFD4FFBB200AFC60FCC848BB" box="[782,854,906,930]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="2" pageNumber="17" unit="m" value="2.5" valueMax="4.0" valueMin="1.0">14 m</quantity>
wide, medium-high gradient, rocky stream in montane evergreen forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB20F7FC44FBCC48DF" box="[1011,1106,942,966]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB2764FC44FB6C48DF" box="[1120,1266,942,966]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.19225">12.19225º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB27F9FC44FA0948DF" box="[1277,1431,942,966]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71494">108.71494º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB2393FC38FF7148F3" box="[151,239,978,1002]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="2" pageNumber="17" unit="m" value="1730.0">1730 m</quantity>
), collected at 23:50 h on
<date id="FFA61000FFD4FFBB210DFC38FD3C48F3" box="[521,674,978,1003]" pageId="2" pageNumber="17" value="2008-05-19">19 May 2008</date>
, in close proximity to
<typeStatus id="54A38862FFD4FFBB2098FC38FB9C48F3" box="[924,1026,978,1002]" pageId="2" pageNumber="17" type="holotype">holotype</typeStatus>
. AMS R 173136, adult male, calling on leaf litter
<quantity id="4CE09B25FFD4FFBB222CFC1CFEF64F17" box="[296,360,1014,1038]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="17" unit="m" value="0.2">0.2 m</quantity>
from
<quantity id="4CE09B25FFD4FFBB22A9FC1CFE6A4F17" box="[429,500,1014,1038]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="17" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">25 m</quantity>
wide, medium-high gradient, rocky stream in montane evergreen forest in Bidoup- Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB2181FBF0FD7A4F2B" box="[645,740,1050,1074]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB21F6FBF0FC1D4F2B" box="[754,899,1050,1074]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.19106">12.19106º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB208AFBF0FBB64F2B" box="[910,1064,1050,1074]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71703">108.71703º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB2737FBF0FB154F2B" box="[1075,1163,1050,1074]" metricMagnitude="3" metricUnit="m" metricValue="1.641" pageId="2" pageNumber="17" unit="m" value="1641.0">1641 m</quantity>
), collected at 21:40 h on
<date id="FFA61000FFD4FFBB2393FBD4FEAF4F4F" box="[151,305,1086,1111]" pageId="2" pageNumber="17" value="2008-05-20">20 May 2008</date>
.
<collectionCode id="ED09AE05FFD4FFBB2238FBD4FE154F4F" box="[316,395,1086,1110]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="2" pageNumber="17">NCSM</collectionCode>
77320, adult female, in water of swampy area off swift, rocky stream in montane evergreen forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB204BFB88FC304F63" box="[847,942,1122,1146]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB20B9FB88FBD14F63" box="[957,1103,1122,1146]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.18644">12.18644º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB275FFB88FB684F63" box="[1115,1270,1122,1146]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71486">108.71486º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB2607FB88FAC54F63" box="[1283,1371,1122,1146]" metricMagnitude="3" metricUnit="m" metricValue="1.627" pageId="2" pageNumber="17" unit="m" value="1627.0">1627 m</quantity>
), collected at 19:50 h on
<date id="FFA61000FFD4FFBB227EFB6CFD8C4F87" box="[378,530,1158,1183]" pageId="2" pageNumber="17" value="2008-05-18">18 May 2008</date>
.
<collectionCode id="ED09AE05FFD4FFBB2119FB6CFDF24F87" box="[541,620,1158,1182]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="2" pageNumber="17">NCSM</collectionCode>
77321, adult male, sitting in upright posture (previously calling?) in leaf litter,
<quantity id="4CE09B25FFD4FFBB23D8FB40FE814FDB" box="[220,287,1194,1218]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="17" unit="m" value="0.1">0.1 m</quantity>
from swift, rocky stream in montane evergreen forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB220FFB24FEF44FFF" box="[267,362,1230,1254]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB227FFB24FD914FFF" box="[379,527,1230,1254]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.18644">12.18644º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB2119FB24FD244FFF" box="[541,698,1230,1254]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71486">108.71486º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB21C3FB24FCBC4FFF" box="[711,802,1230,1254]" metricMagnitude="3" metricUnit="m" metricValue="1.627" pageId="2" pageNumber="17" unit="m" value="1627.0">1627 m</quantity>
), collected at 21:45 h on
<date id="FFA61000FFD4FFBB274FFB24FB774FFF" box="[1099,1257,1230,1255]" pageId="2" pageNumber="17" value="2008-05-18">18 May 2008</date>
.
<collectionCode id="ED09AE05FFD4FFBB27F2FB24FADB4FFF" box="[1270,1349,1230,1254]" httpUri="http://grbio.org/cool/aasf-rm4s" name="North Carolina Museum of Natural Sciences" pageId="2" pageNumber="17">NCSM</collectionCode>
77322, metamorph, on clay bank
<quantity id="4CE09B25FFD4FFBB22B8FB18FE634E13" box="[444,509,1266,1290]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="17" unit="m" value="0.5">0.5 m</quantity>
from
<quantity id="4CE09B25FFD4FFBB2146FB18FD174E13" box="[578,649,1266,1290]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="17" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">25 m</quantity>
wide, medium-high gradient, rocky stream in montane evergreen forest in Bidoup-Nui Ba National Park, Lam Dong Province,
<collectingCountry id="F30F7650FFD4FFBB2044FAFCFC014E37" box="[832,927,1302,1326]" name="Vietnam" pageId="2" pageNumber="17">Vietnam</collectingCountry>
(
<geoCoordinate id="EE2C5007FFD4FFBB20B5FAFCFBDB4E37" box="[945,1093,1302,1326]" direction="north" orientation="latitude" pageId="2" pageNumber="17" precision="1" value="12.19106">12.19106º N</geoCoordinate>
,
<geoCoordinate id="EE2C5007FFD4FFBB2757FAFCFB6E4E37" box="[1107,1264,1302,1326]" direction="east" orientation="longitude" pageId="2" pageNumber="17" precision="1" value="108.71703">108.71703º E</geoCoordinate>
,
<quantity id="4CE09B25FFD4FFBB27FAFAFCFAC74E37" box="[1278,1369,1302,1326]" metricMagnitude="3" metricUnit="m" metricValue="1.641" pageId="2" pageNumber="17" unit="m" value="1641.0">1641 m</quantity>
), collected at 21:30 h on
<date id="FFA61000FFD4FFBB2278FAD0FD8B4E4B" box="[380,533,1338,1363]" pageId="2" pageNumber="17" value="2008-05-20">20 May 2008</date>
. All specimens were collected by J. J. L. Rowley, Hoang D. H., Le T. T. D., and Tran T. A. D.
</paragraph>
</subSubSection>
<subSubSection id="C302654BFFD4FFBB23C1FA69FB6B4E83" box="[197,1269,1410,1435]" pageId="2" pageNumber="17" type="etymology">
<paragraph id="8BA736C0FFD4FFBB23C1FA69FB6B4E83" blockId="2.[151,1437,582,2011]" box="[197,1269,1410,1435]" pageId="2" pageNumber="17">
<emphasis id="B96CEAD2FFD4FFBB23C1FA69FED54E82" bold="true" box="[197,331,1411,1435]" pageId="2" pageNumber="17">Etymology.</emphasis>
specific epithet is in reference to the
<typeStatus id="54A38862FFD4FFBB21EBFA69FC814E82" box="[751,799,1411,1435]" pageId="2" pageNumber="17">type</typeStatus>
locality of Bidoup-Nui Ba National Park.
</paragraph>
</subSubSection>
<subSubSection id="C302654BFFD4FFBB23C1FA4CFE044C3F" pageId="2" pageNumber="17" type="diagnosis">
<paragraph id="8BA736C0FFD4FFBB23C1FA4CFE044C3F" blockId="2.[151,1437,582,2011]" pageId="2" pageNumber="17">
<emphasis id="B96CEAD2FFD4FFBB23C1FA4CFEA14EA6" bold="true" box="[197,319,1446,1471]" pageId="2" pageNumber="17">Diagnosis.</emphasis>
Assigned to the genus
<taxonomicName id="4C184D43FFD4FFBB2142FA4CFD204EA7" box="[582,702,1446,1470]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B96CEAD2FFD4FFBB2142FA4CFD204EA7" box="[582,702,1446,1470]" italics="true" pageId="2" pageNumber="17">Leptolalax</emphasis>
</taxonomicName>
on the basis of the following: small size, rounded finger tips, the presence of an elevated inner palmar tubercle not continuous to the thumb, presence of macroglands on body (including supra-axillary, pectoral, femoral and ventrolateral glands), vomerine teeth absent, tubercles on eyelids, anterior tip of snout with vertical white bar (
<bibRefCitation id="EF894B31FFD4FFBB2191F9F8FCB24D33" author="Dubois" box="[661,812,1554,1578]" pageId="2" pageNumber="17" refString="Dubois, A. (1983) Note preliminaire sur le genre Leptolalax Dubois, 1980 (Amphibiens, Anoures), avec diagnose d'une espece novella du Vietnam. Alytes, 2, 147 - 153." type="journal article" year="1983">Dubois 1983</bibRefCitation>
;
<bibRefCitation id="EF894B31FFD4FFBB203DF9F8FB864D33" author="Lathrop" box="[825,1048,1554,1579]" pageId="2" pageNumber="17" refString="Lathrop, A., Murphy, R. W., Orlov, N. &amp; Ho, C. T. (1998) Two new species of Leptolalax (Anura: Megophryidae) from northern Vietnam. Amphibia-Reptilia, 19, 253 - 267." type="journal article" year="1998">
Lathrop
<emphasis id="B96CEAD2FFD4FFBB209FF9FEFC534D32" box="[923,973,1555,1579]" italics="true" pageId="2" pageNumber="17">et al</emphasis>
. 1998
</bibRefCitation>
; Delorme
<emphasis id="B96CEAD2FFD4FFBB2794F9FEFB5D4D32" box="[1168,1219,1555,1579]" italics="true" pageId="2" pageNumber="17">et al</emphasis>
. 2006).
<taxonomicName id="4C184D43FFD4FFBB2620F9F9FE834D57" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="17" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFD4FFBB2620F9F9FE834D57" italics="true" pageId="2" pageNumber="17">Leptolalax bidoupensis</emphasis>
</taxonomicName>
is distinguished from its congeners by a combination of (1) a dark brownish red ventral surface with white speckling on entire ventral surface including throat, arms and legs, often forming distinct marbling on chest and belly, (2) small size (
<quantity id="4CE09B25FFD4FFBB22A9F994FDCF4D8F" box="[429,593,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" metricValueMax="2.46" metricValueMin="2.36" pageId="2" pageNumber="17" unit="mm" value="24.1" valueMax="24.6" valueMin="23.6">23.624.6 mm</quantity>
in four adult males and
<quantity id="4CE09B25FFD4FFBB2059F994FC614D8F" box="[861,1023,1662,1686]" metricMagnitude="-2" metricUnit="m" metricValue="2.9299999999999997" metricValueMax="2.94" metricValueMin="2.92" pageId="2" pageNumber="17" unit="mm" value="29.299999999999997" valueMax="29.4" valueMin="29.2">29.229.4 mm</quantity>
in two adult females), (3) bicoloured iris (coppery red upper half, fading to pale silver ventrally), (4) a mostly smooth skin texture with no skin ridges, and (5) relatively short tibia (male TIB:SVL 0.440.46). The male advertisement call of the new species, consisting of 69 single-pulsed notes with a dominant frequency of 1.93.8 kHz, is also unique among
<emphasis id="B96CEAD2FFD4FFBB27A7F901FA914C1A" box="[1187,1295,1771,1795]" italics="true" pageId="2" pageNumber="17">Leptolala</emphasis>
x species for which calls are known.
</paragraph>
</subSubSection>
<subSubSection id="C302654BFFD4FFB323C1F8D8FAB149A3" lastPageId="10" lastPageNumber="25" pageId="2" pageNumber="17" type="description">
<paragraph id="8BA736C0FFD4FFBA23C1F8D8FCA14925" blockId="2.[151,1437,582,2011]" lastBlockId="3.[151,1437,151,572]" lastPageId="3" lastPageNumber="18" pageId="2" pageNumber="17">
<emphasis id="B96CEAD2FFD4FFBB23C1F8D8FE7A4C52" bold="true" box="[197,484,1842,1867]" pageId="2" pageNumber="17">
Description of
<typeStatus id="54A38862FFD4FFBB2272F8D9FE414C52" box="[374,479,1843,1867]" pageId="2" pageNumber="17" type="holotype">holotype</typeStatus>
.
</emphasis>
Head slightly longer than wide; snout bluntly rounded in dorsal view and in profile, projecting slightly beyond margin of the lower jaw; nostril closer to tip of snout than eye; canthus rostralis distinct, gently rounded; lores sloping; vertical pupil; eye diameter smaller than snout length; tympanum distinct, round, diameter smaller than that of the eye; tympanic rim elevated relative to skin of temporal region; vomerine teeth absent; pineal ocellus absent; vocal sac openings oval, located posteriolaterally on floor of mouth; tongue long, moderate width, with slight notch at posterior tip; raised supratympanic ridge running from eye towards axillary gland. Tips of fingers rounded, not swollen; relative finger lengths I &lt;II = IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, round inner palmar tubercle distinctly separated from small, laterally compressed outer palmar tubercle; no finger webbing or lateral fringes. Tips of toes like fingers; relative toe length I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent, replaced by dermal ridges, distinct on second, third, fourth and fifth toes; large, oval inner metatarsal tubercle pronounced, outer metatarsal tubercle absent; basal webbing; weak lateral fringes. Tibia relatively short and stout, width approximately one third of length, tibia 46% of snout-vent length; tibiotarsal articulation reaches eye. Skin on dorsum mostly smooth, with fine, scattered tubercles concentrated on eyelids, ventrolateral surfaces and on upper surfaces on tibiotarsus; ventral skin smooth; pectoral gland oval,
<quantity id="4CE09B25FFD5FFBA27FBFE5DFACA4AD6" box="[1279,1364,439,463]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
diameter; femoral gland oval,
<quantity id="4CE09B25FFD5FFBA22B3FE36FD914AED" box="[439,527,476,500]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="18" unit="mm" value="0.8">0.8 mm</quantity>
diameter, on posteroventral surface of thigh, closer to knee than to vent; supraaxillary gland raised,
<quantity id="4CE09B25FFD5FFBA228CFE15FE40490E" box="[392,478,511,535]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="18" unit="mm" value="1.2">1.2 mm</quantity>
diameter. Ventrolateral glands present, dorsolaterally compressed, forming an incomplete line that is difficult to distinguish from white marbling.
</paragraph>
<caption id="DF676648FFD5FFBA2393FD86FE8549B9" ID-Table-UUID="DF676648FFD5FFBA2393FD86FE8549B9" httpUri="http://table.plazi.org/id/DF676648FFD5FFBA2393FD86FE8549B9" pageId="3" pageNumber="18" targetBox="[151,1436,741,1654]" targetIsTable="true" targetPageId="3">
<paragraph id="8BA736C0FFD5FFBA2393FD86FE8549B9" blockId="3.[151,1436,619,721]" pageId="3" pageNumber="18">
<emphasis id="B96CEAD2FFD5FFBA2393FD86FE9B4998" bold="true" box="[151,261,620,641]" pageId="3" pageNumber="18">TABLE 1.</emphasis>
Measurements (mm) of
<taxonomicName id="4C184D43FFD5FFBA2106FD86FD70499B" box="[514,750,620,642]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="18" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD5FFBA2106FD86FD70499B" box="[514,750,620,642]" italics="true" pageId="3" pageNumber="18">Leptolalax bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD5FFBA21F1FD86FCD94998" box="[757,839,620,641]" pageId="3" pageNumber="18" rank="species">
<emphasis id="B96CEAD2FFD5FFBA21F1FD86FCD94998" bold="true" box="[757,839,620,641]" pageId="3" pageNumber="18">sp. nov.</emphasis>
</taxonomicNameLabel>
Abbreviations defined in text. Tympana were indistinct in metamorphs.
</paragraph>
</caption>
<paragraph id="8BA736C0FFD5FFBA224FFD51FA8449C8" blockId="3.[151,1436,619,721]" box="[331,1306,699,721]" pageId="3" pageNumber="18">Males Females Metamorphs</paragraph>
<paragraph id="8BA736C0FFD5FFBA224FFD0FFE9A4D6F" pageId="3" pageNumber="18">
<table id="F918C460FFD500462393FD0FFA024D6F" box="[151,1436,741,1654]" gridcols="9" gridrows="20" pageId="3" pageNumber="18">
<tr id="35283482FFD500462393FD0FFA02484E" box="[151,1436,741,855]" gridrow="0" pageId="3" pageNumber="18" rowspan-0="1">
<th id="76F95DFEFFD50046224FFD0FFE3D484E" box="[331,419,741,855]" gridcol="1" gridrow="0" pageId="3" pageNumber="18">AMS R 173133*</th>
<th id="76F95DFEFFD5004622E8FD0FFDA4484E" box="[492,570,741,855]" gridcol="2" gridrow="0" pageId="3" pageNumber="18">UNS 00101/ AMS R 173135</th>
<th id="76F95DFEFFD50046217DFD0FFD59484E" box="[633,711,741,855]" gridcol="3" gridrow="0" pageId="3" pageNumber="18">AMS R 173136</th>
<th id="76F95DFEFFD500462002FD0FFCD0484E" box="[774,846,741,855]" gridcol="4" gridrow="0" pageId="3" pageNumber="18">NCSM 77321</th>
<th id="76F95DFEFFD50046207BFD0FFC50484E" box="[895,974,741,855]" gridcol="5" gridrow="0" pageId="3" pageNumber="18">AMS R 173134</th>
<th id="76F95DFEFFD500462708FD0FFBCA484E" box="[1036,1108,741,855]" gridcol="6" gridrow="0" pageId="3" pageNumber="18">NCSM 77320</th>
<th id="76F95DFEFFD50046279EFD0FFB76484E" box="[1178,1256,741,855]" gridcol="7" gridrow="0" pageId="3" pageNumber="18">UNS 00102/ AMS R 173137</th>
<th id="76F95DFEFFD500462623FD0FFA02484E" box="[1319,1436,741,855]" gridcol="8" gridrow="0" pageId="3" pageNumber="18">NCSM 77322</th>
</tr>
<tr id="35283482FFD500462393FC80FA024899" box="[151,1436,874,896]" gridrow="1" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FC80FE884899" box="[151,278,874,896]" gridcol="0" gridrow="1" pageId="3" pageNumber="18">SVL</th>
<td id="76F95DFEFFD50046224FFC80FE3D4899" box="[331,419,874,896]" gridcol="1" gridrow="1" pageId="3" pageNumber="18">23.6</td>
<td id="76F95DFEFFD5004622E8FC80FDA44899" box="[492,570,874,896]" gridcol="2" gridrow="1" pageId="3" pageNumber="18">24.3</td>
<td id="76F95DFEFFD50046217DFC80FD594899" box="[633,711,874,896]" gridcol="3" gridrow="1" pageId="3" pageNumber="18">24.6</td>
<td id="76F95DFEFFD500462002FC80FCD04899" box="[774,846,874,896]" gridcol="4" gridrow="1" pageId="3" pageNumber="18">24.6</td>
<td id="76F95DFEFFD50046207BFC80FC504899" box="[895,974,874,896]" gridcol="5" gridrow="1" pageId="3" pageNumber="18">29.4</td>
<td id="76F95DFEFFD500462708FC80FBCA4899" box="[1036,1108,874,896]" gridcol="6" gridrow="1" pageId="3" pageNumber="18">29.2</td>
<td id="76F95DFEFFD50046279EFC80FB764899" box="[1178,1256,874,896]" gridcol="7" gridrow="1" pageId="3" pageNumber="18">17.3</td>
<td id="76F95DFEFFD500462623FC80FA024899" box="[1319,1436,874,896]" gridcol="8" gridrow="1" pageId="3" pageNumber="18">17.1</td>
</tr>
<tr id="35283482FFD500462393FC79FA0248B3" box="[151,1436,915,938]" gridrow="2" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FC79FE8848B3" box="[151,278,915,938]" gridcol="0" gridrow="2" pageId="3" pageNumber="18">HDL</th>
<td id="76F95DFEFFD50046224FFC79FE3D48B3" box="[331,419,915,938]" gridcol="1" gridrow="2" pageId="3" pageNumber="18">9.2</td>
<td id="76F95DFEFFD5004622E8FC79FDA448B3" box="[492,570,915,938]" gridcol="2" gridrow="2" pageId="3" pageNumber="18">9.2</td>
<td id="76F95DFEFFD50046217DFC79FD5948B3" box="[633,711,915,938]" gridcol="3" gridrow="2" pageId="3" pageNumber="18">9.6</td>
<td id="76F95DFEFFD500462002FC79FCD048B3" box="[774,846,915,938]" gridcol="4" gridrow="2" pageId="3" pageNumber="18">9.6</td>
<td id="76F95DFEFFD50046207BFC79FC5048B3" box="[895,974,915,938]" gridcol="5" gridrow="2" pageId="3" pageNumber="18">10.1</td>
<td id="76F95DFEFFD500462708FC79FBCA48B3" box="[1036,1108,915,938]" gridcol="6" gridrow="2" pageId="3" pageNumber="18">10.5</td>
<td id="76F95DFEFFD50046279EFC79FB7648B3" box="[1178,1256,915,938]" gridcol="7" gridrow="2" pageId="3" pageNumber="18">5.4</td>
<td id="76F95DFEFFD500462623FC79FA0248B3" box="[1319,1436,915,938]" gridcol="8" gridrow="2" pageId="3" pageNumber="18">5.7</td>
</tr>
<tr id="35283482FFD500462393FC57FA0248CA" box="[151,1436,957,979]" gridrow="3" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FC57FE8848CA" box="[151,278,957,979]" gridcol="0" gridrow="3" pageId="3" pageNumber="18">HDW</th>
<td id="76F95DFEFFD50046224FFC57FE3D48CA" box="[331,419,957,979]" gridcol="1" gridrow="3" pageId="3" pageNumber="18">8.9</td>
<td id="76F95DFEFFD5004622E8FC57FDA448CA" box="[492,570,957,979]" gridcol="2" gridrow="3" pageId="3" pageNumber="18">8.8</td>
<td id="76F95DFEFFD50046217DFC57FD5948CA" box="[633,711,957,979]" gridcol="3" gridrow="3" pageId="3" pageNumber="18">9.2</td>
<td id="76F95DFEFFD500462002FC57FCD048CA" box="[774,846,957,979]" gridcol="4" gridrow="3" pageId="3" pageNumber="18">9.0</td>
<td id="76F95DFEFFD50046207BFC57FC5048CA" box="[895,974,957,979]" gridcol="5" gridrow="3" pageId="3" pageNumber="18">9.6</td>
<td id="76F95DFEFFD500462708FC57FBCA48CA" box="[1036,1108,957,979]" gridcol="6" gridrow="3" pageId="3" pageNumber="18">9.8</td>
<td id="76F95DFEFFD50046279EFC57FB7648CA" box="[1178,1256,957,979]" gridcol="7" gridrow="3" pageId="3" pageNumber="18">5.5</td>
<td id="76F95DFEFFD500462623FC57FA0248CA" box="[1319,1436,957,979]" gridcol="8" gridrow="3" pageId="3" pageNumber="18">5.5</td>
</tr>
<tr id="35283482FFD500462393FC0CFA0248E5" box="[151,1436,998,1020]" gridrow="4" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FC0CFE8848E5" box="[151,278,998,1020]" gridcol="0" gridrow="4" pageId="3" pageNumber="18">SNT</th>
<td id="76F95DFEFFD50046224FFC0CFE3D48E5" box="[331,419,998,1020]" gridcol="1" gridrow="4" pageId="3" pageNumber="18">3.6</td>
<td id="76F95DFEFFD5004622E8FC0CFDA448E5" box="[492,570,998,1020]" gridcol="2" gridrow="4" pageId="3" pageNumber="18">3.5</td>
<td id="76F95DFEFFD50046217DFC0CFD5948E5" box="[633,711,998,1020]" gridcol="3" gridrow="4" pageId="3" pageNumber="18">3.6</td>
<td id="76F95DFEFFD500462002FC0CFCD048E5" box="[774,846,998,1020]" gridcol="4" gridrow="4" pageId="3" pageNumber="18">3.6</td>
<td id="76F95DFEFFD50046207BFC0CFC5048E5" box="[895,974,998,1020]" gridcol="5" gridrow="4" pageId="3" pageNumber="18">3.8</td>
<td id="76F95DFEFFD500462708FC0CFBCA48E5" box="[1036,1108,998,1020]" gridcol="6" gridrow="4" pageId="3" pageNumber="18">3.8</td>
<td id="76F95DFEFFD50046279EFC0CFB7648E5" box="[1178,1256,998,1020]" gridcol="7" gridrow="4" pageId="3" pageNumber="18">2.3</td>
<td id="76F95DFEFFD500462623FC0CFA0248E5" box="[1319,1436,998,1020]" gridcol="8" gridrow="4" pageId="3" pageNumber="18">2.3</td>
</tr>
<tr id="35283482FFD500462393FBFAFA024F3F" box="[151,1436,1040,1062]" gridrow="5" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FBFAFE884F3F" box="[151,278,1040,1062]" gridcol="0" gridrow="5" pageId="3" pageNumber="18">EYE</th>
<td id="76F95DFEFFD50046224FFBFAFE3D4F3F" box="[331,419,1040,1062]" gridcol="1" gridrow="5" pageId="3" pageNumber="18">2.5</td>
<td id="76F95DFEFFD5004622E8FBFAFDA44F3F" box="[492,570,1040,1062]" gridcol="2" gridrow="5" pageId="3" pageNumber="18">2.6</td>
<td id="76F95DFEFFD50046217DFBFAFD594F3F" box="[633,711,1040,1062]" gridcol="3" gridrow="5" pageId="3" pageNumber="18">2.8</td>
<td id="76F95DFEFFD500462002FBFAFCD04F3F" box="[774,846,1040,1062]" gridcol="4" gridrow="5" pageId="3" pageNumber="18">2.8</td>
<td id="76F95DFEFFD50046207BFBFAFC504F3F" box="[895,974,1040,1062]" gridcol="5" gridrow="5" pageId="3" pageNumber="18">2.8</td>
<td id="76F95DFEFFD500462708FBFAFBCA4F3F" box="[1036,1108,1040,1062]" gridcol="6" gridrow="5" pageId="3" pageNumber="18">2.7</td>
<td id="76F95DFEFFD50046279EFBFAFB764F3F" box="[1178,1256,1040,1062]" gridcol="7" gridrow="5" pageId="3" pageNumber="18">1.6</td>
<td id="76F95DFEFFD500462623FBFAFA024F3F" box="[1319,1436,1040,1062]" gridcol="8" gridrow="5" pageId="3" pageNumber="18">1.7</td>
</tr>
<tr id="35283482FFD500462393FBD0FA024F56" box="[151,1436,1082,1103]" gridrow="6" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FBD0FE884F56" box="[151,278,1082,1103]" gridcol="0" gridrow="6" pageId="3" pageNumber="18">IOD</th>
<td id="76F95DFEFFD50046224FFBD0FE3D4F56" box="[331,419,1082,1103]" gridcol="1" gridrow="6" pageId="3" pageNumber="18">2.9</td>
<td id="76F95DFEFFD5004622E8FBD0FDA44F56" box="[492,570,1082,1103]" gridcol="2" gridrow="6" pageId="3" pageNumber="18">3</td>
<td id="76F95DFEFFD50046217DFBD0FD594F56" box="[633,711,1082,1103]" gridcol="3" gridrow="6" pageId="3" pageNumber="18">2.9</td>
<td id="76F95DFEFFD500462002FBD0FCD04F56" box="[774,846,1082,1103]" gridcol="4" gridrow="6" pageId="3" pageNumber="18">2.9</td>
<td id="76F95DFEFFD50046207BFBD0FC504F56" box="[895,974,1082,1103]" gridcol="5" gridrow="6" pageId="3" pageNumber="18">3.1</td>
<td id="76F95DFEFFD500462708FBD0FBCA4F56" box="[1036,1108,1082,1103]" gridcol="6" gridrow="6" pageId="3" pageNumber="18">3.1</td>
<td id="76F95DFEFFD50046279EFBD0FB764F56" box="[1178,1256,1082,1103]" gridcol="7" gridrow="6" pageId="3" pageNumber="18">2.1</td>
<td id="76F95DFEFFD500462623FBD0FA024F56" box="[1319,1436,1082,1103]" gridcol="8" gridrow="6" pageId="3" pageNumber="18">2.4</td>
</tr>
<tr id="35283482FFD500462393FB88FA024F61" box="[151,1436,1122,1144]" gridrow="7" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FB88FE884F61" box="[151,278,1122,1144]" gridcol="0" gridrow="7" pageId="3" pageNumber="18">TMP</th>
<td id="76F95DFEFFD50046224FFB88FE3D4F61" box="[331,419,1122,1144]" gridcol="1" gridrow="7" pageId="3" pageNumber="18">1.4</td>
<td id="76F95DFEFFD5004622E8FB88FDA44F61" box="[492,570,1122,1144]" gridcol="2" gridrow="7" pageId="3" pageNumber="18">1.5</td>
<td id="76F95DFEFFD50046217DFB88FD594F61" box="[633,711,1122,1144]" gridcol="3" gridrow="7" pageId="3" pageNumber="18">1.9</td>
<td id="76F95DFEFFD500462002FB88FCD04F61" box="[774,846,1122,1144]" gridcol="4" gridrow="7" pageId="3" pageNumber="18">1.5</td>
<td id="76F95DFEFFD50046207BFB88FC504F61" box="[895,974,1122,1144]" gridcol="5" gridrow="7" pageId="3" pageNumber="18">1.8</td>
<td id="76F95DFEFFD500462708FB88FBCA4F61" box="[1036,1108,1122,1144]" gridcol="6" gridrow="7" pageId="3" pageNumber="18">1.6</td>
<td id="76F95DFEFFD50046279EFB88FB764F61" box="[1178,1256,1122,1144]" gridcol="7" gridrow="7" pageId="3" pageNumber="18"></td>
<td id="76F95DFEFFD500462623FB88FA024F61" box="[1319,1436,1122,1144]" gridcol="8" gridrow="7" pageId="3" pageNumber="18"></td>
</tr>
<tr id="35283482FFD500462393FB61FA024FBB" box="[151,1436,1163,1186]" gridrow="8" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FB61FE884FBB" box="[151,278,1163,1186]" gridcol="0" gridrow="8" pageId="3" pageNumber="18">TEY</th>
<td id="76F95DFEFFD50046224FFB61FE3D4FBB" box="[331,419,1163,1186]" gridcol="1" gridrow="8" pageId="3" pageNumber="18">1.3</td>
<td id="76F95DFEFFD5004622E8FB61FDA44FBB" box="[492,570,1163,1186]" gridcol="2" gridrow="8" pageId="3" pageNumber="18">0.4</td>
<td id="76F95DFEFFD50046217DFB61FD594FBB" box="[633,711,1163,1186]" gridcol="3" gridrow="8" pageId="3" pageNumber="18">0.9</td>
<td id="76F95DFEFFD500462002FB61FCD04FBB" box="[774,846,1163,1186]" gridcol="4" gridrow="8" pageId="3" pageNumber="18">1.0</td>
<td id="76F95DFEFFD50046207BFB61FC504FBB" box="[895,974,1163,1186]" gridcol="5" gridrow="8" pageId="3" pageNumber="18">0.6</td>
<td id="76F95DFEFFD500462708FB61FBCA4FBB" box="[1036,1108,1163,1186]" gridcol="6" gridrow="8" pageId="3" pageNumber="18">1.5</td>
<td id="76F95DFEFFD50046279EFB61FB764FBB" box="[1178,1256,1163,1186]" gridcol="7" gridrow="8" pageId="3" pageNumber="18"></td>
<td id="76F95DFEFFD500462623FB61FA024FBB" box="[1319,1436,1163,1186]" gridcol="8" gridrow="8" pageId="3" pageNumber="18"></td>
</tr>
<tr id="35283482FFD500462393FB5FFA024FD2" box="[151,1436,1205,1227]" gridrow="9" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FB5FFE884FD2" box="[151,278,1205,1227]" gridcol="0" gridrow="9" pageId="3" pageNumber="18">TIB</th>
<td id="76F95DFEFFD50046224FFB5FFE3D4FD2" box="[331,419,1205,1227]" gridcol="1" gridrow="9" pageId="3" pageNumber="18">10.4</td>
<td id="76F95DFEFFD5004622E8FB5FFDA44FD2" box="[492,570,1205,1227]" gridcol="2" gridrow="9" pageId="3" pageNumber="18">10.7</td>
<td id="76F95DFEFFD50046217DFB5FFD594FD2" box="[633,711,1205,1227]" gridcol="3" gridrow="9" pageId="3" pageNumber="18">11.2</td>
<td id="76F95DFEFFD500462002FB5FFCD04FD2" box="[774,846,1205,1227]" gridcol="4" gridrow="9" pageId="3" pageNumber="18">11.3</td>
<td id="76F95DFEFFD50046207BFB5FFC504FD2" box="[895,974,1205,1227]" gridcol="5" gridrow="9" pageId="3" pageNumber="18">12.2</td>
<td id="76F95DFEFFD500462708FB5FFBCA4FD2" box="[1036,1108,1205,1227]" gridcol="6" gridrow="9" pageId="3" pageNumber="18">11.3</td>
<td id="76F95DFEFFD50046279EFB5FFB764FD2" box="[1178,1256,1205,1227]" gridcol="7" gridrow="9" pageId="3" pageNumber="18">7.6</td>
<td id="76F95DFEFFD500462623FB5FFA024FD2" box="[1319,1436,1205,1227]" gridcol="8" gridrow="9" pageId="3" pageNumber="18">7.9</td>
</tr>
<tr id="35283482FFD500462393FB34FA024FED" box="[151,1436,1246,1268]" gridrow="10" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FB34FE884FED" box="[151,278,1246,1268]" gridcol="0" gridrow="10" pageId="3" pageNumber="18">ML</th>
<td id="76F95DFEFFD50046224FFB34FE3D4FED" box="[331,419,1246,1268]" gridcol="1" gridrow="10" pageId="3" pageNumber="18">5.2</td>
<td id="76F95DFEFFD5004622E8FB34FDA44FED" box="[492,570,1246,1268]" gridcol="2" gridrow="10" pageId="3" pageNumber="18">5.5</td>
<td id="76F95DFEFFD50046217DFB34FD594FED" box="[633,711,1246,1268]" gridcol="3" gridrow="10" pageId="3" pageNumber="18">5.5</td>
<td id="76F95DFEFFD500462002FB34FCD04FED" box="[774,846,1246,1268]" gridcol="4" gridrow="10" pageId="3" pageNumber="18">5.2</td>
<td id="76F95DFEFFD50046207BFB34FC504FED" box="[895,974,1246,1268]" gridcol="5" gridrow="10" pageId="3" pageNumber="18">6.4</td>
<td id="76F95DFEFFD500462708FB34FBCA4FED" box="[1036,1108,1246,1268]" gridcol="6" gridrow="10" pageId="3" pageNumber="18">5.5</td>
<td id="76F95DFEFFD50046279EFB34FB764FED" box="[1178,1256,1246,1268]" gridcol="7" gridrow="10" pageId="3" pageNumber="18">3.8</td>
<td id="76F95DFEFFD500462623FB34FA024FED" box="[1319,1436,1246,1268]" gridcol="8" gridrow="10" pageId="3" pageNumber="18">4.1</td>
</tr>
<tr id="35283482FFD500462393FAE2FA024E07" box="[151,1436,1288,1310]" gridrow="11" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FAE2FE884E07" box="[151,278,1288,1310]" gridcol="0" gridrow="11" pageId="3" pageNumber="18">PL</th>
<td id="76F95DFEFFD50046224FFAE2FE3D4E07" box="[331,419,1288,1310]" gridcol="1" gridrow="11" pageId="3" pageNumber="18">9.9</td>
<td id="76F95DFEFFD5004622E8FAE2FDA44E07" box="[492,570,1288,1310]" gridcol="2" gridrow="11" pageId="3" pageNumber="18">10.3</td>
<td id="76F95DFEFFD50046217DFAE2FD594E07" box="[633,711,1288,1310]" gridcol="3" gridrow="11" pageId="3" pageNumber="18">10.3</td>
<td id="76F95DFEFFD500462002FAE2FCD04E07" box="[774,846,1288,1310]" gridcol="4" gridrow="11" pageId="3" pageNumber="18">9.8</td>
<td id="76F95DFEFFD50046207BFAE2FC504E07" box="[895,974,1288,1310]" gridcol="5" gridrow="11" pageId="3" pageNumber="18">11.9</td>
<td id="76F95DFEFFD500462708FAE2FBCA4E07" box="[1036,1108,1288,1310]" gridcol="6" gridrow="11" pageId="3" pageNumber="18">10.9</td>
<td id="76F95DFEFFD50046279EFAE2FB764E07" box="[1178,1256,1288,1310]" gridcol="7" gridrow="11" pageId="3" pageNumber="18">6.8</td>
<td id="76F95DFEFFD500462623FAE2FA024E07" box="[1319,1436,1288,1310]" gridcol="8" gridrow="11" pageId="3" pageNumber="18">7.2</td>
</tr>
<tr id="35283482FFD500462393FAD8FA024E5E" box="[151,1436,1330,1351]" gridrow="12" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FAD8FE884E5E" box="[151,278,1330,1351]" gridcol="0" gridrow="12" pageId="3" pageNumber="18">F1L</th>
<td id="76F95DFEFFD50046224FFAD8FE3D4E5E" box="[331,419,1330,1351]" gridcol="1" gridrow="12" pageId="3" pageNumber="18">2.0</td>
<td id="76F95DFEFFD5004622E8FAD8FDA44E5E" box="[492,570,1330,1351]" gridcol="2" gridrow="12" pageId="3" pageNumber="18">2.0</td>
<td id="76F95DFEFFD50046217DFAD8FD594E5E" box="[633,711,1330,1351]" gridcol="3" gridrow="12" pageId="3" pageNumber="18">2.3</td>
<td id="76F95DFEFFD500462002FAD8FCD04E5E" box="[774,846,1330,1351]" gridcol="4" gridrow="12" pageId="3" pageNumber="18">2</td>
<td id="76F95DFEFFD50046207BFAD8FC504E5E" box="[895,974,1330,1351]" gridcol="5" gridrow="12" pageId="3" pageNumber="18">2.3</td>
<td id="76F95DFEFFD500462708FAD8FBCA4E5E" box="[1036,1108,1330,1351]" gridcol="6" gridrow="12" pageId="3" pageNumber="18">2.3</td>
<td id="76F95DFEFFD50046279EFAD8FB764E5E" box="[1178,1256,1330,1351]" gridcol="7" gridrow="12" pageId="3" pageNumber="18">1.1</td>
<td id="76F95DFEFFD500462623FAD8FA024E5E" box="[1319,1436,1330,1351]" gridcol="8" gridrow="12" pageId="3" pageNumber="18">1.4</td>
</tr>
<tr id="35283482FFD500462393FAB0FA024E69" box="[151,1436,1370,1392]" gridrow="13" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FAB0FE884E69" box="[151,278,1370,1392]" gridcol="0" gridrow="13" pageId="3" pageNumber="18">F2L</th>
<td id="76F95DFEFFD50046224FFAB0FE3D4E69" box="[331,419,1370,1392]" gridcol="1" gridrow="13" pageId="3" pageNumber="18">2.3</td>
<td id="76F95DFEFFD5004622E8FAB0FDA44E69" box="[492,570,1370,1392]" gridcol="2" gridrow="13" pageId="3" pageNumber="18">2.5</td>
<td id="76F95DFEFFD50046217DFAB0FD594E69" box="[633,711,1370,1392]" gridcol="3" gridrow="13" pageId="3" pageNumber="18">2.5</td>
<td id="76F95DFEFFD500462002FAB0FCD04E69" box="[774,846,1370,1392]" gridcol="4" gridrow="13" pageId="3" pageNumber="18">2.3</td>
<td id="76F95DFEFFD50046207BFAB0FC504E69" box="[895,974,1370,1392]" gridcol="5" gridrow="13" pageId="3" pageNumber="18">2.5</td>
<td id="76F95DFEFFD500462708FAB0FBCA4E69" box="[1036,1108,1370,1392]" gridcol="6" gridrow="13" pageId="3" pageNumber="18">2.5</td>
<td id="76F95DFEFFD50046279EFAB0FB764E69" box="[1178,1256,1370,1392]" gridcol="7" gridrow="13" pageId="3" pageNumber="18">1.3</td>
<td id="76F95DFEFFD500462623FAB0FA024E69" box="[1319,1436,1370,1392]" gridcol="8" gridrow="13" pageId="3" pageNumber="18">1.6</td>
</tr>
<tr id="35283482FFD500462393FA69FA024E80" box="[151,1436,1411,1433]" gridrow="14" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FA69FE884E80" box="[151,278,1411,1433]" gridcol="0" gridrow="14" pageId="3" pageNumber="18">F3L</th>
<td id="76F95DFEFFD50046224FFA69FE3D4E80" box="[331,419,1411,1433]" gridcol="1" gridrow="14" pageId="3" pageNumber="18">3.8</td>
<td id="76F95DFEFFD5004622E8FA69FDA44E80" box="[492,570,1411,1433]" gridcol="2" gridrow="14" pageId="3" pageNumber="18">4.1</td>
<td id="76F95DFEFFD50046217DFA69FD594E80" box="[633,711,1411,1433]" gridcol="3" gridrow="14" pageId="3" pageNumber="18">4.3</td>
<td id="76F95DFEFFD500462002FA69FCD04E80" box="[774,846,1411,1433]" gridcol="4" gridrow="14" pageId="3" pageNumber="18">4.3</td>
<td id="76F95DFEFFD50046207BFA69FC504E80" box="[895,974,1411,1433]" gridcol="5" gridrow="14" pageId="3" pageNumber="18">5.0</td>
<td id="76F95DFEFFD500462708FA69FBCA4E80" box="[1036,1108,1411,1433]" gridcol="6" gridrow="14" pageId="3" pageNumber="18">4.5</td>
<td id="76F95DFEFFD50046279EFA69FB764E80" box="[1178,1256,1411,1433]" gridcol="7" gridrow="14" pageId="3" pageNumber="18">2.7</td>
<td id="76F95DFEFFD500462623FA69FA024E80" box="[1319,1436,1411,1433]" gridcol="8" gridrow="14" pageId="3" pageNumber="18">2.9</td>
</tr>
<tr id="35283482FFD500462393FA47FA024EDA" box="[151,1436,1453,1475]" gridrow="15" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FA47FE884EDA" box="[151,278,1453,1475]" gridcol="0" gridrow="15" pageId="3" pageNumber="18">TIB:SVL</th>
<td id="76F95DFEFFD50046224FFA47FE3D4EDA" box="[331,419,1453,1475]" gridcol="1" gridrow="15" pageId="3" pageNumber="18">0.44</td>
<td id="76F95DFEFFD5004622E8FA47FDA44EDA" box="[492,570,1453,1475]" gridcol="2" gridrow="15" pageId="3" pageNumber="18">0.44</td>
<td id="76F95DFEFFD50046217DFA47FD594EDA" box="[633,711,1453,1475]" gridcol="3" gridrow="15" pageId="3" pageNumber="18">0.45</td>
<td id="76F95DFEFFD500462002FA47FCD04EDA" box="[774,846,1453,1475]" gridcol="4" gridrow="15" pageId="3" pageNumber="18">0.46</td>
<td id="76F95DFEFFD50046207BFA47FC504EDA" box="[895,974,1453,1475]" gridcol="5" gridrow="15" pageId="3" pageNumber="18">0.41</td>
<td id="76F95DFEFFD500462708FA47FBCA4EDA" box="[1036,1108,1453,1475]" gridcol="6" gridrow="15" pageId="3" pageNumber="18">0.39</td>
<td id="76F95DFEFFD50046279EFA47FB764EDA" box="[1178,1256,1453,1475]" gridcol="7" gridrow="15" pageId="3" pageNumber="18">0.44</td>
<td id="76F95DFEFFD500462623FA47FA024EDA" box="[1319,1436,1453,1475]" gridcol="8" gridrow="15" pageId="3" pageNumber="18">0.46</td>
</tr>
<tr id="35283482FFD500462393FA3CFA024EF5" box="[151,1436,1494,1516]" gridrow="16" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393FA3CFE884EF5" box="[151,278,1494,1516]" gridcol="0" gridrow="16" pageId="3" pageNumber="18">HDL:HDW</th>
<td id="76F95DFEFFD50046224FFA3CFE3D4EF5" box="[331,419,1494,1516]" gridcol="1" gridrow="16" pageId="3" pageNumber="18">1.04</td>
<td id="76F95DFEFFD5004622E8FA3CFDA44EF5" box="[492,570,1494,1516]" gridcol="2" gridrow="16" pageId="3" pageNumber="18">1.05</td>
<td id="76F95DFEFFD50046217DFA3CFD594EF5" box="[633,711,1494,1516]" gridcol="3" gridrow="16" pageId="3" pageNumber="18">1.04</td>
<td id="76F95DFEFFD500462002FA3CFCD04EF5" box="[774,846,1494,1516]" gridcol="4" gridrow="16" pageId="3" pageNumber="18">1.07</td>
<td id="76F95DFEFFD50046207BFA3CFC504EF5" box="[895,974,1494,1516]" gridcol="5" gridrow="16" pageId="3" pageNumber="18">1.06</td>
<td id="76F95DFEFFD500462708FA3CFBCA4EF5" box="[1036,1108,1494,1516]" gridcol="6" gridrow="16" pageId="3" pageNumber="18">1.07</td>
<td id="76F95DFEFFD50046279EFA3CFB764EF5" box="[1178,1256,1494,1516]" gridcol="7" gridrow="16" pageId="3" pageNumber="18">1.00</td>
<td id="76F95DFEFFD500462623FA3CFA024EF5" box="[1319,1436,1494,1516]" gridcol="8" gridrow="16" pageId="3" pageNumber="18">1.02</td>
</tr>
<tr id="35283482FFD500462393F9EAFA024D0F" box="[151,1436,1536,1558]" gridrow="17" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393F9EAFE884D0F" box="[151,278,1536,1558]" gridcol="0" gridrow="17" pageId="3" pageNumber="18">HDL:SVL</th>
<td id="76F95DFEFFD50046224FF9EAFE3D4D0F" box="[331,419,1536,1558]" gridcol="1" gridrow="17" pageId="3" pageNumber="18">0.36</td>
<td id="76F95DFEFFD5004622E8F9EAFDA44D0F" box="[492,570,1536,1558]" gridcol="2" gridrow="17" pageId="3" pageNumber="18">0.37</td>
<td id="76F95DFEFFD50046217DF9EAFD594D0F" box="[633,711,1536,1558]" gridcol="3" gridrow="17" pageId="3" pageNumber="18">0.39</td>
<td id="76F95DFEFFD500462002F9EAFCD04D0F" box="[774,846,1536,1558]" gridcol="4" gridrow="17" pageId="3" pageNumber="18">0.39</td>
<td id="76F95DFEFFD50046207BF9EAFC504D0F" box="[895,974,1536,1558]" gridcol="5" gridrow="17" pageId="3" pageNumber="18">0.34</td>
<td id="76F95DFEFFD500462708F9EAFBCA4D0F" box="[1036,1108,1536,1558]" gridcol="6" gridrow="17" pageId="3" pageNumber="18">0.36</td>
<td id="76F95DFEFFD50046279EF9EAFB764D0F" box="[1178,1256,1536,1558]" gridcol="7" gridrow="17" pageId="3" pageNumber="18">0.31</td>
<td id="76F95DFEFFD500462623F9EAFA024D0F" box="[1319,1436,1536,1558]" gridcol="8" gridrow="17" pageId="3" pageNumber="18">0.33</td>
</tr>
<tr id="35283482FFD500462393F9C3FA024D59" box="[151,1436,1577,1600]" gridrow="18" pageId="3" pageNumber="18">
<th id="76F95DFEFFD500462393F9C3FE884D59" box="[151,278,1577,1600]" gridcol="0" gridrow="18" pageId="3" pageNumber="18">Weight (g)</th>
<td id="76F95DFEFFD50046224FF9C3FE3D4D59" box="[331,419,1577,1600]" gridcol="1" gridrow="18" pageId="3" pageNumber="18">1.1</td>
<td id="76F95DFEFFD5004622E8F9C3FDA44D59" box="[492,570,1577,1600]" gridcol="2" gridrow="18" pageId="3" pageNumber="18">1.2</td>
<td id="76F95DFEFFD50046217DF9C3FD594D59" box="[633,711,1577,1600]" gridcol="3" gridrow="18" pageId="3" pageNumber="18">1.4</td>
<td id="76F95DFEFFD500462002F9C3FCD04D59" box="[774,846,1577,1600]" gridcol="4" gridrow="18" pageId="3" pageNumber="18">1.2</td>
<td id="76F95DFEFFD50046207BF9C3FC504D59" box="[895,974,1577,1600]" gridcol="5" gridrow="18" pageId="3" pageNumber="18">2.2</td>
<td id="76F95DFEFFD500462708F9C3FBCA4D59" box="[1036,1108,1577,1600]" gridcol="6" gridrow="18" pageId="3" pageNumber="18">1.4</td>
<td id="76F95DFEFFD50046279EF9C3FB764D59" box="[1178,1256,1577,1600]" gridcol="7" gridrow="18" pageId="3" pageNumber="18">0.5</td>
<td id="76F95DFEFFD500462623F9C3FA024D59" box="[1319,1436,1577,1600]" gridcol="8" gridrow="18" pageId="3" pageNumber="18">0.6</td>
</tr>
<tr id="35283482FFD500462393F98AFA024D6F" box="[151,1436,1632,1654]" gridrow="19" pageId="3" pageNumber="18" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76F95DFEFFD500462393F98AFE884D6F" box="[151,278,1632,1654]" gridcol="0" gridrow="19" pageId="3" pageNumber="18">*holotype.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BA736C0FFD5FFBD23C1F977FBA74BEE" blockId="3.[151,1436,1693,2005]" lastBlockId="4.[151,1437,151,248]" lastPageId="4" lastPageNumber="19" pageId="3" pageNumber="18">
<emphasis id="B96CEAD2FFD5FFBA23C1F977FE624DAF" bold="true" box="[197,508,1693,1718]" pageId="3" pageNumber="18">
Colour of
<typeStatus id="54A38862FFD5FFBA2245F977FE364DAC" box="[321,424,1693,1717]" pageId="3" pageNumber="18" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dorsal surface brown with distinct darker brown markings; V-shaped interorbital marking, W-shaped marking between axillae and inverted V-shaped, irregular marking above sacrum; fine, pale greyish-blue tubercles on upper eyelids, snout, lateral margins of dorsum and upper surfaces of legs; blackish brown patch on upper lip under eye; blackish brown line along canthus rostralis through eye, and continuing below supratympanic ridge, terminating above axilla, encompassing nare and most of tympanum; transverse dark brown bars on dorsal surface of limbs; large, black blotch on posterior flank joining with inverted V-shaped marking above sacrum and numerous, smaller black spots on sides from groin to axilla; elbow and upper arms without dark bars but with distinct copper colouration; fingers and toes with transverse barring. Dark brownish red ventral surface with white speckling on entire ventral surface including throat, arms and legs, forming distinct marbling on chest and belly; ventral margin of throat bearing broken row of slightly larger white spots. Supra-axillary gland copper; femoral glands white; pectoral glands white, lined with dark brown. Iris bicoloured, with upper half coppery red, fading to pale silver in lower half; minute, black reticulations throughout.
</paragraph>
<caption id="DF676648FFD2FFBD2393FA6EFC5C4EA0" httpUri="https://zenodo.org/record/277017/files/figure.png" pageId="4" pageNumber="19" targetBox="[151,1436,298,1391]" targetPageId="4">
<paragraph id="8BA736C0FFD2FFBD2393FA6EFC5C4EA0" blockId="4.[151,1436,1412,1466]" pageId="4" pageNumber="19">
<emphasis id="B96CEAD2FFD2FFBD2393FA6EFE8D4E83" bold="true" box="[151,275,1412,1435]" pageId="4" pageNumber="19">FIGURE 1.</emphasis>
Collection site of
<taxonomicName id="4C184D43FFD2FFBD22CAFA6FFD244E82" box="[462,698,1413,1435]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="19" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD2FFBD22CAFA6FFD244E82" box="[462,698,1413,1435]" italics="true" pageId="4" pageNumber="19">Leptolalax bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD2FFBD21C5FA6FFC8F4E83" box="[705,785,1413,1434]" pageId="4" pageNumber="19" rank="species">
<emphasis id="B96CEAD2FFD2FFBD21C5FA6FFC8F4E83" bold="true" box="[705,785,1413,1434]" pageId="4" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
from Bidoup-Nui Ba National Park (black star) and type localities of
<taxonomicName id="4C184D43FFD2FFBD23D8FA4EFE3B4EA3" box="[220,421,1444,1466]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="19" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFD2FFBD23D8FA4EFE3B4EA3" box="[220,421,1444,1466]" italics="true" pageId="4" pageNumber="19">Leptolalax applebyi</emphasis>
</taxonomicName>
(white circle) and
<taxonomicName id="4C184D43FFD2FFBD2160FA4EFCBA4EA3" box="[612,804,1444,1466]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="19" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFD2FFBD2160FA4EFCBA4EA3" box="[612,804,1444,1466]" italics="true" pageId="4" pageNumber="19">Leptolalax melicus</emphasis>
</taxonomicName>
(black circles).
</paragraph>
</caption>
<paragraph id="8BA736C0FFD2FFBD23C1FA0CFAA64D3B" blockId="4.[151,1436,1510,2002]" pageId="4" pageNumber="19">
<emphasis id="B96CEAD2FFD2FFBD23C1FA0CFDEF4EE6" bold="true" box="[197,625,1510,1535]" pageId="4" pageNumber="19">
Colour of
<typeStatus id="54A38862FFD2FFBD2240FA0DFE354EE6" box="[324,427,1511,1535]" pageId="4" pageNumber="19" type="holotype">holotype</typeStatus>
in preservative.
</emphasis>
Dorsum dark brown with slightly paler limbs. Ventral surface brown, with white speckling and marbling. Macroglands white. Patterns on dorsal surface less distinct (
<figureCitation id="13232A45FFD2FFBD27CDF9E0FAB44D3B" box="[1225,1322,1546,1570]" captionStart="FIGURE 3" captionStartId="6.[151,250,1434,1457]" captionTargetBox="[159,1428,207,1398]" captionTargetId="figure@6.[151,1436,193,1413]" captionTargetPageId="6" captionText="FIGURE 3. (A) Dorsal, (B) ventral and (C) lateral view of preserved holotype (AMS R 173133) of L. bidoupensis sp. nov. Scale bars = 5 mm." httpUri="https://zenodo.org/record/277019/files/figure.png" pageId="4" pageNumber="19">Figure 3</figureCitation>
).
</paragraph>
<paragraph id="8BA736C0FFD2FFBD23C1F9C5FCE34D73" blockId="4.[151,1436,1510,2002]" pageId="4" pageNumber="19">
<emphasis id="B96CEAD2FFD2FFBD23C1F9C5FEE54D5E" bold="true" box="[197,379,1583,1607]" pageId="4" pageNumber="19">Measurements.</emphasis>
<typeStatus id="54A38862FFD2FFBD2280F9C4FE6F4D5F" box="[388,497,1582,1606]" pageId="4" pageNumber="19" type="holotype">Holotype</typeStatus>
: SVL 23.6, HDL 9.2, HDW 8.9, SNT 3.6, EYE 2.5, IOD 2.9, TMP 1.4, TEY 1.3, TIB 10.4, ML 5.2, PL 9.9, F1L 2.0, F2L 2.3, F3L 3.8, weight
<quantity id="4CE09B25FFD2FFBD2048F9B8FCE34D73" box="[844,893,1618,1642]" metricMagnitude="-3" metricUnit="kg" metricValue="1.1" pageId="4" pageNumber="19" unit="g" value="1.1">1.1g</quantity>
</paragraph>
<paragraph id="8BA736C0FFD2FFBC23C1F99CFF244A7D" blockId="4.[151,1436,1510,2002]" lastBlockId="5.[151,1436,151,356]" lastPageId="5" lastPageNumber="20" pageId="4" pageNumber="19">
<emphasis id="B96CEAD2FFD2FFBD23C1F99CFEA74D96" bold="true" box="[197,313,1654,1679]" pageId="4" pageNumber="19">Variation.</emphasis>
Specimens vary slightly in colour in life. AMS R 173135 and NCSM 77320 have a paler brown dorsum compared to the
<typeStatus id="54A38862FFD2FFBD22AAF970FD8D4DAB" box="[430,531,1690,1714]" pageId="4" pageNumber="19" type="holotype">holotype</typeStatus>
, and dorsal patterns lined with diffuse, paler brown. AMS R 173136 and NCSM 77321 also have paler brown dorsal surfaces, but with only indistinct, diffuse dorsal markings; in NCSM 77321 the only distinct dark markings are over the tympanum, below the supratympanic ridge and as a single patch on the flank; AMS R 173136 has larger and more numerous brownish black patches on the flanks. AMS R 173134 and UNS 00101/AMS R 173135 have more mottled dorsal patterns. The ventral surfaces of specimens in the
<typeStatus id="54A38862FFD2FFBD2622F8C1FAC84C5A" box="[1318,1366,1835,1859]" pageId="4" pageNumber="19">type</typeStatus>
series vary in their degree of white speckling and in the presence or absence of white marbling on parts of the chest and belly; UNS 00101/AMS R 173135 and NCSM 7732077321 have less intense white speckling on ventral surfaces and only slight white marbling on parts of their chest; AMS R 173133 has slightly less intense white speckling and no white marbling. Pectoral glands are more distinct in specimens with less marbling. Ventrolateral glands are most easily detected in the
<typeStatus id="54A38862FFD3FFBC2297FF7DFE6B4BB6" box="[403,501,151,175]" pageId="5" pageNumber="20" type="holotype">holotype</typeStatus>
AMS R 173133, and more widely spaced and difficult to distinguish from white ventral patterning on all other specimens in the
<typeStatus id="54A38862FFD3FFBC21C7FF57FD6D4BCC" box="[707,755,189,213]" pageId="5" pageNumber="20">type</typeStatus>
series. UNS 00102/AMS R 173137 and NCSM 77322 are metamorphs with incompletely reabsorbed tails (tail length
<quantity id="4CE09B25FFD3FFBC204CFF35FC124BEE" box="[840,908,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="20" unit="mm" value="2.0">2 mm</quantity>
in UNS 00101/AMS R 173137 and
<quantity id="4CE09B25FFD3FFBC263CFF35FAE24BEE" box="[1336,1404,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="20" unit="mm" value="8.0">8 mm</quantity>
in NCSM 77322), almost uniformly dark brown dorsal surfaces, dark brown ventral surfaces with white speckling, and indistinct tympana. Iris colouration varies slightly in the balance of coppery red versus pale silver (
<figureCitation id="13232A45FFD3FFBC261EFECDFAE64A26" box="[1306,1400,295,319]" captionStart="FIGURE 5" captionStartId="11.[151,250,1213,1236]" captionTargetBox="[152,1436,339,1191]" captionTargetId="figure@11.[151,1436,339,1192]" captionTargetPageId="11" captionText="FIGURE 5. Iris colouration in life of (A) Leptolalax bidoupensis sp. nov. NCSM 77320, (B) Leptolalax bidoupensis sp. nov. AMS R 173134, (C) Leptolalax bidoupensis sp. nov. AMS R 173133, (D E) Leptolalax applebyi, and (F) Leptolalax melicus MVZ 258199." httpUri="https://zenodo.org/record/277021/files/figure.png" pageId="5" pageNumber="20">Figure 5</figureCitation>
A C).
</paragraph>
<caption id="DF676648FFD3FFBC2393F851FA984CE9" httpUri="https://zenodo.org/record/277018/files/figure.png" pageId="5" pageNumber="20" targetBox="[151,1436,409,1957]" targetPageId="5">
<paragraph id="8BA736C0FFD3FFBC2393F851FA984CE9" blockId="5.[151,1436,1979,2033]" pageId="5" pageNumber="20">
<emphasis id="B96CEAD2FFD3FFBC2393F851FE8A4CC8" bold="true" box="[151,276,1979,2002]" pageId="5" pageNumber="20">FIGURE 2.</emphasis>
(A) Lateral view of male holotype (AMS R 173133) of
<taxonomicName id="4C184D43FFD3FFBC205AF857FC6D4CCB" box="[862,1011,1980,2002]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="20" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD3FFBC205AF857FC6D4CCB" box="[862,1011,1980,2002]" italics="true" pageId="5" pageNumber="20">L. bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD3FFBC20FFF856FBD34CCB" box="[1019,1101,1980,2002]" pageId="5" pageNumber="20" rank="species">
<emphasis id="B96CEAD2FFD3FFBC20FFF856FBD94CC8" bold="true" box="[1019,1095,1980,2001]" pageId="5" pageNumber="20">sp. nov</emphasis>
.
</taxonomicNameLabel>
in life, and (B) ventral views of gravid female paratype (AMS R 173134) and male holotype (AMS R 173133) of
<taxonomicName id="4C184D43FFD3FFBC20D6F836FBF84CE8" box="[978,1126,2011,2033]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="20" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD3FFBC20D6F836FBF84CE8" box="[978,1126,2011,2033]" italics="true" pageId="5" pageNumber="20">L. bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD3FFBC2768F831FB234CE9" box="[1132,1213,2011,2032]" pageId="5" pageNumber="20" rank="species">
<emphasis id="B96CEAD2FFD3FFBC2768F831FB234CE9" bold="true" box="[1132,1213,2011,2032]" pageId="5" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
in life.
</paragraph>
</caption>
<caption id="DF676648FFD0FFBF2393FA70FEC54ED6" httpUri="https://zenodo.org/record/277019/files/figure.png" pageId="6" pageNumber="21" targetBox="[159,1428,207,1398]" targetPageId="6">
<paragraph id="8BA736C0FFD0FFBF2393FA70FEC54ED6" blockId="6.[151,1435,1434,1487]" pageId="6" pageNumber="21">
<emphasis id="B96CEAD2FFD0FFBF2393FA70FE8A4EA9" bold="true" box="[151,276,1434,1457]" pageId="6" pageNumber="21">FIGURE 3.</emphasis>
(A) Dorsal, (B) ventral and (C) lateral view of preserved holotype (AMS R 173133) of
<taxonomicName id="4C184D43FFD0FFBF27A8FA71FADF4EA9" box="[1196,1345,1434,1456]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="21" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD0FFBF27A8FA71FADF4EA9" box="[1196,1345,1434,1456]" italics="true" pageId="6" pageNumber="21">L. bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD0FFBF264DFA71FA054EA9" box="[1353,1435,1435,1456]" pageId="6" pageNumber="21" rank="species">
<emphasis id="B96CEAD2FFD0FFBF264DFA71FA084EA9" bold="true" box="[1353,1430,1435,1456]" pageId="6" pageNumber="21">sp. nov</emphasis>
.
</taxonomicNameLabel>
Scale bars = 5 mm.
</paragraph>
</caption>
<paragraph id="8BA736C0FFD0FFBF23C1FA11FE314C65" blockId="6.[151,1437,1531,1916]" pageId="6" pageNumber="21">
<emphasis id="B96CEAD2FFD0FFBF23C1FA11FE334D0D" bold="true" box="[197,429,1531,1556]" pageId="6" pageNumber="21">Advertisement call.</emphasis>
Call descriptions are based on the calls of the
<typeStatus id="54A38862FFD0FFBF20BDFA11FB804D0A" box="[953,1054,1531,1555]" pageId="6" pageNumber="21" type="holotype">holotype</typeStatus>
, taken at 19.0 ºC ambient temperature. Calls were an average of 479 ms in duration and consisted of 69 (usually 7) singly pulsed notes, or clicks, of variable (113 ms) duration, repeated at a rate of 1821 notes per second (
<tableCitation id="C69A037BFFD0FFBF20F8F9A9FBCB4D42" box="[1020,1109,1603,1627]" captionStart="TABLE 2" captionStartId="7.[151,235,152,173]" captionTargetBox="[151,1436,232,824]" captionTargetPageId="7" captionText="TABLE 2. Measurements of advertisement call parameters for Leptolalax bidoupensis sp. nov. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/DF676648FFD1FFBE2393FF72FEFD4BD5" pageId="6" pageNumber="21" tableUuid="DF676648FFD1FFBE2393FF72FEFD4BD5">Table 2</tableCitation>
,
<figureCitation id="13232A45FFD0FFBF2760F9A9FB4F4D42" box="[1124,1233,1603,1627]" captionStart="FIGURE 4" captionStartId="7.[151,250,1879,1902]" captionTargetBox="[157,1436,913,1853]" captionTargetId="figure@7.[151,1436,905,1858]" captionTargetPageId="7" captionText="FIGURE 4. Advertisement call of Leptolalax bidoupensis sp. nov. (holotype AMS R 173133) recorded at ambient air temperature of 19.0 º C. (A) 48 s waveform of relative amplitude over time for 11 calls, (B) waveform and (C) corresponding spectrogram of single representative call containing eight notes, expanded from section shown in A, and (D) power spectrum (relative amplitude vs. frequency) of fourth note shown in B C." httpUri="https://zenodo.org/record/277020/files/figure.png" pageId="6" pageNumber="21">Figures 4</figureCitation>
A, C). Calls were highly amplitude modulated, with amplitude peaking towards the middle of the call. The dominant frequency was 2.32.6 kHz, and harmonics were weak or absent. Slight frequency modulation was present within each call, with lower amplitude notes, particularly the last note, often slightly higher or lower frequency. Calls were repeated at a rate of approximately 0.2 calls per second, and had an average intercall interval of 4 s. Call repetition rate, intercall interval, and the number of notes per call varied slightly within calling bouts and among individuals (
<tableCitation id="C69A037BFFD0FFBF260EF91DFAFF4C16" box="[1290,1377,1783,1807]" captionStart="TABLE 2" captionStartId="7.[151,235,152,173]" captionTargetBox="[151,1436,232,824]" captionTargetPageId="7" captionText="TABLE 2. Measurements of advertisement call parameters for Leptolalax bidoupensis sp. nov. Parameter values are given as means (and ranges)." httpUri="http://table.plazi.org/id/DF676648FFD1FFBE2393FF72FEFD4BD5" pageId="6" pageNumber="21" tableUuid="DF676648FFD1FFBE2393FF72FEFD4BD5">Table 2</tableCitation>
, Figure 4). To the human ear, the advertisement call of
<taxonomicName id="4C184D43FFD0FFBF21D7F8F7FCE94C2D" box="[723,887,1820,1844]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="21" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFD0FFBF21D7F8F7FCE94C2D" box="[723,887,1820,1844]" italics="true" pageId="6" pageNumber="21">L. bidoupensis</emphasis>
</taxonomicName>
is a slow rasping, similar to an orthopteran, with each note distinctly discernable. The call is extremely faint and difficult to locate, even when compared to other species within the genus.
</paragraph>
<caption id="DF676648FFD1FFBE2393FF72FEFD4BD5" ID-Table-UUID="DF676648FFD1FFBE2393FF72FEFD4BD5" httpUri="http://table.plazi.org/id/DF676648FFD1FFBE2393FF72FEFD4BD5" pageId="7" pageNumber="22" targetBox="[151,1436,232,824]" targetIsTable="true" targetPageId="7">
<paragraph id="8BA736C0FFD1FFBE2393FF72FEFD4BD5" blockId="7.[151,1436,152,204]" pageId="7" pageNumber="22">
<emphasis id="B96CEAD2FFD1FFBE2393FF72FE9B4BB4" bold="true" box="[151,261,152,173]" pageId="7" pageNumber="22">TABLE 2.</emphasis>
Measurements of advertisement call parameters for
<taxonomicName id="4C184D43FFD1FFBE201BFF72FB924BB7" box="[799,1036,152,174]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="22" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD1FFBE201BFF72FB924BB7" box="[799,1036,152,174]" italics="true" pageId="7" pageNumber="22">Leptolalax bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFD1FFBE2717FF72FBFA4BB4" box="[1043,1124,152,173]" pageId="7" pageNumber="22" rank="species">
<emphasis id="B96CEAD2FFD1FFBE2717FF72FBFA4BB4" bold="true" box="[1043,1124,152,173]" pageId="7" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
Parameter values are given as means (and ranges).
</paragraph>
</caption>
<paragraph id="8BA736C0FFD1FFBE22E5FF02FF624821" pageId="7" pageNumber="22">
<table id="F918C460FFD100462393FF02FA024821" box="[151,1436,232,824]" gridcols="4" gridrows="13" pageId="7" pageNumber="22">
<tr id="35283482FFD100462393FF02FA024A05" box="[151,1436,232,284]" gridrow="0" pageId="7" pageNumber="22" rowspan-0="1">
<th id="76F95DFEFFD1004622E4FF02FCDC4A05" box="[480,834,232,284]" gridcol="1" gridrow="0" pageId="7" pageNumber="22">AMS R 173133* UNS 00101/ AMS R 173135</th>
<th id="76F95DFEFFD100462066FF02FB9C4A05" box="[866,1026,232,284]" gridcol="2" gridrow="0" pageId="7" pageNumber="22">AMS R 173136</th>
<th id="76F95DFEFFD100462727FF02FA024A05" box="[1059,1436,232,284]" gridcol="3" gridrow="0" pageId="7" pageNumber="22">Non-vouchered 1 Non-vouchered 2</th>
</tr>
<tr id="35283482FFD100462393FEDAFA024A5F" box="[151,1436,304,326]" gridrow="1" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FEDAFE5F4A5F" box="[151,449,304,326]" gridcol="0" gridrow="1" pageId="7" pageNumber="22">Number of calls</th>
<td id="76F95DFEFFD1004622E4FEDAFCDC4A5F" box="[480,834,304,326]" gridcol="1" gridrow="1" pageId="7" pageNumber="22">10 10</td>
<td id="76F95DFEFFD100462066FEDAFB9C4A5F" box="[866,1026,304,326]" gridcol="2" gridrow="1" pageId="7" pageNumber="22">10</td>
<td id="76F95DFEFFD100462727FEDAFA024A5F" box="[1059,1436,304,326]" gridcol="3" gridrow="1" pageId="7" pageNumber="22">8 10</td>
</tr>
<tr id="35283482FFD100462393FEB3FA024A76" box="[151,1436,345,367]" gridrow="2" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FEB3FE5F4A76" box="[151,449,345,367]" gridcol="0" gridrow="2" pageId="7" pageNumber="22">Number of notes</th>
<td id="76F95DFEFFD1004622E4FEB3FCDC4A76" box="[480,834,345,367]" gridcol="1" gridrow="2" pageId="7" pageNumber="22">85 53</td>
<td id="76F95DFEFFD100462066FEB3FB9C4A76" box="[866,1026,345,367]" gridcol="2" gridrow="2" pageId="7" pageNumber="22">67</td>
<td id="76F95DFEFFD100462727FEB3FA024A76" box="[1059,1436,345,367]" gridcol="3" gridrow="2" pageId="7" pageNumber="22">50 72</td>
</tr>
<tr id="35283482FFD100462393FE68FA024A80" box="[151,1436,386,409]" gridrow="3" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FE68FE5F4A80" box="[151,449,386,409]" gridcol="0" gridrow="3" pageId="7" pageNumber="22">Call duration (ms)</th>
<td id="76F95DFEFFD1004622E4FE68FCDC4A80" box="[480,834,386,409]" gridcol="1" gridrow="3" pageId="7" pageNumber="22">479 (326 543) 320 (278 343)</td>
<td id="76F95DFEFFD100462066FE68FB9C4A80" box="[866,1026,386,409]" gridcol="2" gridrow="3" pageId="7" pageNumber="22">334 (234 391)</td>
<td id="76F95DFEFFD100462727FE68FA024A80" box="[1059,1436,386,409]" gridcol="3" gridrow="3" pageId="7" pageNumber="22">394 (380 411) 401 (277 463)</td>
</tr>
<tr id="35283482FFD100462393FE41FA024ADB" box="[151,1436,427,450]" gridrow="4" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FE41FE5F4ADB" box="[151,449,427,450]" gridcol="0" gridrow="4" pageId="7" pageNumber="22">Call repetition rate (calls/s)</th>
<td id="76F95DFEFFD1004622E4FE41FCDC4ADB" box="[480,834,427,450]" gridcol="1" gridrow="4" pageId="7" pageNumber="22">0.22 0.17</td>
<td id="76F95DFEFFD100462066FE41FB9C4ADB" box="[866,1026,427,450]" gridcol="2" gridrow="4" pageId="7" pageNumber="22">0.13</td>
<td id="76F95DFEFFD100462727FE41FA024ADB" box="[1059,1436,427,450]" gridcol="3" gridrow="4" pageId="7" pageNumber="22">0.14 0.1</td>
</tr>
<tr id="35283482FFD100462393FE3FFA024913" box="[151,1436,469,522]" gridrow="5" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FE3FFE5F4913" box="[151,449,469,522]" gridcol="0" gridrow="5" pageId="7" pageNumber="22">Intercall interval (ms)</th>
<td id="76F95DFEFFD1004622E4FE3FFCDC4913" box="[480,834,469,522]" gridcol="1" gridrow="5" pageId="7" pageNumber="22">4014 (3590 5537 (4393 4521) 7430)</td>
<td id="76F95DFEFFD100462066FE3FFB9C4913" box="[866,1026,469,522]" gridcol="2" gridrow="5" pageId="7" pageNumber="22">7071 (4984 13106)</td>
<td id="76F95DFEFFD100462727FE3FFA024913" box="[1059,1436,469,522]" gridcol="3" gridrow="5" pageId="7" pageNumber="22">6689 9511 (5153 8492) (7560 12284)</td>
</tr>
<tr id="35283482FFD100462393FDF7FA02492A" box="[151,1436,541,563]" gridrow="6" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FDF7FE5F492A" box="[151,449,541,563]" gridcol="0" gridrow="6" pageId="7" pageNumber="22">Notes/call</th>
<td id="76F95DFEFFD1004622E4FDF7FCDC492A" box="[480,834,541,563]" gridcol="1" gridrow="6" pageId="7" pageNumber="22">8.2 (6 9) 6.7 (6 7)</td>
<td id="76F95DFEFFD100462066FDF7FB9C492A" box="[866,1026,541,563]" gridcol="2" gridrow="6" pageId="7" pageNumber="22">6.7 (5 8)</td>
<td id="76F95DFEFFD100462727FDF7FA02492A" box="[1059,1436,541,563]" gridcol="3" gridrow="6" pageId="7" pageNumber="22">7 (7) 7.2 (5 8)</td>
</tr>
<tr id="35283482FFD100462393FDACFA024944" box="[151,1436,582,605]" gridrow="7" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FDACFE5F4944" box="[151,449,582,605]" gridcol="0" gridrow="7" pageId="7" pageNumber="22">Note duration (ms)</th>
<td id="76F95DFEFFD1004622E4FDACFCDC4944" box="[480,834,582,605]" gridcol="1" gridrow="7" pageId="7" pageNumber="22">8.4 (1 13) 6.3 (2 9)</td>
<td id="76F95DFEFFD100462066FDACFB9C4944" box="[866,1026,582,605]" gridcol="2" gridrow="7" pageId="7" pageNumber="22">8.3 (4 11)</td>
<td id="76F95DFEFFD100462727FDACFA024944" box="[1059,1436,582,605]" gridcol="3" gridrow="7" pageId="7" pageNumber="22">7.9 (4 12) 7.4 (4 11)</td>
</tr>
<tr id="35283482FFD100462393FD9AFA02499F" box="[151,1436,624,646]" gridrow="8" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FD9AFE5F499F" box="[151,449,624,646]" gridcol="0" gridrow="8" pageId="7" pageNumber="22">Internote interval (ms)</th>
<td id="76F95DFEFFD1004622E4FD9AFCDC499F" box="[480,834,624,646]" gridcol="1" gridrow="8" pageId="7" pageNumber="22">55 (18 70) 49 (42 59)</td>
<td id="76F95DFEFFD100462066FD9AFB9C499F" box="[866,1026,624,646]" gridcol="2" gridrow="8" pageId="7" pageNumber="22">49 (9 76)</td>
<td id="76F95DFEFFD100462727FD9AFA02499F" box="[1059,1436,624,646]" gridcol="3" gridrow="8" pageId="7" pageNumber="22">56 (46 76) 55 (22 66)</td>
</tr>
<tr id="35283482FFD100462393FD73FA0249A9" box="[151,1436,665,688]" gridrow="9" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FD73FE5F49A9" box="[151,449,665,688]" gridcol="0" gridrow="9" pageId="7" pageNumber="22">Note repetition rate (notes/s)</th>
<td id="76F95DFEFFD1004622E4FD73FCDC49A9" box="[480,834,665,688]" gridcol="1" gridrow="9" pageId="7" pageNumber="22">18 21 21</td>
<td id="76F95DFEFFD100462066FD73FB9C49A9" box="[866,1026,665,688]" gridcol="2" gridrow="9" pageId="7" pageNumber="22">14 21</td>
<td id="76F95DFEFFD100462727FD73FA0249A9" box="[1059,1436,665,688]" gridcol="3" gridrow="9" pageId="7" pageNumber="22">18 18</td>
</tr>
<tr id="35283482FFD100462393FD28FA0249C0" box="[151,1436,706,729]" gridrow="10" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FD28FE5F49C0" box="[151,449,706,729]" gridcol="0" gridrow="10" pageId="7" pageNumber="22">Dominant frequency (kHz)</th>
<td id="76F95DFEFFD1004622E4FD28FCDC49C0" box="[480,834,706,729]" gridcol="1" gridrow="10" pageId="7" pageNumber="22">2.1(2.3 2.6) 2.1 (2.1 3.6)</td>
<td id="76F95DFEFFD100462066FD28FB9C49C0" box="[866,1026,706,729]" gridcol="2" gridrow="10" pageId="7" pageNumber="22">2.5 (1.9 3.4)</td>
<td id="76F95DFEFFD100462727FD28FA0249C0" box="[1059,1436,706,729]" gridcol="3" gridrow="10" pageId="7" pageNumber="22">2.0 (1.9 2.3) 2.5 (2.3 3.8)</td>
</tr>
<tr id="35283482FFD100462393FD06FA02481B" box="[151,1436,748,770]" gridrow="11" pageId="7" pageNumber="22">
<th id="76F95DFEFFD100462393FD06FE5F481B" box="[151,449,748,770]" gridcol="0" gridrow="11" pageId="7" pageNumber="22">Temperature (°C)</th>
<td id="76F95DFEFFD1004622E4FD06FCDC481B" box="[480,834,748,770]" gridcol="1" gridrow="11" pageId="7" pageNumber="22">19.0 20.4</td>
<td id="76F95DFEFFD100462066FD06FB9C481B" box="[866,1026,748,770]" gridcol="2" gridrow="11" pageId="7" pageNumber="22">21.0</td>
<td id="76F95DFEFFD100462727FD06FA02481B" box="[1059,1436,748,770]" gridcol="3" gridrow="11" pageId="7" pageNumber="22">20.2 20.4</td>
</tr>
<tr id="35283482FFD100462393FCC8FA024821" box="[151,1436,802,824]" gridrow="12" pageId="7" pageNumber="22" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76F95DFEFFD100462393FCC8FE5F4821" box="[151,449,802,824]" gridcol="0" gridrow="12" pageId="7" pageNumber="22">*holotype</th>
</tr>
</table>
</paragraph>
<caption id="DF676648FFD1FFBE2393F8BDFD5D4CD0" httpUri="https://zenodo.org/record/277020/files/figure.png" pageId="7" pageNumber="22" targetBox="[157,1436,913,1853]" targetPageId="7">
<paragraph id="8BA736C0FFD1FFBE2393F8BDFD5D4CD0" blockId="7.[151,1437,1879,1993]" pageId="7" pageNumber="22">
<emphasis id="B96CEAD2FFD1FFBE2393F8BDFE8D4C74" bold="true" box="[151,275,1879,1902]" pageId="7" pageNumber="22">FIGURE 4.</emphasis>
Advertisement call of
<taxonomicName id="4C184D43FFD1FFBE22F8F8BDFD794C74" box="[508,743,1879,1901]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="22" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFD1FFBE22F8F8BDFD794C74" box="[508,743,1879,1901]" italics="true" pageId="7" pageNumber="22">Leptolalax bidoupensis</emphasis>
</taxonomicName>
<emphasis id="B96CEAD2FFD1FFBE21EAF8B2FCA14C74" bold="true" box="[750,831,1880,1901]" pageId="7" pageNumber="22">
<taxonomicNameLabel id="A25F57A9FFD1FFBE21EAF8B2FCA14C74" box="[750,831,1880,1901]" pageId="7" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype AMS R 173133) recorded at ambient air temperature of 19.0º C. (A) 48 s waveform of relative amplitude over time for 11 calls, (B) waveform and (C) corresponding spectrogram of single representative call containing eight notes, expanded from section shown in A, and (D) power spectrum (relative amplitude vs. frequency) of fourth note shown in BC.
</paragraph>
</caption>
<paragraph id="8BA736C0FFDEFFB123C1FF7DFB754A9E" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
<emphasis id="B96CEAD2FFDEFFB123C1FF7DFE544BA9" bold="true" box="[197,458,151,176]" pageId="8" pageNumber="23">Sequence divergence.</emphasis>
Uncorrected sequence divergences between
<taxonomicName id="4C184D43FFDEFFB120DBFF73FB164BA9" box="[991,1160,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB120DBFF73FB164BA9" box="[991,1160,152,176]" italics="true" pageId="8" pageNumber="23">L. bidoupensis</emphasis>
</taxonomicName>
(NCSM 7732077321, AMS R 173133173134) and
<taxonomicName id="4C184D43FFDEFFB122F3FF57FDE64BCD" box="[503,632,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDEFFB122F3FF57FDE64BCD" box="[503,632,188,212]" italics="true" pageId="8" pageNumber="23">L. applebyi</emphasis>
</taxonomicName>
(
<typeStatus id="54A38862FFDEFFB1218DFF56FD734BCD" box="[649,749,188,212]" pageId="8" pageNumber="23" type="holotype">holotype</typeStatus>
AMS R 171703,
<typeStatus id="54A38862FFDEFFB120C4FF57FBBE4BCC" box="[960,1056,189,213]" pageId="8" pageNumber="23" type="paratype">paratype</typeStatus>
AMS R 171704) collected from approximately
<quantity id="4CE09B25FFDEFFB12244FF35FE0A4BEE" box="[320,404,223,247]" metricMagnitude="5" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="23" unit="km" value="350.0">350 km</quantity>
away were 9.3% at the 16S rRNA gene. Uncorrected sequence divergences between
<taxonomicName id="4C184D43FFDEFFB12655FF0BFF674A05" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB12655FF0BFF674A05" italics="true" pageId="8" pageNumber="23">L. bidoupensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D43FFDEFFB1222BFEEFFE3D4A05" box="[303,419,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDEFFB1222BFEEFFE3D4A05" box="[303,419,260,284]" italics="true" pageId="8" pageNumber="23">L. melicus</emphasis>
</taxonomicName>
(MVZ 258197258199) collected from approximately
<quantity id="4CE09B25FFDEFFB12714FEEEFBFA4A05" box="[1040,1124,260,284]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="23" unit="km" value="300.0">300 km</quantity>
away were 9.6% at the 16S rRNA gene. This degree of pairwise divergence in the 16S rRNA gene in frogs usually represents differentiation at the species level (Vences
<emphasis id="B96CEAD2FFDEFFB122B8FEA7FE694A7D" box="[444,503,332,356]" italics="true" pageId="8" pageNumber="23">et al.</emphasis>
2005), and is greater than that between
<taxonomicName id="4C184D43FFDEFFB120C2FEA7FBD84A7D" box="[966,1094,332,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDEFFB120C2FEA7FBD84A7D" box="[966,1094,332,356]" italics="true" pageId="8" pageNumber="23">L. applebyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D43FFDEFFB12784FEA7FB6A4A7D" box="[1152,1268,332,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDEFFB12784FEA7FB6A4A7D" box="[1152,1268,332,356]" italics="true" pageId="8" pageNumber="23">L. melicus</emphasis>
</taxonomicName>
(6.6%). There was no intraspecific variation in this gene fragment for
<emphasis id="B96CEAD2FFDEFFB12000FE9BFBA44A91" box="[772,1082,368,392]" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB12000FE9BFC354A91" box="[772,939,368,392]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">L. bidoupensis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB120B2FE9BFBA84A91" box="[950,1078,368,392]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C184D43FFDEFFB12775FE9BFB7D4A91" box="[1137,1251,368,392]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDEFFB12775FE9BFB7D4A91" box="[1137,1251,368,392]" italics="true" pageId="8" pageNumber="23">L. melicus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA736C0FFDEFFB123C1FE7EFE12490E" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
<emphasis id="B96CEAD2FFDEFFB123C1FE7EFEB74AB5" bold="true" box="[197,297,404,428]" pageId="8" pageNumber="23">Ecology.</emphasis>
All specimens of the new species were found in montane evergreen forest between
<quantity id="4CE09B25FFDEFFB127D5FE7EFAF54AB5" box="[1233,1387,404,428]" metricMagnitude="3" metricUnit="m" metricValue="1.675" metricValueMax="1.73" metricValueMin="1.6199999999999999" pageId="8" pageNumber="23" unit="m" value="1675.0" valueMax="1730.0" valueMin="1620.0">16201730 m</quantity>
elevation. Males were observed calling on stream banks, less than
<quantity id="4CE09B25FFDEFFB12075FE5DFC2C4AD6" box="[881,946,439,463]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="23" unit="m" value="0.5">0.5 m</quantity>
from small (&lt;
<quantity id="4CE09B25FFDEFFB12766FE5DFB104AD6" box="[1122,1166,439,463]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="23" unit="m" value="5.0">5 m</quantity>
wide) rocky streams in May and July. The new species was not heard or observed during surveys in the month of March, when conditions were cooler and drier.
</paragraph>
<paragraph id="8BA736C0FFDEFFB123C1FDCCFB8F49EB" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
<emphasis id="B96CEAD2FFDEFFB123C1FDCCFE254926" bold="true" box="[197,443,550,575]" pageId="8" pageNumber="23">Conservation status.</emphasis>
<taxonomicName id="4C184D43FFDEFFB122C7FDCDFD574926" box="[451,713,551,575]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB122C7FDCDFD574926" box="[451,713,551,575]" italics="true" pageId="8" pageNumber="23">Leptolalax bidoupensis</emphasis>
</taxonomicName>
is known only from an area of approximately
<quantity id="4CE09B25FFDEFFB127E4FDCDFA844926" box="[1248,1306,551,575]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="23" unit="km" value="1.0">1 km</quantity>
<quantity id="4CE09B25FFDEFFB1261FFDCEFADF4926" box="[1307,1345,548,575]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="23" unit="in" value="2.0">2 in</quantity>
Bidoup Nui Ba National Park. The actual distribution of the new species is unknown but probably extends to adjacent forested areas in the Langbian Plateau, including Chu Yang
<collectingCountry id="F30F7650FFDEFFB1201AFD85FCDD499E" box="[798,835,623,647]" name="China" pageId="8" pageNumber="23">Sin</collectingCountry>
National Park in Dak Lak Province, and Phuoc Binh National Park in Ninh Thuan Province, both of which contain montane evergreen forest at similar elevations and continuous with Bidoup-Nui Ba National Park. Given the available information, we suggest the species should be considered Data Deficient following IUCNs Red List categories (
<bibRefCitation id="EF894B31FFDEFFB1207FFD30FB9A49EB" author="IUCN" box="[891,1028,730,754]" pageId="8" pageNumber="23" refString="IUCN. (2001) IUCN Red List Categories and Criteria: Version 3.1. IUCN Species Survival Commission. IUCN, Gland, Switzerland and Cambridge, UK. ii + 30 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8BA736C0FFDEFFB123C1FD14FEB54FBB" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
<emphasis id="B96CEAD2FFDEFFB123C1FD14FEF5480F" bold="true" box="[197,363,766,791]" pageId="8" pageNumber="23">Comparisons.</emphasis>
<taxonomicName id="4C184D43FFDEFFB12273FD15FD1C480E" box="[375,642,767,791]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB12273FD15FD1C480E" box="[375,642,767,791]" italics="true" pageId="8" pageNumber="23">Leptolalax bidoupensis</emphasis>
</taxonomicName>
is distinguished from all other species in the genus except for
<taxonomicName id="4C184D43FFDEFFB12681FCEAFF664822" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDEFFB12681FCEAFF664822" italics="true" pageId="8" pageNumber="23">L. applebyi</emphasis>
</taxonomicName>
by having a dark brownish red ventral surface with white speckling on entire ventral surface including throat, arms and legs, forming distinct marbling on chest and belly (
<taxonomicName id="4C184D43FFDEFFB1209CFCA2FB9E4846" box="[920,1024,840,863]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B96CEAD2FFDEFFB1209CFCA2FB9E4846" box="[920,1024,840,863]" italics="true" pageId="8" pageNumber="23">L. aereus</emphasis>
</taxonomicName>
,
<emphasis id="B96CEAD2FFDEFFB1270AFCA2FF45489A" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB1270AFCA2FBE54846" box="[1038,1147,839,863]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="alpinis">L. alpinis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1278CFCA2FB6C4846" box="[1160,1266,840,863]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB127F9FCA2FAE44846" box="[1277,1402,839,863]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bourreti">L. bourreti</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12681FCA2FF45489A" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
</emphasis>
,
<emphasis id="B96CEAD2FFDEFFB123EFFC86FCB0489A" box="[235,814,875,899]" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB123EFFC86FE17489A" box="[235,393,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="fuliginosus">L. fuliginosus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12291FC86FD93489A" box="[405,525,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1211FFC86FD15489A" box="[539,651,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1219CFC86FCB0489A" box="[664,814,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
</emphasis>
,
<taxonomicName id="4C184D43FFDEFFB1203AFC86FC24489A" box="[830,954,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B96CEAD2FFDEFFB1203AFC86FC24489A" box="[830,954,875,899]" italics="true" pageId="8" pageNumber="23">L. lateralis</emphasis>
</taxonomicName>
,
<emphasis id="B96CEAD2FFDEFFB120CFFC86FEBA48BE" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB120CFFC86FB8F489A" box="[971,1041,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1271BFC86FB50489A" box="[1055,1230,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB127DFFC86FAE7489A" box="[1243,1401,875,899]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="oshanensis">L. oshanensis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12681FC86FEBA48BE" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
</emphasis>
,
<emphasis id="B96CEAD2FFDEFFB12236FC7AFD4E48BE" box="[306,720,911,935]" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB12236FC7AFE0B48BE" box="[306,405,912,935]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB122A4FC7AFE6748BE" box="[416,505,911,935]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12100FC7AFDFE48BE" box="[516,608,912,935]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1216FFC7AFD4E48BE" box="[619,720,911,935]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tamdil">L. tamdil</taxonomicName>
</emphasis>
and
<taxonomicName id="4C184D43FFDEFFB12002FC7AFC1148BE" box="[774,911,911,935]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CEAD2FFDEFFB12002FC7AFC1148BE" box="[774,911,911,935]" italics="true" pageId="8" pageNumber="23">L. tuberosus</emphasis>
</taxonomicName>
have mostly white or pale grey/brown venters, with or without dark spots or mottling;
<taxonomicName id="4C184D43FFDEFFB12148FC5EFD2048D2" box="[588,702,948,971]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B96CEAD2FFDEFFB12148FC5EFD2048D2" box="[588,702,948,971]" italics="true" pageId="8" pageNumber="23">L. croceus</emphasis>
</taxonomicName>
has a bright orange belly;
<taxonomicName id="4C184D43FFDEFFB120E1FC5EFBC648D2" box="[997,1112,947,971]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDEFFB120E1FC5EFBC648D2" box="[997,1112,947,971]" italics="true" pageId="8" pageNumber="23">L. melicus</emphasis>
</taxonomicName>
has an off-white to pale pink ventral surface with diffuse dark brown blotches and distinct white speckling on the chest, belly and throat;
<taxonomicName id="4C184D43FFDEFFB12655FC32FF4A4F0A" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CEAD2FFDEFFB12655FC32FF4A4F0A" italics="true" pageId="8" pageNumber="23">L. pluvialis</emphasis>
</taxonomicName>
has a dirty white/grey venter with dark brown/grey marbling, and uniform pale dirty white/grey throat with pale speckling only around the margins;
<taxonomicName id="4C184D43FFDEFFB1216DFBCAFCBC4F2E" box="[617,802,1055,1079]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="B96CEAD2FFDEFFB1216DFBCAFCBC4F2E" box="[617,802,1055,1079]" italics="true" pageId="8" pageNumber="23">L. melanoleucus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D43FFDEFFB12058FBCAFBB44F2E" box="[860,1066,1056,1079]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B96CEAD2FFDEFFB12058FBCAFBB44F2E" box="[860,1066,1056,1079]" italics="true" pageId="8" pageNumber="23">L. ventripunctatus</emphasis>
</taxonomicName>
display large patches of distinct black and white marbling,
<taxonomicName id="4C184D43FFDEFFB122C0FBAEFDD14F42" box="[452,591,1091,1115]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CEAD2FFDEFFB122C0FBAEFDD14F42" box="[452,591,1091,1115]" italics="true" pageId="8" pageNumber="23">L. heteropus</emphasis>
</taxonomicName>
has a grey venter, speckled with black;
<taxonomicName id="4C184D43FFDEFFB12710FBAEFB184F42" box="[1044,1158,1092,1115]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CEAD2FFDEFFB12710FBAEFB184F42" box="[1044,1158,1092,1115]" italics="true" pageId="8" pageNumber="23">L. maurus</emphasis>
</taxonomicName>
has a black or dark grey brown venter, with indistinct small light areas, and
<taxonomicName id="4C184D43FFDEFFB121D5FB82FCBD4F66" box="[721,803,1127,1151]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B96CEAD2FFDEFFB121D5FB82FCBD4F66" box="[721,803,1127,1151]" italics="true" pageId="8" pageNumber="23">L. kecil</emphasis>
</taxonomicName>
has a uniformly dark venter with large, dark orange pectoral glands).
</paragraph>
<paragraph id="8BA736C0FFDEFFB123C1FB45FB954D83" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
The small size of
<taxonomicName id="4C184D43FFDEFFB12296FB45FD064FDE" box="[402,664,1199,1223]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB12296FB45FD064FDE" box="[402,664,1199,1223]" italics="true" pageId="8" pageNumber="23">Leptolalax bidoupensis</emphasis>
</taxonomicName>
(
<quantity id="4CE09B25FFDEFFB121ACFB45FCD04FDE" box="[680,846,1199,1223]" metricMagnitude="-2" metricUnit="m" metricValue="2.41" metricValueMax="2.46" metricValueMin="2.36" pageId="8" pageNumber="23" unit="mm" value="24.1" valueMax="24.6" valueMin="23.6">23.624.6 mm</quantity>
in four adult males and
<quantity id="4CE09B25FFDEFFB12763FB45FA954FDE" box="[1127,1291,1199,1223]" metricMagnitude="-2" metricUnit="m" metricValue="2.9299999999999997" metricValueMax="2.94" metricValueMin="2.92" pageId="8" pageNumber="23" unit="mm" value="29.299999999999997" valueMax="29.4" valueMin="29.2">29.229.4 mm</quantity>
in two adult females) further distinguishes it from the larger
<taxonomicName id="4C184D43FFDEFFB121AAFB3EFC8B4FF2" box="[686,789,1236,1259]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B96CEAD2FFDEFFB121AAFB3EFC8B4FF2" box="[686,789,1236,1259]" italics="true" pageId="8" pageNumber="23">L. arayai</emphasis>
</taxonomicName>
(male
<quantity id="4CE09B25FFDEFFB12067FB38FC544FF3" box="[867,970,1234,1258]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="8" pageNumber="23" unit="mm" value="29.6">29.6 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB120D9FB3EFBC84FF2" box="[989,1110,1235,1259]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B96CEAD2FFDEFFB120D9FB3EFBC84FF2" box="[989,1110,1235,1259]" italics="true" pageId="8" pageNumber="23">L. bourreti</emphasis>
</taxonomicName>
(male
<quantity id="4CE09B25FFDEFFB127A7FB38FA974FF3" box="[1187,1289,1234,1258]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="8" pageNumber="23" unit="mm" value="36.2">36.2 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB12676FB38FF774E16" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="8" pageNumber="23" unit="mm" value="43.5" valueMax="45.0" valueMin="42.0">42 45 mm</quantity>
).
<taxonomicName id="4C184D43FFDEFFB123F8FB12FEC34E16" box="[252,349,1271,1295]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B96CEAD2FFDEFFB123F8FB12FEC34E16" box="[252,349,1271,1295]" italics="true" pageId="8" pageNumber="23">L. dringi</emphasis>
</taxonomicName>
(males
<quantity id="4CE09B25FFDEFFB122B1FB1DFDC74E16" box="[437,601,1271,1295]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.03" metricValueMin="2.87" pageId="8" pageNumber="23" unit="mm" value="29.5" valueMax="30.3" valueMin="28.7">28.730.3 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB121B2FB1DFC804E16" box="[694,798,1271,1295]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="8" pageNumber="23" unit="mm" value="37.5">37.5 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB12034FB12FC564E16" box="[816,968,1271,1295]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B96CEAD2FFDEFFB12034FB12FC564E16" box="[816,968,1271,1295]" italics="true" pageId="8" pageNumber="23">L. fuliginosus</emphasis>
</taxonomicName>
(males 28.230.0 mm),
<taxonomicName id="4C184D43FFDEFFB127DCFB12FAD54E16" box="[1240,1355,1271,1295]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B96CEAD2FFDEFFB127DCFB12FAD54E16" box="[1240,1355,1271,1295]" italics="true" pageId="8" pageNumber="23">L. gracilis</emphasis>
</taxonomicName>
(males
<quantity id="4CE09B25FFDEFFB12393FAF0FE8A4E2B" box="[151,276,1306,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.6" metricValueMin="3.0" pageId="8" pageNumber="23" unit="mm" value="33.0" valueMax="36.0" valueMin="30.0">3036 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB1222CFAF6FE0A4E2A" box="[296,404,1307,1331]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B96CEAD2FFDEFFB1222CFAF6FE0A4E2A" box="[296,404,1307,1331]" italics="true" pageId="8" pageNumber="23">L. hamidi</emphasis>
</taxonomicName>
(males
<quantity id="4CE09B25FFDEFFB122E9FAF0FD0D4E2B" box="[493,659,1306,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.13" metricValueMin="2.87" pageId="8" pageNumber="23" unit="mm" value="30.0" valueMax="31.3" valueMin="28.7">28.731.3 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB121F9FAF0FC3B4E2B" box="[765,933,1306,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.9450000000000003" metricValueMax="4.279999999999999" metricValueMin="3.6100000000000003" pageId="8" pageNumber="23" unit="mm" value="39.45" valueMax="42.8" valueMin="36.1">36.142.8 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB120BCFAF6FBDA4E2A" box="[952,1092,1307,1331]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CEAD2FFDEFFB120BCFAF6FBDA4E2A" box="[952,1092,1307,1331]" italics="true" pageId="8" pageNumber="23">L. heteropus</emphasis>
</taxonomicName>
(male
<quantity id="4CE09B25FFDEFFB12797FAF0FB784E2B" box="[1171,1254,1306,1330]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="23" unit="mm" value="33.0">33 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB127FEFAF6FA024E2A" box="[1274,1436,1307,1331]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="kajangensis">
<emphasis id="B96CEAD2FFDEFFB127FEFAF6FA024E2A" box="[1274,1436,1307,1331]" italics="true" pageId="8" pageNumber="23">L. kajangensis</emphasis>
</taxonomicName>
(males
<quantity id="4CE09B25FFDEFFB123EEFAD5FEF74E4E" box="[234,361,1343,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" metricValueMax="3.5" metricValueMin="3.4" pageId="8" pageNumber="23" unit="mm" value="34.5" valueMax="35.0" valueMin="34.0">3435 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB1227BFAAAFDB44E4E" box="[383,554,1343,1367]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="B96CEAD2FFDEFFB1227BFAAAFDB44E4E" box="[383,554,1343,1367]" italics="true" pageId="8" pageNumber="23">L. nahangensis</emphasis>
</taxonomicName>
(male
<quantity id="4CE09B25FFDEFFB12179FAD5FD784E4E" box="[637,742,1343,1367]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="8" pageNumber="23" unit="mm" value="40.8">40.8 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB121F8FAAAFC374E4E" box="[764,937,1343,1367]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B96CEAD2FFDEFFB121F8FAAAFC374E4E" box="[764,937,1343,1367]" italics="true" pageId="8" pageNumber="23">L. pelodytoides</emphasis>
</taxonomicName>
(male ~
<quantity id="4CE09B25FFDEFFB1270DFAD5FBC34E4E" box="[1033,1117,1343,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="23" unit="mm" value="30.0">30 mm</quantity>
, female ~
<quantity id="4CE09B25FFDEFFB127CBFAD5FABA4E4E" box="[1231,1316,1343,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="23" unit="mm" value="37.0">37 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB1263DFAAAFA024E4E" box="[1337,1436,1344,1367]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CEAD2FFDEFFB1263DFAAAFA024E4E" box="[1337,1436,1344,1367]" italics="true" pageId="8" pageNumber="23">L. pictus</emphasis>
</taxonomicName>
(males
<quantity id="4CE09B25FFDEFFB123EEFA88FEF84E63" box="[234,358,1378,1402]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="8" pageNumber="23" unit="mm" value="32.5" valueMax="34.0" valueMin="31.0">3134 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB122CCFA88FD824E63" box="[456,540,1378,1402]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="23" unit="mm" value="47.0">47 mm</quantity>
),
<taxonomicName id="4C184D43FFDEFFB12134FA8EFD124E62" box="[560,652,1380,1403]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B96CEAD2FFDEFFB12134FA8EFD124E62" box="[560,652,1380,1403]" italics="true" pageId="8" pageNumber="23">L. sungi</emphasis>
</taxonomicName>
(males
<quantity id="4CE09B25FFDEFFB121E3FA88FC104E63" box="[743,910,1378,1402]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.27" metricValueMin="4.83" pageId="8" pageNumber="23" unit="mm" value="50.5" valueMax="52.7" valueMin="48.3">48.352.7 mm</quantity>
, females 56.758.9), and
<taxonomicName id="4C184D43FFDEFFB127B0FA8EFA844E62" box="[1204,1306,1379,1403]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B96CEAD2FFDEFFB127B0FA8EFA844E62" box="[1204,1306,1379,1403]" italics="true" pageId="8" pageNumber="23">L. tamdil</emphasis>
</taxonomicName>
(male
<quantity id="4CE09B25FFDEFFB1266FFA88FF5B4E86" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="8" pageNumber="23" unit="mm" value="32.3">32.3 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB12222FA6DFE114E86" box="[294,399,1415,1439]" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="8" pageNumber="23" unit="mm" value="31.8">31.8 mm</quantity>
). The remaining, small-sized congeners may have overlapping body-sizes for either or both sexes (
<taxonomicName id="4C184D43FFDEFFB123E0FA46FED34EDA" box="[228,333,1452,1475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B96CEAD2FFDEFFB123E0FA46FED34EDA" box="[228,333,1452,1475]" italics="true" pageId="8" pageNumber="23">L. aereus</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12299FA40FDDC4EDB" box="[413,578,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.8899999999999997" metricValueMin="2.51" pageId="8" pageNumber="23" unit="mm" value="27.0" valueMax="28.9" valueMin="25.1">25.128.9 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB121AAFA40FCCA4EDB" box="[686,852,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="3.285" metricValueMax="3.86" metricValueMin="2.71" pageId="8" pageNumber="23" unit="mm" value="32.85" valueMax="38.6" valueMin="27.1">27.138.6 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB12064FA46FC544EDA" box="[864,970,1451,1475]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="B96CEAD2FFDEFFB12064FA46FC544EDA" box="[864,970,1451,1475]" italics="true" pageId="8" pageNumber="23">L. alpinis</emphasis>
</taxonomicName>
males 24.0
<quantity id="4CE09B25FFDEFFB1275EFA40FB5F4EDB" box="[1114,1217,1450,1474]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="8" pageNumber="23" unit="mm" value="26.4">26.4 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB12628FA40FF584EFE" metricMagnitude="-2" metricUnit="m" metricValue="3.21" metricValueMax="3.25" metricValueMin="3.17" pageId="8" pageNumber="23" unit="mm" value="32.1" valueMax="32.5" valueMin="31.7">31.732.5 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB123D5FA3AFED04EFE" box="[209,334,1487,1511]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDEFFB123D5FA3AFED04EFE" box="[209,334,1487,1511]" italics="true" pageId="8" pageNumber="23">L. applebyi</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12298FA25FDDE4EFE" box="[412,576,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="2.02" metricValueMax="2.08" metricValueMin="1.96" pageId="8" pageNumber="23" unit="mm" value="20.2" valueMax="20.8" valueMin="19.6">19.620.8 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB121ADFA25FC8E4EFE" box="[681,784,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="2.17" pageId="8" pageNumber="23" unit="mm" value="21.7">21.7 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB1201EFA3AFC134EFE" box="[794,909,1488,1511]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B96CEAD2FFDEFFB1201EFA3AFC134EFE" box="[794,909,1488,1511]" italics="true" pageId="8" pageNumber="23">L. croceus</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB120D8FA25FB1F4EFE" box="[988,1153,1487,1511]" metricMagnitude="-2" metricUnit="m" metricValue="2.475" metricValueMax="2.73" metricValueMin="2.2199999999999998" pageId="8" pageNumber="23" unit="mm" value="24.75" valueMax="27.3" valueMin="22.2">22.227.3 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB12788FA3AFB404EFE" box="[1164,1246,1487,1511]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B96CEAD2FFDEFFB12788FA3AFB404EFE" box="[1164,1246,1487,1511]" italics="true" pageId="8" pageNumber="23">L. kecil</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12628FA25FF5B4D13" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" metricValueMax="2.05" metricValueMin="1.93" pageId="8" pageNumber="23" unit="mm" value="19.9" valueMax="20.5" valueMin="19.3">19.320.5 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB1222DFA18FE0D4D13" box="[297,403,1522,1546]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="8" pageNumber="23" unit="mm" value="20.5">20.5 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB122A5FA1EFDA94D12" box="[417,567,1523,1547]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="khasiorum">
<emphasis id="B96CEAD2FFDEFFB122A5FA1EFDA94D12" box="[417,567,1523,1547]" italics="true" pageId="8" pageNumber="23">L. khasiorum</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12188FA18FCAD4D13" box="[652,819,1522,1546]" metricMagnitude="-2" metricUnit="m" metricValue="2.59" metricValueMax="2.73" metricValueMin="2.45" pageId="8" pageNumber="23" unit="mm" value="25.9" valueMax="27.3" valueMin="24.5">24.527.3 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB120A7FA18FBD54D13" box="[931,1099,1522,1546]" metricMagnitude="-2" metricUnit="m" metricValue="3.235" metricValueMax="3.35" metricValueMin="3.12" pageId="8" pageNumber="23" unit="mm" value="32.35" valueMax="33.5" valueMin="31.2">31.233.5 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB1275EFA1EFB494D12" box="[1114,1239,1523,1547]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B96CEAD2FFDEFFB1275EFA1EFB494D12" box="[1114,1239,1523,1547]" italics="true" pageId="8" pageNumber="23">L. lateralis</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12628FA18FF5B4D36" metricMagnitude="-2" metricUnit="m" metricValue="2.76" metricValueMax="2.83" metricValueMin="2.69" pageId="8" pageNumber="23" unit="mm" value="27.6" valueMax="28.3" valueMin="26.9">26.928.3 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB12223F9FDFEE54D36" box="[295,379,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="8" pageNumber="23" unit="mm" value="36.0">36 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB12283F9F2FE524D36" box="[391,460,1559,1583]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="liui">
<emphasis id="B96CEAD2FFDEFFB12283F9F2FE524D36" box="[391,460,1559,1583]" italics="true" pageId="8" pageNumber="23">L. liui</emphasis>
</taxonomicName>
males 23.0
<quantity id="4CE09B25FFDEFFB1215AF9FDFD5B4D36" box="[606,709,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="2.87" pageId="8" pageNumber="23" unit="mm" value="28.7">28.7 mm</quantity>
, females
<quantity id="4CE09B25FFDEFFB12036F9FDFC474D36" box="[818,985,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="2.56" metricValueMax="2.81" metricValueMin="2.31" pageId="8" pageNumber="23" unit="mm" value="25.6" valueMax="28.1" valueMin="23.1">23.128.1 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB120E2F9F2FBC44D36" box="[998,1114,1560,1583]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CEAD2FFDEFFB120E2F9F2FBC44D36" box="[998,1114,1560,1583]" italics="true" pageId="8" pageNumber="23">L. maurus</emphasis>
</taxonomicName>
male
<quantity id="4CE09B25FFDEFFB127A5F9FDFA974D36" box="[1185,1289,1559,1583]" metricMagnitude="-2" metricUnit="m" metricValue="2.61" pageId="8" pageNumber="23" unit="mm" value="26.1">26.1 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB1266FF9FDFF584D4B" metricMagnitude="-2" metricUnit="m" metricValue="3.18" pageId="8" pageNumber="23" unit="mm" value="31.8">31.8 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB123D5F9D6FE164D4A" box="[209,392,1595,1619]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="B96CEAD2FFDEFFB123D5F9D6FE164D4A" box="[209,392,1595,1619]" italics="true" pageId="8" pageNumber="23">L. melanoleucus</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB122D3F9D0FDE54D4B" box="[471,635,1594,1618]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" metricValueMax="2.88" metricValueMin="2.66" pageId="8" pageNumber="23" unit="mm" value="27.7" valueMax="28.8" valueMin="26.6">26.628.8 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB121DDF9D0FCA14D4B" box="[729,831,1594,1618]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="8" pageNumber="23" unit="mm" value="32.7">32.7 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB1204EF9D6FC234D4A" box="[842,957,1595,1619]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDEFFB1204EF9D6FC234D4A" box="[842,957,1595,1619]" italics="true" pageId="8" pageNumber="23">L. melicus</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB1270FF9D0FB2F4D4B" box="[1035,1201,1594,1618]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" metricValueMax="2.27" metricValueMin="1.95" pageId="8" pageNumber="23" unit="mm" value="21.1" valueMax="22.7" valueMin="19.5">19.522.7 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB127B8F9D6FACA4D4A" box="[1212,1364,1595,1619]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CEAD2FFDEFFB127B8F9D6FACA4D4A" box="[1212,1364,1595,1619]" italics="true" pageId="8" pageNumber="23">L. oshanensis</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12393F9B5FEA04D6E" box="[151,318,1631,1655]" metricMagnitude="-2" metricUnit="m" metricValue="2.8649999999999998" metricValueMax="3.07" metricValueMin="2.66" pageId="8" pageNumber="23" unit="mm" value="28.65" valueMax="30.7" valueMin="26.6">26.630.7 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB122A7F9B5FD934D6E" box="[419,525,1631,1655]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="8" pageNumber="23" unit="mm" value="31.6">31.6 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB1211FF98AFD024D6E" box="[539,668,1631,1655]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CEAD2FFDEFFB1211FF98AFD024D6E" box="[539,668,1631,1655]" italics="true" pageId="8" pageNumber="23">L. pluvialis</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB121F5F9B5FC044D6E" box="[753,922,1631,1655]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.23" metricValueMin="2.13" pageId="8" pageNumber="23" unit="mm" value="21.8" valueMax="22.3" valueMin="21.3">21.322.3 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB120ACF98AFB9C4D6E" box="[936,1026,1631,1655]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="solus">
<emphasis id="B96CEAD2FFDEFFB120ACF98AFB9C4D6E" box="[936,1026,1631,1655]" italics="true" pageId="8" pageNumber="23">L. solus</emphasis>
</taxonomicName>
male
<quantity id="4CE09B25FFDEFFB12748F9B5FB284D6E" box="[1100,1206,1631,1655]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="8" pageNumber="23" unit="mm" value="27.6">27.6 mm</quantity>
;
<taxonomicName id="4C184D43FFDEFFB127C0F98AFACF4D6E" box="[1220,1361,1631,1655]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CEAD2FFDEFFB127C0F98AFACF4D6E" box="[1220,1361,1631,1655]" italics="true" pageId="8" pageNumber="23">L. tuberosus</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDEFFB12393F968FEA24D83" box="[151,316,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="2.695" metricValueMax="2.95" metricValueMin="2.44" pageId="8" pageNumber="23" unit="mm" value="26.95" valueMax="29.5" valueMin="24.4">24.429.5 mm</quantity>
, female
<quantity id="4CE09B25FFDEFFB1229EF968FD9F4D83" box="[410,513,1666,1690]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="8" pageNumber="23" unit="mm" value="30.2">30.2 mm</quantity>
; and
<taxonomicName id="4C184D43FFDEFFB12138F96EFC964D82" box="[572,776,1668,1691]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B96CEAD2FFDEFFB12138F96EFC964D82" box="[572,776,1668,1691]" italics="true" pageId="8" pageNumber="23">L. ventripunctatus</emphasis>
</taxonomicName>
males 25.528.0 mm).
</paragraph>
<paragraph id="8BA736C0FFDEFFB123C1F942FCEF4C8E" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
In having a bicoloured iris, with the upper half coppery red and the lower half fading to pale silver,
<taxonomicName id="4C184D43FFDEFFB1264BF942FF674DFA" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB1264BF942FF674DFA" italics="true" pageId="8" pageNumber="23">L. bidoupensis</emphasis>
</taxonomicName>
differs from at least
<emphasis id="B96CEAD2FFDEFFB122E2F926FE124C1E" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB122E2F926FDCC4DFA" box="[486,594,1740,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="aereus">L. aereus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12159F926FD414DFA" box="[605,735,1739,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB121EFF926FCFC4DFA" box="[747,866,1740,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="croceus">L. croceus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1206AF926FB8A4DFA" box="[878,1044,1739,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1271BF926FBEB4DFA" box="[1055,1141,1739,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12785F926FB684DFA" box="[1153,1270,1740,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12606F926FAE74DFA" box="[1282,1401,1739,1763]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">L. melicus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12681F926FEB84C1E" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12235F91AFE124C1E" box="[305,396,1776,1799]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C184D43FFDEFFB122C6F91AFDD24C1E" box="[450,588,1775,1799]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CEAD2FFDEFFB122C6F91AFDD24C1E" box="[450,588,1775,1799]" italics="true" pageId="8" pageNumber="23">L. tuberosus</emphasis>
</taxonomicName>
, all of which have uniform iris colouration under black reticulations (
<tableCitation id="C69A037BFFDEFFB12658F905FF394C33" captionStart="TABLE 3" captionStartId="9.[151,235,296,317]" captionTargetBox="[151,1413,345,552]" captionTargetPageId="9" captionText="TABLE 3. Iris colouration in life for species within the genus Leptolalax." httpUri="http://table.plazi.org/id/DF676648FFDFFFB02393FEC2FC1F4A27" pageId="8" pageNumber="23" tableUuid="DF676648FFDFFFB02393FEC2FC1F4A27">Table 3</tableCitation>
;
<figureCitation id="13232A45FFDEFFB123B0F8F8FE8A4C33" box="[180,276,1810,1834]" captionStart="FIGURE 5" captionStartId="11.[151,250,1213,1236]" captionTargetBox="[152,1436,339,1191]" captionTargetId="figure@11.[151,1436,339,1192]" captionTargetPageId="11" captionText="FIGURE 5. Iris colouration in life of (A) Leptolalax bidoupensis sp. nov. NCSM 77320, (B) Leptolalax bidoupensis sp. nov. AMS R 173134, (C) Leptolalax bidoupensis sp. nov. AMS R 173133, (D E) Leptolalax applebyi, and (F) Leptolalax melicus MVZ 258199." httpUri="https://zenodo.org/record/277021/files/figure.png" pageId="8" pageNumber="23">Figure 5</figureCitation>
).
<taxonomicName id="4C184D43FFDEFFB1222EF8F9FDAE4C32" box="[298,560,1811,1835]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB1222EF8F9FDAE4C32" box="[298,560,1811,1835]" italics="true" pageId="8" pageNumber="23">Leptolalax bidoupensis</emphasis>
</taxonomicName>
is also differentiated from
<emphasis id="B96CEAD2FFDEFFB12065F8FEFDC04C56" italics="true" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB12065F8FEFC554C32" box="[865,971,1812,1835]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="arayai">L. arayai</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB120D2F8FEFBD34C32" box="[982,1101,1812,1835]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="croceus">L. croceus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1275DF8FEFB6E4C32" box="[1113,1264,1811,1835]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB127FFF8FEFAE44C32" box="[1275,1402,1811,1835]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB12681F8FEFF704C56" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB123FDF8D2FECD4C56" box="[249,339,1847,1871]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB1225BF8D2FE564C56" box="[351,456,1847,1871]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tamdil">L. tamdil</taxonomicName>
,
<taxonomicName id="4C184D43FFDEFFB122D7F8D2FDC04C56" box="[467,606,1847,1871]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="tuberosus">L. tuberosus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C184D43FFDEFFB12191F8D2FCFF4C56" box="[661,865,1848,1871]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B96CEAD2FFDEFFB12191F8D2FCFF4C56" box="[661,865,1848,1871]" italics="true" pageId="8" pageNumber="23">L. ventripunctatus</emphasis>
</taxonomicName>
in having mostly smooth (versus tuberculate) skin texture with no skin ridges, and from
<taxonomicName id="4C184D43FFDEFFB1213CF8B6FD064C6A" box="[568,664,1884,1907]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CEAD2FFDEFFB1213CF8B6FD064C6A" box="[568,664,1884,1907]" italics="true" pageId="8" pageNumber="23">L. pictus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D43FFDEFFB121C9F8B6FCD44C6A" box="[717,842,1883,1907]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CEAD2FFDEFFB121C9F8B6FCD44C6A" box="[717,842,1883,1907]" italics="true" pageId="8" pageNumber="23">L. pluvialis</emphasis>
</taxonomicName>
in having relatively short tibia (male TIB:SVL 0.44 0.46, versus
<quantity id="4CE09B25FFDEFFB12220F895FE2F4C8E" box="[292,433,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.4732000000000003" metricValueMax="1.5493999999999999" metricValueMin="1.3970000000000002" pageId="8" pageNumber="23" unit="in" value="0.58" valueMax="0.61" valueMin="0.55">0.550.61 in</quantity>
<taxonomicName id="4C184D43FFDEFFB122BCF86AFD874C8E" box="[440,537,1920,1943]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CEAD2FFDEFFB122BCF86AFD874C8E" box="[440,537,1920,1943]" italics="true" pageId="8" pageNumber="23">L. pictus</emphasis>
</taxonomicName>
and
<quantity id="4CE09B25FFDEFFB1214BF895FD424C8E" box="[591,732,1919,1943]" metricMagnitude="-2" metricUnit="m" metricValue="1.3716" metricValueMax="1.4224" metricValueMin="1.3208" pageId="8" pageNumber="23" unit="in" value="0.54" valueMax="0.56" valueMin="0.52">0.520.56 in</quantity>
<taxonomicName id="4C184D43FFDEFFB121E7F86AFCFE4C8E" box="[739,864,1919,1943]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B96CEAD2FFDEFFB121E7F86AFCFE4C8E" box="[739,864,1919,1943]" italics="true" pageId="8" pageNumber="23">L. pluvialis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BA736C0FFDEFFB123C1F849FA024CC6" blockId="8.[151,1437,151,2015]" pageId="8" pageNumber="23">
<taxonomicName id="4C184D43FFDEFFB123C1F849FE534CA2" box="[197,461,1955,1979]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDEFFB123C1F849FE534CA2" box="[197,461,1955,1979]" italics="true" pageId="8" pageNumber="23">Leptolalax bidoupensis</emphasis>
</taxonomicName>
can be further differentiated from the two most morphologically similar species,
<taxonomicName id="4C184D43FFDEFFB12681F84EFF664CC6" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDEFFB12681F84EFF664CC6" italics="true" pageId="8" pageNumber="23">L. applebyi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D43FFDEFFB12234F822FE3A4CC6" box="[304,420,1991,2015]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="23" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDEFFB12234F822FE3A4CC6" box="[304,420,1991,2015]" italics="true" pageId="8" pageNumber="23">L. melicus</emphasis>
</taxonomicName>
, by having slight lateral fringing on feet, and by being approximately 19% larger than
<emphasis id="B96CEAD2FFDEFFB12681F822FA024CC6" box="[1413,1436,1992,2015]" italics="true" pageId="8" pageNumber="23">L.</emphasis>
</paragraph>
<paragraph id="8BA736C0FFDFFFB02393FF72FBC04BEE" blockId="9.[151,1436,151,248]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB02393FF72FF664BA9" box="[151,248,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDFFFB02393FF72FF664BA9" box="[151,248,152,176]" italics="true" pageId="9" pageNumber="24">applebyi</emphasis>
</taxonomicName>
(Wilcoxin rank-sum test,
<emphasis id="B96CEAD2FFDFFFB02125FF73FDD54BA9" box="[545,587,153,176]" italics="true" pageId="9" pageNumber="24">W=</emphasis>
16,
<emphasis id="B96CEAD2FFDFFFB0217BFF73FD134BA9" box="[639,653,153,176]" italics="true" pageId="9" pageNumber="24">p</emphasis>
= 0.0284; N=8) and 14% larger than
<taxonomicName id="4C184D43FFDFFFB02738FF73FB2F4BA9" box="[1084,1201,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDFFFB02738FF73FB2F4BA9" box="[1084,1201,152,176]" italics="true" pageId="9" pageNumber="24">L. melicus</emphasis>
</taxonomicName>
(Wilcoxin rank-sum test,
<emphasis id="B96CEAD2FFDFFFB023C8FF57FF684BCD" box="[204,246,189,212]" italics="true" pageId="9" pageNumber="24">W=</emphasis>
28,
<emphasis id="B96CEAD2FFDFFFB0222CFF57FEA84BCD" box="[296,310,189,212]" italics="true" pageId="9" pageNumber="24">p</emphasis>
= 0.0106; N=11). In life,
<taxonomicName id="4C184D43FFDFFFB0215DFF57FD634BCD" box="[601,765,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDFFFB0215DFF57FD634BCD" box="[601,765,188,212]" italics="true" pageId="9" pageNumber="24">L. bidoupensis</emphasis>
</taxonomicName>
also weighs more than either species (
<taxonomicName id="4C184D43FFDFFFB027B4FF57FACD4BCD" box="[1200,1363,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDFFFB027B4FF57FACD4BCD" box="[1200,1363,188,212]" italics="true" pageId="9" pageNumber="24">L. bidoupensis</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDFFFB02393FF35FE9C4BE1" box="[151,258,223,248]" metricMagnitude="-3" metricUnit="kg" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="9" pageNumber="24" unit="g" value="1.25" valueMax="1.4" valueMin="1.1">1.11.4 g</quantity>
, N=4;
<taxonomicName id="4C184D43FFDFFFB0224AFF0BFE524BE1" box="[334,460,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDFFFB0224AFF0BFE524BE1" box="[334,460,224,248]" italics="true" pageId="9" pageNumber="24">L. applebyi</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDFFFB02118FF35FD194BE1" box="[540,647,223,248]" metricMagnitude="-4" metricUnit="kg" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="9" pageNumber="24" unit="g" value="0.85" valueMax="0.9" valueMin="0.8">0.80.9 g</quantity>
, N=7;
<taxonomicName id="4C184D43FFDFFFB021D0FF0BFCD94BE1" box="[724,839,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDFFFB021D0FF0BFCD94BE1" box="[724,839,224,248]" italics="true" pageId="9" pageNumber="24">L. melicus</emphasis>
</taxonomicName>
males
<quantity id="4CE09B25FFDFFFB02092FF35FB914BE1" box="[918,1039,223,248]" metricMagnitude="-4" metricUnit="kg" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="9" pageNumber="24" unit="g" value="0.725" valueMax="0.75" valueMin="0.7">0.70.75 g</quantity>
, N=2).
</paragraph>
<caption id="DF676648FFDFFFB02393FEC2FC1F4A27" ID-Table-UUID="DF676648FFDFFFB02393FEC2FC1F4A27" box="[151,897,296,318]" httpUri="http://table.plazi.org/id/DF676648FFDFFFB02393FEC2FC1F4A27" pageId="9" pageNumber="24" targetBox="[151,1413,345,552]" targetIsTable="true" targetPageId="9">
<paragraph id="8BA736C0FFDFFFB02393FEC2FC1F4A27" blockId="9.[151,897,296,318]" box="[151,897,296,318]" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB02393FEC2FE9A4A24" bold="true" box="[151,260,296,317]" pageId="9" pageNumber="24">TABLE 3.</emphasis>
Iris colouration in life for species within the genus
<taxonomicName id="4C184D43FFDFFFB0200BFEC2FCE54A27" box="[783,891,296,318]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="B96CEAD2FFDFFFB0200BFEC2FCE54A27" box="[783,891,296,318]" italics="true" pageId="9" pageNumber="24">Leptolalax</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA736C0FFDFFFB0239BFEB3FA1A4931" pageId="9" pageNumber="24">
<table id="F918C460FFDF00462393FEB3FA1B4931" box="[151,1413,345,552]" gridcols="3" gridrows="4" pageId="9" pageNumber="24">
<tr id="35283482FFDF00462393FEB3FA1B4A97" box="[151,1413,345,398]" gridrow="0" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FEB3FE834A97" box="[151,285,345,398]" gridcol="0" gridrow="0" pageId="9" pageNumber="24">Species</th>
<th id="76F95DFEFFDF004622ABFEB3FC6C4A97" box="[431,1010,345,398]" gridcol="1" gridrow="0" pageId="9" pageNumber="24">Iris bi- Iris colouration coloured</th>
<th id="76F95DFEFFDF004627B7FEB3FA1B4A97" box="[1203,1413,345,398]" gridcol="2" gridrow="0" pageId="9" pageNumber="24">Source</th>
</tr>
<tr id="35283482FFDF00462393FE4BFA1B4AAE" box="[151,1413,417,439]" gridrow="1" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FE4BFE834AAE" box="[151,285,417,439]" gridcol="0" gridrow="1" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFE48FE974AAE" box="[159,265,418,439]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFE48FF614AAE" box="[159,255,418,439]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="aereus">L. aereus</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFE4BFC6C4AAE" box="[431,1010,417,439]" gridcol="1" gridrow="1" pageId="9" pageNumber="24">No Bronze with minute, black reticulations.</td>
<td id="76F95DFEFFDF004627B7FE4BFA1B4AAE" box="[1203,1413,417,439]" gridcol="2" gridrow="1" pageId="9" pageNumber="24">
Rowley
<emphasis id="B96CEAD2FFDFFFB0260CFE48FAAA4AAE" box="[1288,1332,417,439]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2010c
</td>
</tr>
<tr id="35283482FFDF00462393FE20FA1B4AE6" box="[151,1413,458,511]" gridrow="2" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FE20FE834AE6" box="[151,285,458,511]" gridcol="0" gridrow="2" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFE21FF614AF9" box="[159,255,458,480]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="B96CEAD2FFDFFFB0239BFE21FF614AF9" box="[159,255,458,480]" italics="true" pageId="9" pageNumber="24">L. alpinis</emphasis>
</taxonomicName>
</th>
<td id="76F95DFEFFDF004622ABFE20FC6C4AE6" box="[431,1010,458,511]" gridcol="1" gridrow="2" pageId="9" pageNumber="24">No? Copper/gold with black reticulations.</td>
<td id="76F95DFEFFDF004627B7FE20FA1B4AE6" box="[1203,1413,458,511]" gridcol="2" gridrow="2" pageId="9" pageNumber="24">
Fei
<emphasis id="B96CEAD2FFDFFFB027DEFE21FA984AF9" box="[1242,1286,458,480]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2010 (from figures on p. 169)
</td>
</tr>
<tr id="35283482FFDF00462393FDF8FA1B4931" box="[151,1413,530,552]" gridrow="3" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FDF8FE834931" box="[151,285,530,552]" gridcol="0" gridrow="3" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFDF9FE834931" box="[159,285,530,552]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFDF9FE8A4931" box="[159,276,530,552]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="applebyi">L. applebyi</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFDF8FC6C4931" box="[431,1010,530,552]" gridcol="1" gridrow="3" pageId="9" pageNumber="24">No Coppery gold to gold with black reticulations.</td>
<td id="76F95DFEFFDF004627B7FDF8FA1B4931" box="[1203,1413,530,552]" gridcol="2" gridrow="3" pageId="9" pageNumber="24">Rowley &amp; Cao 2009</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFDD7FA1B4948" blockId="9.[159,1413,571,1037]" box="[159,1413,571,594]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFDD7FE4A494B" authority="Yes" authorityName="Yes" box="[159,468,573,594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B96CEAD2FFDFFFB0239BFDD7FF62494B" box="[159,252,573,594]" italics="true" pageId="9" pageNumber="24">L. arayai</emphasis>
Yes
</taxonomicName>
Upper 100120° bright red-brown, rest whitish grey to light
<bibRefCitation id="EF894B31FFDFFFB027B7FDD1FA1B4948" author="Malkmus" box="[1203,1413,571,594]" pageId="9" pageNumber="24" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffman, P. &amp; Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North Borneo). ARG Ganter Verlag Kommanditgesellschaft, Ruggell. 424 pp." type="book" year="2002">
Malkmus
<emphasis id="B96CEAD2FFDFFFB0261DFDD7FADB494B" box="[1305,1349,572,594]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2002
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0211BFDB0FC344969" blockId="9.[159,1413,571,1037]" box="[543,938,602,624]" pageId="9" pageNumber="24">greyish green, with black reticulations.</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFD6FFC8E49A1" blockId="9.[159,1413,571,1037]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFD6FFEAD4983" box="[159,307,644,666]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="bidoupensis" status="sp. nov.">
<emphasis id="B96CEAD2FFDFFFB0239BFD6FFEAD4983" box="[159,307,644,666]" italics="true" pageId="9" pageNumber="24">L. bidoupensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25F57A9FFDFFFB0223DFD6EFE194980" box="[313,391,644,665]" pageId="9" pageNumber="24" rank="species">
<emphasis id="B96CEAD2FFDFFFB0223DFD6EFE1C4980" bold="true" box="[313,386,644,665]" pageId="9" pageNumber="24">sp. nov</emphasis>
.
</taxonomicNameLabel>
* Yes Upper half coppery red, lower half fading to pale silver. Exten- Present paper sive black reticulations.
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFD27FB5E49FB" blockId="9.[159,1413,571,1037]" box="[159,1216,716,738]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFD27FE9249FB" box="[159,268,716,738]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B96CEAD2FFDFFFB0239BFD27FE9249FB" box="[159,268,716,738]" italics="true" pageId="9" pageNumber="24">L. bourreti</emphasis>
</taxonomicName>
??
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFD1CFAE14813" blockId="9.[159,1413,571,1037]" box="[159,1407,757,779]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFD1CFDD54812" authority="No Pale" authorityName="No Pale" box="[159,587,757,779]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B96CEAD2FFDFFFB0239BFD1CFE8A4812" box="[159,276,758,779]" italics="true" pageId="9" pageNumber="24">L. croceus*</emphasis>
No Pale
</taxonomicName>
gold with distinct, black reticulations encircling the pupil.
<bibRefCitation id="EF894B31FFDFFFB027B0FD1FFAE14813" author="Rowley" box="[1204,1407,757,779]" pageId="9" pageNumber="24" refString="Rowley, J. J. L., Hoang, D. H., Le, T. T. D., Dau, Q. V. &amp; Cao, T. T. (2010 a) A new species of Leptolalax (Anura: Megophryidae) from Vietnam and further information on Leptolalax tuberosus. Zootaxa, 2660, 33 - 45." type="journal article" year="2010" yearSuffix="a">
Rowley
<emphasis id="B96CEAD2FFDFFFB0260CFD1CFAAA4812" box="[1288,1332,757,779]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2010a
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFCCAFAE3484A" blockId="9.[159,1413,571,1037]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFCCAFE4A482D" authority="Yes" authorityName="Yes" box="[159,468,799,821]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B96CEAD2FFDFFFB0239BFCCAFF66482C" box="[159,248,799,821]" italics="true" pageId="9" pageNumber="24">L. dringi</emphasis>
Yes
</taxonomicName>
Upper section reddish, rest pale golden grey. Black reticula- Inger &amp; Stuebing tions. 2005 (from Fig. 19)
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFC82FAAC4865" blockId="9.[159,1413,571,1037]" box="[159,1330,870,893]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFC82FE4A4865" authority="Yes" authorityName="Yes" box="[159,468,871,893]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B96CEAD2FFDFFFB0239BFC82FEA84864" box="[159,310,871,893]" italics="true" pageId="9" pageNumber="24">L. fuliginosus*</emphasis>
Yes
</taxonomicName>
Upper half reddish orange, lower half gold. Black reticulations.
<bibRefCitation id="EF894B31FFDFFFB027B7FC8CFAAC4865" author="Matsui" box="[1203,1330,870,892]" pageId="9" pageNumber="24" refString="Matsui, M. (2006) Three new species of Leptolalax from Thailand (Amphibia, Anura, Megophryidae). Zoological Science, 23, 821 - 830." type="journal article" year="2006">Matsui 2006</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFC7BFAE248DC" blockId="9.[159,1413,571,1037]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFC7BFE4A48BF" authority="Yes" authorityName="Yes" box="[159,468,912,934]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B96CEAD2FFDFFFB0239BFC7BFE9948BF" box="[159,263,912,934]" italics="true" pageId="9" pageNumber="24">L. gracilis</emphasis>
Yes
</taxonomicName>
Upper section reddish orange, rest pale golden grey. Black Inger &amp; Stuebing reticulations. 2005 (from Fig. 20)
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFC33FAD34FA8" blockId="9.[159,1413,571,1037]" lastBlockId="9.[159,1357,1179,1232]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFC33FE4A48F7" authority="Yes" authorityName="Yes" box="[159,468,984,1006]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B96CEAD2FFDFFFB0239BFC33FE9E48F7" box="[159,256,984,1006]" italics="true" pageId="9" pageNumber="24">L. hamidi</emphasis>
Yes
</taxonomicName>
Upper half reddish, rest pale golden grey, with black reticula- Inger &amp; Stuebing tions. 2005 (from Fig. 21)
<taxonomicName id="4C184D43FFDFFFB0239BFB77FE4A4FAB" authority="Yes" authorityName="Yes" box="[159,468,1180,1202]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="khasiorum">
<emphasis id="B96CEAD2FFDFFFB0239BFB77FEAF4FAB" box="[159,305,1180,1202]" italics="true" pageId="9" pageNumber="24">L. khasiorum*</emphasis>
Yes
</taxonomicName>
Upper third bright orange, rest yellowish cream. Black reticula-
<bibRefCitation id="EF894B31FFDFFFB027B7FB77FAD34FA8" author="Das" box="[1203,1357,1180,1202]" pageId="9" pageNumber="24" refString="Sengupta, S., Sailo, S., Lalremsanga, H. T., Das, A. &amp; Das, I. (2010) A new species of Leptolalax (Anura: Megophryidae) from Mizoram, North-eastern India, Zootaxa, 2406, 57 - 68." type="journal article" year="2010">
Das
<emphasis id="B96CEAD2FFDFFFB027E5FB77FA8A4FAB" box="[1249,1300,1180,1202]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
2010
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFBCBFAF54F91" pageId="9" pageNumber="24">
<table id="F918C460FFDF00462393FBCAFA1B4F91" box="[151,1413,1056,1160]" gridcols="4" gridrows="3" pageId="9" pageNumber="24">
<tr id="35283482FFDF00462393FBCAFA1B4F2F" box="[151,1413,1056,1078]" gridrow="0" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FBCAFEA04F2F" box="[151,318,1056,1078]" gridcol="0" gridrow="0" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFBCBFEB74F2F" box="[159,297,1056,1078]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFBCBFE814F2F" box="[159,287,1056,1078]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
*
</emphasis>
</th>
<th id="76F95DFEFFDF004622ABFBCAFE504F2F" box="[431,462,1056,1078]" gridcol="1" gridrow="0" pageId="9" pageNumber="24">?</th>
<th id="76F95DFEFFDF0046211BFBCAFC3C4F2F" box="[543,930,1056,1078]" gridcol="2" gridrow="0" pageId="9" pageNumber="24">?</th>
<th id="76F95DFEFFDF004627B7FBCAFA1B4F2F" box="[1203,1413,1056,1078]" gridcol="3" gridrow="0" pageId="9" pageNumber="24"></th>
</tr>
<tr id="35283482FFDF00462393FBA3FA1B4F46" box="[151,1413,1097,1119]" gridrow="1" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FBA3FEA04F46" box="[151,318,1097,1119]" gridcol="0" gridrow="1" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFBA0FEA04F46" box="[159,318,1097,1119]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFBA0FEAA4F46" box="[159,308,1097,1119]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFBA3FE504F46" box="[431,462,1097,1119]" gridcol="1" gridrow="1" pageId="9" pageNumber="24">No</td>
<td id="76F95DFEFFDF0046211BFBA3FC3C4F46" box="[543,930,1097,1119]" gridcol="2" gridrow="1" pageId="9" pageNumber="24">Metallic gold with black reticulations.</td>
<td id="76F95DFEFFDF004627B7FBA3FA1B4F46" box="[1203,1413,1097,1119]" gridcol="3" gridrow="1" pageId="9" pageNumber="24">
Grismer
<emphasis id="B96CEAD2FFDFFFB02608FBA0FADE4F46" box="[1292,1344,1097,1119]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
2004
</td>
</tr>
<tr id="35283482FFDF00462393FB98FA1B4F91" box="[151,1413,1138,1160]" gridrow="2" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FB98FEA04F91" box="[151,318,1138,1160]" gridcol="0" gridrow="2" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFB99FF684F91" box="[159,246,1138,1160]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFB99FF724F91" box="[159,236,1138,1160]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFB98FE504F91" box="[431,462,1138,1160]" gridcol="1" gridrow="2" pageId="9" pageNumber="24">No</td>
<td id="76F95DFEFFDF0046211BFB98FC3C4F91" box="[543,930,1138,1160]" gridcol="2" gridrow="2" pageId="9" pageNumber="24">Dark red.</td>
<td id="76F95DFEFFDF004627B7FB98FA1B4F91" box="[1203,1413,1138,1160]" gridcol="3" gridrow="2" pageId="9" pageNumber="24">
Matsui
<emphasis id="B96CEAD2FFDFFFB02604FB99FAAD4F91" box="[1280,1331,1138,1160]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
2009
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0211BFB50FDC94FC9" blockId="9.[159,1357,1179,1232]" box="[543,599,1210,1232]" pageId="9" pageNumber="24">tions.</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BFB0FFAEB4C05" pageId="9" pageNumber="24">
<table id="F918C460FFDF00462393FB0EFA1B4C04" box="[151,1413,1252,1821]" gridcols="4" gridrows="12" pageId="9" pageNumber="24">
<tr id="35283482FFDF00462393FB0EFA1B4FE3" box="[151,1413,1252,1274]" gridrow="0" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FB0EFECC4FE3" box="[151,338,1252,1274]" gridcol="0" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFB0FFE914FE3" box="[159,271,1252,1274]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B96CEAD2FFDFFFB0239BFB0FFE914FE3" box="[159,271,1252,1274]" italics="true" pageId="9" pageNumber="24">L. lateralis</emphasis>
</taxonomicName>
</th>
<th id="76F95DFEFFDF004622ABFB0EFE474FE3" box="[431,473,1252,1274]" gridcol="1" gridrow="0" pageId="9" pageNumber="24">?</th>
<th id="76F95DFEFFDF0046211BFB0EFBE44FE3" box="[543,1146,1252,1274]" gridcol="2" gridrow="0" pageId="9" pageNumber="24">?</th>
<th id="76F95DFEFFDF004627B7FB0EFA1B4FE3" box="[1203,1413,1252,1274]" gridcol="3" gridrow="0" pageId="9" pageNumber="24"></th>
</tr>
<tr id="35283482FFDF00462393FAE7FA1B4E3A" box="[151,1413,1293,1315]" gridrow="1" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FAE7FECC4E3A" box="[151,338,1293,1315]" gridcol="0" gridrow="1" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFAE4FF424E3A" box="[159,220,1293,1315]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="liui">
<emphasis id="B96CEAD2FFDFFFB0239BFAE4FF424E3A" box="[159,220,1293,1315]" italics="true" pageId="9" pageNumber="24">L. liui</emphasis>
</taxonomicName>
</th>
<td id="76F95DFEFFDF004622ABFAE7FE474E3A" box="[431,473,1293,1315]" gridcol="1" gridrow="1" pageId="9" pageNumber="24">?</td>
<td id="76F95DFEFFDF0046211BFAE7FBE44E3A" box="[543,1146,1293,1315]" gridcol="2" gridrow="1" pageId="9" pageNumber="24">?</td>
<td id="76F95DFEFFDF004627B7FAE7FA1B4E3A" box="[1203,1413,1293,1315]" gridcol="3" gridrow="1" pageId="9" pageNumber="24"></td>
</tr>
<tr id="35283482FFDF00462393FADCFA1B4E54" box="[151,1413,1334,1357]" gridrow="2" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FADCFECC4E54" box="[151,338,1334,1357]" gridcol="0" gridrow="2" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFAD2FE984E54" box="[159,262,1336,1357]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CEAD2FFDFFFB0239BFAD2FE984E54" box="[159,262,1336,1357]" italics="true" pageId="9" pageNumber="24">L. maurus</emphasis>
</taxonomicName>
</th>
<td id="76F95DFEFFDF004622ABFADCFE474E54" box="[431,473,1334,1357]" gridcol="1" gridrow="2" pageId="9" pageNumber="24">No</td>
<td id="76F95DFEFFDF0046211BFADCFBE44E54" box="[543,1146,1334,1357]" gridcol="2" gridrow="2" pageId="9" pageNumber="24">Reddish brown.</td>
<td id="76F95DFEFFDF004627B7FADCFA1B4E54" box="[1203,1413,1334,1357]" gridcol="3" gridrow="2" pageId="9" pageNumber="24">
Malkmus
<emphasis id="B96CEAD2FFDFFFB0261DFAD2FADB4E54" box="[1305,1349,1335,1357]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2002
</td>
</tr>
<tr id="35283482FFDF00462393FA8AFA1B4E6F" box="[151,1413,1376,1398]" gridrow="3" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FA8AFECC4E6F" box="[151,338,1376,1398]" gridcol="0" gridrow="3" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFA8BFECC4E6F" box="[159,338,1376,1398]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFA8BFED64E6F" box="[159,328,1376,1398]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFA8AFE474E6F" box="[431,473,1376,1398]" gridcol="1" gridrow="3" pageId="9" pageNumber="24">Yes</td>
<td id="76F95DFEFFDF0046211BFA8AFBE44E6F" box="[543,1146,1376,1398]" gridcol="2" gridrow="3" pageId="9" pageNumber="24">Upper half orange, lower half silver. Black reticulations.</td>
<td id="76F95DFEFFDF004627B7FA8AFA1B4E6F" box="[1203,1413,1376,1398]" gridcol="3" gridrow="3" pageId="9" pageNumber="24">Matsui 2006</td>
</tr>
<tr id="35283482FFDF00462393FA63FA1B4E86" box="[151,1413,1417,1439]" gridrow="4" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FA63FECC4E86" box="[151,338,1417,1439]" gridcol="0" gridrow="4" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFA60FE8D4E86" box="[159,275,1417,1439]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFA60FE944E86" box="[159,266,1417,1439]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="melicus">L. melicus</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFA63FE474E86" box="[431,473,1417,1439]" gridcol="1" gridrow="4" pageId="9" pageNumber="24">No</td>
<td id="76F95DFEFFDF0046211BFA63FBE44E86" box="[543,1146,1417,1439]" gridcol="2" gridrow="4" pageId="9" pageNumber="24">Dark gold. Black reticulations.</td>
<td id="76F95DFEFFDF004627B7FA63FA1B4E86" box="[1203,1413,1417,1439]" gridcol="3" gridrow="4" pageId="9" pageNumber="24">
Rowley
<emphasis id="B96CEAD2FFDFFFB0260CFA60FAA24E86" box="[1288,1340,1417,1439]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
2010b
</td>
</tr>
<tr id="35283482FFDF00462393FA58FA1B4ED1" box="[151,1413,1458,1480]" gridrow="5" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FA58FECC4ED1" box="[151,338,1458,1480]" gridcol="0" gridrow="5" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BFA59FEDB4ED1" box="[159,325,1458,1480]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFA59FEA54ED1" box="[159,315,1458,1480]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABFA58FE474ED1" box="[431,473,1458,1480]" gridcol="1" gridrow="5" pageId="9" pageNumber="24">No</td>
<td id="76F95DFEFFDF0046211BFA58FBE44ED1" box="[543,1146,1458,1480]" gridcol="2" gridrow="5" pageId="9" pageNumber="24">Gold with black reticulations.</td>
<td id="76F95DFEFFDF004627B7FA58FA1B4ED1" box="[1203,1413,1458,1480]" gridcol="3" gridrow="5" pageId="9" pageNumber="24">
Lathrop
<emphasis id="B96CEAD2FFDFFFB0260EFA59FAA34ED1" box="[1290,1341,1458,1480]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
1998
</td>
</tr>
<tr id="35283482FFDF00462393FA31FA1B4D08" box="[151,1413,1499,1553]" gridrow="6" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393FA31FECC4D08" box="[151,338,1499,1553]" gridcol="0" gridrow="6" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BFA37FEB74EEB" box="[159,297,1500,1522]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CEAD2FFDFFFB0239BFA37FEB74EEB" box="[159,297,1500,1522]" italics="true" pageId="9" pageNumber="24">L. oshanensis</emphasis>
</taxonomicName>
</th>
<td id="76F95DFEFFDF004622ABFA31FE474D08" box="[431,473,1499,1553]" gridcol="1" gridrow="6" pageId="9" pageNumber="24">No?</td>
<td id="76F95DFEFFDF0046211BFA31FBE44D08" box="[543,1146,1499,1553]" gridcol="2" gridrow="6" pageId="9" pageNumber="24">Copper/gold with black reticulations.</td>
<td id="76F95DFEFFDF004627B7FA31FA1B4D08" box="[1203,1413,1499,1553]" gridcol="3" gridrow="6" pageId="9" pageNumber="24">
Fei
<emphasis id="B96CEAD2FFDFFFB027DEFA37FA984EEB" box="[1242,1286,1500,1522]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2010 (from figures on p. 172)
</td>
</tr>
<tr id="35283482FFDF00462393F9CEFA1B4D23" box="[151,1413,1572,1594]" gridrow="7" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F9CEFECC4D23" box="[151,338,1572,1594]" gridcol="0" gridrow="7" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF9CFFEA44D23" box="[159,314,1572,1594]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B96CEAD2FFDFFFB0239BF9CFFEA44D23" box="[159,314,1572,1594]" italics="true" pageId="9" pageNumber="24">L. pelodytoides</emphasis>
</taxonomicName>
</th>
<td id="76F95DFEFFDF004622ABF9CEFE474D23" box="[431,473,1572,1594]" gridcol="1" gridrow="7" pageId="9" pageNumber="24">?</td>
<td id="76F95DFEFFDF0046211BF9CEFBE44D23" box="[543,1146,1572,1594]" gridcol="2" gridrow="7" pageId="9" pageNumber="24">?</td>
<td id="76F95DFEFFDF004627B7F9CEFA1B4D23" box="[1203,1413,1572,1594]" gridcol="3" gridrow="7" pageId="9" pageNumber="24"></td>
</tr>
<tr id="35283482FFDF00462393F9A7FA1B4D98" box="[151,1413,1613,1665]" gridrow="8" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F9A7FECC4D98" box="[151,338,1613,1665]" gridcol="0" gridrow="8" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BF9A4FE9D4D7A" box="[159,259,1614,1635]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF9A4FF674D7A" box="[159,249,1614,1635]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABF9A7FE474D98" box="[431,473,1613,1665]" gridcol="1" gridrow="8" pageId="9" pageNumber="24">Yes</td>
<td id="76F95DFEFFDF0046211BF9A7FBE44D98" box="[543,1146,1613,1665]" gridcol="2" gridrow="8" pageId="9" pageNumber="24">Upper 120° with segments of dark brown, rest golden-grey. Black reticulations</td>
<td id="76F95DFEFFDF004627B7F9A7FA1B4D98" box="[1203,1413,1613,1665]" gridcol="3" gridrow="8" pageId="9" pageNumber="24">
Malkmus
<emphasis id="B96CEAD2FFDFFFB0261DF9A4FADB4D7A" box="[1305,1349,1613,1635]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2002
</td>
</tr>
<tr id="35283482FFDF00462393F97FFA1B4DD3" box="[151,1413,1685,1738]" gridrow="9" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F97FFECC4DD3" box="[151,338,1685,1738]" gridcol="0" gridrow="9" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BF97CFE804DB2" box="[159,286,1685,1707]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF97CFE8A4DB2" box="[159,276,1685,1707]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABF97FFE474DD3" box="[431,473,1685,1738]" gridcol="1" gridrow="9" pageId="9" pageNumber="24">No?</td>
<td id="76F95DFEFFDF0046211BF97FFBE44DD3" box="[543,1146,1685,1738]" gridcol="2" gridrow="9" pageId="9" pageNumber="24">Brown?</td>
<td id="76F95DFEFFDF004627B7F97FFA1B4DD3" box="[1203,1413,1685,1738]" gridcol="3" gridrow="9" pageId="9" pageNumber="24">
Ohler
<emphasis id="B96CEAD2FFDFFFB027F7F97CFAB94DB2" box="[1267,1319,1685,1707]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
2000 (from Fig. 2)
</td>
</tr>
<tr id="35283482FFDF00462393F937FA1B4DEA" box="[151,1413,1757,1779]" gridrow="10" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F937FECC4DEA" box="[151,338,1757,1779]" gridcol="0" gridrow="10" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BF934FF644DEA" box="[159,250,1757,1779]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF934FF6E4DEA" box="[159,240,1757,1779]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="solus">L. solus</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABF937FE474DEA" box="[431,473,1757,1779]" gridcol="1" gridrow="10" pageId="9" pageNumber="24">Yes</td>
<td id="76F95DFEFFDF0046211BF937FBE44DEA" box="[543,1146,1757,1779]" gridcol="2" gridrow="10" pageId="9" pageNumber="24">Upper half dark red, lower half dark brown</td>
<td id="76F95DFEFFDF004627B7F937FA1B4DEA" box="[1203,1413,1757,1779]" gridcol="3" gridrow="10" pageId="9" pageNumber="24">Matsui 2006</td>
</tr>
<tr id="35283482FFDF00462393F8ECFA1B4C04" box="[151,1413,1798,1821]" gridrow="11" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F8ECFECC4C04" box="[151,338,1798,1821]" gridcol="0" gridrow="11" pageId="9" pageNumber="24">
<emphasis id="B96CEAD2FFDFFFB0239BF8E2FF634C05" box="[159,253,1799,1821]" italics="true" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF8E2FF6D4C05" box="[159,243,1799,1821]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
*
</emphasis>
</th>
<td id="76F95DFEFFDF004622ABF8ECFE474C04" box="[431,473,1798,1821]" gridcol="1" gridrow="11" pageId="9" pageNumber="24">No</td>
<td id="76F95DFEFFDF0046211BF8ECFBE44C04" box="[543,1146,1798,1821]" gridcol="2" gridrow="11" pageId="9" pageNumber="24">Gold-green. Black reticulations only at edges</td>
<td id="76F95DFEFFDF004627B7F8ECFA1B4C04" box="[1203,1413,1798,1821]" gridcol="3" gridrow="11" pageId="9" pageNumber="24">
Lathrop
<emphasis id="B96CEAD2FFDFFFB0260EF8E2FAA34C04" box="[1290,1341,1799,1821]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
1998
</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BF8DBFA1A4C5C" blockId="9.[159,1412,1840,1892]" box="[159,1412,1840,1862]" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF8DBFE4A4C5F" authority="Yes" authorityName="Yes" box="[159,468,1840,1862]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B96CEAD2FFDFFFB0239BF8DBFE994C5F" box="[159,263,1840,1862]" italics="true" pageId="9" pageNumber="24">L. tamdil*</emphasis>
Yes
</taxonomicName>
Upper third bright orange, rest greyish cream, edged with black
<bibRefCitation id="EF894B31FFDFFFB027B7F8DAFA1A4C5C" author="Sengupta" box="[1203,1412,1840,1862]" pageId="9" pageNumber="24" refString="Sengupta, S., Sailo, S., Lalremsanga, H. T., Das, A. &amp; Das, I. (2010) A new species of Leptolalax (Anura: Megophryidae) from Mizoram, North-eastern India, Zootaxa, 2406, 57 - 68." type="journal article" year="2010">
Sengupta
<emphasis id="B96CEAD2FFDFFFB0261CF8DBFAD54C5F" box="[1304,1355,1840,1862]" italics="true" pageId="9" pageNumber="24">et al.</emphasis>
2010
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDFFFB0211BF8A4FD3D4C7D" blockId="9.[159,1412,1840,1892]" box="[543,675,1870,1892]" pageId="9" pageNumber="24">reticulations.</paragraph>
<paragraph id="8BA736C0FFDFFFB0239BF893FE6C4CF6" pageId="9" pageNumber="24">
<table id="F918C460FFDF00462393F892FA1B4CF6" box="[151,1413,1912,2031]" gridcols="3" gridrows="3" pageId="9" pageNumber="24">
<tr id="35283482FFDF00462393F892FA1B4C97" box="[151,1413,1912,1934]" gridrow="0" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F892FE6C4C97" box="[151,498,1912,1934]" gridcol="0" gridrow="0" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF893FE504C97" authority="No" authorityName="No" box="[159,462,1912,1934]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CEAD2FFDFFFB0239BF893FE824C97" box="[159,284,1912,1934]" italics="true" pageId="9" pageNumber="24">L. tuberosus</emphasis>
No
</taxonomicName>
</th>
<th id="76F95DFEFFDF0046211BF892FC584C97" box="[543,966,1912,1934]" gridcol="1" gridrow="0" pageId="9" pageNumber="24">Pale gold with minute black reticulations.</th>
<th id="76F95DFEFFDF004627B7F892FA1B4C97" box="[1203,1413,1912,1934]" gridcol="2" gridrow="0" pageId="9" pageNumber="24">
Rowley
<emphasis id="B96CEAD2FFDFFFB0260CF893FAAA4C97" box="[1288,1332,1912,1934]" italics="true" pageId="9" pageNumber="24">et al</emphasis>
. 2010a
</th>
</tr>
<tr id="35283482FFDF00462393F848FA1B4CAE" box="[151,1413,1954,1975]" gridrow="1" pageId="9" pageNumber="24">
<th id="76F95DFEFFDF00462393F848FE6C4CAE" box="[151,498,1954,1975]" gridcol="0" gridrow="1" pageId="9" pageNumber="24">
<taxonomicName id="4C184D43FFDFFFB0239BF848FEC94CAE" box="[159,343,1954,1975]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="24" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B96CEAD2FFDFFFB0239BF848FEC94CAE" box="[159,343,1954,1975]" italics="true" pageId="9" pageNumber="24">L. ventripunctatus</emphasis>
</taxonomicName>
?
</th>
<td id="76F95DFEFFDF0046211BF848FC584CAE" box="[543,966,1954,1975]" gridcol="1" gridrow="1" pageId="9" pageNumber="24">?</td>
<td id="76F95DFEFFDF004627B7F848FA1B4CAE" box="[1203,1413,1954,1975]" gridcol="2" gridrow="1" pageId="9" pageNumber="24"></td>
</tr>
<tr id="35283482FFDF00462393F833FA1B4CF6" box="[151,1413,2009,2031]" gridrow="2" pageId="9" pageNumber="24" rowspan-1="1" rowspan-2="1">
<th id="76F95DFEFFDF00462393F833FE6C4CF6" box="[151,498,2009,2031]" gridcol="0" gridrow="2" pageId="9" pageNumber="24">* documented for type specimens.</th>
</tr>
</table>
</paragraph>
<caption id="DF676648FFDCFFB32373FDEFFB614905" box="[119,1279,516,541]" pageId="10" pageNumber="25">
<paragraph id="8BA736C0FFDCFFB32373FDEFFB614905" blockId="10.[119,1279,516,541]" box="[119,1279,516,541]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB32373FDEFFF744904" bold="true" box="[119,234,517,541]" pageId="10" pageNumber="25">TABLE 4.</emphasis>
Summary of male advertisement call parameters for species of
<taxonomicName id="4C184D43FFDCFFB3208BFDEEFB9F4904" box="[911,1025,516,541]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B96CEAD2FFDCFFB3208BFDEEFB9F4904" box="[911,1025,516,541]" italics="true" pageId="10" pageNumber="25">Leptolalax</emphasis>
</taxonomicName>
with documented calls.
</paragraph>
</caption>
<paragraph id="8BA736C0FFDCFFB3238AFDBDF89C4972" blockId="10.[142,1930,599,1689]" box="[142,1794,599,619]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB3238AFDBDF89C4972" bold="true" box="[142,1794,599,619]" pageId="10" pageNumber="25">Species Notes/ call Call Call Variable note Pulses/note Note duration Dominant Frequency Temp ºC Source</emphasis>
</paragraph>
<paragraph id="8BA736C0FFDCFFB322C0FD94FA71498B" blockId="10.[142,1930,599,1689]" box="[452,1519,638,658]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB322C0FD94FA71498B" bold="true" box="[452,1519,638,658]" pageId="10" pageNumber="25">duration repetition structure (ms) frequency modulation</emphasis>
</paragraph>
<paragraph id="8BA736C0FFDCFFB322C0FD4CFAB149A3" blockId="10.[142,1930,599,1689]" box="[452,1327,678,698]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB322C0FD4CFAB149A3" bold="true" box="[452,1327,678,698]" pageId="10" pageNumber="25">(ms) rate (calls/s) (kHz)</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C302654BFFDCFFB2238AFD27FBD54A05" lastPageId="11" lastPageNumber="26" pageId="10" pageNumber="25" type="reference_group">
<paragraph id="8BA736C0FFDCFFB3238AFD27FBB948B1" blockId="10.[142,1930,599,1689]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB3238AFD27FF4B49FB" box="[142,213,717,738]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFD27FF5249FB" ID-CoL="3TQPB" box="[142,204,717,738]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="aereus">aereus</taxonomicName>
*
</emphasis>
<date id="FFA61000FFDCFFB3223FFD27FE4749FB" box="[315,473,717,738]" pageId="10" pageNumber="25" value="1913-02-16">216 13</date>
130 2.48.0 Slight
<date id="FFA61000FFDCFFB32088FD27FBEF49FB" box="[908,1137,717,738]" pageId="10" pageNumber="25" value="1930-03-01" valueMax="1930-03-02" valueMin="1930-03-01">12 330</date>
6.27.9 No 22.425.7
<bibRefCitation id="EF894B31FFDCFFB325C5FD27F8E649FB" author="Rowley" box="[1729,1912,717,738]" pageId="10" pageNumber="25" refString="Rowley, J. J. L., Stuart, B. L., Richards, S. J., Phimmachak, S. &amp; Sivongxay, N. (2010 c) A new species of Leptolalax (Anura: Megophryidae) from Laos. Zootaxa, 2681, 35 - 46." type="journal article" year="2010" yearSuffix="c">
Rowley
<emphasis id="B96CEAD2FFDCFFB32408FD27F8A549FB" box="[1804,1851,717,738]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2010c
</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFD1EFF534810" box="[142,205,756,777]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="alpinis">
<emphasis id="B96CEAD2FFDCFFB3238AFD1EFF534810" box="[142,205,756,777]" italics="true" pageId="10" pageNumber="25">alpinis</emphasis>
</taxonomicName>
9.45±2.73 270±69? No? 1? 3 6.7±0.1 No? 16
<bibRefCitation id="EF894B31FFDCFFB325C5FD1FF8DA4813" author="Xu" box="[1729,1860,756,778]" pageId="10" pageNumber="25" refString="Xu, J-X., Xie, F., Jiang, J-P., Mo, Y-M. &amp; Zheng, Z-H. (2005) The acoustic features of the mating call of 12 anuran species. Chinese Journal of Zoology, 40, 12 - 19." type="journal article" year="2005">
Xu
<emphasis id="B96CEAD2FFDCFFB325E7FD1EF88F4810" box="[1763,1809,756,777]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2005
</bibRefCitation>
<emphasis id="B96CEAD2FFDCFFB3238AFCF6FF764828" box="[142,232,796,817]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFCF6FF7E4828" ID-CoL="3TQPD" box="[142,224,796,817]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="applebyi">applebyi</taxonomicName>
*
</emphasis>
45 280441 1.7 No
<date id="FFA61000FFDCFFB32088FCF6FBEE4828" box="[908,1136,796,817]" pageId="10" pageNumber="25" value="1932-04-01" valueMax="1932-04-05" valueMin="1932-04-01">15 432</date>
4.04.3 No 21.5
<bibRefCitation id="EF894B31FFDCFFB325C5FCF6F8E24828" author="Rowley" box="[1729,1916,796,817]" pageId="10" pageNumber="25" refString="Rowley, J. J. L, &amp; Cao, T. T. (2009) A new species of Leptolalax (Anura: Megophryidae) from central Vietnam. Zootaxa, 2198, 51 - 60." type="journal article" year="2009">Rowley &amp; Cao 2009</bibRefCitation>
<emphasis id="B96CEAD2FFDCFFB3238AFCA9FF4B4841" box="[142,213,835,856]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFCA9FF534841" ID-CoL="6PGC3" box="[142,205,835,856]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="arayai">arayai</taxonomicName>
*
</emphasis>
2 29 9.09.3 Note 1 shorter 1? 5.45.8 Slight decrease 17.4
<bibRefCitation id="EF894B31FFDCFFB325C4FCAEF8F94840" author="Matsui" box="[1728,1895,835,857]" pageId="10" pageNumber="25" refString="Matsui, M. (1997) Call characteristics of Malaysian Leptolalax with a description of two new species (Anura: Pelobatidae). Copeia, 1997, 158 - 165." type="journal article" year="1997">
Matsui
<emphasis id="B96CEAD2FFDCFFB32402FCA9F8AB4841" box="[1798,1845,835,856]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
1997
</bibRefCitation>
<emphasis id="B96CEAD2FFDCFFB3238AFC81FE994899" box="[142,263,875,896]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFC81FF604899" ID-CoL="3TQPG" box="[142,254,875,896]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="bidoupensis">bidoupensis</taxonomicName>
*
</emphasis>
69 277543 0.10.2 No
<date id="FFA61000FFDCFFB32088FC81FBEE4899" box="[908,1136,875,896]" pageId="10" pageNumber="25" value="1913-01-01">1 113</date>
1.93.8 Very slight 1921 Present paper
<emphasis id="B96CEAD2FFDCFFB3238AFC78FF4148BE" box="[142,223,914,935]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFC78FF4848BE" ID-CoL="3TQPK" box="[142,214,914,935]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="croceus">croceus</taxonomicName>
*
</emphasis>
131 (variable
</paragraph>
<paragraph id="8BA736C0FFDCFFB3223FFC50F8F64F8C" blockId="10.[142,1930,599,1689]" pageId="10" pageNumber="25">
46 164249 0.10.7 Yes note
<typeStatus id="54A38862FFDCFFB320BDFC50FC7F48D6" box="[953,993,954,975]" pageId="10" pageNumber="25">type</typeStatus>
) Variable 4.63.0 No 21.625.1
<bibRefCitation id="EF894B31FFDCFFB325C4FC50F8E748D6" author="Rowley" box="[1728,1913,954,975]" pageId="10" pageNumber="25" refString="Rowley, J. J. L., Hoang, D. H., Le, T. T. D., Dau, Q. V. &amp; Cao, T. T. (2010 a) A new species of Leptolalax (Anura: Megophryidae) from Vietnam and further information on Leptolalax tuberosus. Zootaxa, 2660, 33 - 45." type="journal article" year="2010" yearSuffix="a">
Rowley
<emphasis id="B96CEAD2FFDCFFB32408FC50F8A548D6" box="[1804,1851,954,975]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2010a
</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFC08FF5848EE" ID-CoL="3TQPL" box="[142,198,994,1015]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="dringi">
<emphasis id="B96CEAD2FFDCFFB3238AFC08FF5848EE" box="[142,198,994,1015]" italics="true" pageId="10" pageNumber="25">dringi</emphasis>
</taxonomicName>
<date id="FFA61000FFDCFFB3223FFC08FE7848EE" box="[315,486,994,1015]" pageId="10" pageNumber="25" value="1947-04-02">24 47</date>
11.011.5 No?? 7.68.1 Strongdecrease 24.3
<bibRefCitation id="EF894B31FFDCFFB325C4FC08F8F648EE" author="Matsui" box="[1728,1896,994,1015]" pageId="10" pageNumber="25" refString="Matsui, M. (1997) Call characteristics of Malaysian Leptolalax with a description of two new species (Anura: Pelobatidae). Copeia, 1997, 158 - 165." type="journal article" year="1997">
Matsui
<emphasis id="B96CEAD2FFDCFFB32402FC08F8B048EE" box="[1798,1838,994,1015]" italics="true" pageId="10" pageNumber="25">et al</emphasis>
. 1997
</bibRefCitation>
<emphasis id="B96CEAD2FFDCFFB3238AFBE3FF604F07" box="[142,254,1033,1054]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFBE3FF684F07" ID-CoL="3TQPQ" box="[142,246,1033,1054]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="fuliginosus">fuliginosus</taxonomicName>
*
</emphasis>
<date id="FFA61000FFDCFFB3223FFBE3FE444F07" box="[315,474,1033,1054]" pageId="10" pageNumber="25" value="1951-09-06">69 51</date>
80 2.9 No??? 2.32.4 No 19.319.6
<bibRefCitation id="EF894B31FFDCFFB325C4FBE3F8AA4F07" author="Matsui" box="[1728,1844,1033,1054]" pageId="10" pageNumber="25" refString="Matsui, M. (2006) Three new species of Leptolalax from Thailand (Amphibia, Anura, Megophryidae). Zoological Science, 23, 821 - 830." type="journal article" year="2006">Matsui2006</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFBDBFF4A4F5F" ID-CoL="3TQPR" box="[142,212,1073,1094]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B96CEAD2FFDCFFB3238AFBDBFF4A4F5F" box="[142,212,1073,1094]" italics="true" pageId="10" pageNumber="25">gracilis</emphasis>
</taxonomicName>
<date id="FFA61000FFDCFFB3223FFBDBFE454F5F" box="[315,475,1073,1094]" pageId="10" pageNumber="25" value="1947-04-03">34 47</date>
6.97.2 No 1?? 2.52.7 Slight 20.0
<bibRefCitation id="EF894B31FFDCFFB325C4FBDBF8F94F5F" author="Matsui" box="[1728,1895,1073,1094]" pageId="10" pageNumber="25" refString="Matsui, M. (1997) Call characteristics of Malaysian Leptolalax with a description of two new species (Anura: Pelobatidae). Copeia, 1997, 158 - 165." type="journal article" year="1997">
Matsui
<emphasis id="B96CEAD2FFDCFFB32402FBDBF8AB4F5F" box="[1798,1845,1073,1094]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
1997
</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFBB2FF504F74" ID-CoL="3TQPS" box="[142,206,1112,1133]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="hamidi">
<emphasis id="B96CEAD2FFDCFFB3238AFBB2FF504F74" box="[142,206,1112,1133]" italics="true" pageId="10" pageNumber="25">hamidi</emphasis>
</taxonomicName>
<date id="FFA61000FFDCFFB3223FFBB2FDDE4F74" box="[315,576,1112,1133]" pageId="10" pageNumber="25" value="1933-02-09" valueMax="1933-03-09" valueMin="1933-02-09">23 33 9</date>
9.3 Note 1 longer??? 6.77.3 Strong decrease 22.9
<bibRefCitation id="EF894B31FFDCFFB325C4FBB2F8F84F74" author="Matsui" box="[1728,1894,1112,1133]" pageId="10" pageNumber="25" refString="Matsui, M. (1997) Call characteristics of Malaysian Leptolalax with a description of two new species (Anura: Pelobatidae). Copeia, 1997, 158 - 165." type="journal article" year="1997">Matsui et al. 1997</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFB6AFF794F8C" ID-CoL="3TQPT" box="[142,231,1152,1173]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="heteropus">
<emphasis id="B96CEAD2FFDCFFB3238AFB6AFF794F8C" box="[142,231,1152,1173]" italics="true" pageId="10" pageNumber="25">heteropus</emphasis>
</taxonomicName>
36 207513? Yes Note 1: c. 20 Note 1: 80 2.8 Slight 21.0
<bibRefCitation id="EF894B31FFDCFFB325C4FB6AF8F64F8C" author="Matsui" box="[1728,1896,1152,1173]" pageId="10" pageNumber="25" refString="Matsui, M. (1997) Call characteristics of Malaysian Leptolalax with a description of two new species (Anura: Pelobatidae). Copeia, 1997, 158 - 165." type="journal article" year="1997">
Matsui
<emphasis id="B96CEAD2FFDCFFB32402FB6AF8B04F8C" box="[1798,1838,1152,1173]" italics="true" pageId="10" pageNumber="25">et al</emphasis>
. 1997
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDCFFB32088FB4DFB5D4FA5" blockId="10.[142,1930,599,1689]" box="[908,1219,1191,1212]" pageId="10" pageNumber="25">Notes 23: 56 Notes 23: 28</paragraph>
<paragraph id="8BA736C0FFDCFFB3238AFB25F8144E12" blockId="10.[142,1930,599,1689]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB3238AFB25FF5A4FFD" box="[142,196,1231,1252]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFB25FF254FFD" ID-CoL="3TQPX" box="[142,187,1231,1252]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="kecil">kecil</taxonomicName>
*
</emphasis>
4 110130 c. 0.2 No? 12? 3.2 Very weak 18.521.4
<bibRefCitation id="EF894B31FFDCFFB325C4FB25F8F94FFD" author="Matsui" box="[1728,1895,1231,1252]" pageId="10" pageNumber="25" refString="Matsui, M., Belabut, D. M., Ahmad, N. &amp; Yong, H-S. (2009) A New species of Leptolalax (Amphibia, Anura, Megophryidae) from peninsular Malaysia. Zoological Science, 26, 243 - 247." type="journal article" year="2009">
Matsui
<emphasis id="B96CEAD2FFDCFFB32402FB25F8AB4FFD" box="[1798,1845,1231,1252]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2009
</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFB1CFF4D4E12" ID-CoL="3TQQ9" box="[142,211,1270,1291]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="maurus">
<emphasis id="B96CEAD2FFDCFFB3238AFB1CFF4D4E12" box="[142,211,1270,1291]" italics="true" pageId="10" pageNumber="25">maurus</emphasis>
</taxonomicName>
4 85 No 1 8 5.2 No?
<bibRefCitation id="EF894B31FFDCFFB325C4FB1CF8144E12" author="Sukumaran" box="[1728,1930,1270,1291]" pageId="10" pageNumber="25" refString="Sukumaran, J., Das, I. &amp; Haas, A. (2010) Descriptions of the advertisement calls of some Bornean frogs. Russian Journal of Herpetology, 17, 189 - 194." type="journal article" year="2010">
Sukumaran
<emphasis id="B96CEAD2FFDCFFB32441FB1CF8144E12" box="[1861,1930,1270,1291]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDCFFB325C4FAF4F9704E2A" blockId="10.[142,1930,599,1689]" box="[1728,1774,1310,1331]" pageId="10" pageNumber="25">2010</paragraph>
<paragraph id="8BA736C0FFDCFFB3238AFAAFF8E44E9B" blockId="10.[142,1930,599,1689]" pageId="10" pageNumber="25">
<emphasis id="B96CEAD2FFDCFFB3238AFAAFFE894E43" box="[142,279,1349,1370]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFAAFFE904E43" ID-CoL="3TQQB" box="[142,270,1349,1370]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="melanoleucus">melanoleucus</taxonomicName>
*
</emphasis>
<date id="FFA61000FFDCFFB3223FFAAFFE424E43" box="[315,476,1349,1370]" pageId="10" pageNumber="25" value="1940-04-03">34 40</date>
63 6.1 No??? 3.13.3 No 23.9
<bibRefCitation id="EF894B31FFDCFFB325C4FAAFF8AA4E43" author="Matsui" box="[1728,1844,1349,1370]" pageId="10" pageNumber="25" refString="Matsui, M. (2006) Three new species of Leptolalax from Thailand (Amphibia, Anura, Megophryidae). Zoological Science, 23, 821 - 830." type="journal article" year="2006">Matsui2006</bibRefCitation>
<emphasis id="B96CEAD2FFDCFFB3238AFA87FF414E9B" box="[142,223,1389,1410]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFA87FF484E9B" ID-CoL="3TQQC" box="[142,214,1389,1410]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="melicus">melicus</taxonomicName>
*
</emphasis>
411 168484 0.70.8 Yes Note 1: 850 Note 1: 24132 2.64.0 No 26.126.2
<bibRefCitation id="EF894B31FFDCFFB325C4FA87F8E44E9B" author="Rowley" box="[1728,1914,1389,1410]" pageId="10" pageNumber="25" refString="Rowley, J. J. L., Stuart, B. L., Neang, T., Emmett, D. A. (2010 b) A new species of Leptolalax (Anura: Megophryidae) from northeastern Cambodia. Zootaxa, 2567, 57 - 68." type="journal article" year="2010" yearSuffix="b">
Rowley
<emphasis id="B96CEAD2FFDCFFB32408FA87F8A54E9B" box="[1804,1851,1389,1410]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2010b
</bibRefCitation>
</paragraph>
<paragraph id="8BA736C0FFDCFFB32088FA7EFB294EB0" blockId="10.[142,1930,599,1689]" box="[908,1207,1428,1449]" pageId="10" pageNumber="25">Clicks: 17 Clicks: 224</paragraph>
<paragraph id="8BA736C0FFDCFFB3238AFA56FC6A4D5E" blockId="10.[142,1930,599,1689]" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AFA56FF6D4EC8" ID-CoL="3TQQH" box="[142,243,1468,1489]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CEAD2FFDCFFB3238AFA56FF6D4EC8" box="[142,243,1468,1489]" italics="true" pageId="10" pageNumber="25">oshanensis</emphasis>
</taxonomicName>
3? c.23 No?? 2967 4.44.6 No? 14
<bibRefCitation id="EF894B31FFDCFFB325C4FA56F8C64EC8" author="Jiang" box="[1728,1880,1468,1489]" pageId="10" pageNumber="25" refString="Jiang, J. - P., Xie, F., Fei, L., Ye, C. &amp; Zheng, M-Q. (2002) Mating calls of six forms of pelobatid in Wawu Mountain national forest park, Sichuan, China (Anura: Pelobatidae). Zoological Research, 23, 89 - 94." type="journal article" year="2002">
Jiang
<emphasis id="B96CEAD2FFDCFFB325F2FA56F8BB4EC8" box="[1782,1829,1468,1489]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2002
</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AFA09FF5B4EE1" ID-CoL="3TQQM" box="[142,197,1507,1528]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CEAD2FFDCFFB3238AFA09FF5B4EE1" box="[142,197,1507,1528]" italics="true" pageId="10" pageNumber="25">pictus</emphasis>
</taxonomicName>
1 30
<date id="FFA61000FFDCFFB322E3FA09FE634EE1" box="[487,509,1507,1528]" pageId="10" pageNumber="25" value="1935-11-13">35</date>
1113 No? na 6.87.2 Strongdecrease 1922
<bibRefCitation id="EF894B31FFDCFFB325C4FA09F8E04EE1" author="Malkmus" box="[1728,1918,1507,1528]" pageId="10" pageNumber="25" refString="Malkmus, R., Manthey, U., Vogel, G., Hoffman, P. &amp; Kosuch, J. (2002) Amphibians and Reptiles of Mount Kinabalu (North Borneo). ARG Ganter Verlag Kommanditgesellschaft, Ruggell. 424 pp." type="book" year="2002">
Malkmus
<emphasis id="B96CEAD2FFDCFFB32418FA09F8DA4EE1" box="[1820,1860,1507,1528]" italics="true" pageId="10" pageNumber="25">et al</emphasis>
. 2002
</bibRefCitation>
<emphasis id="B96CEAD2FFDCFFB3238AF9E1FF564D39" box="[142,200,1547,1568]" italics="true" pageId="10" pageNumber="25">
<taxonomicName id="4C184D43FFDCFFB3238AF9E1FF214D39" ID-CoL="3TQQV" box="[142,191,1547,1568]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="solus">solus</taxonomicName>
*
</emphasis>
48 130260 2.12.3 No?? 3.13.2 No 24.224.3
<bibRefCitation id="EF894B31FFDCFFB325C4F9E1F8AA4D39" author="Matsui" box="[1728,1844,1547,1568]" pageId="10" pageNumber="25" refString="Matsui, M. (2006) Three new species of Leptolalax from Thailand (Amphibia, Anura, Megophryidae). Zoological Science, 23, 821 - 830." type="journal article" year="2006">Matsui2006</bibRefCitation>
<taxonomicName id="4C184D43FFDCFFB3238AF9D8FC6A4D5E" ID-CoL="3TQR3" authority="Indistinctly" authorityName="Indistinctly" box="[142,1012,1586,1607]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="10" pageNumber="25" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B96CEAD2FFDCFFB3238AF9D8FF784D5E" box="[142,230,1586,1607]" italics="true" pageId="10" pageNumber="25">tuberosus</emphasis>
Indistinctly
</taxonomicName>
</paragraph>
<paragraph id="8BA736C0FFDCFFB3223FF9B0FE644D80" blockId="10.[142,1930,599,1689]" pageId="10" pageNumber="25">
1 5478 0.10.5 No pulsed na 2.62.8 No 22.524.5
<bibRefCitation id="EF894B31FFDCFFB325C4F9B0F8E74D76" author="Rowley" box="[1728,1913,1626,1647]" pageId="10" pageNumber="25" refString="Rowley, J. J. L., Hoang, D. H., Le, T. T. D., Dau, Q. V. &amp; Cao, T. T. (2010 a) A new species of Leptolalax (Anura: Megophryidae) from Vietnam and further information on Leptolalax tuberosus. Zootaxa, 2660, 33 - 45." type="journal article" year="2010" yearSuffix="a">
Rowley
<emphasis id="B96CEAD2FFDCFFB32408F9B0F8A54D76" box="[1804,1851,1626,1647]" italics="true" pageId="10" pageNumber="25">et al.</emphasis>
2010a
</bibRefCitation>
* documented from
<typeStatus id="54A38862FFDCFFB3225CF96EFE1D4D80" box="[344,387,1668,1689]" pageId="10" pageNumber="25">type</typeStatus>
specimens.
</paragraph>
<paragraph id="8BA736C0FFDDFFB223C1FF7DFBD54A05" blockId="11.[151,1436,151,284]" pageId="11" pageNumber="26">
The advertisement call of
<taxonomicName id="4C184D43FFDDFFB222E8FF73FD0E4BA9" ID-CoL="3TQPG" box="[492,656,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDDFFB222E8FF73FD0E4BA9" box="[492,656,152,176]" italics="true" pageId="11" pageNumber="26">L. bidoupensis</emphasis>
</taxonomicName>
differs in terms of structure and/or frequency from all other
<taxonomicName id="4C184D43FFDDFFB22639FF72FF274BCD" ID-CoL="5D4M" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="genus">
<emphasis id="B96CEAD2FFDDFFB22639FF72FF274BCD" italics="true" pageId="11" pageNumber="26">Leptolalax</emphasis>
</taxonomicName>
with described calls (
<tableCitation id="C69A037BFFDDFFB222B0FF56FD904BCD" box="[436,526,188,212]" captionStart="TABLE 4" captionStartId="10.[119,208,517,541]" captionText="TABLE 4. Summary of male advertisement call parameters for species of Leptolalax with documented calls." pageId="11" pageNumber="26">Table 4</tableCitation>
). In addition, uncorrected sequence divergences between
<taxonomicName id="4C184D43FFDDFFB227A4FF57FAD84BCD" ID-CoL="3TQPG" box="[1184,1350,188,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B96CEAD2FFDDFFB227A4FF57FAD84BCD" box="[1184,1350,188,212]" italics="true" pageId="11" pageNumber="26">L. bidoupensis</emphasis>
</taxonomicName>
and all homologous 16S rRNA sequences available on GenBank (from individuals assigned to
<taxonomicName id="4C184D43FFDDFFB2277AFF0BFB634BE1" ID-CoL="3TQPD" box="[1150,1277,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B96CEAD2FFDDFFB2277AFF0BFB634BE1" box="[1150,1277,224,248]" italics="true" pageId="11" pageNumber="26">L. applebyi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D43FFDDFFB22609FF0BFAEB4BE1" ID-CoL="6PGC3" box="[1293,1397,225,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="arayai">
<emphasis id="B96CEAD2FFDDFFB22609FF0BFAEB4BE1" box="[1293,1397,225,248]" italics="true" pageId="11" pageNumber="26">L. arayai</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D43FFDDFFB22681FF0BFF6C4A05" ID-CoL="3TQPJ" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B96CEAD2FFDDFFB22681FF0BFF6C4A05" italics="true" pageId="11" pageNumber="26">L. bourreti</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D43FFDDFFB223FBFEEFFEDC4A05" ID-CoL="3TQQ3" box="[255,322,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="liui">
<emphasis id="B96CEAD2FFDDFFB223FBFEEFFEDC4A05" box="[255,322,260,284]" italics="true" pageId="11" pageNumber="26">L. liui</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D43FFDDFFB22254FEEFFE5D4A05" ID-CoL="3TQQC" box="[336,451,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B96CEAD2FFDDFFB22254FEEFFE5D4A05" box="[336,451,260,284]" italics="true" pageId="11" pageNumber="26">L. melicus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D43FFDDFFB222D6FEEFFDF44A05" ID-CoL="3TQQH" box="[466,618,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B96CEAD2FFDDFFB222D6FEEFFDF44A05" box="[466,618,260,284]" italics="true" pageId="11" pageNumber="26">L. oshanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C184D43FFDDFFB2217CFEEFFCBC4A05" ID-CoL="3TQQK" box="[632,802,260,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B96CEAD2FFDDFFB2217CFEEFFCBC4A05" box="[632,802,260,284]" italics="true" pageId="11" pageNumber="26">L. pelodytoides</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C184D43FFDDFFB22064FEEFFC5F4A05" ID-CoL="3TQQM" box="[864,961,261,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="11" pageNumber="26" phylum="Chordata" rank="species" species="pictus">
<emphasis id="B96CEAD2FFDDFFB22064FEEFFC5F4A05" box="[864,961,261,284]" italics="true" pageId="11" pageNumber="26">L. pictus</emphasis>
</taxonomicName>
) were&gt;9%.
</paragraph>
</subSubSection>
</treatment>
</document>