514 lines
65 KiB
XML
514 lines
65 KiB
XML
<document id="CA46B58F0A5604F9C445491751797B5F" ID-DOI="10.5281/zenodo.176897" ID-GBIF-Dataset="85773b3a-9274-4a30-ad96-ae4b27cc0ea5" ID-ISSN="1175-5326" ID-Zenodo-Dep="176897" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459842485466" checkinUser="plazi" docAuthor="Alonso, María R. & Ibáñez, Miguel" docDate="2007" docId="03B187BAC718FFD1A69B267A6CF5FA53" docLanguage="en" docName="zt01482p023.pdf" docOrigin="Zootaxa 1482" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hemicycla (Hemicycla) fulgida Alonso & Ibáñez, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="15" masterDocId="FF88FFC2C710FFDFA60C21216F69FF98" masterDocTitle="Anatomy and function of the penial twin papillae system of the Helicinae (Gastropoda: Helicoidea: Helicidae) and description of two new, small Hemicycla species from the laurel forest of the Canary Islands" masterLastPageNumber="23" masterPageNumber="1" pageNumber="9" updateTime="1698224473241" updateUser="plazi">
|
||
<mods:mods id="B492CAD6B61713526CE8E1DD53021166" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="D143AC085FF7BA80434FE29251516A79">
|
||
<mods:title id="B8FDB033CDBD9D916147759E2C215104">Anatomy and function of the penial twin papillae system of the Helicinae (Gastropoda: Helicoidea: Helicidae) and description of two new, small Hemicycla species from the laurel forest of the Canary Islands</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="50B0ECCD532E4124C9831EBB89848CDA" type="personal">
|
||
<mods:role id="C44D2A87FCA23984A7FDEC746514ED49">
|
||
<mods:roleTerm id="004EA75A52AF131021982FE8E3A0F8D5">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4C3A06ED378E795C1CB3DB040CA76B3F">Alonso, María R.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="117422990A2A8BA329C2B99C2A257879" type="personal">
|
||
<mods:role id="2AF782AC0E06BF67D7FE1624568421A6">
|
||
<mods:roleTerm id="53D22AAF12532C754FD4FEEF2FB0C750">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6B840ACC75F79F9901A74D29BDA08A67">Ibáñez, Miguel</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="508BD899F15C5BB494F3C30259DBFBC2">text</mods:typeOfResource>
|
||
<mods:relatedItem id="02CB49ED02410307090A12B7CCCC1026" type="host">
|
||
<mods:titleInfo id="989BAC7E70E4BCEA25DC646EE4D44D18">
|
||
<mods:title id="52989D6529474E39F08AFD16828CA8F9">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C39A33D58E5E8528D4C1FCBD749FA72B">
|
||
<mods:date id="935F4A9F806B0B639910C70E8515CF48">2007</mods:date>
|
||
<mods:detail id="DBCDB4468AE26B8E01B98CE9C91ABCAB" type="volume">
|
||
<mods:number id="81DB0DC4117ED98D3AAF9A449B85A9C8">1482</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="348B944CDF7F1741BEC57FD5A9AA8919" unit="page">
|
||
<mods:start id="ED7BDA9B2F27A18A7747EC1A214B73A4">1</mods:start>
|
||
<mods:end id="E7FB7AA61AF52BBE3C660BD75D9E7357">23</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="04AEE8BF6D3951EBB44E961668C875C2">journal article</mods:classification>
|
||
<mods:identifier id="79235D81EDEC5494CBC73C3B464881E6" type="DOI">10.5281/zenodo.176897</mods:identifier>
|
||
<mods:identifier id="7BF3640AA205A2C1A8A512CA421F222E" type="GBIF-Dataset">85773b3a-9274-4a30-ad96-ae4b27cc0ea5</mods:identifier>
|
||
<mods:identifier id="5746FA6CEA9AF1800383ABD36FC951BF" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="ECCE5A4B1598A24F4B7F08441B3E22AF" type="Zenodo-Dep">176897</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03B187BAC718FFD1A69B267A6CF5FA53" ID-DOI="http://doi.org/10.5281/zenodo.5612786" ID-GBIF-Taxon="119360616" ID-Zenodo-Dep="5612786" LSID="urn:lsid:plazi:treatment:03B187BAC718FFD1A69B267A6CF5FA53" httpUri="http://treatment.plazi.org/id/03B187BAC718FFD1A69B267A6CF5FA53" lastPageId="14" lastPageNumber="15" pageId="8" pageNumber="9">
|
||
<subSubSection id="C3026527C718FFD7A69B267A6D1CF8EC" box="[151,629,1882,1909]" pageId="8" pageNumber="9" type="nomenclature">
|
||
<paragraph id="8BA736ACC718FFD7A69B267A6D1CF8EC" blockId="8.[151,629,1882,1909]" box="[151,629,1882,1909]" pageId="8" pageNumber="9">
|
||
<heading id="D0EF81C0C718FFD7A69B267A6D1CF8EC" bold="true" box="[151,629,1882,1909]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
|
||
<emphasis id="B96CEABEC718FFD7A69B267A6D1CF8EC" bold="true" box="[151,629,1882,1909]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C184D2FC718FFD7A69B267A6D79F8ED" ID-CoL="3KMWK" box="[151,528,1882,1909]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov." subGenus="Hemicycla">
|
||
<emphasis id="B96CEABEC718FFD7A69B267A6E71F8ED" bold="true" box="[151,280,1883,1909]" italics="true" pageId="8" pageNumber="9">Hemicycla</emphasis>
|
||
(
|
||
<emphasis id="B96CEABEC718FFD7A724267A6EC0F8ED" bold="true" box="[296,425,1883,1909]" italics="true" pageId="8" pageNumber="9">Hemicycla</emphasis>
|
||
)
|
||
<emphasis id="B96CEABEC718FFD7A7B6267A6D79F8ED" bold="true" box="[442,528,1883,1909]" italics="true" pageId="8" pageNumber="9">fulgida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C718FFD7A414267B6D1CF8EC" box="[536,629,1882,1908]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3026527C718FFD6A69B26846DD5F9C6" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="materials_examined">
|
||
<paragraph id="8BA736ACC718FFD7A69B26846A6BF873" blockId="8.[151,1282,1957,2027]" pageId="8" pageNumber="9">
|
||
<emphasis id="B96CEABEC718FFD7A69B26846E5CF827" bold="true" box="[151,309,1957,1983]" pageId="8" pageNumber="9">
|
||
<typeStatus id="54A3880EC718FFD7A69B26846FBDF827" box="[151,212,1957,1983]" pageId="8" pageNumber="9">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
. Pista de Las Yedras, Anaga massif, Tenerife (UTM: 28
|
||
<collectionCode id="ED09AE69C718FFD7A5C926876C97F858" box="[965,1022,1958,1984]" httpUri="http://grbio.org/cool/bha5-99ng" name="Royal College of Surgeons" pageId="8" pageNumber="9">RCS</collectionCode>
|
||
<quantity id="4CE09B49C718FFD7A5F226876BF8F858" box="[1022,1169,1958,1984]" metricMagnitude="3" metricUnit="m" metricValue="7.456899999999999" pageId="8" pageNumber="9" unit="m" value="7456.9">7456, 900 m</quantity>
|
||
altitude).
|
||
<emphasis id="B96CEABEC718FFD7A6CB26F16E23F872" bold="true" box="[199,330,2000,2026]" pageId="8" pageNumber="9">
|
||
<typeStatus id="54A3880EC718FFD7A6CB26F16E50F872" box="[199,313,2000,2026]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
|
||
(
|
||
</emphasis>
|
||
<figureCitation id="13232A29C718FFD7A74626F06EE4F873" box="[330,397,2001,2027]" captionStart="FIGURE 8" captionStartId="12.[151,255,1235,1259]" captionTargetBox="[151,1434,194,1211]" captionTargetId="figure@12.[151,1436,194,1212]" captionTargetPageId="12" captionText="FIGURE 8. Shells from Tenerife. A. Hemicycla fulgida sp. nov., holotype. B. Hemicycla invernicata from Montaña Cabeza de Toro pine forest, near Las Lagunetas (La Esperanza forests). C. Hemicycla consobrina from Las Mercedes laurel forest." httpUri="https://zenodo.org/record/176904/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
|
||
A).
|
||
<collectionCode id="ED09AE69C718FFD7A7B026F06D65F873" box="[444,524,2001,2027]" pageId="8" pageNumber="9">TFMC</collectionCode>
|
||
(MT 0394); leg. M.R. Alonso and M. Ibáñez,
|
||
<date id="FFA6106CC718FFD7A22026F06B97F873" box="[1068,1278,1997,2027]" pageId="8" pageNumber="9" value="1993-01-15">15th January 1993</date>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF676624C719FFD6A69B248F6B49FA70" httpUri="https://zenodo.org/record/176902/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1436,155,1440]" targetPageId="9">
|
||
<paragraph id="8BA736ACC719FFD6A69B248F6B49FA70" blockId="9.[151,1435,1454,1512]" pageId="9" pageNumber="10">
|
||
<emphasis id="B96CEABEC719FFD6A69B248F6E70FA5E" bold="true" box="[151,281,1454,1478]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
|
||
Shell body whorl details. A–C.
|
||
<taxonomicName id="4C184D2FC719FFD6A462248E6C23FA5E" box="[622,842,1455,1478]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC719FFD6A462248E6C23FA5E" box="[622,842,1455,1478]" italics="true" pageId="9" pageNumber="10">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C719FFD6A55C248F6CCCFA5E" box="[848,933,1454,1478]" pageId="9" pageNumber="10" rank="species">
|
||
<emphasis id="B96CEABEC719FFD6A55C248F6CCCFA5E" bold="true" box="[848,933,1454,1478]" pageId="9" pageNumber="10">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype. D–F.
|
||
<taxonomicName id="4C184D2FC719FFD6A25B248E6A3CFA5E" box="[1111,1365,1455,1478]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC719FFD6A25B248E6A3CFA5E" box="[1111,1365,1455,1478]" italics="true" pageId="9" pageNumber="10">Hemicycla perraudierei</emphasis>
|
||
</taxonomicName>
|
||
. A, D, dorsal view. B, E, frontal view. C, F, ventral view. The sells are the same as in Fig. 4.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736ACC719FFD6A6CB273D6DD5F9C6" blockId="9.[151,1437,1564,2030]" pageId="9" pageNumber="10">
|
||
<typeStatus id="54A3880EC719FFD6A6CB273D6E2DF9AE" box="[199,324,1564,1590]" pageId="9" pageNumber="10" type="paratype">
|
||
<emphasis id="B96CEABEC719FFD6A6CB273D6E2DF9AE" bold="true" box="[199,324,1564,1590]" pageId="9" pageNumber="10">Paratypes</emphasis>
|
||
</typeStatus>
|
||
. 24
|
||
<typeStatus id="54A3880EC719FFD6A771273D6E84F9AE" box="[381,493,1564,1590]" pageId="9" pageNumber="10" type="paratype">paratypes</typeStatus>
|
||
(4 preserved in ethanol specimens and 20 shells, collected between
|
||
<date id="FFA6106CC719FFD6A321273D6FBFF9C6" pageId="9" pageNumber="10" value="1982" valueMax="2006">1982 and 2006</date>
|
||
),
|
||
<collectionCode id="ED09AE69C719FFD6A6E627656E53F9C6" box="[234,314,1604,1630]" pageId="9" pageNumber="10">TFMC</collectionCode>
|
||
(1
|
||
<typeStatus id="54A3880EC719FFD6A76D27656EA3F9C6" box="[353,458,1604,1630]" pageId="9" pageNumber="10" type="paratype">paratype</typeStatus>
|
||
: MT 0395) and AIT.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3026527C719FFD6A6CB274D6DD0F936" pageId="9" pageNumber="10" type="etymology">
|
||
<paragraph id="8BA736ACC719FFD6A6CB274D6DD0F936" blockId="9.[151,1437,1564,2030]" pageId="9" pageNumber="10">
|
||
<emphasis id="B96CEABEC719FFD6A6CB274D6E25F91E" bold="true" box="[199,332,1644,1670]" pageId="9" pageNumber="10">Etymology</emphasis>
|
||
. The name
|
||
<emphasis id="B96CEABEC719FFD6A7D4274D6D22F91D" box="[472,587,1644,1669]" italics="true" pageId="9" pageNumber="10">
|
||
“
|
||
<taxonomicName id="4C184D2FC719FFD6A7EF274D6D2BF91D" box="[483,578,1644,1669]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="fulgida">fulgida</taxonomicName>
|
||
”
|
||
</emphasis>
|
||
derives from the feminine of the Spanish adjective “fúlgido” and refers to the brilliant, radiant appearance of the shell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3026527C719FFD6A6CB279D6FAEF8BE" pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph id="8BA736ACC719FFD6A6CB279D6FAEF8BE" blockId="9.[151,1437,1564,2030]" pageId="9" pageNumber="10">
|
||
<emphasis id="B96CEABEC719FFD6A6CB279D6E91F94E" bold="true" box="[199,504,1724,1750]" pageId="9" pageNumber="10">Distribution and habitat</emphasis>
|
||
(
|
||
<figureCitation id="13232A29C719FFD6A40B279D6D25F94E" box="[519,588,1724,1750]" captionStart="FIGURE 1" captionStartId="6.[151,255,1385,1409]" captionTargetBox="[161,1421,476,1348]" captionTargetId="figure@6.[151,1436,469,1358]" captionTargetPageId="6" captionText="FIGURE 1. Geographical distribution of Hemicycla laurijona sp. nov. and Hemicycla fulgida sp. nov. The blue arrow indicates approximately the area of the Hemicycla invernicata type locality. Symbols represent 1 x 1 km UTM squares." httpUri="https://zenodo.org/record/176898/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
|
||
). The species is endemic to Tenerife. It occurs at an altitude of
|
||
<quantity id="4CE09B49C719FFD6A339279D6FD8F966" metricMagnitude="2" metricUnit="m" metricValue="8.15" metricValueMax="9.8" metricValueMin="6.5" pageId="9" pageNumber="10" unit="m" value="815.0" valueMax="980.0" valueMin="650.0">650–980 m</quantity>
|
||
, living between the very abundant dead leaves of the laurel forest from the Anaga massif (northeast Tenerife).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3026527C719FFD4A6CB26156B96F87C" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8BA736ACC719FFD6A6CB26156B92F876" blockId="9.[151,1437,1564,2030]" pageId="9" pageNumber="10">
|
||
<emphasis id="B96CEABEC719FFD6A6CB26156E31F8D6" bold="true" box="[199,344,1844,1870]" pageId="9" pageNumber="10">Description</emphasis>
|
||
. Body brown coloured (
|
||
<figureCitation id="13232A29C719FFD6A47826156DD0F8D6" box="[628,697,1844,1870]" captionStart="FIGURE 3. A" captionStartId="7.[151,255,1863,1887]" captionTargetBox="[151,1435,1331,1839]" captionTargetId="figure@7.[151,1436,1331,1839]" captionTargetPageId="7" captionText="FIGURE 3. A. Hemicycla laurijona sp. nov., paratype from Puntas Coloradas, La Gomera. B. Hemicycla fulgida sp. nov., paratype from El Bailadero, Tenerife." httpUri="https://zenodo.org/record/176900/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
|
||
B), sole pale-brown. The shell (Table 2;
|
||
<figureCitation id="13232A29C719FFD6A28226156BB7F8D6" box="[1166,1246,1844,1870]" captionStart="FIGURE 3. A" captionStartId="7.[151,255,1863,1887]" captionTargetBox="[151,1435,1331,1839]" captionTargetId="figure@7.[151,1436,1331,1839]" captionTargetPageId="7" captionText="FIGURE 3. A. Hemicycla laurijona sp. nov., paratype from Puntas Coloradas, La Gomera. B. Hemicycla fulgida sp. nov., paratype from El Bailadero, Tenerife." httpUri="https://zenodo.org/record/176900/files/figure.png" pageId="9" pageNumber="10">Figs. 3</figureCitation>
|
||
B, 8A) is imperforate, depressed globular but slightly conical in the first whorls, with 3¾–4¼ convex whorls and well marked sutures. The aperture is oblique and rounded, without angulations. The peristome is reflected in a white lip (corneous-yellowish above) which has a width of about
|
||
<quantity id="4CE09B49C719FFD6A529268D6C00F85E" box="[805,873,1964,1990]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="10" unit="mm" value="1.0">1 mm</quantity>
|
||
and covers the umbilicus. The shell has a small, weak and wide radial costulation combined in the body whorl with a malleate surface sculpture.
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71AFFD5A6CB21B96D59FECA" blockId="10.[151,1437,152,738]" pageId="10" pageNumber="11">
|
||
The shell surface (
|
||
<figureCitation id="13232A29C71AFFD5A79121B96E8BFF2A" box="[413,482,152,178]" captionStart="FIGURE 9" captionStartId="13.[151,255,1574,1598]" captionTargetBox="[151,1434,194,1550]" captionTargetId="figure@13.[151,1435,194,1550]" captionTargetPageId="13" captionText="FIGURE 9. Shell body whorl details. A, B. Hemicycla fulgida sp. nov., holotype. C, D. Hemicycla invernicata. E, F. Hemicycla consobrina. A, C, E, dorsal view. B, D, F, frontal view." httpUri="https://zenodo.org/record/176905/files/figure.png" pageId="10" pageNumber="11">Fig. 9</figureCitation>
|
||
A,B) is glossy, brilliant and pale-brown to yellowish in colour. In some specimens there are four spiral, darker bands, two dorsal wide and frequently interrupted and two ventral narrower and more uniform. In many specimens the bands are wide, but interrupted forming irregular patches. These patches sometimes fuse with those of the neighbouring bands (dorsal or ventral), giving the appearance of irregular patches arranged radially.
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71AFFD5A6CB20416B69FD7A" blockId="10.[151,1437,152,738]" pageId="10" pageNumber="11">
|
||
Genital system (
|
||
<figureCitation id="13232A29C71AFFD5A78920416E8DFEE2" box="[389,484,352,378]" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="10" pageNumber="11">Figs. 11</figureCitation>
|
||
A, 12;
|
||
<specimenCount id="9D1EFD25C71AFFD5A43820416DADFEE2" box="[564,708,352,378]" pageId="10" pageNumber="11" type="generic">4 specimens</specimenCount>
|
||
dissected): Atrium short. The bursa copulatrix complex in the female genital system shows a moderately short diverticulum, shorter than the bursa copulatrix duct; the latter is also shorter than the slender, not swollen distally common stalk. The bursa copulatrix is rather small, globular. The female duct seemingly concludes in the common stalk duct (
|
||
<figureCitation id="13232A29C71AFFD5A5CA20F96B4DFE6A" box="[966,1060,472,498]" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="10" pageNumber="11">Figs. 11</figureCitation>
|
||
A, 12A) but in fact this duct and the vagina are close and they open up in the atrium, together with the dart sac, each independent of the others (Fig. 12B). The dart sac is accompanied by a pair of branched tubular mucus glands. The dart (
|
||
<figureCitation id="13232A29C71AFFD5A30023096A0BFDDA" box="[1292,1378,552,578]" captionStart="FIGURE 13" captionStartId="17.[151,255,1424,1448]" captionTargetBox="[152,1434,194,1399]" captionTargetId="figure@17.[152,1436,194,1400]" captionTargetPageId="17" captionText="FIGURE 13. Hemicycla fulgida sp. nov., paratype from El Bailadero (Tenerife). A. Spermatophore in the stalk of the bursa copulatrix complex. B. An undigested part of the spermatophore. C. Dart; bd, bursa duct; c, crown; d, diverticulum; st, stalk." httpUri="https://zenodo.org/record/176908/files/figure.png" pageId="10" pageNumber="11">Fig. 13</figureCitation>
|
||
C) is long (about 4.3 mm), slightly curved (almost straight), spear-head shaped, and has four lateral wings. Its transverse section resembles a normal cross (relationship width/depth = about 1.3) and is similar overall to that of some species of
|
||
<taxonomicName id="4C184D2FC71AFFD5A75023816EF2FD21" box="[348,411,672,697]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B96CEABEC71AFFD5A75023816EF2FD21" box="[348,411,672,697]" italics="true" pageId="10" pageNumber="11">Helix</emphasis>
|
||
</taxonomicName>
|
||
, such as
|
||
<taxonomicName id="4C184D2FC71AFFD5A40723816C38FD22" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[523,849,672,698]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="pomatia">
|
||
<emphasis id="B96CEABEC71AFFD5A40723816DF9FD21" box="[523,656,672,697]" italics="true" pageId="10" pageNumber="11">H. pomatia</emphasis>
|
||
Linnaeus, 1758
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C184D2FC71AFFD5A55223816BCDFD22" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[862,1188,672,698]" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="lucorum">
|
||
<emphasis id="B96CEABEC71AFFD5A55223816C8FFD21" box="[862,998,672,697]" italics="true" pageId="10" pageNumber="11">H. lucorum</emphasis>
|
||
Linnaeus, 1758
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71AFFD5A2EC23816FBBFD7A" authority="Born, 1778" authorityName="Born" authorityYear="1778" class="Gastropoda" family="Helicidae" genus="Helix" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="aperta">
|
||
<emphasis id="B96CEABEC71AFFD5A2EC23816A38FD21" box="[1248,1361,672,697]" italics="true" pageId="10" pageNumber="11">H. aperta</emphasis>
|
||
Born, 1778
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF894B5DC71AFFD5A6EC23E96EA7FD7A" author="Meisenheimer" box="[224,462,712,738]" pageId="10" pageNumber="22" refString="Meisenheimer, J. (1912) Die Weinbergschnecke Helix pomatia L. In: Ziegler, H. E. & Woltereck, R. (Eds.), Monographien einheimischer Tiere, 4. Verlag von Dr. Werner Klinkhardt, Leipzig, pp. 1 - 140." type="journal article" year="1912">Meisenheimer 1912</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF894B5DC71AFFD5A7D623E96DA2FD7A" author="Giusti" box="[474,715,712,738]" pageId="10" pageNumber="21" refString="Giusti, F. & Lepri, A. (1981) Aspetti morfologici ed etologici dell'accoppiamento in alcune specie della famiglia Helicidae (Gastropoda: Pulmonata). In: Giusti F. (Ed.), Atti del IV ° Congresso della Societa Malacologica Italiana (Siena 6 - 9 Ottobre, 1978), pp. 11 - 71." type="journal article" year="1981">Giusti & Lepri 1981</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF894B5DC71AFFD5A4D423E96C9AFD7A" author="Giusti" box="[728,1011,712,738]" pageId="10" pageNumber="21" refString="Giusti, F. & Andreini, S. (1988) Morphological and ethological aspects of mating in two species of the family Helicidae (Gastropoda: Pulmonata): Theba pisana (Muller) and Helix aperta Born. Monitore Zoologico Italiano (Nuova Serie), 22, 331 - 363." type="journal article" year="1988">Giusti & Andreini 1988</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71AFFD5A69B26716E6BF84A" blockId="10.[151,1436,1872,2002]" pageId="10" pageNumber="11">
|
||
<emphasis id="B96CEABEC71AFFD5A69B26716E70F8F0" bold="true" box="[151,281,1872,1896]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
|
||
Genital system and details. A–C.
|
||
<taxonomicName id="4C184D2FC71AFFD5A48D26706C35F8F0" box="[641,860,1873,1896]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC71AFFD5A48D26706C35F8F0" box="[641,860,1873,1896]" italics="true" pageId="10" pageNumber="11">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C71AFFD5A56E26716CDEF8F0" box="[866,951,1872,1896]" pageId="10" pageNumber="11" rank="species">
|
||
<emphasis id="B96CEABEC71AFFD5A56E26716CDEF8F0" bold="true" box="[866,951,1872,1896]" pageId="10" pageNumber="11">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54A3880EC71AFFD5A5C826716B43F8FF" box="[964,1066,1872,1895]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
|
||
from the
|
||
<typeStatus id="54A3880EC71AFFD5A29C26716BD7F8FF" box="[1168,1214,1872,1895]" pageId="10" pageNumber="11">type</typeStatus>
|
||
locality. D–F.
|
||
<emphasis id="B96CEABEC71AFFD5A35526706E0BF812" italics="true" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C184D2FC71AFFD5A35526706E36F812" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="perraudierei">Hemicycla perraudierei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
from Las Casillas, El Hierro. A, B, D, E. The whole genital system. C, F. Anatomy of the distal genital system; co, contact organ; e, epiphallus; go, genital orifice; pp
|
||
<emphasis id="B96CEABEC71AFFD5A55B26856C37F829" bold="true" box="[855,862,1956,1969]" pageId="10" pageNumber="11">1</emphasis>
|
||
, proximal penial papilla; pp
|
||
<emphasis id="B96CEABEC71AFFD5A28226856BFCF829" bold="true" box="[1166,1173,1956,1969]" pageId="10" pageNumber="11">2</emphasis>
|
||
, distal penial papilla; ri, ring zone.
|
||
</paragraph>
|
||
<caption id="DF676624C71BFFD4A69B25D36D96FA0C" httpUri="https://zenodo.org/record/176903/files/figure.png" pageId="11" pageNumber="12" targetBox="[232,1344,185,1251]" targetPageId="11">
|
||
<paragraph id="8BA736ACC71BFFD4A69B25D36D96FA0C" blockId="11.[151,1436,1266,1428]" pageId="11" pageNumber="12">
|
||
<emphasis id="B96CEABEC71BFFD4A69B25D36E71FA92" bold="true" box="[151,280,1266,1290]" pageId="11" pageNumber="12">FIGURE 7.</emphasis>
|
||
Scatter plots of some shell measurements of the species studied. 1,
|
||
<taxonomicName id="4C184D2FC71BFFD4A5EF25D36BBAFA91" box="[995,1235,1266,1289]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71BFFD4A5EF25D36BBAFA91" box="[995,1235,1266,1289]" italics="true" pageId="11" pageNumber="12">Hemicycla consobrina</emphasis>
|
||
</taxonomicName>
|
||
; 2,
|
||
<taxonomicName id="4C184D2FC71BFFD4A2F725D36F99FAB4" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71BFFD4A2F725D36F99FAB4" italics="true" pageId="11" pageNumber="12">Hemicycla invernicata</emphasis>
|
||
</taxonomicName>
|
||
; 3,
|
||
<taxonomicName id="4C184D2FC71BFFD4A71224346E8FFAB4" box="[286,486,1301,1324]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov.">
|
||
<emphasis id="B96CEABEC71BFFD4A71224346E8FFAB4" box="[286,486,1301,1324]" italics="true" pageId="11" pageNumber="12">Hemicycla fulgida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C71BFFD4A7E324356D2EFAB4" box="[495,583,1300,1324]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B96CEABEC71BFFD4A7E324356D2EFAB4" bold="true" box="[495,583,1300,1324]" pageId="11" pageNumber="12">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
; 4,
|
||
<taxonomicName id="4C184D2FC71BFFD4A47A24346C1EFAB4" box="[630,887,1301,1324]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC71BFFD4A47A24346C1EFAB4" box="[630,887,1301,1324]" italics="true" pageId="11" pageNumber="12">Hemicycla perraudierei</emphasis>
|
||
</taxonomicName>
|
||
; 5,
|
||
<taxonomicName id="4C184D2FC71BFFD4A5AA24346BECFAB4" box="[934,1157,1301,1324]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC71BFFD4A5AA24346BECFAB4" box="[934,1157,1301,1324]" italics="true" pageId="11" pageNumber="12">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C71BFFD4A28324356B8EFAB4" box="[1167,1255,1300,1324]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B96CEABEC71BFFD4A28324356B8EFAB4" bold="true" box="[1167,1255,1300,1324]" pageId="11" pageNumber="12">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
BP, body whorl frontal perimeter; BS, body whorl frontal surface (plane view); D1, maximum shell diameter; D2, shell diameter perpendicular to D1; FP, shell frontal perimeter; FS, shell frontal surface (plane view); SH, shell height; SP, shell perimeter (dorsal plane view); SS, shell surface (dorsal plane view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736ACC71BFFD4A6CB24E36B63F9E4" blockId="11.[151,1437,1474,2020]" pageId="11" pageNumber="12">
|
||
The male portion of the genital system shows a penial complex with a short flagellum, similar in length to the sum of the penis and epiphallus. The penis sheath is very thin, translucent and poorly visible. The retractor muscle has an epiphallar insertion and the penis has a system of twin papillae with a penial chamber between them (Fig. 12B). The “ring zone” is visible from the exterior (
|
||
<figureCitation id="13232A29C71BFFD4A59D271B6C9BF9CC" box="[913,1010,1594,1620]" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="11" pageNumber="12">Figs. 11</figureCitation>
|
||
A, 12A). A well-developed contact organ is located between the distal penial papilla and the atrium (Fig. 12B).
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71BFFD4A6CB27AB6B18F96C" blockId="11.[151,1437,1474,2020]" pageId="11" pageNumber="12">
|
||
A specimen had a spermatophore partially digested in the common stalk of the bursa copulatrix system (
|
||
<figureCitation id="13232A29C71BFFD4A69327936F87F954" box="[159,238,1714,1740]" captionStart="FIGURE 13" captionStartId="17.[151,255,1424,1448]" captionTargetBox="[152,1434,194,1399]" captionTargetId="figure@17.[152,1436,194,1400]" captionTargetPageId="17" captionText="FIGURE 13. Hemicycla fulgida sp. nov., paratype from El Bailadero (Tenerife). A. Spermatophore in the stalk of the bursa copulatrix complex. B. An undigested part of the spermatophore. C. Dart; bd, bursa duct; c, crown; d, diverticulum; st, stalk." httpUri="https://zenodo.org/record/176908/files/figure.png" pageId="11" pageNumber="12">Fig 13</figureCitation>
|
||
A,B). This undigested part is a long, corneous sheet spirally rolled around the axis, forming a lengthwise open, filiform tube (diameter less than ½ mm) folded inside the receiving duct.
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71BFFD4A6CB26236B96F87C" blockId="11.[151,1437,1474,2020]" pageId="11" pageNumber="12">Another specimen everted the distal genital system during the fixation process (Fig. 14). Namely, the atrium and the entire penis were evaginated, whereas the distal end of the female part of the genital system was protruded. The boundary of the evagination has been studied by dissecting the penial evagination: the distal epiphallar folds are located immediately behind the hole of the proximal penial papilla, which is at the end of the evaginated penis (Fig. 14C,D). The contact organ is at the base of the evaginated penis, immediately adjacent to a protuberant ring which corresponds to the “ring zone” of the non-evaginated penis.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF676624C71CFFD3A69B25F26E77FAA8" httpUri="https://zenodo.org/record/176904/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1434,194,1211]" targetPageId="12">
|
||
<paragraph id="8BA736ACC71CFFD3A69B25F26E77FAA8" blockId="12.[151,1436,1235,1328]" pageId="12" pageNumber="13">
|
||
<emphasis id="B96CEABEC71CFFD3A69B25F26E72FB73" bold="true" box="[151,283,1235,1259]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
|
||
Shells from Tenerife. A.
|
||
<taxonomicName id="4C184D2FC71CFFD3A43525F56C68FB73" box="[569,769,1236,1259]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov.">
|
||
<emphasis id="B96CEABEC71CFFD3A43525F56C68FB73" box="[569,769,1236,1259]" italics="true" pageId="12" pageNumber="13">Hemicycla fulgida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C71CFFD3A50625F26C0BFB73" box="[778,866,1235,1259]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="B96CEABEC71CFFD3A50625F26C0BFB73" bold="true" box="[778,866,1235,1259]" pageId="12" pageNumber="13">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype. B.
|
||
<taxonomicName id="4C184D2FC71CFFD3A20E25F56B9EFB73" box="[1026,1271,1236,1259]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71CFFD3A20E25F56B9EFB73" box="[1026,1271,1236,1259]" italics="true" pageId="12" pageNumber="13">Hemicycla invernicata</emphasis>
|
||
</taxonomicName>
|
||
from Montaña Cabeza de Toro pine forest, near Las Lagunetas (La Esperanza forests). C.
|
||
<taxonomicName id="4C184D2FC71CFFD3A5C125D66BA9FA96" box="[973,1216,1271,1294]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71CFFD3A5C125D66BA9FA96" box="[973,1216,1271,1294]" italics="true" pageId="12" pageNumber="13">Hemicycla consobrina</emphasis>
|
||
</taxonomicName>
|
||
from Las Mercedes laurel forest.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3026527C71CFFD1A6CA247F6CF5FA53" lastPageId="14" lastPageNumber="15" pageId="12" pageNumber="13" type="discussion">
|
||
<paragraph id="8BA736ACC71CFFD3A6CA247F6DB3F979" blockId="12.[151,1436,1374,2001]" pageId="12" pageNumber="13">
|
||
<emphasis id="B96CEABEC71CFFD3A6CA247F6E29FAE0" bold="true" box="[198,320,1374,1400]" pageId="12" pageNumber="13">Remarks.</emphasis>
|
||
Regarding the shell, the nearest species to
|
||
<taxonomicName id="4C184D2FC71CFFD3A53B247E6B67FAE0" box="[823,1038,1375,1400]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A53B247E6B67FAE0" box="[823,1038,1375,1400]" italics="true" pageId="12" pageNumber="13">Hemicycla fulgida</emphasis>
|
||
</taxonomicName>
|
||
are
|
||
<emphasis id="B96CEABEC71CFFD3A24C247E6A6BFAE0" box="[1088,1282,1375,1400]" italics="true" pageId="12" pageNumber="13">H. consobrina—</emphasis>
|
||
also from the Anaga laurel forest and similar areas (
|
||
<figureCitation id="13232A29C71CFFD3A45724A66DC4FA39" box="[603,685,1415,1441]" captionStart="FIGURE 8" captionStartId="12.[151,255,1235,1259]" captionTargetBox="[151,1434,194,1211]" captionTargetId="figure@12.[151,1436,194,1212]" captionTargetPageId="12" captionText="FIGURE 8. Shells from Tenerife. A. Hemicycla fulgida sp. nov., holotype. B. Hemicycla invernicata from Montaña Cabeza de Toro pine forest, near Las Lagunetas (La Esperanza forests). C. Hemicycla consobrina from Las Mercedes laurel forest." httpUri="https://zenodo.org/record/176904/files/figure.png" pageId="12" pageNumber="13">Figs. 8</figureCitation>
|
||
C, 9E,F, 10)—and
|
||
<taxonomicName id="4C184D2FC71CFFD3A58124A66B5EFA38" box="[909,1079,1415,1440]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71CFFD3A58124A66B5EFA38" box="[909,1079,1415,1440]" italics="true" pageId="12" pageNumber="13">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13232A29C71CFFD3A24524A66BF2FA39" box="[1097,1179,1415,1441]" captionStart="FIGURE 8" captionStartId="12.[151,255,1235,1259]" captionTargetBox="[151,1434,194,1211]" captionTargetId="figure@12.[151,1436,194,1212]" captionTargetPageId="12" captionText="FIGURE 8. Shells from Tenerife. A. Hemicycla fulgida sp. nov., holotype. B. Hemicycla invernicata from Montaña Cabeza de Toro pine forest, near Las Lagunetas (La Esperanza forests). C. Hemicycla consobrina from Las Mercedes laurel forest." httpUri="https://zenodo.org/record/176904/files/figure.png" pageId="12" pageNumber="13">Figs. 8</figureCitation>
|
||
B, 9C,D), whose
|
||
<typeStatus id="54A3880EC71CFFD3A36524A66AF2FA39" box="[1385,1435,1415,1441]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality is: "Ténérife, environs de Sta-Cruz. Dans les creux d'arbres dans les bosquets à la Esperanza, près des Lagunes" [correct name: Las Lagunetas, about
|
||
<quantity id="4CE09B49C71CFFD3A4CA24F66C49FA69" box="[710,800,1495,1521]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="13" unit="m" value="1400.0">1400 m</quantity>
|
||
altitude, including a pine forest; it is the area marked with the blue arrow in
|
||
<figureCitation id="13232A29C71CFFD3A7B424DE6D49F981" box="[440,544,1535,1561]" captionStart="FIGURE 1" captionStartId="6.[151,255,1385,1409]" captionTargetBox="[161,1421,476,1348]" captionTargetId="figure@6.[151,1436,469,1358]" captionTargetPageId="6" captionText="FIGURE 1. Geographical distribution of Hemicycla laurijona sp. nov. and Hemicycla fulgida sp. nov. The blue arrow indicates approximately the area of the Hemicycla invernicata type locality. Symbols represent 1 x 1 km UTM squares." httpUri="https://zenodo.org/record/176898/files/figure.png" pageId="12" pageNumber="13">Figure 1</figureCitation>
|
||
]. The last two species are similar in appearance and
|
||
<bibRefCitation id="EF894B5DC71CFFD3A2B524DE6AF5F981" box="[1209,1436,1535,1561]" pageId="12" pageNumber="20" refString="Bank, R. A., Groh, K. & Ripken, T. E. J. (2002) Catalogue and bibliography of the non-marine Mollusca of Macaronesia (with colour-plates 14 - 26). In: Falkner, M., Groh, K. & Speight, M. C. D. (Eds.), Collectanea Malacologica- Festschrift fur Gerhard Falkner. Conchbooks, Hackenheim, 89 - 235." type="book chapter">
|
||
Bank
|
||
<emphasis id="B96CEABEC71CFFD3A30F24DE6A52F980" box="[1283,1339,1535,1560]" italics="true" pageId="12" pageNumber="13">et al</emphasis>
|
||
. (2002)
|
||
</bibRefCitation>
|
||
grouped
|
||
<taxonomicName id="4C184D2FC71CFFD3A6F227066ECEF9D8" box="[254,423,1575,1600]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71CFFD3A6F227066ECEF9D8" box="[254,423,1575,1600]" italics="true" pageId="12" pageNumber="13">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<emphasis id="B96CEABEC71CFFD3A48B27066C5EF9D8" box="[647,823,1575,1600]" italics="true" pageId="12" pageNumber="13">
|
||
<taxonomicName id="4C184D2FC71CFFD3A48B27066C5AF9D8" box="[647,819,1575,1600]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="consobrina">H. consobrina</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
However, they differ in shell and genital characters. The aperture of the
|
||
<taxonomicName id="4C184D2FC71CFFD3A78C276E6D41F9F0" box="[384,552,1615,1640]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71CFFD3A78C276E6D41F9F0" box="[384,552,1615,1640]" italics="true" pageId="12" pageNumber="13">H. consobrina</emphasis>
|
||
</taxonomicName>
|
||
shell was described by
|
||
<bibRefCitation id="EF894B5DC71CFFD3A54E276E6B6AF9F1" author="Mousson" box="[834,1027,1615,1641]" pageId="12" pageNumber="22" refString="Mousson, A. (1872) Revision de la faune malacologique des Canaries. Neue Denkschriften der allgemeinen schweizerischen Gesellschaft fur die gesammten Naturwissenschaften, 25 (1), I - V, 1 - 176 pp., 6 pl." type="journal article" year="1872">Mousson (1872)</bibRefCitation>
|
||
as “obtusément triangulaire” (
|
||
<figureCitation id="13232A29C71CFFD3A361276E6FDCF909" captionStart="FIGURE 10" captionStartId="14.[151,255,1238,1262]" captionTargetBox="[151,1435,194,1214]" captionTargetId="figure@14.[151,1436,194,1214]" captionTargetPageId="14" captionText="FIGURE 10. Shell aperture details of the Hemicycla consobrina specimen in different views (A – D), showing the “ small callosity ” (yellow arrows) and the form “ obtusément triangulaire ” described by Mousson. A. Aperture accurately in plane view." httpUri="https://zenodo.org/record/176906/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
A), with a very small callosity (
|
||
<figureCitation id="13232A29C71CFFD3A42227566DEBF909" box="[558,642,1655,1681]" captionStart="FIGURE 10" captionStartId="14.[151,255,1238,1262]" captionTargetBox="[151,1435,194,1214]" captionTargetId="figure@14.[151,1436,194,1214]" captionTargetPageId="14" captionText="FIGURE 10. Shell aperture details of the Hemicycla consobrina specimen in different views (A – D), showing the “ small callosity ” (yellow arrows) and the form “ obtusément triangulaire ” described by Mousson. A. Aperture accurately in plane view." httpUri="https://zenodo.org/record/176906/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
A–D, yellow arrows) in the same position as the smaller tooth-like callosity of the two present in the shell aperture of
|
||
<taxonomicName id="4C184D2FC71CFFD3A4E927BE6B22F921" authority="Lamarck, 1822" authorityName="Lamarck" authorityYear="1822" box="[741,1099,1695,1721]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="bidentalis">
|
||
<emphasis id="B96CEABEC71CFFD3A4E927BE6C12F920" box="[741,891,1695,1720]" italics="true" pageId="12" pageNumber="13">H. bidentalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF894B5DC71CFFD3A58727BE6B2BF921" author="Lamarck" box="[907,1090,1695,1721]" pageId="12" pageNumber="22" refString="Lamarck, J. B. de (1822) Histoire naturelle des animaux sans vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres et la citation des principales especes qui s'y rapportent; precedee d'une Introduction offrant la determination des caracteres essentiels de l'Animal, sa distinction du vegetal et des autres corps naturels, enfin, l'Exposition des Principes fondamentaux de la Zoologie. J. B. Bailliere Libr., Paris, 6 (2), 232 pp." type="book" year="1822">Lamarck, 1822</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, a well-known species living in the same area as
|
||
<taxonomicName id="4C184D2FC71CFFD3A77727E66E9BF978" box="[379,498,1735,1760]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A77727E66E9BF978" box="[379,498,1735,1760]" italics="true" pageId="12" pageNumber="13">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71CFFD3A42727E66DBAF978" box="[555,723,1735,1760]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71CFFD3A42727E66DBAF978" box="[555,723,1735,1760]" italics="true" pageId="12" pageNumber="13">H. consobrina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71CFFD3A6CB27CE6A53F819" blockId="12.[151,1436,1374,2001]" pageId="12" pageNumber="13">
|
||
The
|
||
<taxonomicName id="4C184D2FC71CFFD3A6F127CE6E1CF890" box="[253,373,1775,1800]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A6F127CE6E1CF890" box="[253,373,1775,1800]" italics="true" pageId="12" pageNumber="13">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
shell dimensions are smaller than those of the other two species mentioned above, which also are more depressed (Figs. 7,8). The shell ornamentation of
|
||
<taxonomicName id="4C184D2FC71CFFD3A58F26366C93F8A8" box="[899,1018,1815,1840]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A58F26366C93F8A8" box="[899,1018,1815,1840]" italics="true" pageId="12" pageNumber="13">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71CFFD3A23F26366BB5F8A8" box="[1075,1244,1815,1840]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71CFFD3A23F26366BB5F8A8" box="[1075,1244,1815,1840]" italics="true" pageId="12" pageNumber="13">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
are very similar (
|
||
<figureCitation id="13232A29C71CFFD3A693261E6F9CF8C1" box="[159,245,1855,1881]" captionStart="FIGURE 10" captionStartId="14.[151,255,1238,1262]" captionTargetBox="[151,1435,194,1214]" captionTargetId="figure@14.[151,1436,194,1214]" captionTargetPageId="14" captionText="FIGURE 10. Shell aperture details of the Hemicycla consobrina specimen in different views (A – D), showing the “ small callosity ” (yellow arrows) and the form “ obtusément triangulaire ” described by Mousson. A. Aperture accurately in plane view." httpUri="https://zenodo.org/record/176906/files/figure.png" pageId="12" pageNumber="13">Fig. 10</figureCitation>
|
||
A–D), whereas the
|
||
<taxonomicName id="4C184D2FC71CFFD3A7D2261E6DE1F8C0" box="[478,648,1855,1880]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71CFFD3A7D2261E6DE1F8C0" box="[478,648,1855,1880]" italics="true" pageId="12" pageNumber="13">H. consobrina</emphasis>
|
||
</taxonomicName>
|
||
shell has a more developed malleation (
|
||
<figureCitation id="13232A29C71CFFD3A266261E6BD8F8C1" box="[1130,1201,1855,1881]" captionStart="FIGURE 9" captionStartId="13.[151,255,1574,1598]" captionTargetBox="[151,1434,194,1550]" captionTargetId="figure@13.[151,1435,194,1550]" captionTargetPageId="13" captionText="FIGURE 9. Shell body whorl details. A, B. Hemicycla fulgida sp. nov., holotype. C, D. Hemicycla invernicata. E, F. Hemicycla consobrina. A, C, E, dorsal view. B, D, F, frontal view." httpUri="https://zenodo.org/record/176905/files/figure.png" pageId="12" pageNumber="13">Fig. 9</figureCitation>
|
||
E–F). Moreover,
|
||
<taxonomicName id="4C184D2FC71CFFD3A373261E6F83F818" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A373261E6F83F818" italics="true" pageId="12" pageNumber="13">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71CFFD3A72F26466EA5F818" box="[291,460,1895,1920]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71CFFD3A72F26466EA5F818" box="[291,460,1895,1920]" italics="true" pageId="12" pageNumber="13">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
differ in their habitat: humid laurel forest and dry pine forest, respectively.
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC71CFFD3A6CB26AE6B48F849" blockId="12.[151,1436,1374,2001]" pageId="12" pageNumber="13">
|
||
The maximum diameter of the
|
||
<taxonomicName id="4C184D2FC71CFFD3A44B26AE6DABF830" box="[583,706,1935,1960]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A44B26AE6DABF830" box="[583,706,1935,1960]" italics="true" pageId="12" pageNumber="13">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
shell is similar to that of
|
||
<taxonomicName id="4C184D2FC71CFFD3A20726AE6BC9F830" box="[1035,1184,1935,1960]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC71CFFD3A20726AE6BC9F830" box="[1035,1184,1935,1960]" italics="true" pageId="12" pageNumber="13">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71CFFD3A2ED26AE6AF5F830" box="[1249,1436,1935,1960]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC71CFFD3A2ED26AE6AF5F830" box="[1249,1436,1935,1960]" italics="true" pageId="12" pageNumber="13">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
although the
|
||
<taxonomicName id="4C184D2FC71CFFD3A73C26966ECEF848" box="[304,423,1975,2000]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="13" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71CFFD3A73C26966ECEF848" box="[304,423,1975,2000]" italics="true" pageId="12" pageNumber="13">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
shell is higher than those of the other species (
|
||
<figureCitation id="13232A29C71CFFD3A5C726966B7AF849" box="[971,1043,1975,2001]" captionStart="FIGURE 7" captionStartId="11.[151,255,1266,1290]" captionTargetBox="[232,1344,185,1251]" captionTargetId="figure@11.[217,1369,176,1264]" captionTargetPageId="11" captionText="FIGURE 7. Scatter plots of some shell measurements of the species studied. 1, Hemicycla consobrina; 2, Hemicycla invernicata; 3, Hemicycla fulgida sp. nov.; 4, Hemicycla perraudierei; 5, Hemicycla laurijona sp. nov. BP, body whorl frontal perimeter; BS, body whorl frontal surface (plane view); D 1, maximum shell diameter; D 2, shell diameter perpendicular to D 1; FP, shell frontal perimeter; FS, shell frontal surface (plane view); SH, shell height; SP, shell perimeter (dorsal plane view); SS, shell surface (dorsal plane view)." httpUri="https://zenodo.org/record/176903/files/figure.png" pageId="12" pageNumber="13">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF676624C71DFFD2A69B27076C33F9F8" httpUri="https://zenodo.org/record/176905/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1434,194,1550]" targetPageId="13">
|
||
<paragraph id="8BA736ACC71DFFD2A69B27076C33F9F8" blockId="13.[151,1437,1574,1632]" pageId="13" pageNumber="14">
|
||
<emphasis id="B96CEABEC71DFFD2A69B27076E75F9A6" bold="true" box="[151,284,1574,1598]" pageId="13" pageNumber="14">FIGURE 9.</emphasis>
|
||
Shell body whorl details. A, B.
|
||
<taxonomicName id="4C184D2FC71DFFD2A48C27066C2FF9A6" box="[640,838,1575,1598]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov.">
|
||
<emphasis id="B96CEABEC71DFFD2A48C27066C2FF9A6" box="[640,838,1575,1598]" italics="true" pageId="13" pageNumber="14">Hemicycla fulgida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C71DFFD2A54327076CCFF9A6" box="[847,934,1574,1598]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="B96CEABEC71DFFD2A54327076CCFF9A6" bold="true" box="[847,934,1574,1598]" pageId="13" pageNumber="14">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype. C, D.
|
||
<taxonomicName id="4C184D2FC71DFFD2A26A27066A30F9A6" box="[1126,1369,1575,1598]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71DFFD2A26A27066A30F9A6" box="[1126,1369,1575,1598]" italics="true" pageId="13" pageNumber="14">Hemicycla invernicata</emphasis>
|
||
</taxonomicName>
|
||
. E, F.
|
||
<taxonomicName id="4C184D2FC71DFFD2A69B27686EE1F9F8" box="[151,392,1609,1632]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71DFFD2A69B27686EE1F9F8" box="[151,392,1609,1632]" italics="true" pageId="13" pageNumber="14">Hemicycla consobrina</emphasis>
|
||
</taxonomicName>
|
||
. A, C, E, dorsal view. B, D, F, frontal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736ACC71DFFD2A6CB27966BEFF871" blockId="13.[151,1437,1719,2025]" pageId="13" pageNumber="14">
|
||
Several other differences were found among
|
||
<emphasis id="B96CEABEC71DFFD2A4D227966B78F948" box="[734,1041,1719,1744]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C184D2FC71DFFD2A4D227966C33F948" box="[734,858,1719,1744]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fulgida">H. fulgida</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C184D2FC71DFFD2A56A27966B78F948" box="[870,1041,1719,1744]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="invernicata">H. invernicata</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C184D2FC71DFFD2A24127966B9EF948" box="[1101,1271,1719,1744]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71DFFD2A24127966B9EF948" box="[1101,1271,1719,1744]" italics="true" pageId="13" pageNumber="14">H. consobrina</emphasis>
|
||
</taxonomicName>
|
||
regarding the genital system (
|
||
<figureCitation id="13232A29C71DFFD2A75927FE6EC4F961" box="[341,429,1759,1785]" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
). The
|
||
<taxonomicName id="4C184D2FC71DFFD2A7F127FE6D1FF960" box="[509,630,1759,1784]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71DFFD2A7F127FE6D1FF960" box="[509,630,1759,1784]" italics="true" pageId="13" pageNumber="14">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
genital system (
|
||
<figureCitation id="13232A29C71DFFD2A53227FE6CFAF961" box="[830,915,1759,1785]" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
A) differs from that of
|
||
<taxonomicName id="4C184D2FC71DFFD2A2A227FE6A33F960" box="[1198,1370,1759,1784]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71DFFD2A2A227FE6A33F960" box="[1198,1370,1759,1784]" italics="true" pageId="13" pageNumber="14">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13232A29C71DFFD2A36027FE6FDAF8B9" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
B) and
|
||
<taxonomicName id="4C184D2FC71DFFD2A70226266ED1F8B8" box="[270,440,1799,1824]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71DFFD2A70226266ED1F8B8" box="[270,440,1799,1824]" italics="true" pageId="13" pageNumber="14">H. consobrina</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13232A29C71DFFD2A7C726266D76F8B9" box="[459,543,1799,1825]" captionStart="FIGURE 11" captionStartId="15.[151,255,1161,1185]" captionTargetBox="[151,1435,344,1141]" captionTargetId="figure@15.[151,1436,344,1143]" captionTargetPageId="15" captionText="FIGURE 11. Genital system. A. Hemicycla fulgida sp. nov., paratype from El Bailadero. B. Hemicycla invernicata from Montaña Cabeza de Toro. C. Hemicycla consobrina from Guamasa; ri, ring zone." httpUri="https://zenodo.org/record/176907/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
|
||
C) in the flagellum length. It is 1/3 longer than that of
|
||
<taxonomicName id="4C184D2FC71DFFD2A2B026266A0EF8B8" box="[1212,1383,1799,1824]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71DFFD2A2B026266A0EF8B8" box="[1212,1383,1799,1824]" italics="true" pageId="13" pageNumber="14">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
and about twice as long as that of
|
||
<emphasis id="B96CEABEC71DFFD2A409260E6DD1F8D0" box="[517,696,1839,1864]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C184D2FC71DFFD2A409260E6DDDF8D0" box="[517,692,1839,1864]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="consobrina">H. consobrina</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The atrium of the latter two species is longer than that of
|
||
<emphasis id="B96CEABEC71DFFD2A373260E6F9BF8E8" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C184D2FC71DFFD2A373260E6F87F8E8" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fulgida">H. fulgida</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The
|
||
<taxonomicName id="4C184D2FC71DFFD2A73A26766ED8F8E8" box="[310,433,1879,1904]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71DFFD2A73A26766ED8F8E8" box="[310,433,1879,1904]" italics="true" pageId="13" pageNumber="14">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
bursa copulatrix stalk is about twice as long, while the diverticulum is about 1/3 shorter than that of
|
||
<taxonomicName id="4C184D2FC71DFFD2A789265E6D59F800" box="[389,560,1919,1944]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="invernicata">
|
||
<emphasis id="B96CEABEC71DFFD2A789265E6D59F800" box="[389,560,1919,1944]" italics="true" pageId="13" pageNumber="14">H. invernicata</emphasis>
|
||
</taxonomicName>
|
||
, in which the diverticulum is slightly longer than the bursa duct. The
|
||
<taxonomicName id="4C184D2FC71DFFD2A373265E6F83F858" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71DFFD2A373265E6F83F858" italics="true" pageId="13" pageNumber="14">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
stalk and the bursa duct are about 1/3 longer than those of
|
||
<emphasis id="B96CEABEC71DFFD2A5B026866B04F858" box="[956,1133,1959,1984]" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C184D2FC71DFFD2A5B026866B03F858" box="[956,1130,1959,1984]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="14" phylum="Mollusca" rank="species" species="consobrina">H. consobrina</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
while the diverticulae of both species are similar in length, and they are shorter than the respective bursa ducts.
|
||
</paragraph>
|
||
<caption id="DF676624C71EFFD1A69B25F76E78FAB7" httpUri="https://zenodo.org/record/176906/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1435,194,1214]" targetPageId="14">
|
||
<paragraph id="8BA736ACC71EFFD1A69B25F76E78FAB7" blockId="14.[151,1436,1238,1327]" pageId="14" pageNumber="15">
|
||
<emphasis id="B96CEABEC71EFFD1A69B25F76E4FFB76" bold="true" box="[151,294,1238,1262]" pageId="14" pageNumber="15">FIGURE 10.</emphasis>
|
||
Shell aperture details of the
|
||
<taxonomicName id="4C184D2FC71EFFD1A45425F66C20FB76" box="[600,841,1239,1262]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="consobrina">
|
||
<emphasis id="B96CEABEC71EFFD1A45425F66C20FB76" box="[600,841,1239,1262]" italics="true" pageId="14" pageNumber="15">Hemicycla consobrina</emphasis>
|
||
</taxonomicName>
|
||
specimen in different views (A–D), showing the “small callosity” (yellow arrows) and the form “obtusément triangulaire” described by Mousson. A. Aperture accurately in plane view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736ACC71EFFD1A69B24406CF5FA53" blockId="14.[151,1436,1377,1483]" pageId="14" pageNumber="15">
|
||
The main difference between the
|
||
<taxonomicName id="4C184D2FC71EFFD1A42124406DCEFAE2" box="[557,679,1377,1402]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71EFFD1A42124406DCEFAE2" box="[557,679,1377,1402]" italics="true" pageId="14" pageNumber="15">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
genital system and that of
|
||
<taxonomicName id="4C184D2FC71EFFD1A5FF24406BEFFAE2" box="[1011,1158,1377,1402]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC71EFFD1A5FF24406BEFFAE2" box="[1011,1158,1377,1402]" italics="true" pageId="14" pageNumber="15">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71EFFD1A2C924406A17FAE2" box="[1221,1406,1377,1402]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC71EFFD1A2C924406A17FAE2" box="[1221,1406,1377,1402]" italics="true" pageId="14" pageNumber="15">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
is related to the flagellum length. The diverticulum of
|
||
<taxonomicName id="4C184D2FC71EFFD1A4F024A86C1AFA3A" box="[764,883,1417,1442]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="fulgida">
|
||
<emphasis id="B96CEABEC71EFFD1A4F024A86C1AFA3A" box="[764,883,1417,1442]" italics="true" pageId="14" pageNumber="15">H. fulgida</emphasis>
|
||
</taxonomicName>
|
||
is also shorter than that of
|
||
<taxonomicName id="4C184D2FC71EFFD1A2B824A86A2FFA3A" box="[1204,1350,1417,1442]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC71EFFD1A2B824A86A2FFA3A" box="[1204,1350,1417,1442]" italics="true" pageId="14" pageNumber="15">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C184D2FC71EFFD1A37324A86E43FA52" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC71EFFD1A37324A86E43FA52" italics="true" pageId="14" pageNumber="15">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
, in comparison with the respective bursa duct length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |