281 lines
33 KiB
XML
281 lines
33 KiB
XML
<document id="28087CDC890E98121A61B01066E9BC06" ID-DOI="10.5281/zenodo.176897" ID-GBIF-Dataset="85773b3a-9274-4a30-ad96-ae4b27cc0ea5" ID-ISSN="1175-5326" ID-Zenodo-Dep="176897" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459842485466" checkinUser="plazi" docAuthor="Alonso, María R. & Ibáñez, Miguel" docDate="2007" docId="03B187BAC715FFD7A69B241E6AF5F965" docLanguage="en" docName="zt01482p023.pdf" docOrigin="Zootaxa 1482" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hemicycla (Hemicycla) laurijona Alonso & Ibáñez, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="9" masterDocId="FF88FFC2C710FFDFA60C21216F69FF98" masterDocTitle="Anatomy and function of the penial twin papillae system of the Helicinae (Gastropoda: Helicoidea: Helicidae) and description of two new, small Hemicycla species from the laurel forest of the Canary Islands" masterLastPageNumber="23" masterPageNumber="1" pageNumber="6" updateTime="1698224473241" updateUser="plazi">
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<mods:mods id="15EB28E4B2FB30304798D333F54D078C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8E85D3B46B679E977260655147067EB0">
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<mods:title id="1374DAFA0CA6EF6B355034E30EC6EC4D">Anatomy and function of the penial twin papillae system of the Helicinae (Gastropoda: Helicoidea: Helicidae) and description of two new, small Hemicycla species from the laurel forest of the Canary Islands</mods:title>
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<mods:name id="95AC78E8CDFFC6CFBA01225A63E6E130" type="personal">
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<mods:role id="F0ED7DF1423DF91EF5B3DF7DA0579F61">
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<mods:roleTerm id="0D9E1029495C163D85CAA25EEE19A0FA">Author</mods:roleTerm>
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<mods:namePart id="AD2D8B382800B67617CB460E7793CD0B">Alonso, María R.</mods:namePart>
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</mods:name>
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<mods:name id="8E552C4A62AE017245E7A7878D4F7B47" type="personal">
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<mods:role id="27D28D870DE09A3A363CA9851AE4FD7D">
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<mods:roleTerm id="B17714DE7F65D55F18572E2E74178428">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="94A26243B0B8E0AAD19E81D0DA83D990">Ibáñez, Miguel</mods:namePart>
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<mods:typeOfResource id="A67F79A32D3A4E00C6B1261A1B4C9D37">text</mods:typeOfResource>
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<mods:title id="FF7DE73A160FEF6590F8F4E922D559D5">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CA783C4AA4115681031873ADD084F793">
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<mods:date id="F1A671E61CD716DCBC1F1DD2A288A5C9">2007</mods:date>
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<mods:detail id="EF7085CA9CBBA5549043B588568F7AF4" type="volume">
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<mods:number id="5FB6D787E13A215642D62C9E12E32061">1482</mods:number>
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<mods:start id="E8958218DCD239AFB672C801F7B944F6">1</mods:start>
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<mods:end id="175CCBE6E6DE927E340EF0520A19E27D">23</mods:end>
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<mods:classification id="4BE3F130864EA22966FE1432F9B1D207">journal article</mods:classification>
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<mods:identifier id="0EA584DD89D685DEDE8552FABB705305" type="DOI">10.5281/zenodo.176897</mods:identifier>
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<mods:identifier id="D2DAB32C77142393A646CBBE243F4EC3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9C9D4E30F22A332068FB7352F57136AA" type="Zenodo-Dep">176897</mods:identifier>
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<treatment id="03B187BAC715FFD7A69B241E6AF5F965" ID-DOI="http://doi.org/10.5281/zenodo.5612784" ID-GBIF-Taxon="119360618" ID-Zenodo-Dep="5612784" LSID="urn:lsid:plazi:treatment:03B187BAC715FFD7A69B241E6AF5F965" httpUri="http://treatment.plazi.org/id/03B187BAC715FFD7A69B241E6AF5F965" lastPageId="8" lastPageNumber="9" pageId="5" pageNumber="6">
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<subSubSection id="C3026527C715FFDAA69B241E6DE6FAC1" box="[151,655,1343,1369]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8BA736ACC715FFDAA69B241E6DE6FAC1" blockId="5.[151,655,1343,1369]" box="[151,655,1343,1369]" pageId="5" pageNumber="6">
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<heading id="D0EF81C0C715FFDAA69B241E6DE6FAC1" bold="true" box="[151,655,1343,1369]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis id="B96CEABEC715FFDAA69B241E6DE6FAC1" bold="true" box="[151,655,1343,1369]" pageId="5" pageNumber="6">
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<taxonomicName id="4C184D2FC715FFDAA69B241E6D42FAC1" ID-CoL="3KMWW" box="[151,555,1343,1369]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov." subGenus="Hemicycla">
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<emphasis id="B96CEABEC715FFDAA69B241E6E71FAC1" bold="true" box="[151,280,1343,1369]" italics="true" pageId="5" pageNumber="6">Hemicycla</emphasis>
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(
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<emphasis id="B96CEABEC715FFDAA724241E6EC0FAC1" bold="true" box="[296,425,1343,1369]" italics="true" pageId="5" pageNumber="6">Hemicycla</emphasis>
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)
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<emphasis id="B96CEABEC715FFDAA7B6241E6D42FAC1" bold="true" box="[442,555,1343,1369]" italics="true" pageId="5" pageNumber="6">laurijona</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57C5C715FFDAA43D241E6DE6FAC1" box="[561,655,1343,1369]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026527C715FFDAA69B24A46E17F9DA" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="8BA736ACC715FFDAA69B24A46BDCFA07" blockId="5.[151,1437,1413,2002]" box="[151,1205,1413,1439]" pageId="5" pageNumber="6">
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<emphasis id="B96CEABEC715FFDAA69B24A46E5CFA07" bold="true" box="[151,309,1413,1439]" pageId="5" pageNumber="6">
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<typeStatus id="54A3880EC715FFDAA69B24A46FBDFA07" box="[151,212,1413,1439]" pageId="5" pageNumber="6">Type</typeStatus>
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locality
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</emphasis>
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. Barranco de Aramaqué, La Gomera (UTM: 28R BS8114,
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<quantity id="4CE09B49C715FFDAA5E024A46B2CFA07" box="[1004,1093,1413,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="6" unit="m" value="1000.0">1000 m</quantity>
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altitude).
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</paragraph>
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<paragraph id="8BA736ACC715FFDAA6CB24916A78FA52" blockId="5.[151,1437,1413,2002]" box="[199,1297,1453,1482]" pageId="5" pageNumber="6">
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<emphasis id="B96CEABEC715FFDAA6CB24916E23FA52" bold="true" box="[199,330,1456,1482]" pageId="5" pageNumber="6">
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<typeStatus id="54A3880EC715FFDAA6CB24916E50FA52" box="[199,313,1456,1482]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
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(
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</emphasis>
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<figureCitation id="13232A29C715FFDAA74624916EE6FA52" box="[330,399,1456,1482]" captionStart="FIGURE 4" captionStartId="8.[151,255,1373,1397]" captionTargetBox="[151,1434,194,1348]" captionTargetId="figure@8.[151,1435,194,1349]" captionTargetPageId="8" captionText="FIGURE 4. Shells. A. Hemicycla laurijona sp. nov., holotype. B. Hemicycla laurijona sp. nov., paratypes from Puntas Coloradas, La Gomera. C. Hemicycla perraudierei, from Las Casillas, El Hierro." httpUri="https://zenodo.org/record/176901/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
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A).
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<collectionCode id="ED09AE69C715FFDAA7B024916D65FA52" box="[444,524,1456,1482]" pageId="5" pageNumber="6">TFMC</collectionCode>
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(MT 0392); leg. M.R. Alonso and M. Ibáñez,
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<date id="FFA6106CC715FFDAA22024916A64FA52" box="[1068,1293,1453,1482]" pageId="5" pageNumber="6" value="1985-02-15">15th February 1985</date>
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.
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</paragraph>
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<paragraph id="8BA736ACC715FFDAA6CB24F96E17F9DA" blockId="5.[151,1437,1413,2002]" pageId="5" pageNumber="6">
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<typeStatus id="54A3880EC715FFDAA6CB24F96E2DFA6A" box="[199,324,1496,1522]" pageId="5" pageNumber="6" type="paratype">
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<emphasis id="B96CEABEC715FFDAA6CB24F96E2DFA6A" bold="true" box="[199,324,1496,1522]" pageId="5" pageNumber="6">Paratypes</emphasis>
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</typeStatus>
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. 45
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<typeStatus id="54A3880EC715FFDAA77124F96E84FA6A" box="[381,493,1496,1522]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
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collected between
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<date id="FFA6106CC715FFDAA4DB24F96CE2FA6A" box="[727,907,1496,1522]" pageId="5" pageNumber="6" value="1985" valueMax="2006">1985 and 2006</date>
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(
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<specimenCount id="9D1EFD25C715FFDAA59224F96B2AFA6A" box="[926,1091,1496,1522]" pageId="5" pageNumber="6" type="generic">14 specimens</specimenCount>
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, bodies preserved in ethanol and 31 shells): CHC (7
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<typeStatus id="54A3880EC715FFDAA7A227216D4AF982" box="[430,547,1536,1562]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
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:
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<specimenCount id="9D1EFD25C715FFDAA42227216DA8F982" box="[558,705,1536,1562]" pageId="5" pageNumber="6" type="generic">6 specimens</specimenCount>
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, bodies preserved in ethanol, and 1 shell);
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<collectionCode id="ED09AE69C715FFDAA2BB27216A6EF982" box="[1207,1287,1536,1562]" pageId="5" pageNumber="6">TFMC</collectionCode>
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(1
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<typeStatus id="54A3880EC715FFDAA32327216AF0F982" box="[1327,1433,1536,1562]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
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: MT 0393) and AIT.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026527C715FFDAA6CB27716CC6F922" pageId="5" pageNumber="6" type="etymology">
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<paragraph id="8BA736ACC715FFDAA6CB27716CC6F922" blockId="5.[151,1437,1413,2002]" pageId="5" pageNumber="6">
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<emphasis id="B96CEABEC715FFDAA6CB27716E25F9F2" bold="true" box="[199,332,1616,1642]" pageId="5" pageNumber="6">Etymology</emphasis>
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. The name
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<emphasis id="B96CEABEC715FFDAA7EE27716D06F9F1" box="[482,623,1616,1641]" italics="true" pageId="5" pageNumber="6">
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“
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<taxonomicName id="4C184D2FC715FFDAA7E027716D0DF9F1" box="[492,612,1616,1641]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="laurijona">laurijona</taxonomicName>
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”
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</emphasis>
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is a combination of the first five letters of the Spanish term “laurisilva” (= laurel forest) and the first four letters of the name “Jonay”, the lover of Gara (The Garajonay National Park, in La Gomera, is named after both mythical lovers.).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026527C715FFDAA6CB27E96C24F8AA" pageId="5" pageNumber="6" type="distribution">
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<paragraph id="8BA736ACC715FFDAA6CB27E96C24F8AA" blockId="5.[151,1437,1413,2002]" pageId="5" pageNumber="6">
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<emphasis id="B96CEABEC715FFDAA6CB27E96E90F97A" bold="true" box="[199,505,1736,1762]" pageId="5" pageNumber="6">Distribution and habitat</emphasis>
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(
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<figureCitation id="13232A29C715FFDAA40527E96D27F97A" box="[521,590,1736,1762]" captionStart="FIGURE 1" captionStartId="6.[151,255,1385,1409]" captionTargetBox="[161,1421,476,1348]" captionTargetId="figure@6.[151,1436,469,1358]" captionTargetPageId="6" captionText="FIGURE 1. Geographical distribution of Hemicycla laurijona sp. nov. and Hemicycla fulgida sp. nov. The blue arrow indicates approximately the area of the Hemicycla invernicata type locality. Symbols represent 1 x 1 km UTM squares." httpUri="https://zenodo.org/record/176898/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
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). The species is endemic to La Gomera. It occurs at an altitude of
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<quantity id="4CE09B49C715FFDAA36D27E96F9BF892" metricMagnitude="2" metricUnit="m" metricValue="9.9" metricValueMax="12.8" metricValueMin="7.0" pageId="5" pageNumber="6" unit="m" value="990.0" valueMax="1280.0" valueMin="700.0">700– 1280 m</quantity>
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in the National Park of Garajonay and its surroundings. The live specimens (
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<figureCitation id="13232A29C715FFDAA29427D16B9FF892" box="[1176,1270,1776,1802]" captionStart="FIGURE 3. A" captionStartId="7.[151,255,1863,1887]" captionTargetBox="[151,1435,1331,1839]" captionTargetId="figure@7.[151,1436,1331,1839]" captionTargetPageId="7" captionText="FIGURE 3. A. Hemicycla laurijona sp. nov., paratype from Puntas Coloradas, La Gomera. B. Hemicycla fulgida sp. nov., paratype from El Bailadero, Tenerife." httpUri="https://zenodo.org/record/176900/files/figure.png" pageId="5" pageNumber="6">Fig. 3A</figureCitation>
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) were mainly found under moss on the trunks of trees in the laurel forest.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3026527C715FFD9A6CB26616A67F86B" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph id="8BA736ACC715FFD9A6CB26616EA6FE3A" blockId="5.[151,1437,1413,2002]" lastBlockId="6.[151,1437,152,418]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<emphasis id="B96CEABEC715FFDAA6CB26616E31F8C2" bold="true" box="[199,344,1856,1882]" pageId="5" pageNumber="6">Description</emphasis>
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. Body dark-brown coloured, sole pale-brown. The shell (Table 2;
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<figureCitation id="13232A29C715FFDAA25626616BC3F8C2" box="[1114,1194,1856,1882]" captionStart="FIGURE 2" captionStartId="7.[151,255,1115,1139]" captionTargetBox="[161,1427,324,1067]" captionTargetId="figure@7.[151,1436,314,1091]" captionTargetPageId="7" captionText="FIGURE 2. Shell drawings. A. Hemicycla laurijona sp. nov. holotype. B. Hemicycla fulgida sp. nov., holotype showing the placement of the measurements obtained. BH, body whorl height (at columella level); BP, body whorl frontal perimeter; BS, body whorl frontal surface (plane view); D 1, maximum shell diameter; D 2, shell diameter perpendicular to D 1; FP, shell frontal perimeter; FS, shell frontal surface (plane view); SH, shell height; SP, shell perimeter (dorsal plane view); SS, shell surface (dorsal plane view)." httpUri="https://zenodo.org/record/176899/files/figure.png" pageId="5" pageNumber="6">Figs. 2</figureCitation>
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A, 3A, 4A,B, 5A–C) is imperforate, slightly depressed globular, with 3¾–4¼ low convex whorls and shallow sutures. The aperture is oblique, without angulations and rounded in the palatal zone. The white peristome is only slightly reflected in a lip in the palatal zone, whereas it is well reflected in the columellar zone, covering the umbilicus. The shell has a small, weak and wide radial costulation combined in the body whorl, with an also weak malleate surface sculpture. The shell surface is glossy, brilliant, with highly variable colouring. Some specimens are pale-brown, others dark brown; some are uniform, slightly reddish and others pale greenish coloured. In some specimens there are four, in others five spiral, darker bands: two-three dorsal bands narrow or wide, sometimes interrupted, and two ventral narrower and more uniform. In many specimens, all the bands are interrupted, forming irregular patches that fuse with those of the neighbouring bands, giving the shell a multicoloured appearance.
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</paragraph>
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<caption id="DF676624C716FFD9A69B24486AE7FA38" httpUri="https://zenodo.org/record/176898/files/figure.png" pageId="6" pageNumber="7" targetBox="[161,1421,476,1348]" targetPageId="6">
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<paragraph id="8BA736ACC716FFD9A69B24486AE7FA38" blockId="6.[151,1436,1385,1440]" pageId="6" pageNumber="7">
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<emphasis id="B96CEABEC716FFD9A69B24486E73FA19" bold="true" box="[151,282,1385,1409]" pageId="6" pageNumber="7">FIGURE 1.</emphasis>
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Geographical distribution of
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<taxonomicName id="4C184D2FC716FFD9A45624486C5EFA18" box="[602,823,1385,1408]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
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<emphasis id="B96CEABEC716FFD9A45624486C5EFA18" box="[602,823,1385,1408]" italics="true" pageId="6" pageNumber="7">Hemicycla laurijona</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25F57C5C716FFD9A53324486CFCFA19" box="[831,917,1385,1409]" pageId="6" pageNumber="7" rank="species">
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||
<emphasis id="B96CEABEC716FFD9A53324486CFCFA19" bold="true" box="[831,917,1385,1409]" pageId="6" pageNumber="7">sp. nov.</emphasis>
|
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</taxonomicNameLabel>
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and
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<taxonomicName id="4C184D2FC716FFD9A5C024486BF8FA18" box="[972,1169,1385,1408]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov.">
|
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<emphasis id="B96CEABEC716FFD9A5C024486BF8FA18" box="[972,1169,1385,1408]" italics="true" pageId="6" pageNumber="7">Hemicycla fulgida</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C716FFD9A29524486B86FA19" box="[1177,1263,1385,1409]" pageId="6" pageNumber="7" rank="species">
|
||
<emphasis id="B96CEABEC716FFD9A29524486B86FA19" bold="true" box="[1177,1263,1385,1409]" pageId="6" pageNumber="7">sp. nov.</emphasis>
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</taxonomicNameLabel>
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The blue arrow indicates approximately the area of the
|
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<taxonomicName id="4C184D2FC716FFD9A43724A86C44FA38" box="[571,813,1417,1440]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="invernicata">
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<emphasis id="B96CEABEC716FFD9A43724A86C44FA38" box="[571,813,1417,1440]" italics="true" pageId="6" pageNumber="7">Hemicycla invernicata</emphasis>
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</taxonomicName>
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type locality. Symbols represent 1 x 1 km UTM squares.
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</paragraph>
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</caption>
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<paragraph id="8BA736ACC716FFD9A6CB24F06C84F913" blockId="6.[151,1437,1489,2035]" pageId="6" pageNumber="7">
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Genital system (Fig. 6A–C;
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<specimenCount id="9D1EFD25C716FFD9A41024F06DC7FA73" box="[540,686,1489,1515]" pageId="6" pageNumber="7" type="generic">3 specimens</specimenCount>
|
||
dissected): Atrium short. The bursa copulatrix complex in the female genital system shows a moderately short diverticulum, slightly longer than the bursa copulatrix duct. The latter is slightly shorter than the common stalk and slightly swollen distally, shortly before entering the proximal vagina. The bursa copulatrix is rather small and globular, with its neck inserted into the diaphragm. The dart sac is accompanied by a pair of branched tubular mucus glands.
|
||
</paragraph>
|
||
<paragraph id="8BA736ACC716FFD9A6CB27B86C6DF8CB" blockId="6.[151,1437,1489,2035]" pageId="6" pageNumber="7">The male portion of the genital system shows a penial complex with a very long flagellum, about 4–5 times longer than the distal male duct (i. e. that part of the penial complex between the retractor muscle insertion and the atrium). The penis sheath is very thin, translucent and poorly visible. The retractor muscle has an epiphallar insertion. The length proportions between the above mentioned male and female ducts are similar to those of the immature specimen shown in Fig. 6B.</paragraph>
|
||
<paragraph id="8BA736ACC716FFD9A6CB26406A67F86B" blockId="6.[151,1437,1489,2035]" pageId="6" pageNumber="7">The penis has a system of twin papillae with a penial chamber between them (Fig. 6C). The penial chamber has pleated walls and is in fact the chamber of the distal penial papilla. The connection between the distal penial papilla and the penis wall is sometimes visible from the exterior (Fig. 6B) and we named it as the “ring zone”. A well developed contact organ is located between the distal penial papilla and the atrium.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3026527C717FFD7A6CA21B66AF5F965" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="discussion">
|
||
<paragraph id="8BA736ACC717FFD8A6CA21B66E8FFE9A" blockId="7.[151,1436,151,258]" pageId="7" pageNumber="8">
|
||
<emphasis id="B96CEABEC717FFD8A6CA21B66E29FF29" bold="true" box="[198,320,151,177]" pageId="7" pageNumber="8">Remarks.</emphasis>
|
||
The species most similar to
|
||
<taxonomicName id="4C184D2FC717FFD8A48221B96C76FF29" box="[654,799,152,177]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC717FFD8A48221B96C76FF29" box="[654,799,152,177]" italics="true" pageId="7" pageNumber="8">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
in shell shape and shell dimensions is
|
||
<taxonomicName id="4C184D2FC717FFD8A2E921B96AF5FF29" box="[1253,1436,152,177]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC717FFD8A2E921B96AF5FF29" box="[1253,1436,152,177]" italics="true" pageId="7" pageNumber="8">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13232A29C717FFD8A69321E16F9BFF42" box="[159,242,192,218]" captionStart="FIGURE 4" captionStartId="8.[151,255,1373,1397]" captionTargetBox="[151,1434,194,1348]" captionTargetId="figure@8.[151,1435,194,1349]" captionTargetPageId="8" captionText="FIGURE 4. Shells. A. Hemicycla laurijona sp. nov., holotype. B. Hemicycla laurijona sp. nov., paratypes from Puntas Coloradas, La Gomera. C. Hemicycla perraudierei, from Las Casillas, El Hierro." httpUri="https://zenodo.org/record/176901/files/figure.png" pageId="7" pageNumber="8">Figs. 4</figureCitation>
|
||
C, 5D–F) from El Hierro, the westernmost, youngest and smallest island of the archipelago, about
|
||
<quantity id="4CE09B49C717FFD8A37321E16FD4FE9A" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="8" unit="km" value="60.0">60 km</quantity>
|
||
distant from La Gomera.
|
||
</paragraph>
|
||
<caption id="DF676624C717FFD8A69B257A6D07FB65" httpUri="https://zenodo.org/record/176899/files/figure.png" pageId="7" pageNumber="8" targetBox="[161,1427,324,1067]" targetPageId="7">
|
||
<paragraph id="8BA736ACC717FFD8A69B257A6D07FB65" blockId="7.[151,1436,1115,1277]" pageId="7" pageNumber="8">
|
||
<emphasis id="B96CEABEC717FFD8A69B257A6E70FBEB" bold="true" box="[151,281,1115,1139]" pageId="7" pageNumber="8">FIGURE 2.</emphasis>
|
||
Shell drawings. A.
|
||
<taxonomicName id="4C184D2FC717FFD8A7E7257A6DAEFBEA" box="[491,711,1115,1138]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC717FFD8A7E7257A6DAEFBEA" box="[491,711,1115,1138]" italics="true" pageId="7" pageNumber="8">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C717FFD8A4C1257A6C4BFBEB" box="[717,802,1115,1139]" pageId="7" pageNumber="8" rank="species">
|
||
<emphasis id="B96CEABEC717FFD8A4C1257A6C4BFBEB" bold="true" box="[717,802,1115,1139]" pageId="7" pageNumber="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype. B.
|
||
<taxonomicName id="4C184D2FC717FFD8A5BE257A6B1FFBEA" box="[946,1142,1115,1138]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov.">
|
||
<emphasis id="B96CEABEC717FFD8A5BE257A6B1FFBEA" box="[946,1142,1115,1138]" italics="true" pageId="7" pageNumber="8">Hemicycla fulgida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C717FFD8A270257A6BB8FBEB" box="[1148,1233,1115,1139]" pageId="7" pageNumber="8" rank="species">
|
||
<emphasis id="B96CEABEC717FFD8A270257A6BB8FBEB" bold="true" box="[1148,1233,1115,1139]" pageId="7" pageNumber="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype showing the placement of the measurements obtained. BH, body whorl height (at columella level); BP, body whorl frontal perimeter; BS, body whorl frontal surface (plane view); D1, maximum shell diameter; D2, shell diameter perpendicular to D1; FP, shell frontal perimeter; FS, shell frontal surface (plane view); SH, shell height; SP, shell perimeter (dorsal plane view); SS, shell surface (dorsal plane view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF676624C717FFD8A69B26666D0AF819" httpUri="https://zenodo.org/record/176900/files/figure.png" pageId="7" pageNumber="8" targetBox="[151,1435,1331,1839]" targetPageId="7">
|
||
<paragraph id="8BA736ACC717FFD8A69B26666D0AF819" blockId="7.[151,1435,1863,1922]" pageId="7" pageNumber="8">
|
||
<emphasis id="B96CEABEC717FFD8A69B26666E72F8C7" bold="true" box="[151,283,1863,1887]" pageId="7" pageNumber="8">FIGURE 3.</emphasis>
|
||
A.
|
||
<taxonomicName id="4C184D2FC717FFD8A74B26696D4CF8C7" box="[327,549,1864,1887]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC717FFD8A74B26696D4CF8C7" box="[327,549,1864,1887]" italics="true" pageId="7" pageNumber="8">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C717FFD8A42226666DECF8C7" box="[558,645,1863,1887]" pageId="7" pageNumber="8" rank="species">
|
||
<emphasis id="B96CEABEC717FFD8A42226666DECF8C7" bold="true" box="[558,645,1863,1887]" pageId="7" pageNumber="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype from Puntas Coloradas, La Gomera. B.
|
||
<taxonomicName id="4C184D2FC717FFD8A2A126696A1AF8C7" box="[1197,1395,1864,1887]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="fulgida" status="sp. nov.">
|
||
<emphasis id="B96CEABEC717FFD8A2A126696A1AF8C7" box="[1197,1395,1864,1887]" italics="true" pageId="7" pageNumber="8">Hemicycla fulgida</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C717FFD8A37026666FAFF81A" pageId="7" pageNumber="8" rank="species">
|
||
<emphasis id="B96CEABEC717FFD8A37026666FAFF81A" bold="true" pageId="7" pageNumber="8">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype from El Bailadero, Tenerife.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF676624C718FFD7A69B247C6C9FFA0F" httpUri="https://zenodo.org/record/176901/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1434,194,1348]" targetPageId="8">
|
||
<paragraph id="8BA736ACC718FFD7A69B247C6C9FFA0F" blockId="8.[151,1436,1373,1431]" pageId="8" pageNumber="9">
|
||
<emphasis id="B96CEABEC718FFD7A69B247C6E70FAED" bold="true" box="[151,281,1373,1397]" pageId="8" pageNumber="9">FIGURE 4.</emphasis>
|
||
Shells. A.
|
||
<taxonomicName id="4C184D2FC718FFD7A79E247F6D07FAED" box="[402,622,1374,1397]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC718FFD7A79E247F6D07FAED" box="[402,622,1374,1397]" italics="true" pageId="8" pageNumber="9">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C718FFD7A479247C6DA2FAED" box="[629,715,1373,1397]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B96CEABEC718FFD7A479247C6DA2FAED" bold="true" box="[629,715,1373,1397]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype. B.
|
||
<taxonomicName id="4C184D2FC718FFD7A568247F6B28FAED" box="[868,1089,1374,1397]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="laurijona" status="sp. nov.">
|
||
<emphasis id="B96CEABEC718FFD7A568247F6B28FAED" box="[868,1089,1374,1397]" italics="true" pageId="8" pageNumber="9">Hemicycla laurijona</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25F57C5C718FFD7A244247C6BF7FAED" box="[1096,1182,1373,1397]" pageId="8" pageNumber="9" rank="species">
|
||
<emphasis id="B96CEABEC718FFD7A244247C6BF7FAED" bold="true" box="[1096,1182,1373,1397]" pageId="8" pageNumber="9">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratypes from Puntas Coloradas, La Gomera. C.
|
||
<emphasis id="B96CEABEC718FFD7A7B924A16DD3FA0F" box="[437,698,1408,1431]" italics="true" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C184D2FC718FFD7A7B924A16DDFFA0F" box="[437,694,1408,1431]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="perraudierei">Hemicycla perraudierei</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
from Las Casillas, El Hierro.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA736ACC718FFD7A6CA24EA6AF5F965" blockId="8.[151,1436,1483,1789]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="4C184D2FC718FFD7A6CA24EA6E31FA7C" box="[198,344,1483,1508]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC718FFD7A6CA24EA6E31FA7C" box="[198,344,1483,1508]" italics="true" pageId="8" pageNumber="9">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
generally has a slightly smaller and more depressed shell than
|
||
<taxonomicName id="4C184D2FC718FFD7A24824EA6B95FA7C" box="[1092,1276,1483,1508]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC718FFD7A24824EA6B95FA7C" box="[1092,1276,1483,1508]" italics="true" pageId="8" pageNumber="9">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
(Figs. 4,5,7), and also a weaker shell malleation (
|
||
<figureCitation id="13232A29C718FFD7A44424D26DFBF995" box="[584,658,1523,1549]" captionStart="FIGURE 5" captionStartId="9.[151,255,1454,1478]" captionTargetBox="[151,1436,155,1440]" captionTargetId="figure@9.[151,1436,155,1440]" captionTargetPageId="9" captionText="FIGURE 5. Shell body whorl details. A – C. Hemicycla laurijona sp. nov., holotype. D – F. Hemicycla perraudierei. A, D, dorsal view. B, E, frontal view. C, F, ventral view. The sells are the same as in Fig. 4." httpUri="https://zenodo.org/record/176902/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
), but the main morphological difference between both species is found in the female part of the genital organs, namely in the length of the stalk of the bursa copulatrix complex. The stalk in
|
||
<taxonomicName id="4C184D2FC718FFD7A76527626E90F9C4" box="[361,505,1603,1628]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC718FFD7A76527626E90F9C4" box="[361,505,1603,1628]" italics="true" pageId="8" pageNumber="9">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
is, in fact, similar in length to that of the bursa copulatrix duct and of the diverticulum (Fig. 6A,B), whereas in
|
||
<taxonomicName id="4C184D2FC718FFD7A41F274A6DA3F91C" box="[531,714,1643,1668]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC718FFD7A41F274A6DA3F91C" box="[531,714,1643,1668]" italics="true" pageId="8" pageNumber="9">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
it is nearly twice as long (Fig. 6D,E). Both species also differ in their microhabitat.
|
||
<taxonomicName id="4C184D2FC718FFD7A7AC27B26D5CF934" box="[416,565,1683,1708]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="laurijona">
|
||
<emphasis id="B96CEABEC718FFD7A7AC27B26D5CF934" box="[416,565,1683,1708]" italics="true" pageId="8" pageNumber="9">H. laurijona</emphasis>
|
||
</taxonomicName>
|
||
lives under moss on the tree trunks in the laurel forest of La Gomera, whereas
|
||
<taxonomicName id="4C184D2FC718FFD7A6F1279A6EDDF94C" box="[253,436,1723,1748]" class="Gastropoda" family="Helicidae" genus="Hemicycla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="perraudierei">
|
||
<emphasis id="B96CEABEC718FFD7A6F1279A6EDDF94C" box="[253,436,1723,1748]" italics="true" pageId="8" pageNumber="9">H. perraudierei</emphasis>
|
||
</taxonomicName>
|
||
was found at a similar altitude (
|
||
<quantity id="4CE09B49C718FFD7A524279A6CD6F94D" box="[808,959,1723,1749]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="11.0" metricValueMin="6.0" pageId="8" pageNumber="9" unit="m" value="850.0" valueMax="1100.0" valueMin="600.0">600–1100 m</quantity>
|
||
), in the upper area of the degraded laurel forests of El Golfo and Tiñor (El Hierro), among the leaves of
|
||
<taxonomicName id="4C184D2FC718FFD7A56327C26CBEF964" box="[879,983,1763,1788]" class="Magnoliopsida" family="Crassulaceae" genus="Aeonium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B96CEABEC718FFD7A56327C26CBEF964" box="[879,983,1763,1788]" italics="true" pageId="8" pageNumber="9">Aeonium</emphasis>
|
||
</taxonomicName>
|
||
sp. and also among fallen dead leaves.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |