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<mods:title id="D3CABFFDE076783B81D4BFABC014B796">The taxonomy of spider crabs of the genera Eurynome, Choniognathus, Seiitaiodes and Kasagia (Crustacea: Brachyura: Majidae) from southwest Indian Ocean</mods:title>
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<mods:namePart id="F22C50BDB67C2E68F2412B72AC701465">Forges, Bertrand Richer De</mods:namePart>
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<mods:namePart id="289D53FCC49B3E782927818CDCCAC0DA">Lee, Bee Yan</mods:namePart>
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<mods:namePart id="C7826D99D4007FB68B4CF8F6B72148B1">Ng, Peter K. L.</mods:namePart>
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<treatment id="03B187B7FF96FFB889B3A3B8FE6F5DB4" ID-DOI="http://doi.org/10.5281/zenodo.5569406" ID-GBIF-Taxon="188638480" ID-Zenodo-Dep="5569406" LSID="urn:lsid:plazi:treatment:03B187B7FF96FFB889B3A3B8FE6F5DB4" httpUri="http://treatment.plazi.org/id/03B187B7FF96FFB889B3A3B8FE6F5DB4" lastPageId="11" lastPageNumber="311" pageId="10" pageNumber="311">
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<heading id="D0EF81CDFF96FFB989B3A3B8FDEC5E74" bold="true" box="[151,524,762,789]" fontSize="11" level="1" pageId="10" pageNumber="311" reason="1">
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<taxonomicName id="4C184D22FF96FFB989B3A3B8FE225E74" ID-CoL="8RW7F" authority="Forges &amp; Lee &amp; Ng, 2021" authorityName="Forges &amp; Lee &amp; Ng" authorityYear="2021" box="[151,450,762,789]" class="Malacostraca" family="Majidae" genus="Choniognathus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B96CEAB3FF96FFB989B3A3B8FE225E74" bold="true" box="[151,450,762,789]" italics="true" pageId="10" pageNumber="311">Choniognathus spinosus</emphasis>
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<taxonomicNameLabel id="A25F57C8FF96FFB988EEA3B9FDEC5E74" box="[458,524,763,789]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C302652AFF96FFB989B3A25DFE955E5B" box="[151,373,799,826]" pageId="10" pageNumber="311" type="description">
<paragraph id="8BA736A1FF96FFB989B3A25DFE955E5B" blockId="10.[151,524,762,826]" box="[151,373,799,826]" pageId="10" pageNumber="311">
(
<figureCitation id="13232A24FF96FFB989BBA25DFEC05E58" box="[159,288,799,825]" captionStart="FIGURE 1" captionStartId="4.[151,250,1781,1805]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-22@4.[151,1436,181,1757]" captionTargetPageId="4" captionText="FIGURE 1. Colour in life. A, Choniognathus spinosus n. sp., holotype female (cl 9.5 mm, cw 7.3 mm) (MNHN-IU-2008- 10297); B, Kasagia elegans (Stebbing, 1921) comb. nov., male (cl 11.8 mm, cw 7.9 mm) (MNHN-IU-2008-10340); C, K. elegans (Stebbing, 1921) comb. nov., female (cl 11.9 mm, cw 7.4 mm) (MNHN-IU-2010-77); D, K. elegans (Stebbing, 1921) comb. nov., female (cl 7.9 mm, cw 4.9 mm) (MNHN-IU-2008-10339); E, K. sudhakari Padate, Manjebrayakath &amp; Ng, 2019, female, station CP3131, MAINBAZA cruise, Mozambique Channel, specimen not found. Photographs: T.-Y. Chan." figureDoi="http://doi.org/10.5281/zenodo.5556343" httpUri="https://zenodo.org/record/5556343/files/figure.png" pageId="10" pageNumber="311">Figures 1A</figureCitation>
,
<figureCitation id="13232A24FF96FFB98808A25DFE8D5E5B" box="[300,365,799,826]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">6AF</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C302652AFF96FFB989B3A224FA705EC4" pageId="10" pageNumber="311" type="materials_examined">
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<emphasis id="B96CEAB3FF96FFB989B3A224FE1D5EE0" bold="true" box="[151,509,870,897]" pageId="10" pageNumber="311">
Material examined.
<typeStatus id="54A38803FF96FFB988A2A225FE175EE0" box="[390,503,871,897]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
:
</emphasis>
ovigerous female (cl
<quantity id="4CE09B44FF96FFB98BCAA225FCB45EE0" box="[750,852,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="10" pageNumber="311" unit="mm" value="10.2">10.2 mm</quantity>
, pcl
<quantity id="4CE09B44FF96FFB98AA3A224FC3F5EE0" box="[903,991,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="10" pageNumber="311" unit="mm" value="9.5">9.5 mm</quantity>
, cw
<quantity id="4CE09B44FF96FFB98D35A225FB885EE0" box="[1041,1128,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="10" pageNumber="311" unit="mm" value="8.3">8.3 mm</quantity>
, bcw
<quantity id="4CE09B44FF96FFB98D8CA225FB1F5EE0" box="[1192,1279,871,897]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="311" unit="mm" value="7.3">7.3 mm</quantity>
) (MNHN-IU-2008-10297), stn CP3132,
<collectingCountry id="F30F7631FF96FFB988E2A2C9FDBA5EC4" box="[454,602,907,933]" name="Mozambique" pageId="10" pageNumber="311">Mozambique</collectingCountry>
Channel, 2511S 3501E,
<quantity id="4CE09B44FF96FFB98AA7A2C9FBE45EC4" box="[899,1028,907,933]" metricMagnitude="2" metricUnit="m" metricValue="1.015" metricValueMax="1.02" metricValueMin="1.01" pageId="10" pageNumber="311" unit="m" value="101.5" valueMax="102.0" valueMin="101.0">101102 m</quantity>
, coll. MAINBAZA,
<date id="FFA61061FF96FFB98DC9A2C9FA6C5EC4" box="[1261,1420,907,933]" pageId="10" pageNumber="311" value="2009-04-10">10 April 2009</date>
.
</paragraph>
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<subSubSection id="C302652AFF96FFB989E3A2ECFD4A58A0" pageId="10" pageNumber="311" type="diagnosis">
<paragraph id="8BA736A1FF96FFB989E3A2ECFD4A58A0" blockId="10.[151,1437,870,2013]" pageId="10" pageNumber="311">
<emphasis id="B96CEAB3FF96FFB989E3A2ECFEA25EA9" bold="true" box="[199,322,942,968]" pageId="10" pageNumber="311">Diagnosis.</emphasis>
Dorsal surface of carapace covered with numerous prominent spines and sharp tubercles; with stiff scattered setae, without dense tomentum which completely covers surface (
<figureCitation id="13232A24FF96FFB98D07A291FB7F5E8C" box="[1059,1183,979,1005]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6A, B</figureCitation>
). Pseudorostral spines short, strongly diverging (
<figureCitation id="13232A24FF96FFB9889BA2B5FDDB5970" box="[447,571,1015,1041]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6A, B</figureCitation>
). Supraocular eave broad with concave margin lined with 2 or 3 spines, not completely covering eye (
<figureCitation id="13232A24FF96FFB988E5A559FDFB5957" box="[449,539,1051,1078]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B</figureCitation>
); postocular spine longer than preocular tooth on eave, with short intercalated spine totally filling space between them (
<figureCitation id="13232A24FF96FFB98B54A57DFD2B593B" box="[624,715,1087,1114]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B</figureCitation>
). Hepatic spine long, oriented outwards (
<figureCitation id="13232A24FF96FFB98D87A57DFB1E593B" box="[1187,1278,1087,1114]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B</figureCitation>
). Gastric area raised, with numerous granules, with 2 tubercles on top; cardiac area raised, covered by prominent granules, with 5 posterior granules tuberculiform, forming blunt spines (
<figureCitation id="13232A24FF96FFB98A05A5C5FC7C59C0" box="[801,924,1159,1185]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B, C</figureCitation>
); branchial area inflated covered with spines, 2 of them longer (
<figureCitation id="13232A24FF96FFB9884EA5E9FE2559A7" box="[362,453,1195,1222]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B</figureCitation>
). Posterior carapace margin with 3 rows of long granules like-spines (
<figureCitation id="13232A24FF96FFB98DC9A5E9FAA859A7" box="[1261,1352,1195,1222]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B</figureCitation>
). Basal antennal article with 4 spines distally, base appressed against infraorbital margin with 2 long spines (
<figureCitation id="13232A24FF96FFB98C16A58DFA6F598B" box="[1330,1423,1231,1258]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6D</figureCitation>
). Buccal frame quadrangular, completely covered by third maxillipeds (
<figureCitation id="13232A24FF96FFB98A80A5B1FBFC586C" box="[932,1052,1267,1293]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6D, E</figureCitation>
). Third maxilliped with merus and ischium covered by scattered tubercles, merus separated from ischium by suture, structures mobile; anteroexternal angle of merus strongly produced, auriculiform (
<figureCitation id="13232A24FF96FFB98B9AA479FCD95834" box="[702,825,1339,1365]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6D, E</figureCitation>
). Cheliped short, dorsal margins and outer surfaces of carpus, merus and chela covered by spines and stiff setae; cutting margins of fingers finely serrulate (
<figureCitation id="13232A24FF96FFB98C3CA41DFA6F5818" box="[1304,1423,1375,1401]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6E, F</figureCitation>
). Ambulatory legs short, setose (
<figureCitation id="13232A24FF96FFB988D0A4C1FDAE58FF" box="[500,590,1411,1438]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6A</figureCitation>
). Pleon with 6 free somites and telson, somite 1 with large median swelling covered by tubercles and granules (
<figureCitation id="13232A24FF96FFB98B07A4E5FD7D58A0" box="[547,669,1447,1473]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302652AFF96FFB989E3A489FD845884" box="[199,612,1482,1509]" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="8BA736A1FF96FFB989E3A489FD845884" blockId="10.[151,1437,870,2013]" box="[199,612,1482,1509]" pageId="10" pageNumber="311">
<materialsCitation id="3B703CFCFF96FFB989E3A489FD845884" ID-GBIF-Occurrence="3383504319" box="[199,612,1482,1509]" country="Mozambique" location="Channel" pageId="10" pageNumber="311" specimenCount="1" typeStatus="holotype">
<emphasis id="B96CEAB3FF96FFB989E3A489FE865885" bold="true" box="[199,358,1482,1509]" pageId="10" pageNumber="311">
<typeStatus id="54A38803FF96FFB989E3A489FEE25884" box="[199,258,1483,1509]" pageId="10" pageNumber="311">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F30F7631FF96FFB98848A489FDE05884" box="[364,512,1483,1509]" name="Mozambique" pageId="10" pageNumber="311">Mozambique</collectingCountry>
<location id="8EC7607AFF96FFB98B23A489FD845884" LSID="urn:lsid:plazi:treatment:03B187B7FF96FFB889B3A3B8FE6F5DB4:8EC7607AFF96FFB98B23A489FD845884" box="[519,612,1483,1509]" country="Mozambique" name="Channel" pageId="10" pageNumber="311">Channel</location>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302652AFF96FFB989E3A4ADFB255B68" box="[199,1221,1519,1545]" pageId="10" pageNumber="311" type="etymology">
<paragraph id="8BA736A1FF96FFB989E3A4ADFB255B68" blockId="10.[151,1437,870,2013]" box="[199,1221,1519,1545]" pageId="10" pageNumber="311">
<emphasis id="B96CEAB3FF96FFB989E3A4ADFEAE5B68" bold="true" box="[199,334,1519,1545]" pageId="10" pageNumber="311">Etymology.</emphasis>
The name
<taxonomicName id="4C184D22FF96FFB988EFA4ADFDCB5B69" authorityName="Forges &amp; Lee &amp; Ng" authorityYear="2021" box="[459,555,1519,1544]" class="Malacostraca" family="Majidae" genus="Choniognathus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B96CEAB3FF96FFB988EFA4ADFDCB5B69" box="[459,555,1519,1544]" italics="true" pageId="10" pageNumber="311">spinosus</emphasis>
</taxonomicName>
alludes to the spiny carapace and pereopods of the species.
</paragraph>
</subSubSection>
<subSubSection id="C302652AFF96FFB889E3A751FE6F5DB4" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311" type="discussion">
<paragraph id="8BA736A1FF96FFB889E3A751FE6F5DB4" blockId="10.[151,1437,870,2013]" lastBlockId="11.[151,1437,151,213]" lastPageId="11" lastPageNumber="312" pageId="10" pageNumber="311">
<emphasis id="B96CEAB3FF96FFB989E3A751FEDB5B4C" bold="true" box="[199,315,1555,1581]" pageId="10" pageNumber="311">Remarks.</emphasis>
<taxonomicName id="4C184D22FF96FFB98866A751FDB35B4D" authority="Forges &amp; Lee &amp; Ng, 2021" authorityName="Forges &amp; Lee &amp; Ng" authorityYear="2021" box="[322,595,1555,1581]" class="Malacostraca" family="Majidae" genus="Choniognathus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B96CEAB3FF96FFB98866A751FDB35B4D" box="[322,595,1555,1581]" italics="true" pageId="10" pageNumber="311">Choniognathus spinosus</emphasis>
</taxonomicName>
<emphasis id="B96CEAB3FF96FFB98B7EA751FD795B4C" bold="true" box="[602,665,1555,1581]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A25F57C8FF96FFB98B7EA751FD795B4C" box="[602,665,1555,1581]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has several unusual features that question its placement in the genus. It differs from the
<typeStatus id="54A38803FF96FFB9884CA775FE785B30" box="[360,408,1591,1617]" pageId="10" pageNumber="311">type</typeStatus>
species,
<taxonomicName id="4C184D22FF96FFB988DBA775FDB55B31" authorityName="Balss" authorityYear="1924" box="[511,597,1591,1617]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB988DBA775FDB55B31" box="[511,597,1591,1617]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
, in the following characters: the pseudorostral spines are clearly divergent (
<figureCitation id="13232A24FF96FFB98984A719FF1B5B17" box="[160,251,1627,1654]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6A</figureCitation>
) (versus directed anteriorly and subparallel in
<taxonomicName id="4C184D22FF96FFB98A05A719FC995B15" authorityName="Balss" authorityYear="1924" box="[801,889,1627,1653]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98A05A719FC995B15" box="[801,889,1627,1653]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
;
<figureCitation id="13232A24FF96FFB98AA8A719FBD05B14" box="[908,1072,1627,1653]" captionStart="FIGURE 7" captionStartId="12.[151,250,1267,1291]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-22@12.[151,1436,181,1244]" captionTargetPageId="12" captionText="FIGURE 7. Choniognathus reini (Balss, 1924).A, B, Holotype of Choniognathus koreensis Rathbun, 1932, female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; C, D, ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; E, female (cl 9.1 mm, cw 6.2 mm) (KPM NH 0104599), Japan. A, C, overall view; B, ventral view of cephalothorax; D, outer view of right chela and carpus; E, dorsal view of carapace (frontal parts and right side denuded). A, B, courtesy T. Naruse." figureDoi="http://doi.org/10.5281/zenodo.5556367" httpUri="https://zenodo.org/record/5556367/files/figure.png" pageId="10" pageNumber="311">Fig. 7A, C, E</figureCitation>
); the supraorbital eave is more expanded with a spinous margin (
<figureCitation id="13232A24FF96FFB98B36A73DFD6B5BF8" box="[530,651,1663,1689]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6A, B</figureCitation>
) (versus narrow with the margin entire in
<taxonomicName id="4C184D22FF96FFB98D43A73DFB5C5BF9" authorityName="Balss" authorityYear="1924" box="[1127,1212,1663,1689]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98D43A73DFB5C5BF9" box="[1127,1212,1663,1689]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
;
<figureCitation id="13232A24FF96FFB98DEFA73DFA845BF8" box="[1227,1380,1663,1689]" captionStart="FIGURE 7" captionStartId="12.[151,250,1267,1291]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-22@12.[151,1436,181,1244]" captionTargetPageId="12" captionText="FIGURE 7. Choniognathus reini (Balss, 1924).A, B, Holotype of Choniognathus koreensis Rathbun, 1932, female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; C, D, ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; E, female (cl 9.1 mm, cw 6.2 mm) (KPM NH 0104599), Japan. A, C, overall view; B, ventral view of cephalothorax; D, outer view of right chela and carpus; E, dorsal view of carapace (frontal parts and right side denuded). A, B, courtesy T. Naruse." figureDoi="http://doi.org/10.5281/zenodo.5556367" httpUri="https://zenodo.org/record/5556367/files/figure.png" pageId="10" pageNumber="311">Fig. 7A, C, E</figureCitation>
); the postocular spine is as long as than the preocular eave tooth, with the intercalated spine totally covering space (
<figureCitation id="13232A24FF96FFB98C4BA7E1FF595B83" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6B</figureCitation>
) (versus postocular spine is distinctly shorter than the preocular eave tooth with the intercalated spine separated from the eave and postorbital spine by narrow clefts (
<figureCitation id="13232A24FF96FFB98BCAA7A9FC685A64" box="[750,904,1771,1797]" captionStart="FIGURE 7" captionStartId="12.[151,250,1267,1291]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-22@12.[151,1436,181,1244]" captionTargetPageId="12" captionText="FIGURE 7. Choniognathus reini (Balss, 1924).A, B, Holotype of Choniognathus koreensis Rathbun, 1932, female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; C, D, ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; E, female (cl 9.1 mm, cw 6.2 mm) (KPM NH 0104599), Japan. A, C, overall view; B, ventral view of cephalothorax; D, outer view of right chela and carpus; E, dorsal view of carapace (frontal parts and right side denuded). A, B, courtesy T. Naruse." figureDoi="http://doi.org/10.5281/zenodo.5556367" httpUri="https://zenodo.org/record/5556367/files/figure.png" pageId="10" pageNumber="311">Fig. 7A, C, E</figureCitation>
); the merus and ischium of the third maxilliped are mobile (
<figureCitation id="13232A24FF96FFB98804A64DFE985A4B" box="[288,376,1807,1834]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6D</figureCitation>
) (versus fused in
<taxonomicName id="4C184D22FF96FFB98B61A64DFD7B5A49" authorityName="Balss" authorityYear="1924" box="[581,667,1807,1833]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98B61A64DFD7B5A49" box="[581,667,1807,1833]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
with the suture very shallow or undiscernible in
<taxonomicName id="4C184D22FF96FFB98DEFA64DFAC15A49" authorityName="Balss" authorityYear="1924" box="[1227,1313,1807,1833]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98DEFA64DFAC15A49" box="[1227,1313,1807,1833]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
;
<figureCitation id="13232A24FF96FFB98C14A64DFF4F5A2C" captionStart="FIGURE 13" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[199,1387,190,1745]" captionTargetId="figure-22@20.[183,1404,181,1756]" captionTargetPageId="20" captionText="FIGURE 13. AC, Eurynome longimana Stimpson, 1857. A, B, female, South Africa; C, male (cl 14.0 mm, cl 10.0 mm), South Africa; DF, Eurynome longimana Stimpson, 1857, male (cl 8.5 mm, cw 5.9 mm) (ZRC 2020.378), Mozambique; G, Choniognathus reini (Balss, 1924), female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; H, C. reini (Balss, 1924), ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; IK, Seiitaoides mirabilis n. sp., holotype male (cl 13.2 mm, cw 7.3 mm) (MNHN-IU-2010-1301), Madagascar.A, dorsal view of carapace and enlarged boletiform tubercle; B, right P5 merus; C, G1; D, I, left G1 (ventral view); E, J, left G1 (dorsal view); F, K, left G2; G, H, left third maxilliped.AC after Barnard (1950: fig. 12ac). G, from original figure by T. Naruse. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5556389" httpUri="https://zenodo.org/record/5556389/files/figure.png" pageId="10" pageNumber="311">Fig. 13G, H</figureCitation>
); the anteroexternal angle of the merus of the third maxilliped is strongly produced (
<figureCitation id="13232A24FF96FFB98D44A671FB575A2F" box="[1120,1207,1843,1870]" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6D</figureCitation>
) (versus lower in
<taxonomicName id="4C184D22FF96FFB98CA7A671FF2B5A11" authorityName="Balss" authorityYear="1924" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98CA7A671FF2B5A11" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
;
<figureCitation id="13232A24FF96FFB989FFA615FE875A10" box="[219,359,1879,1905]" captionStart="FIGURE 13" captionStartId="20.[151,250,1779,1803]" captionTargetBox="[199,1387,190,1745]" captionTargetId="figure-22@20.[183,1404,181,1756]" captionTargetPageId="20" captionText="FIGURE 13. AC, Eurynome longimana Stimpson, 1857. A, B, female, South Africa; C, male (cl 14.0 mm, cl 10.0 mm), South Africa; DF, Eurynome longimana Stimpson, 1857, male (cl 8.5 mm, cw 5.9 mm) (ZRC 2020.378), Mozambique; G, Choniognathus reini (Balss, 1924), female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; H, C. reini (Balss, 1924), ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; IK, Seiitaoides mirabilis n. sp., holotype male (cl 13.2 mm, cw 7.3 mm) (MNHN-IU-2010-1301), Madagascar.A, dorsal view of carapace and enlarged boletiform tubercle; B, right P5 merus; C, G1; D, I, left G1 (ventral view); E, J, left G1 (dorsal view); F, K, left G2; G, H, left third maxilliped.AC after Barnard (1950: fig. 12ac). G, from original figure by T. Naruse. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5556389" httpUri="https://zenodo.org/record/5556389/files/figure.png" pageId="10" pageNumber="311">Fig. 13G, H</figureCitation>
); and the outer surface of the merus, carpus and chela are covered with prominent spines (
<figureCitation id="13232A24FF96FFB98C4BA615FF165AF4" captionStart="FIGURE 6" captionStartId="11.[151,250,1752,1776]" captionTargetBox="[151,1436,237,1729]" captionTargetId="figure-42@11.[151,1436,237,1729]" captionTargetPageId="11" captionText="FIGURE 6. Choniognathus spinosus n. sp., holotype female (cl 10.2 mm, cw 8.3 mm) (MNHN-IU-2008-10297), Mozambique. A, overall view; B, dorsal view of carapace (denuded); C, lateral view of cephalothorax; D, ventral view showing antennules, antennae, epistome, buccal cavity and third maxillipeds; E, ventral view of cephalothorax; F, view of right chela and carpus." figureDoi="http://doi.org/10.5281/zenodo.5556363" httpUri="https://zenodo.org/record/5556363/files/figure.png" pageId="10" pageNumber="311">Fig. 6A, E, F</figureCitation>
) (versus almost smooth in
<taxonomicName id="4C184D22FF96FFB98B06A639FD965AF5" authorityName="Balss" authorityYear="1924" box="[546,630,1915,1941]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98B06A639FD965AF5" box="[546,630,1915,1941]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
;
<figureCitation id="13232A24FF96FFB98BA0A639FD3C5AF4" box="[644,732,1915,1941]" captionStart="FIGURE 7" captionStartId="12.[151,250,1267,1291]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-22@12.[151,1436,181,1244]" captionTargetPageId="12" captionText="FIGURE 7. Choniognathus reini (Balss, 1924).A, B, Holotype of Choniognathus koreensis Rathbun, 1932, female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; C, D, ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; E, female (cl 9.1 mm, cw 6.2 mm) (KPM NH 0104599), Japan. A, C, overall view; B, ventral view of cephalothorax; D, outer view of right chela and carpus; E, dorsal view of carapace (frontal parts and right side denuded). A, B, courtesy T. Naruse." figureDoi="http://doi.org/10.5281/zenodo.5556367" httpUri="https://zenodo.org/record/5556367/files/figure.png" pageId="10" pageNumber="311">Fig. 7D</figureCitation>
). In addition, while
<taxonomicName id="4C184D22FF96FFB98A9FA639FBEF5AF5" authorityName="Balss" authorityYear="1924" box="[955,1039,1915,1941]" class="Malacostraca" family="Majidae" genus="Eurynome" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="reini">
<emphasis id="B96CEAB3FF96FFB98A9FA639FBEF5AF5" box="[955,1039,1915,1941]" italics="true" pageId="10" pageNumber="311">C. reini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C184D22FF96FFB98D60A639FB005AF4" baseAuthorityName="Baker" baseAuthorityYear="1906" box="[1092,1248,1915,1941]" class="Malacostraca" family="Majidae" genus="Choniognathus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="B96CEAB3FF96FFB98D60A639FB005AF4" box="[1092,1248,1915,1941]" italics="true" pageId="10" pageNumber="311">C. granulosus</emphasis>
</taxonomicName>
(cf.
<bibRefCitation id="EF894B50FF96FFB98C37A639FF225AD8" author="Baker, W. H." pageId="10" pageNumber="311" pagination="104 - 117" refId="ref15522" refString="Baker, W. H. (1906) Notes on South Australian decapod Crustacea. Part IV. Transactions of the Royal Society of South Australia, 30, 104 - 117, pls. 1 - 3." type="journal article" year="1906">Baker 1906: 108</bibRefCitation>
) have short ambulatory legs with all the dorsal surfaces of the carapace and pereiopods completely covered by a short dense tomentum (
<figureCitation id="13232A24FF96FFB98890A681FDEC5ABC" box="[436,524,1987,2013]" captionStart="FIGURE 7" captionStartId="12.[151,250,1267,1291]" captionTargetBox="[151,1436,181,1244]" captionTargetId="figure-22@12.[151,1436,181,1244]" captionTargetPageId="12" captionText="FIGURE 7. Choniognathus reini (Balss, 1924).A, B, Holotype of Choniognathus koreensis Rathbun, 1932, female (cl 9.1 mm, cw 6.0 mm) (USNM 48204), Korea; C, D, ovigerous female (cl 6.3 mm, cw 4.0 mm) (ZRC 2020.373), Philippines; E, female (cl 9.1 mm, cw 6.2 mm) (KPM NH 0104599), Japan. A, C, overall view; B, ventral view of cephalothorax; D, outer view of right chela and carpus; E, dorsal view of carapace (frontal parts and right side denuded). A, B, courtesy T. Naruse." figureDoi="http://doi.org/10.5281/zenodo.5556367" httpUri="https://zenodo.org/record/5556367/files/figure.png" pageId="10" pageNumber="311">Fig. 7E</figureCitation>
), this is not the case with
<taxonomicName id="4C184D22FF96FFB98A17A681FC545ABD" authority="Forges &amp; Lee &amp; Ng, 2021" authorityName="Forges &amp; Lee &amp; Ng" authorityYear="2021" box="[819,948,1987,2013]" class="Malacostraca" family="Majidae" genus="Choniognathus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="spinosus" status="sp. nov.">
<emphasis id="B96CEAB3FF96FFB98A17A681FC545ABD" box="[819,948,1987,2013]" italics="true" pageId="10" pageNumber="311">C. spinosus</emphasis>
</taxonomicName>
<emphasis id="B96CEAB3FF96FFB98A98A681FC1C5ABC" bold="true" box="[956,1020,1987,2013]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A25F57C8FF96FFB98A98A681FC1C5ABC" box="[956,1020,1987,2013]" pageId="10" pageNumber="311" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the carapace and pereopods are just covered with short stiff setae, which while dense, do not completely obscure the outline and surfaces like the other two species (
<figureCitation id="13232A24FF97FFB8880CA1F9FE615DB4" box="[296,385,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1781,1805]" captionTargetBox="[151,1436,181,1757]" captionTargetId="figure-22@4.[151,1436,181,1757]" captionTargetPageId="4" captionText="FIGURE 1. Colour in life. A, Choniognathus spinosus n. sp., holotype female (cl 9.5 mm, cw 7.3 mm) (MNHN-IU-2008- 10297); B, Kasagia elegans (Stebbing, 1921) comb. nov., male (cl 11.8 mm, cw 7.9 mm) (MNHN-IU-2008-10340); C, K. elegans (Stebbing, 1921) comb. nov., female (cl 11.9 mm, cw 7.4 mm) (MNHN-IU-2010-77); D, K. elegans (Stebbing, 1921) comb. nov., female (cl 7.9 mm, cw 4.9 mm) (MNHN-IU-2008-10339); E, K. sudhakari Padate, Manjebrayakath &amp; Ng, 2019, female, station CP3131, MAINBAZA cruise, Mozambique Channel, specimen not found. Photographs: T.-Y. Chan." figureDoi="http://doi.org/10.5281/zenodo.5556343" httpUri="https://zenodo.org/record/5556343/files/figure.png" pageId="11" pageNumber="312">Fig. 1A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>