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<document ID-DOI="http://doi.org/10.5281/zenodo.6461215" ID-GBIF-Dataset="0cd8eeae-9f43-4947-9c9d-402bf45c1a15" ID-Zenodo-Dep="6461215" LSID="urn:lsid:zoobank.org:act:F092C8CB-A78E-46E8-9CF2-0FEFE201450E" approvalRequired="1" approvalRequired_for_document="1" checkinTime="1649940404519" checkinUser="jonas" docAuthor="Ramage, Thibault &amp; Löbl, Ivan" docDate="2022" docId="03B1879EFFCE502059EF2CF1C117FC0B" docLanguage="en" docName="BullSocEntFra.127.1.55-60.pdf" docOrigin="Bulletin de la Société entomologique de France 127 (1)" docTitle="Scaphisoma jacqi Lobl &amp; Ramage 2022, n. sp." docType="treatment" docUuid="F092C8CB-A78E-46E8-9CF2-0FEFE201450E" docUuidSource="ZooBank" docVersion="4" lastPageId="4" lastPageNumber="59" masterDocId="FF88FFE6FFCC5024584B2D43C058FF8D" masterDocTitle="A new species of Scaphisoma Leach from the Society Islands with commentary on Staphylinidae of the French Polynesia (Coleoptera, Staphylinidae, Scaphidiinae)" masterLastPageNumber="60" masterPageNumber="55" pageId="2" pageNumber="57" updateTime="1649941245975" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Scaphisoma Leach from the Society Islands with commentary on Staphylinidae of the French Polynesia (Coleoptera, Staphylinidae, Scaphidiinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ramage, Thibault</mods:namePart>
<mods:affiliation>9 quartier de la Glacière, F 29900 Concarneau, France</mods:affiliation>
<mods:nameIdentifier type="email">thibault.ramage@hotmail.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Löbl, Ivan</mods:namePart>
<mods:affiliation>Muséum dhistoire naturelle, Case postale 6434, CH 1211 Genève, Suisse</mods:affiliation>
<mods:nameIdentifier type="email">ivan.lobl@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Bulletin de la Société entomologique de France</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-03-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>127</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>55</mods:start>
<mods:end>60</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.32475/bsef_2224</mods:identifier>
<mods:identifier type="GBIF-Dataset">0cd8eeae-9f43-4947-9c9d-402bf45c1a15</mods:identifier>
<mods:identifier type="Zenodo-Dep">6461191</mods:identifier>
<mods:identifier type="ZooBank">EBBF29FA-1CDA-446A-8D76-966C6C5B50B7</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6461215" ID-Zenodo-Dep="6461215" LSID="urn:lsid:zoobank.org:act:F092C8CB-A78E-46E8-9CF2-0FEFE201450E" httpUri="http://treatment.plazi.org/id/03B1879EFFCE502059EF2CF1C117FC0B" lastPageId="4" lastPageNumber="59" pageId="2" pageNumber="57">
<subSubSection box="[420,865,434,459]" pageId="2" pageNumber="57" type="nomenclature">
<paragraph blockId="2.[144,865,434,490]" box="[420,865,434,459]" pageId="2" pageNumber="57">
<heading bold="true" box="[420,865,434,459]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="57" reason="0">
<taxonomicName authority="Lobl &amp; Ramage" authorityName="Lobl &amp; Ramage" authorityYear="2022" box="[420,794,434,459]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="jacqi" status="n. sp.">
<emphasis bold="true" box="[420,613,434,458]" italics="true" pageId="2" pageNumber="57">Scaphisoma jacqi</emphasis>
Löbl &amp; Ramage
</taxonomicName>
,
<emphasis bold="true" box="[805,865,434,458]" pageId="2" pageNumber="57">
<taxonomicNameLabel box="[805,865,434,458]" pageId="2" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[144,711,471,490]" pageId="2" pageNumber="57" type="multiple">
<paragraph blockId="2.[144,865,434,490]" box="[144,711,471,490]" pageId="2" pageNumber="57">http://zoobank.org/F092C8CB-A78E-46E8-9CF2-0FEFE201450E</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="57" type="materials_examined">
<paragraph blockId="2.[144,1142,515,604]" pageId="2" pageNumber="57">
<materialsCitation collectingDate="2021-03-01" collectingDateMax="2021-03-08" collectingDateMin="2021-03-01" collectingMethod="Malaise trap" collectionCode="MHNG" collectorName="Frederic Jacq" country="French Polynesia" elevation="57" latitude="-16.847557" location="Faaroa Valley" longLatPrecision="1" longitude="-151.42992" pageId="2" pageNumber="57" specimenCode="MHNG-ENTO 0093802" specimenCount="1" specimenCount-male="1" stateProvince="Society Islands" typeStatus="holotype">
<typeStatus box="[197,309,515,539]" pageId="2" pageNumber="57">HOLOTYPE</typeStatus>
:
<specimenCount box="[334,356,516,539]" pageId="2" pageNumber="57" type="male"></specimenCount>
,
<collectingCountry box="[377,576,515,539]" name="French Polynesia" pageId="2" pageNumber="57" value="French Polynesia">French Polynesia</collectingCountry>
,
<collectingRegion box="[596,771,515,539]" pageId="2" pageNumber="57">Society Islands</collectingRegion>
,
<collectingRegion box="[793,958,515,539]" pageId="2" pageNumber="57">Raiatea Island</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:03B1879EFFCE502059EF2CF1C117FC0B:8EC76053FFCE50265B982F47C42AFD96" box="[979,1138,515,539]" country="French Polynesia" latitude="-16.847557" longLatPrecision="1" longitude="-151.42992" name="Faaroa Valley" pageId="2" pageNumber="57" stateProvince="Society Islands">Faaroa Valley</location>
,
<geoCoordinate box="[144,293,547,571]" degrees="16" direction="south" minutes="50" orientation="latitude" pageId="2" pageNumber="57" precision="1" seconds="51.2" value="-16.847557">16°5051.2S</geoCoordinate>
<geoCoordinate box="[300,474,547,571]" degrees="151" direction="west" minutes="25" orientation="longitude" pageId="2" pageNumber="57" precision="1" seconds="47.7" value="-151.42992">151°2547.7W</geoCoordinate>
,
<quantity box="[484,540,547,571]" metricMagnitude="1" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="57" unit="m" value="57.0">
<elevation box="[484,540,547,571]" metricMagnitude="1" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="57" unit="m" value="57.0">57 m</elevation>
</quantity>
,
<collectingMethod box="[550,685,547,571]" pageId="2" pageNumber="57">Malaise trap</collectingMethod>
,
<date box="[695,826,547,571]" pageId="2" pageNumber="57" value="2021-03-01" valueMax="2021-03-08" valueMin="2021-03-01">
<collectingDate box="[695,826,547,571]" pageId="2" pageNumber="57" value="2021-03-01" valueMax="2021-03-08" valueMin="2021-03-01">1-8.III.2021</collectingDate>
</date>
,
<emphasis box="[836,1031,547,571]" italics="true" pageId="2" pageNumber="57">
leg.
<collectorName box="[882,1031,547,571]" pageId="2" pageNumber="57">Frédéric Jacq</collectorName>
</emphasis>
;
<specimenCode pageId="2" pageNumber="57">MHNG-ENTO 0093802</specimenCode>
(
<collectionCode box="[328,416,580,604]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="2" pageNumber="57">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6461195" ID-Zenodo-Dep="6461195" httpUri="https://zenodo.org/record/6461195/files/figure.png" pageId="2" pageNumber="57" startId="2.[144,175,1668,1687]" targetBox="[282,984,637,1648]" targetPageId="2">
<paragraph blockId="2.[144,1141,1668,1713]" pageId="2" pageNumber="57">
<emphasis bold="true" box="[144,213,1668,1687]" pageId="2" pageNumber="57">Fig. 3-5</emphasis>
.
<taxonomicName authority="Ramage &amp; Löbl, 2022" authorityName="Ramage &amp; Löbl" authorityYear="2022" box="[242,394,1668,1687]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="jacqi" status="n. sp.">
<emphasis box="[242,394,1668,1687]" italics="true" pageId="2" pageNumber="57">Scaphisoma jacqi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[400,448,1669,1688]" pageId="2" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
, aedeagus.
<emphasis bold="true" box="[564,575,1668,1687]" pageId="2" pageNumber="57">3</emphasis>
, Median lobe and parameres in dorsal view, scale = 0.1 mm.
<emphasis bold="true" box="[1125,1136,1668,1687]" pageId="2" pageNumber="57">4</emphasis>
, Aedeagus in lateral view, same scale.
<emphasis bold="true" box="[483,494,1694,1713]" pageId="2" pageNumber="57">5</emphasis>
, Internal sac in dorsal view, scale = 0.05 mm.
</paragraph>
</caption>
<subSubSection lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58" type="description">
<paragraph blockId="3.[144,1142,197,960]" pageId="3" pageNumber="58">
<emphasis bold="true" box="[197,312,197,218]" italics="true" pageId="3" pageNumber="58">Description</emphasis>
. Length
<quantity box="[422,512,197,218]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="3" pageNumber="58" unit="mm" value="1.28">1.28 mm</quantity>
, width
<quantity box="[587,677,197,218]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="3" pageNumber="58" unit="mm" value="0.87">0.87 mm</quantity>
. Head and pronotum blackish (
<figureCitation box="[988,1062,197,219]" captionStart="Fig" captionStartId="1.[269,300,1694,1713]" captionTargetBox="[164,1127,574,1668]" captionTargetId="figure-134@1.[144,1142,574,1685]" captionTargetPageId="1" captionText="Fig. 1-2. Scaphisoma jacqi n. sp. 1, Dorsal view. 2, Lateral view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6461193" httpUri="https://zenodo.org/record/6461193/files/figure.png" pageId="3" pageNumber="58">fig. 1-2</figureCitation>
). Elytra blackish along bases, suture, lateral margins and on transverse area posterior of mid-length and anterior apical sixth. Area anterior of blackish transverse band dark reddish brown, apical sixth yellowish. Venter of thorax and ventrite I blackish, following abdominal segments and appendages light, ochraceous or yellowish.
</paragraph>
<paragraph blockId="3.[144,1142,197,960]" pageId="3" pageNumber="58">
<emphasis box="[197,290,346,367]" italics="true" pageId="3" pageNumber="58">Pronotum</emphasis>
not microsculptured, with lateral margins evenly arcuate, lateral margins carinae visible in dorsal view, lateral striae finely punctate, discal punctation dense and fine, consisting of well delimited punctures much smaller than puncture intervals. Exposed tip of scutellum minute, triangular. Elytra not microsculptured, moderately narrowed apically, with lateral margins nearly evenly rounded, lateral margins striae impunctate near bases, densely punctate posterior basal third of lateral length; apical margins rounded, inner apical angles situated in level with outer apical angles; sutural margin not raised, sutural striae starting at level of scutellar tip, straight, gradually converging apically; adsutural areas flat, combined
<quantity box="[144,231,552,574]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="58" unit="mm" value="0.16">0.16 mm</quantity>
wide at level of scutellar tip, each with three irregular puncture rows in basal third reduced to a single puncture row in apical third. Hind wings fully developed. Hypomeron not microsculptured, smooth, lacking traces of punctation. Mesepimeron nearly four times as long as wide and slightly longer than interval to mesocoxa. Median part of metaventrite slightly convex, lacking impressions. Lateral areas of metaventrite not microsculptured, without antecoxal puncture row, sparsely and very finely punctate. Submesocoxal area about
<quantity box="[477,563,701,723]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="58" unit="mm" value="0.05">0.05 mm</quantity>
, as fourth of interval to metacoxa. Submesocoxal line arcuate, fairly coarsely punctate. Metanepisternum flat, narrowed anteriad, with slightly sinuate suture, not microsculptured, with punctation as that on sides of metaventrite. Tibiae straight. Abdomen with transversely striate microsculpture. Ventrite I as finely and sparsely punctate as lateral parts of metaventrite; submetacoxal area about
<quantity box="[311,400,819,841]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="58" unit="mm" value="0.08">0.08 mm</quantity>
, as two thirds of interval to apical margin; submetacoxal line convex, fairly coarsely punctate.
</paragraph>
<paragraph blockId="3.[144,1142,197,960]" lastBlockId="4.[144,1143,197,902]" lastPageId="4" lastPageNumber="59" pageId="3" pageNumber="58">
<emphasis box="[197,356,879,900]" italics="true" pageId="3" pageNumber="58">Male characters</emphasis>
. Protarsomeres I to III distinctly widened, narrower than apices of tibiae. Mesotarsomeres not widened.Apical process of ventrite VI triangular, acute,
<quantity box="[818,902,908,930]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="58" unit="mm" value="0.02">0.02 mm</quantity>
long.Aedeagus (
<figureCitation box="[1064,1134,908,930]" captionStart="Fig" captionStartId="2.[144,175,1668,1687]" captionTargetBox="[282,984,637,1648]" captionTargetId="figure-150@2.[265,1021,629,1659]" captionTargetPageId="2" captionText="Fig. 3-5. Scaphisoma jacqi n. sp., aedeagus. 3, Median lobe and parameres in dorsal view, scale = 0.1 mm. 4, Aedeagus in lateral view, same scale. 5, Internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6461195" httpUri="https://zenodo.org/record/6461195/files/figure.png" pageId="3" pageNumber="58">fig. 3-5</figureCitation>
)
<quantity box="[144,229,938,959]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="58" unit="mm" value="0.49">0.49 mm</quantity>
long. Basal bulb large, about twice as long as apical process, with robust, ventrally prominent articular process. Ventral branch of apical process sinuate, in lateral view, lacking denticles. Parameres sinuate in lateral view, arcuate and narrowed posterior of mid-length in dorsal view. Internal sac with two apical bunches of long, straight denticles, single large, straight mesal denticle and two basal vesicles containing dense, curved denticles appearing in dorsal view as irregularly striate structures.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6461199" ID-Zenodo-Dep="6461199" box="[427,854,1694,1713]" httpUri="https://zenodo.org/record/6461199/files/figure.png" pageId="3" pageNumber="58" startId="3.[427,458,1694,1713]" targetBox="[144,1142,1001,1686]" targetPageId="3">
<paragraph blockId="3.[427,854,1694,1713]" box="[427,854,1694,1713]" pageId="3" pageNumber="58">
<emphasis bold="true" box="[427,479,1694,1713]" pageId="3" pageNumber="58">Fig. 6</emphasis>
. The site in Faaroa Valley, Raiatea Island.
</paragraph>
</caption>
<subSubSection pageId="4" pageNumber="59" type="etymology">
<paragraph blockId="4.[144,1143,197,902]" pageId="4" pageNumber="59">
<emphasis bold="true" box="[197,298,327,349]" italics="true" pageId="4" pageNumber="59">Etymology</emphasis>
. The specific epithet refers to the family name of the collector of the species, Frédéric Jacq, a great naturalist and photographer, and for his huge contribution to the knowledge of the entomofauna of French Polynesia. The species epithet is to be treated as a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="59" type="biology_ecology">
<paragraph blockId="4.[144,1143,197,902]" pageId="4" pageNumber="59">
<emphasis bold="true" box="[197,280,428,452]" italics="true" pageId="4" pageNumber="59">Habitat</emphasis>
. The Malaise trap (
<figureCitation box="[519,600,429,453]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[427,458,1694,1713]" captionStartId-1="4.[408,438,1694,1713]" captionTargetBox-0="[144,1142,1001,1686]" captionTargetBox-1="[145,1141,945,1682]" captionTargetId-0="figure-369@3.[144,1142,1001,1686]" captionTargetId-1="figure-307@4.[144,1142,943,1686]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 6. The site in Faaroa Valley, Raiatea Island." captionText-1="Fig. 7. Maps of French Polynesia and Raiatea Island." figureDoi-0="http://doi.org/10.5281/zenodo.6461199" figureDoi-1="http://doi.org/10.5281/zenodo.6461201" httpUri-0="https://zenodo.org/record/6461199/files/figure.png" httpUri-1="https://zenodo.org/record/6461201/files/figure.png" pageId="4" pageNumber="59">fig. 6-7</figureCitation>
) was set along an abandoned trail, in a
<taxonomicName authority="(L.) Fryxell" authorityName="Fryxell" baseAuthorityName="L." class="Magnoliopsida" family="Malvaceae" genus="Talipariti" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="tiliaceum">
<emphasis italics="true" pageId="4" pageNumber="59">Talipariti tiliaceum</emphasis>
(L.) Fryxell
</taxonomicName>
secondarized forest, degraded with
<taxonomicName authority="(L.) Skeels" authorityName="Skeels" baseAuthorityName="L." box="[752,1054,463,487]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="cumini">
<emphasis box="[752,929,463,487]" italics="true" pageId="4" pageNumber="59">Syzygium cumini</emphasis>
(L.) Skeels
</taxonomicName>
,
<taxonomicName authority="Thunb." authorityName="Thunb." class="Magnoliopsida" family="Primulaceae" genus="Ardisia" kingdom="Plantae" order="Ericales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="elliptica">
<emphasis italics="true" pageId="4" pageNumber="59">Ardisia elliptica</emphasis>
Thunb.
</taxonomicName>
, and
<taxonomicName authority="L." authorityName="L." box="[382,635,497,521]" class="Magnoliopsida" family="Passifloraceae" genus="Passiflora" kingdom="Plantae" order="Malpighiales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="laurifolia">
<emphasis box="[382,601,497,521]" italics="true" pageId="4" pageNumber="59">Passiflora laurifolia</emphasis>
L.
</taxonomicName>
The herbaceous understory was composed of
<taxonomicName authority="(L.) Pruski" authorityName="Pruski" baseAuthorityName="L." box="[144,531,531,555]" class="Magnoliopsida" family="Asteraceae" genus="Sphagneticola" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="trilobata">
<emphasis box="[144,399,531,555]" italics="true" pageId="4" pageNumber="59">Sphagneticola trilobata</emphasis>
(L.) Pruski
</taxonomicName>
,
<taxonomicName box="[545,691,531,555]" class="Polypodiopsida" family="Thelypteridaceae" genus="Christella" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis box="[545,686,531,555]" italics="true" pageId="4" pageNumber="59">Christella sp</emphasis>
.
</taxonomicName>
,
<taxonomicName authority="(L.) Roscoe ex Sm." authorityName="Sm." baseAuthorityName="L." box="[709,1137,531,555]" class="Liliopsida" family="Zingiberaceae" genus="Zingiber" kingdom="Plantae" order="Zingiberales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="zerumbet">
<emphasis box="[709,908,531,555]" italics="true" pageId="4" pageNumber="59">Zingiber zerumbet</emphasis>
(L.) Roscoe ex Sm.
</taxonomicName>
,
<taxonomicName authority="(G. Forst.) C. Presl" authorityName="C. Presl" baseAuthorityName="G. Forst." box="[144,572,565,589]" class="Polypodiopsida" family="Nephrolepidaceae" genus="Nephrolepis" kingdom="Plantae" order="Polypodiales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="hirsutula">
<emphasis box="[144,376,565,589]" italics="true" pageId="4" pageNumber="59">Nephrolepis hirsutula</emphasis>
(G.Forst.) C.Presl
</taxonomicName>
and
<taxonomicName authority="(L.) Desv." authorityName="Desv." baseAuthorityName="L." box="[624,965,565,589]" class="Liliopsida" family="Poaceae" genus="Centotheca" kingdom="Plantae" order="Poales" pageId="4" pageNumber="59" phylum="Tracheophyta" rank="species" species="lappacea">
<emphasis box="[624,850,565,589]" italics="true" pageId="4" pageNumber="59">Centotheca lappacea</emphasis>
(L.) Desv.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="59" type="discussion">
<paragraph blockId="4.[144,1143,197,902]" pageId="4" pageNumber="59">
<emphasis bold="true" box="[197,314,606,630]" italics="true" pageId="4" pageNumber="59">Comments</emphasis>
. The species is a member of the
<taxonomicName authorityName="Reitter" authorityYear="1877" box="[676,982,606,630]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis box="[676,982,606,630]" italics="true" pageId="4" pageNumber="59">Scaphisoma haemorrhoidale</emphasis>
</taxonomicName>
group, defined by the aedeagi symmetrical with a trifid apical process, narrow, simple parameres usually lacking lobes and a complex internal sac lacking flagellum (
<bibRefCitation author="LOBL I." box="[727,858,675,701]" journalOrPublisher="Revue suisse de zoologie" pageId="4" pageNumber="59" pagination="727 - 799" part="77" refId="ref2569" refString="LOBL I., 1970. - Revision der palaarktischen Arten der Gattungen Scaphisoma Leach und Caryoscapha Ganglbauer der Tribus Scaphisomini (Col. Scaphidiidae). Revue suisse de zoologie, 77: 727 - 799. https: // doi. org / 10.5962 / bhl. part. 75925" title="Revision der palaarktischen Arten der Gattungen Scaphisoma Leach und Caryoscapha Ganglbauer der Tribus Scaphisomini (Col. Scaphidiidae)" type="journal article" year="1970">LÖBL, 1970</bibRefCitation>
). The aedeagal characters suggest relationships with
<taxonomicName authority="Scott, 1908" authorityName="Scott" authorityYear="1908" box="[437,689,708,733]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="perkinsi">
<emphasis box="[437,552,708,732]" italics="true" pageId="4" pageNumber="59">S. perkinsi</emphasis>
Scott, 1908
</taxonomicName>
, described from Hawaii, redescribed and subsequently recorded from the Bonin (Ogasawara) Islands (
<bibRefCitation author="LOBL I." box="[810,940,743,769]" journalOrPublisher="Insects of Micronesia" pageId="4" pageNumber="59" pagination="69 - 80" part="15" refId="ref2621" refString="LOBL I., 1981. - Insects of Micronesia Coleoptera Scaphidiidae. Insects of Micronesia, 15: 69 - 80." title="Insects of Micronesia Coleoptera Scaphidiidae" type="journal article" year="1981">LÖBL, 1981</bibRefCitation>
). The new species may be readily distinguished from
<taxonomicName authorityName="Scott" authorityYear="1908" box="[526,640,776,800]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="perkinsi">
<emphasis box="[526,640,776,800]" italics="true" pageId="4" pageNumber="59">S. perkinsi</emphasis>
</taxonomicName>
by the pattern of elytral colour (the latter has elytra uniformly black between bases and the light apical area) and by the internal sac of the aedeagus bearing an elongate mesal denticle absent from
<taxonomicName authorityName="Scott" authorityYear="1908" box="[761,874,844,868]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="perkinsi">
<emphasis box="[761,874,844,868]" italics="true" pageId="4" pageNumber="59">S. perkinsi</emphasis>
</taxonomicName>
, and lacking an elongate V-shaped sclerite.
</paragraph>
</subSubSection>
</treatment>
</document>