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<document id="B33F4598B171AB45CF51C333D3D8E43B" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFF29D124BA5077CDDFB2534" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retrorsioides bammerti Shear &amp; Marek 2021, new species" docType="treatment" docVersion="7" lastPageNumber="108" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="108" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F4C20C6F76EB837CB40FD4BD4CDE38BE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EC10C4B8B984F4E158AF000A1BC235D5">
<mods:title id="D74AE1EDB5A164811C22E05A96D66ECC">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
</mods:titleInfo>
<mods:name id="B0B4164BA173A6436F7EE29F24E178DD" type="personal">
<mods:role id="ABF7017D60D26D978094CA0F6D97DC70">
<mods:roleTerm id="B86DD7601AB3794E89850BFDB4BA9761">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DEF72AA1C9C0276A82F2E4F0BD54C2AF">Shear, William A.</mods:namePart>
<mods:affiliation id="BE536283AB2C7ED96271A052EF9DF8C6">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
</mods:name>
<mods:name id="004B493FF825AD88DA31965C59B5CDD9" type="personal">
<mods:role id="E13C8ED0C76CB15081FD7617683FA7F6">
<mods:roleTerm id="CAC5121D4E0362976CF98BA9E17C88D9">Author</mods:roleTerm>
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<mods:namePart id="91AA40D5F6F95FB072DAD53CC28AE793">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="1E31E46FF3BD6856DE68E3281BC38C92" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="257C411BF77E90C7C68DB249EE847EB0">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="21BD819BC98F545F1249E9945BC77436" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:title id="0A81FE67939266F1D974E57A23B30CDC">Zootaxa</mods:title>
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<mods:part id="7076A3E989124D6A581FB1333D875833">
<mods:date id="EF3C8B522B0BDD87CA0776C8B13ED146">2021</mods:date>
<mods:detail id="2AA5271481C43E81EF8D118E1880BB4B" type="pubDate">
<mods:number id="B14F181F115EFD63D4380842957DFC3D">2021-05-24</mods:number>
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<mods:detail id="3ADFBCB66E545C0BEFCFC65BB3D33B74" type="volume">
<mods:number id="5DA482104C2B1B9B9154B7C2ADD76B70">4975</mods:number>
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<mods:start id="123630D8E296780F965AE049C829B5AF">81</mods:start>
<mods:end id="73425B268232BFB7DE0BE18B0F672D7D">126</mods:end>
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<mods:classification id="8569A8EF8BE0042EBA5261ECC4C37BED">journal article</mods:classification>
<mods:identifier id="1415892F1D5D004D84D8806627B621CC" type="CLB-Dataset">6194</mods:identifier>
<mods:identifier id="ADFF2D87EF7725FA1578994BFF3173C5" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
<mods:identifier id="651BA7CF8EC479198EE321E8D3565319" type="GBIF-Dataset">529d48db-4299-4e2d-9aaf-6c4178c08b18</mods:identifier>
<mods:identifier id="F3AD5134EED27ADB18F6069FAC08C1AD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2470013BCDEECA86FD71F3C6D21AF5B3" type="Zenodo-Dep">4804909</mods:identifier>
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<treatment id="03B18797FFF29D124BA5077CDDFB2534" ID-DOI="http://doi.org/10.5281/zenodo.4923603" ID-GBIF-Taxon="182644898" ID-Zenodo-Dep="4923603" LSID="urn:lsid:plazi:treatment:03B18797FFF29D124BA5077CDDFB2534" httpUri="http://treatment.plazi.org/id/03B18797FFF29D124BA5077CDDFB2534" lastPageNumber="108" pageId="27" pageNumber="108">
<subSubSection id="C302650AFFF29D124BA5077CDA5F222D" box="[151,796,582,609]" pageId="27" pageNumber="108" type="nomenclature">
<paragraph id="8BA73681FFF29D124BA5077CDA5F222D" blockId="27.[151,796,582,645]" box="[151,796,582,609]" pageId="27" pageNumber="108">
<heading id="D0EF81EDFFF29D124BA5077CDA5F222D" bold="true" box="[151,796,582,609]" fontSize="11" level="1" pageId="27" pageNumber="108" reason="1">
<emphasis id="B96CEA93FFF29D124BA5077CDA5F222D" bold="true" box="[151,796,582,609]" pageId="27" pageNumber="108">
<taxonomicName id="4C184D02FFF29D124BA5077CDB3C222C" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,639,582,609]" class="Diplopoda" family="Polydesmidae" genus="Retrorsioides" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="108" phylum="Arthropoda" rank="species" species="bammerti" status="sp. nov.">
<emphasis id="B96CEA93FFF29D124BA5077CD8F1222C" bold="true" box="[151,434,582,609]" italics="true" pageId="27" pageNumber="108">Retrorsioides bammerti</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFF29D1249BE077DDA5F222D" box="[652,796,582,609]" pageId="27" pageNumber="108" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BA50751DCC722BC" pageId="27" pageNumber="108" type="description">
<paragraph id="8BA73681FFF29D124BA50751D80F22C8" blockId="27.[151,796,582,645]" box="[151,332,619,645]" pageId="27" pageNumber="108">
<figureCitation id="13232A04FFF29D124BA50751D85422C8" box="[151,279,619,645]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Figs 76–78</figureCitation>
,
<figureCitation id="13232A04FFF29D124A100751D80F22C8" box="[290,332,619,645]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">130</figureCitation>
</paragraph>
<paragraph id="8BA73681FFF29D124BA50789DCC722BC" blockId="27.[151,1437,691,1401]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BA50789D9A72280" bold="true" box="[151,228,691,717]" pageId="27" pageNumber="108">Types.</emphasis>
<materialsCitation id="3B703CDCFFF29D124BDE0789DCC222BC" ID-GBIF-Occurrence="3128519331" collectingDate="2004-11-14" collectionCode="CAS" collectorName="W. Leonard. All" county="Thurston Co." elevation="87" location="Bob Bammert Grove" municipality="Male" pageId="27" pageNumber="108" specimenCount="2" stateProvince="Washington" typeStatus="holotype">
<collectingMunicipality id="6BC3ACFBFFF29D124BDE0789D8652280" box="[236,294,691,717]" pageId="27" pageNumber="108">Male</collectingMunicipality>
<typeStatus id="54A38823FFF29D124A1F0789D8CC2280" box="[301,399,691,717]" pageId="27" pageNumber="108" type="holotype">holotype</typeStatus>
and male
<typeStatus id="54A38823FFF29D1249340789DB322280" box="[518,625,691,717]" pageId="27" pageNumber="108" type="paratype">paratypes</typeStatus>
from
<location id="8EC7605AFFF29D12498A0789DAEB2280" LSID="urn:lsid:plazi:treatment:03B18797FFF29D124BA5077CDDFB2534:8EC7605AFFF29D12498A0789DAEB2280" box="[696,936,691,717]" county="Thurston Co." municipality="Male" name="Bob Bammert Grove" pageId="27" pageNumber="108" stateProvince="Washington">Bob Bammert Grove</location>
,
<location id="8EC7605AFFF29D1248860789DDDB2280" LSID="urn:lsid:plazi:treatment:03B18797FFF29D124BA5077CDDFB2534:8EC7605AFFF29D1248860789DDDB2280" box="[948,1176,691,717]" county="Thurston Co." municipality="Male" name="Capitol State Forest" pageId="27" pageNumber="108" stateProvince="Washington">Capitol State Forest</location>
,
<elevation id="0035D1B2FFF29D124F960789DC522280" box="[1188,1297,691,717]" metricMagnitude="1" metricUnit="m" metricValue="8.74776" pageId="27" pageNumber="108" unit="ft" value="287.0">
<quantity id="4CE09B64FFF29D124F960789DDA42280" box="[1188,1255,691,717]" metricMagnitude="1" metricUnit="m" metricValue="8.74776" pageId="27" pageNumber="108" unit="ft" value="287.0">287 ft</quantity>
asl
</elevation>
, 46.8960°, - 123.0963°,
<collectingCounty id="62C64E0DFFF29D124A2B07EDD8EC22BC" box="[281,431,727,753]" pageId="27" pageNumber="108">Thurston Co.</collectingCounty>
,
<collectingRegion id="49DCF863FFF29D124A8807EDDB0122BC" box="[442,578,727,753]" country="United States of America" name="Washington" pageId="27" pageNumber="108">Washington</collectingRegion>
, collected
<date id="FFA61041FFF29D12498B07EDDAD122BC" box="[697,914,727,753]" pageId="27" pageNumber="108" value="2004-11-14">
<collectingDate id="EFE2E9A9FFF29D12498B07EDDAD122BC" box="[697,914,727,753]" pageId="27" pageNumber="108" value="2004-11-14">14 November 2004</collectingDate>
</date>
by
<collectorName id="26ED5357FFF29D12488907EDDD2C22BC" box="[955,1135,727,753]" pageId="27" pageNumber="108">W. Leonard. All</collectorName>
types deposited in
<collectionCode id="ED09AE44FFF29D124E7507EDDCC222BC" box="[1351,1409,727,753]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="27" pageNumber="108" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BF507C0DA6F2374" pageId="27" pageNumber="108" type="diagnosis">
<paragraph id="8BA73681FFF29D124BF507C0DA6F2374" blockId="27.[151,1437,691,1401]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF507C0D8012359" bold="true" box="[199,322,762,788]" pageId="27" pageNumber="108">Diagnosis.</emphasis>
The sigmoid terminal zone of the gonopod (
<emphasis id="B96CEA93FFF29D12480807C1DA142358" bold="true" box="[826,855,763,789]" pageId="27" pageNumber="108">tz,</emphasis>
<figureCitation id="13232A04FFF29D12486D07C1DAFD2358" box="[863,958,763,789]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) and the basal position of the pulvillus (
<emphasis id="B96CEA93FFF29D124EB407C1DCDF2358" bold="true" box="[1414,1436,763,789]" pageId="27" pageNumber="108">p,</emphasis>
<figureCitation id="13232A04FFF29D124BA50625D9B52374" box="[151,246,799,825]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) set this species apart from all others in the genus.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BF50679DA8723CC" pageId="27" pageNumber="108" type="etymology">
<paragraph id="8BA73681FFF29D124BF50679DA8723CC" blockId="27.[151,1437,691,1401]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF50679D80D2310" bold="true" box="[199,334,835,861]" pageId="27" pageNumber="108">Etymology.</emphasis>
The species name recognizes Robert (Bob) Bammert (1926–2019), noted
<collectingRegion id="49DCF863FFF29D124F900679DC642310" box="[1186,1319,835,861]" country="United States of America" name="Washington" pageId="27" pageNumber="108">Washington</collectingRegion>
conservationist for whom the
<typeStatus id="54A38823FFF29D124AB0065DD8F123CC" box="[386,434,871,897]" pageId="27" pageNumber="108">type</typeStatus>
locality, a grove of old-growth trees, is named.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BF506B0DD6C2488" pageId="27" pageNumber="108" type="description">
<paragraph id="8BA73681FFF29D124BF506B0DD6C2488" blockId="27.[151,1437,691,1401]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF506B0D81A23E9" bold="true" box="[199,345,906,932]" pageId="27" pageNumber="108">Description.</emphasis>
Male
<typeStatus id="54A38823FFF29D124A9106B1DB4A23E8" box="[419,521,907,933]" pageId="27" pageNumber="108" type="holotype">holotype</typeStatus>
. Length about 4.0 mm, greatest width
<quantity id="4CE09B64FFF29D1248F306B0DD6923E8" box="[961,1066,906,933]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="27" pageNumber="108" unit="mm" value="0.5">0.50 mm</quantity>
. Head densely setose, cuticle alveolate (
<figureCitation id="13232A04FFF29D124BC50695D80F2387" box="[247,332,943,970]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Fig. 76</figureCitation>
). Collum (
<figureCitation id="13232A04FFF29D124AF10695DB572387" box="[451,532,943,970]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Fig. 76</figureCitation>
) with anterior marginal row of 14 setae. Anterior metazonites with three rows of setae (
<figureCitation id="13232A04FFF29D124BED06E9D87723A3" box="[223,308,979,1006]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Fig. 76</figureCitation>
), transitioning to four rows (
<figureCitation id="13232A04FFF29D12494B06E9DB8F23A0" box="[633,716,979,1005]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Fig. 77</figureCitation>
) about ring 5, posteriormost segments with five rows (
<figureCitation id="13232A04FFF29D124E0806E9DCCC23A0" box="[1338,1423,979,1005]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Fig. 78</figureCitation>
); setal tubercles becoming almost obsolete on midbody to posterior rings. Alveolate cuticle absent from metazonites posterior to collum. Epiproct (
<figureCitation id="13232A04FFF29D1249100121DB352478" box="[546,630,1051,1077]" captionStart="FIGURES 73–78" captionStartId="28.[151,264,1813,1837]" captionTargetBox="[151,1436,182,1786]" captionTargetId="figure-24@28.[151,1436,181,1789]" captionTargetPageId="28" captionText="FIGURES 73–78. Species of Retrorsioides, n.gen. 73–74. Retrorsioides kittitas, n. sp. 73. Gonopods in situ, posteriolateral view. 74. Same, posterior view. 75. Gonopod tip, lateral view. SEM stub WS32-17. 76–78. Retrorsioides bammerti, n. sp. 76. Head and anterior rings, dorsal view. 77. Midbody ring, dorsal view. 78. Pygidium, ventral view. SEM stub WS33-1. Abbreviations: ac, acropodite; adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process." figureDoi="http://doi.org/10.5281/zenodo.4805007" httpUri="https://zenodo.org/record/4805007/files/figure.png" pageId="27" pageNumber="108">Fig. 78</figureCitation>
) swollen, short, bent ventrally, without alveolate cuticle. Anterior legs crassate, tarsi with sphaerotrichomes. Gonopod with oblong, inflated prefemorite (
<emphasis id="B96CEA93FFF29D124F070104DD162415" bold="true" box="[1077,1109,1086,1112]" pageId="27" pageNumber="108">pf,</emphasis>
<figureCitation id="13232A04FFF29D124F6E0105DDF82414" box="[1116,1211,1087,1113]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) similar to previous species. Acropodite short, stout. Pulvillus (
<emphasis id="B96CEA93FFF29D12494A0159DBCD2430" bold="true" box="[632,654,1123,1149]" pageId="27" pageNumber="108">p,</emphasis>
<figureCitation id="13232A04FFF29D1249A70159DBB72430" box="[661,756,1123,1149]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) in basal third of acropodite, pulvillar process (
<emphasis id="B96CEA93FFF29D124E3A0159DC6C2430" bold="true" box="[1288,1327,1123,1149]" pageId="27" pageNumber="108">pp,</emphasis>
<figureCitation id="13232A04FFF29D124E040159DCD72430" box="[1334,1428,1123,1149]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) long, acute, directed distally. Anteriorly directed process (
<emphasis id="B96CEA93FFF29D12481401BDDA1824EC" bold="true" box="[806,859,1159,1185]" pageId="27" pageNumber="108">adp,</emphasis>
<figureCitation id="13232A04FFF29D12485001BDDA8224EC" box="[866,961,1159,1185]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) rounded, tapering slightly, without teeth. Terminal zone (
<emphasis id="B96CEA93FFF29D124A7A0191D8262488" bold="true" box="[328,357,1195,1221]" pageId="27" pageNumber="108">tz,</emphasis>
<figureCitation id="13232A04FFF29D124A5E0191D8892488" box="[364,458,1195,1221]" captionStart="FIGURES 127–132" captionStartId="41.[151,264,1305,1329]" captionTargetBox="[163,1429,185,1275]" captionTargetId="figure-17@41.[151,1436,181,1281]" captionTargetPageId="41" captionText="FIGURES 127–132. Drawings of gonopods of polydesmid species. 127. Tip of gonopod of Retrorsioides castellum, n. sp., mesal view. 128. Right gonopod of Retrorsioides linnensis, n. sp., lateral view. 129. Left gonopod of Retrorsioides kittitas, n. sp., lateral view. 130. Right gonopod of Retrorsioides bannerti, n. sp., lateral view. 131. Right gonopod of Rowlandesmus millicoma, n. sp., mesal view. 132. Right gonopod of Rowlandesmus dentogonopus, n. sp., lateral view. Abbreviations: adp, anteriorly directed process; p, pulvillus; pf, prefemorite; pp, pulvillar process; tz, terminal zone." figureDoi="http://doi.org/10.5281/zenodo.4805077" httpUri="https://zenodo.org/record/4805077/files/figure.png" pageId="27" pageNumber="108">Fig. 130</figureCitation>
) not bifurcate, sigmoidally curved. Females unknown.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFF29D124BF501F4DDFB2534" pageId="27" pageNumber="108" type="distribution">
<paragraph id="8BA73681FFF29D124BF501F4DD1E24A4" blockId="27.[151,1437,691,1401]" box="[199,1117,1230,1257]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF501F4D82224A5" bold="true" box="[199,353,1230,1256]" pageId="27" pageNumber="108">Distribution.</emphasis>
Northwestern
<collectingRegion id="49DCF863FFF29D12493401F5DBC824A4" box="[518,651,1231,1257]" country="United States of America" name="Washington" pageId="27" pageNumber="108">Washington</collectingRegion>
in Thurston, Pacific and Lewis Counties.
</paragraph>
<paragraph id="8BA73681FFF29D124BF501C9DAED2518" blockId="27.[151,1437,691,1401]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF501C9D8712540" bold="true" box="[199,306,1267,1293]" pageId="27" pageNumber="108">Records:</emphasis>
<collectingRegion id="49DCF863FFF29D124A0B01C9D8BB2540" box="[313,504,1267,1293]" country="United States of America" name="Washington" pageId="27" pageNumber="108">WASHINGTON</collectingRegion>
:
<emphasis id="B96CEA93FFF29D12493001C9DB322540" box="[514,625,1267,1293]" italics="true" pageId="27" pageNumber="108">Lewis Co.</emphasis>
: Stillman Basin,
<quantity id="4CE09B64FFF29D12480001C9DA392540" box="[818,890,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.8968192" pageId="27" pageNumber="108" unit="mi" value="1.8">1.8 mi</quantity>
on Weyerhauser 4200 from road W4000, 2116 ft asl, 48.4956°, -123.2134°,
<date id="FFA61041FFF29D124AF3002DDBCA257C" box="[449,649,1303,1329]" pageId="27" pageNumber="108" value="2004-12-04">4 December 2004</date>
, W. Leonard, C. Richart, mm.
<emphasis id="B96CEA93FFF29D1248EB002DDD15257C" box="[985,1110,1303,1329]" italics="true" pageId="27" pageNumber="108">Pacific Co.</emphasis>
:
<quantity id="4CE09B64FFF29D124F56002DDDE8257C" box="[1124,1195,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.7702784" pageId="27" pageNumber="108" unit="mi" value="1.1">1.1 mi</quantity>
south of SR6 on Trap Creek Road, B-line, 46.5432°, -123/6152°, W. Leonard, C. Richart, m.
</paragraph>
<paragraph id="8BA73681FFF29D124BF50065DDFB2534" blockId="27.[151,1437,691,1401]" box="[199,1208,1375,1401]" pageId="27" pageNumber="108">
<emphasis id="B96CEA93FFF29D124BF50065D84B2534" bold="true" box="[199,264,1375,1401]" pageId="27" pageNumber="108">Note:</emphasis>
At the Stillman Basin locality, this species is syntopic with
<emphasis id="B96CEA93FFF29D12489A0065DDFB2534" box="[936,1208,1375,1401]" italics="true" pageId="27" pageNumber="108">
<taxonomicName id="4C184D02FFF29D12489A0065DDF72534" authorityName="Shear &amp; Marek" authorityYear="2021" box="[936,1204,1375,1401]" class="Diplopoda" family="Polydesmidae" genus="Retrorsia" kingdom="Animalia" order="Polydesmida" pageId="27" pageNumber="108" phylum="Arthropoda" rank="species" species="simplicissima">Retrorsia simplicissima</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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