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<document id="E8146459CB6B474D0F350D3895172A43" ID-CLB-Dataset="6194" ID-DOI="10.11646/zootaxa.4975.1.3" ID-GBIF-Dataset="529d48db-4299-4e2d-9aaf-6c4178c08b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="4804909" ID-ZooBank="DB7C9028-3EDF-454F-88D0-336624AD1DC4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622016306479" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2021" docId="03B18797FFE09D034BA50229DBB2219D" docLanguage="en" docName="zootaxa.4975.1.3.pdf" docOrigin="Zootaxa 4975 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bidentogon buttensis Shear &amp; Marek 2021, new species" docType="treatment" docVersion="7" lastPageNumber="91" masterDocId="FF88FFEFFFE99D094B32053AD943204D" masterDocTitle="Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)" masterLastPageNumber="126" masterPageNumber="81" pageNumber="90" updateTime="1698991024886" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E867648F57C61C8BE30D6044918FFCB9">Three new genera and eighteen new species of miniature polydesmid millipedes from the northwestern United States (Diplopoda, Polydesmida, Polydesmidae)</mods:title>
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<mods:name id="CF155293618D9C5BEAD5C92F85A6A48E" type="personal">
<mods:role id="C10CB63E76D8487DFEEF3741D7C0B908">
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<mods:namePart id="63D60A6704D1E9A5973D506C40DE38BD">Shear, William A.</mods:namePart>
<mods:affiliation id="2688D376D41F8747B1BDC9130F6991E3">Professor Emeritus, Department of Biology, Hampden-Sydney College, Hampden-Sydney VA 23943; present address 1950 Price Drive, Farmville VA 23901.</mods:affiliation>
</mods:name>
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<mods:role id="AED67B72F06C0DFDDC62999C70C23DD3">
<mods:roleTerm id="66D007DEE090D163D083C5C0FA40E414">Author</mods:roleTerm>
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<mods:namePart id="8AA8294390D758D6758782E46EFB70D3">Marek, Paul E.</mods:namePart>
<mods:nameIdentifier id="89C0E1D9DA47DB6B52E47A97D94B506F" type="ORCID">0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation id="1878E294BC8B0565D7F3A3087A06D338">Associate Professor, Department of Entomology, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061. pmarek @ vt. edu; https: // orcid. org / 0000 - 0002 - 7048 - 2514</mods:affiliation>
<mods:nameIdentifier id="D68CCFD0289EB342996039812A61A589" type="email">pmarek@vt.edu</mods:nameIdentifier>
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<mods:title id="9FB55EBF4AB4EE4E6040972AD9D1BDF1">Zootaxa</mods:title>
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<mods:date id="653946D1A361414A66D4CA52E63967D3">2021</mods:date>
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<mods:number id="16B9B65847021D78D262AE0F9AFBF7A3">2021-05-24</mods:number>
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<mods:number id="61C72478E8726EE0A96FE205C7EBD479">4975</mods:number>
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<mods:identifier id="3B77408786A34C65CF462E93BB0EC937" type="DOI">10.11646/zootaxa.4975.1.3</mods:identifier>
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<treatment id="03B18797FFE09D034BA50229DBB2219D" ID-DOI="http://doi.org/10.5281/zenodo.4804974" ID-GBIF-Taxon="182644914" ID-Zenodo-Dep="4804974" LSID="urn:lsid:plazi:treatment:03B18797FFE09D034BA50229DBB2219D" httpUri="http://treatment.plazi.org/id/03B18797FFE09D034BA50229DBB2219D" lastPageId="10" lastPageNumber="91" pageId="9" pageNumber="90">
<subSubSection id="C302650AFFE09D004BA50229DA432760" box="[151,768,1811,1838]" pageId="9" pageNumber="90" type="nomenclature">
<paragraph id="8BA73681FFE09D004BA50229DA432760" blockId="9.[151,768,1811,1873]" box="[151,768,1811,1838]" pageId="9" pageNumber="90">
<heading id="D0EF81EDFFE09D004BA50229DA432760" bold="true" box="[151,768,1811,1838]" fontSize="11" level="1" pageId="9" pageNumber="90" reason="1">
<emphasis id="B96CEA93FFE09D004BA50229DA432760" bold="true" box="[151,768,1811,1838]" pageId="9" pageNumber="90">
<taxonomicName id="4C184D02FFE09D004BA50229DB212760" authority="Shear &amp; Marek" authorityName="Shear &amp; Marek" authorityYear="2021" box="[151,610,1811,1838]" class="Diplopoda" family="Polydesmidae" genus="Bidentogon" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="90" phylum="Arthropoda" rank="species" species="buttensis" status="sp. nov.">
<emphasis id="B96CEA93FFE09D004BA50229D8D42763" bold="true" box="[151,407,1811,1838]" italics="true" pageId="9" pageNumber="90">Bidentogon buttensis</emphasis>
Shear &amp; Marek
</taxonomicName>
,
<taxonomicNameLabel id="A25F57E8FFE09D0049420229DA432760" box="[624,768,1811,1837]" pageId="9" pageNumber="90" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE09D004BA5020DD856271C" box="[151,277,1847,1873]" pageId="9" pageNumber="90" type="description">
<paragraph id="8BA73681FFE09D004BA5020DD856271C" blockId="9.[151,768,1811,1873]" box="[151,277,1847,1873]" pageId="9" pageNumber="90">
<figureCitation id="13232A04FFE09D004BA5020DD856271C" box="[151,277,1847,1873]" captionStart="FIGURES 1316" captionStartId="10.[151,264,1502,1526]" captionTargetBox="[151,1436,488,1476]" captionTargetId="figure-154@10.[151,1436,488,1478]" captionTargetPageId="10" captionText="FIGURES 1316. Bidentogon buttensis, n. sp. 13. Head and anterior rings, dorsal view. 14. Midbody ring, dorsal view. 15. Gonopods in situ, ventral view. 16. Gonopod tips, ventral view. Abbreviations: ac, acropodite; pf, prefemorite; sm, solenomere; tz, terminal zone. SEM stub WS33-9." figureDoi="http://doi.org/10.5281/zenodo.4804923" httpUri="https://zenodo.org/record/4804923/files/figure.png" pageId="9" pageNumber="90">Figs 1316</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE09D004BA50245DBA827AC" pageId="9" pageNumber="90" type="materials_examined">
<paragraph id="8BA73681FFE09D004BA50245DBA827AC" blockId="9.[151,1436,1919,2018]" pageId="9" pageNumber="90">
<emphasis id="B96CEA93FFE09D004BA50245D99A27D4" bold="true" box="[151,217,1919,1945]" pageId="9" pageNumber="90">Type.</emphasis>
<materialsCitation id="3B703CDCFFE09D004BED0245DBA427AC" ID-GBIF-Occurrence="3128519334" collectingDate="1956-02-09" collectionCode="R" collectorName="Male" county="Butte Co." location="Oroville" municipality="Stringtown Hill" pageId="9" pageNumber="90" specimenCount="1" stateProvince="California" typeStatus="holotype">
<collectorName id="26ED5357FFE09D004BED0245D85A27D4" box="[223,281,1919,1945]" pageId="9" pageNumber="90">Male</collectorName>
<typeStatus id="54A38823FFE09D004A2C0245D8C327D4" box="[286,384,1919,1945]" pageId="9" pageNumber="90" type="holotype">holotype</typeStatus>
from near
<collectingMunicipality id="6BC3ACFBFFE09D004ACA0245DBEB27D4" box="[504,680,1919,1945]" pageId="9" pageNumber="90">Stringtown Hill</collectingMunicipality>
, NE of
<location id="8EC7605AFFE09D0049C80245DA1927D4" LSID="urn:lsid:plazi:treatment:03B18797FFE09D034BA50229DBB2219D:8EC7605AFFE09D0049C80245DA1927D4" box="[762,858,1919,1945]" county="Butte Co." municipality="Stringtown Hill" name="Oroville" pageId="9" pageNumber="90" stateProvince="California">Oroville</location>
,
<collectingCounty id="62C64E0DFFE09D00485602BADA9227D4" box="[868,977,1919,1946]" pageId="9" pageNumber="90">Butte Co.</collectingCounty>
,
<collectingRegion id="49DCF863FFE09D0048E90245DD0C27D4" box="[987,1103,1919,1945]" country="United States of America" name="California" pageId="9" pageNumber="90">California</collectingRegion>
, collected
<date id="FFA61041FFE09D004FF60245DC3927D7" box="[1220,1402,1919,1946]" pageId="9" pageNumber="90" value="1956-02-09">
<collectingDate id="EFE2E9A9FFE09D004FF60245DC3927D7" box="[1220,1402,1919,1946]" pageId="9" pageNumber="90" value="1956-02-09">9 February 1956</collectingDate>
</date>
by
<collectionCode id="ED09AE44FFE09D004BA5029ED9EE27F3" box="[151,173,1956,1982]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="90">R</collectionCode>
.
<collectorName id="26ED5357FFE09D004B850299D87927F0" box="[183,314,1955,1981]" pageId="9" pageNumber="90">O. Schuster</collectorName>
, deposited in the
<collectingRegion id="49DCF863FFE09D004AC80299DBCA27F0" box="[506,649,1955,1981]" country="United States of America" name="Florida" pageId="9" pageNumber="90">Florida State</collectingRegion>
<location id="8EC7605AFFE09D0049BD0299DAE427F0" LSID="urn:lsid:plazi:treatment:03B18797FFE09D034BA50229DBB2219D:8EC7605AFFE09D0049BD0299DAE427F0" box="[655,935,1955,1981]" name="Collection of Arthropods" pageId="9" pageNumber="90" stateProvince="Florida">Collection of Arthropods</location>
,
<collectorName id="26ED5357FFE09D0048800299DD7727F0" box="[946,1076,1955,1981]" pageId="9" pageNumber="90">Gainesville</collectorName>
,
<collectingRegion id="49DCF863FFE09D004F0D0299DDD727F0" box="[1087,1172,1955,1981]" country="United States of America" name="Florida" pageId="9" pageNumber="90">Florida</collectingRegion>
.
<collectorName id="26ED5357FFE09D004FAE0299DD8B27F0" box="[1180,1224,1955,1981]" pageId="9" pageNumber="90">The</collectorName>
dissected
<typeStatus id="54A38823FFE09D004E080299DCDF27F0" box="[1338,1436,1955,1981]" pageId="9" pageNumber="90" type="holotype">holotype</typeStatus>
is mounted on SEM stub WS33-9, deposited in
<collectionCode id="ED09AE44FFE09D00499F02FDDBA427AC" box="[685,743,1991,2017]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="9" pageNumber="90" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE39D034BF505ACDBD420B5" pageId="10" pageNumber="91" type="diagnosis">
<paragraph id="8BA73681FFE39D034BF505ACDBD420B5" blockId="10.[151,1437,150,465]" pageId="10" pageNumber="91">
<emphasis id="B96CEA93FFE39D034BF505ACD80120FD" bold="true" box="[199,322,150,176]" pageId="10" pageNumber="91">Diagnosis.</emphasis>
The solenomere (
<emphasis id="B96CEA93FFE39D03492305ACDB7020FD" bold="true" box="[529,563,150,176]" pageId="10" pageNumber="91">sm</emphasis>
,
<figureCitation id="13232A04FFE39D03497005ACDBD720FC" box="[578,660,150,177]" captionStart="FIGURES 1316" captionStartId="10.[151,264,1502,1526]" captionTargetBox="[151,1436,488,1476]" captionTargetId="figure-154@10.[151,1436,488,1478]" captionTargetPageId="10" captionText="FIGURES 1316. Bidentogon buttensis, n. sp. 13. Head and anterior rings, dorsal view. 14. Midbody ring, dorsal view. 15. Gonopods in situ, ventral view. 16. Gonopod tips, ventral view. Abbreviations: ac, acropodite; pf, prefemorite; sm, solenomere; tz, terminal zone. SEM stub WS33-9." figureDoi="http://doi.org/10.5281/zenodo.4804923" httpUri="https://zenodo.org/record/4804923/files/figure.png" pageId="10" pageNumber="91">Fig. 16</figureCitation>
) is short and has a low tooth subdistally, nearly appearing bifid; the short terminal zone (
<emphasis id="B96CEA93FFE39D034A4D0580D8DF2099" bold="true" box="[383,412,186,212]" pageId="10" pageNumber="91">tz,</emphasis>
<figureCitation id="13232A04FFE39D034A900580D8B12098" box="[418,498,186,213]" captionStart="FIGURES 1316" captionStartId="10.[151,264,1502,1526]" captionTargetBox="[151,1436,488,1476]" captionTargetId="figure-154@10.[151,1436,488,1478]" captionTargetPageId="10" captionText="FIGURES 1316. Bidentogon buttensis, n. sp. 13. Head and anterior rings, dorsal view. 14. Midbody ring, dorsal view. 15. Gonopods in situ, ventral view. 16. Gonopod tips, ventral view. Abbreviations: ac, acropodite; pf, prefemorite; sm, solenomere; tz, terminal zone. SEM stub WS33-9." figureDoi="http://doi.org/10.5281/zenodo.4804923" httpUri="https://zenodo.org/record/4804923/files/figure.png" pageId="10" pageNumber="91">Fig. 16</figureCitation>
) is slightly expanded and also has a low subterminal tooth. There are 16 setae in the anterior marginal row of the collum (
<figureCitation id="13232A04FFE39D03490405E4DBC920B5" box="[566,650,222,248]" captionStart="FIGURES 1316" captionStartId="10.[151,264,1502,1526]" captionTargetBox="[151,1436,488,1476]" captionTargetId="figure-154@10.[151,1436,488,1478]" captionTargetPageId="10" captionText="FIGURES 1316. Bidentogon buttensis, n. sp. 13. Head and anterior rings, dorsal view. 14. Midbody ring, dorsal view. 15. Gonopods in situ, ventral view. 16. Gonopod tips, ventral view. Abbreviations: ac, acropodite; pf, prefemorite; sm, solenomere; tz, terminal zone. SEM stub WS33-9." figureDoi="http://doi.org/10.5281/zenodo.4804923" httpUri="https://zenodo.org/record/4804923/files/figure.png" pageId="10" pageNumber="91">Fig. 13</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE39D034BF50438DDA32151" box="[199,1248,258,285]" pageId="10" pageNumber="91" type="etymology">
<paragraph id="8BA73681FFE39D034BF50438DDA32151" blockId="10.[151,1437,150,465]" box="[199,1248,258,285]" pageId="10" pageNumber="91">
<emphasis id="B96CEA93FFE39D034BF50438D80D2151" bold="true" box="[199,334,258,284]" pageId="10" pageNumber="91">Etymology.</emphasis>
The species name, an adjective, is from the
<typeStatus id="54A38823FFE39D03480D0439DA2C2150" box="[831,879,259,285]" pageId="10" pageNumber="91">type</typeStatus>
locality in Butte Co.,
<collectingRegion id="49DCF863FFE39D034F550438DD9F2151" box="[1127,1244,258,284]" country="United States of America" name="California" pageId="10" pageNumber="91">California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE39D034BF5041CDA5E21E1" pageId="10" pageNumber="91" type="description">
<paragraph id="8BA73681FFE39D034BF5041CDA5E21E1" blockId="10.[151,1437,150,465]" pageId="10" pageNumber="91">
<emphasis id="B96CEA93FFE39D034BF5041CD81A210D" bold="true" box="[199,345,294,320]" pageId="10" pageNumber="91">Description.</emphasis>
Male
<typeStatus id="54A38823FFE39D034AAD041CDB46210D" box="[415,517,294,320]" pageId="10" pageNumber="91" type="holotype">holotype</typeStatus>
. Length about
<quantity id="4CE09B64FFE39D03499B041CDA44210C" box="[681,775,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="10" pageNumber="91" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">56 mm</quantity>
, maximum width
<quantity id="4CE09B64FFE39D0348E2041CDD6A210C" box="[976,1065,294,321]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="91" unit="mm" value="0.7">0.7 mm</quantity>
. Nonsexual characters typical for the genus. Femora of third legpair expanded, nearly spherical. Gonopod prefemora typically flattened (
<emphasis id="B96CEA93FFE39D034E240470DC752129" bold="true" box="[1302,1334,330,356]" pageId="10" pageNumber="91">pf,</emphasis>
<figureCitation id="13232A04FFE39D034E0F0470DCCC2128" box="[1341,1423,330,357]" captionStart="FIGURES 1316" captionStartId="10.[151,264,1502,1526]" captionTargetBox="[151,1436,488,1476]" captionTargetId="figure-154@10.[151,1436,488,1478]" captionTargetPageId="10" captionText="FIGURES 1316. Bidentogon buttensis, n. sp. 13. Head and anterior rings, dorsal view. 14. Midbody ring, dorsal view. 15. Gonopods in situ, ventral view. 16. Gonopod tips, ventral view. Abbreviations: ac, acropodite; pf, prefemorite; sm, solenomere; tz, terminal zone. SEM stub WS33-9." figureDoi="http://doi.org/10.5281/zenodo.4804923" httpUri="https://zenodo.org/record/4804923/files/figure.png" pageId="10" pageNumber="91">Fig. 15</figureCitation>
). Acropodite (
<emphasis id="B96CEA93FFE39D034A150454D80A21C5" bold="true" box="[295,329,366,392]" pageId="10" pageNumber="91">ac,</emphasis>
<figureCitation id="13232A04FFE39D034A600454D8E721C4" box="[338,420,366,393]" captionStart="FIGURES 1316" captionStartId="10.[151,264,1502,1526]" captionTargetBox="[151,1436,488,1476]" captionTargetId="figure-154@10.[151,1436,488,1478]" captionTargetPageId="10" captionText="FIGURES 1316. Bidentogon buttensis, n. sp. 13. Head and anterior rings, dorsal view. 14. Midbody ring, dorsal view. 15. Gonopods in situ, ventral view. 16. Gonopod tips, ventral view. Abbreviations: ac, acropodite; pf, prefemorite; sm, solenomere; tz, terminal zone. SEM stub WS33-9." figureDoi="http://doi.org/10.5281/zenodo.4804923" httpUri="https://zenodo.org/record/4804923/files/figure.png" pageId="10" pageNumber="91">Fig. 15</figureCitation>
) short, distally broadened, solenomere curved, nearly as long as terminal zone. Terminal zone with very low subterminal tooth. Females unknown.
</paragraph>
</subSubSection>
<subSubSection id="C302650AFFE39D034BF5048CDBB2219D" box="[199,753,438,465]" pageId="10" pageNumber="91" type="distribution">
<paragraph id="8BA73681FFE39D034BF5048CDBB2219D" blockId="10.[151,1437,150,465]" box="[199,753,438,465]" pageId="10" pageNumber="91">
<emphasis id="B96CEA93FFE39D034BF5048CD822219D" bold="true" box="[199,353,438,464]" pageId="10" pageNumber="91">Distribution.</emphasis>
Known only from the
<typeStatus id="54A38823FFE39D034952048DDBD3219C" box="[608,656,439,465]" pageId="10" pageNumber="91">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
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