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<document id="1E3847801E612C3B874D4B9E819937E8" ID-CLB-Dataset="95771" ID-DOI="10.11646/zootaxa.5165.2.8" ID-GBIF-Dataset="6da23e13-9225-433c-bb30-11578e6bdc84" ID-ISSN="1175-5326" ID-Zenodo-Dep="6831866" ID-ZooBank="34847ACE-E93D-4FD2-BA3E-CA86F8A45FC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657785003881" checkinUser="plazi" docAuthor="Datto-Liberato, Felipe H. &amp; Guillermo-Ferreira, Rhainer" docDate="2022" docId="03B18783FF8FAC4F689FFDF40472FF76" docLanguage="en" docName="zootaxa.5165.2.8.pdf" docOrigin="Zootaxa 5165 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="292" masterDocId="FF88FFFBFF8DAC4A6808FC66010AFFA3" masterDocTitle="A new species of Gomphoides Selys, 1854 (Odonata: Gomphidae) from the Environmental Protection Area of the Uberaba River, Brazil" masterLastPageNumber="293" masterPageNumber="287" pageNumber="289" updateTime="1699359029666" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F1BD02FFCC17D2D6A96D2F355A7BE557" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F0FE018CE61DECC227A5A94E9D2A04A8">
<mods:title id="1D0BAE74BCDC61A75CBDDB07C51311E5">A new species of Gomphoides Selys, 1854 (Odonata: Gomphidae) from the Environmental Protection Area of the Uberaba River, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="01583552A27CAA8064BBDA68170E9BD0" type="personal">
<mods:role id="03AD641940223BC10B64C558AA6524B2">
<mods:roleTerm id="0C431495BE3A7BDD20FED2B9072EAB36">Author</mods:roleTerm>
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<mods:namePart id="CACAB01187776B41C43C43E1C8B3E910">Datto-Liberato, Felipe H.</mods:namePart>
<mods:nameIdentifier id="F35A58E70E39DE4B6A6F65EF16CF847E" type="ORCID">0000-0002-4622-1592</mods:nameIdentifier>
<mods:affiliation id="6CA0271D648E0CC5B15F87AD757A48E6">Graduate Program in Entomology, Department of Biology, University of São Paulo (USP), Ribeirão Preto, Brazil. felipe. datto. liberato @ gmail. com; https: // orcid. org / 0000 - 0002 - 4622 - 1592 &amp; Lestes Lab., Federal University of Triângulo Mineiro, Uberaba, MG, Brazil.</mods:affiliation>
<mods:nameIdentifier id="8EB1AE74A70945F25076E6005DF2FA2D" type="email">felipe.datto.liberato@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="4058B5693305CB32D10F162451347D76" type="personal">
<mods:role id="F6DB9E1F61322ECD101A621E97B7E16E">
<mods:roleTerm id="5A778333056E176DED75E131AE9E475B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="981473EA4545F443F793D34576C8CD86">Guillermo-Ferreira, Rhainer</mods:namePart>
<mods:nameIdentifier id="36FC8C406EA701065457EB78968262BE" type="ORCID">0000-0002-4622-1592</mods:nameIdentifier>
<mods:affiliation id="9F8ADEC058ABF425A24DC35D22BEDA07">Graduate Program in Entomology, Department of Biology, University of São Paulo (USP), Ribeirão Preto, Brazil. felipe. datto. liberato @ gmail. com; https: // orcid. org / 0000 - 0002 - 4622 - 1592 &amp; Lestes Lab., Federal University of Triângulo Mineiro, Uberaba, MG, Brazil.</mods:affiliation>
<mods:nameIdentifier id="5E1E47308068D4DF0F9BA59B9104ACBB" type="email">felipe.datto.liberato@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="93CF4686843EC2326E473BE8279798AA">text</mods:typeOfResource>
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<mods:title id="690D95FAD3072ACD7944F94F01A48681">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="A49872425C63356D48D69FC0A4903739">
<mods:date id="AE00408B4296842FC5B844DCD9BE5AB1">2022</mods:date>
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<mods:number id="B3D0D7D73A10ED592A06BE5E91A7D980">2022-07-14</mods:number>
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<mods:detail id="193FB01E4ADA331316B5AF549ED8EAD8" type="volume">
<mods:number id="11531E27CE7EDF8292648F8BB0DD6DE4">5165</mods:number>
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<mods:number id="0D7F60D1E1D19DFF453B067DA1DE256E">2</mods:number>
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<mods:classification id="14484C092773E78210A20A002733756D">journal article</mods:classification>
<mods:identifier id="416549D99B4149665A8277A83FFFC3AA" type="CLB-Dataset">95771</mods:identifier>
<mods:identifier id="50AC0D68948BAD2FD7A11A89B74AE17F" type="DOI">10.11646/zootaxa.5165.2.8</mods:identifier>
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<mods:identifier id="D6EF0C15EB4F56D6EC0123D590197D99" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B18783FF8FAC4F689FFDF40472FF76" ID-DOI="http://doi.org/10.5281/zenodo.6831859" ID-GBIF-Taxon="197517063" ID-Zenodo-Dep="6831859" LSID="urn:lsid:plazi:treatment:03B18783FF8FAC4F689FFDF40472FF76" httpUri="http://treatment.plazi.org/id/03B18783FF8FAC4F689FFDF40472FF76" lastPageId="5" lastPageNumber="292" pageId="2" pageNumber="289">
<subSubSection id="C302651EFF8FAC48689FFDF40019FDBA" pageId="2" pageNumber="289" type="nomenclature">
<paragraph id="8BA73695FF8FAC48689FFDF402BEFE0E" blockId="2.[151,948,402,429]" box="[151,948,402,429]" pageId="2" pageNumber="289">
<heading id="D0EF81F9FF8FAC48689FFDF402BEFE0E" bold="true" box="[151,948,402,429]" fontSize="11" level="1" pageId="2" pageNumber="289" reason="1">
<emphasis id="B96CEA87FF8FAC48689FFDF402BEFE0E" bold="true" box="[151,948,402,429]" pageId="2" pageNumber="289">
<taxonomicName id="4C184D16FF8FAC48689FFDF40240FE0F" ID-CoL="9X62N" authority="Datto-Liberato &amp; Guillermo-Ferreira" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" box="[151,842,402,429]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="289" phylum="Arthropoda" rank="species" species="davi" status="sp. nov.">
<emphasis id="B96CEA87FF8FAC48689FFDF40063FE0E" bold="true" box="[151,361,402,429]" italics="true" pageId="2" pageNumber="289">Gomphoides davi</emphasis>
Datto-Liberato &amp; Guillermo-Ferreira
</taxonomicName>
,
<taxonomicNameLabel id="A25F57FCFF8FAC486B5EFDF502BEFE0E" box="[854,948,403,429]" pageId="2" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA73695FF8FAC48689FFDBC0019FDBA" blockId="2.[151,1436,474,681]" pageId="2" pageNumber="289">
<emphasis id="B96CEA87FF8FAC48689FFDBC0089FE57" bold="true" box="[151,387,474,500]" pageId="2" pageNumber="289">Specimen examined</emphasis>
.
<materialsCitation id="3B703CC8FF8FAC486998FDBD0019FDBA" ID-GBIF-Occurrence="3862677301" collectingDate="2022-03-18" collectorName="Liberato, F. H. D." country="Brazil" elevation="873" location="Limo" municipality="Uberaba" pageId="2" pageNumber="289" specimenCount="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="54A38837FF8FAC486998FDBD00F2FE56" box="[400,504,475,501]" pageId="2" pageNumber="289" type="holotype">Holotype</typeStatus>
(
<specimenCount id="9D1EFD1CFF8FAC486A00FDBD0326FE56" box="[520,556,475,501]" pageId="2" pageNumber="289" type="male">1♂</specimenCount>
,
<collectingCountry id="F30F7605FF8FAC486A3EFDBD0394FE56" box="[566,670,475,501]" name="Brazil" pageId="2" pageNumber="289">BRAZIL</collectingCountry>
,
<collectingRegion id="49DCF877FF8FAC486AA0FDBD024BFE56" box="[680,833,475,501]" country="Brazil" name="Minas Gerais" pageId="2" pageNumber="289">Minas Gerais</collectingRegion>
:
<collectingMunicipality id="6BC3ACEFFF8FAC486B42FDBD02A3FE56" box="[842,937,475,501]" pageId="2" pageNumber="289">Uberaba</collectingMunicipality>
(
<location id="8EC7604EFF8FAC486BBEFDBD02FEFE56" LSID="urn:lsid:plazi:treatment:03B18783FF8FAC4F689FFDF40472FF76:8EC7604EFF8FAC486BBEFDBD02FEFE56" box="[950,1012,475,501]" country="Brazil" municipality="Uberaba" name="Limo" pageId="2" pageNumber="289" stateProvince="Minas Gerais">Limo</location>
stream,
<quantity id="4CE09B70FF8FAC486C59FDBD059FFE56" box="[1105,1173,475,501]" metricMagnitude="2" metricUnit="m" metricValue="8.73" pageId="2" pageNumber="289" unit="m" value="873.0">
<elevation id="0035D1A6FF8FAC486C59FDBD059FFE56" box="[1105,1173,475,501]" metricMagnitude="2" metricUnit="m" metricValue="8.73" pageId="2" pageNumber="289" unit="m" value="873.0">873m</elevation>
</quantity>
),
<date id="FFA61055FF8FAC486CAFFDBD0427FE56" box="[1191,1325,475,501]" pageId="2" pageNumber="289" value="2022-03-18">
<collectingDate id="EFE2E9BDFF8FAC486CAFFDBD0427FE56" box="[1191,1325,475,501]" pageId="2" pageNumber="289" value="2022-03-18">18-III-2022</collectingDate>
</date>
,
<collectorName id="26ED5343FF8FAC486D3EFDBD01E8FDBA" pageId="2" pageNumber="289">Liberato, F.H.D.</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C302651EFF8FAC4868CFFE450471FD9E" box="[199,1403,547,573]" pageId="2" pageNumber="289" type="etymology">
<paragraph id="8BA73695FF8FAC4868CFFE450471FD9E" blockId="2.[151,1436,474,681]" box="[199,1403,547,573]" pageId="2" pageNumber="289">
<emphasis id="B96CEA87FF8FAC4868CFFE450042FD9E" bold="true" box="[199,328,547,573]" pageId="2" pageNumber="289">Etymology</emphasis>
. This species is named after Rhainer Guillermo Ferreira’s beloved son, Davi Vilarinho Ferreira.
</paragraph>
</subSubSection>
<subSubSection id="C302651EFF8FAC4968CFFE200094FD52" lastPageId="3" lastPageNumber="290" pageId="2" pageNumber="289" type="description">
<paragraph id="8BA73695FF8FAC4868CFFE2003B4FDC2" blockId="2.[151,1436,474,681]" box="[199,702,582,609]" pageId="2" pageNumber="289">
<emphasis id="B96CEA87FF8FAC4868CFFE200300FDC2" bold="true" box="[199,522,582,609]" pageId="2" pageNumber="289">
Description of the
<typeStatus id="54A38837FF8FAC4869ABFE210300FDC2" box="[419,522,583,609]" pageId="2" pageNumber="289" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="13232A10FF8FAC486A10FE2103B8FDC2" box="[536,690,583,609]" captionStart-0="FIGURE 4" captionStart-1="FIGURES 5–9" captionStart-2="FIGURE 10–16" captionStartId-0="2.[151,250,1924,1948]" captionStartId-1="3.[151,264,1546,1570]" captionStartId-2="4.[151,250,1879,1903]" captionTargetBox-0="[151,1436,702,1898]" captionTargetBox-1="[158,1427,784,1516]" captionTargetBox-2="[238,1350,414,1843]" captionTargetId-0="figure-168@2.[151,1436,702,1898]" captionTargetId-1="figure-280@3.[151,1436,777,1523]" captionTargetId-2="figure-125@4.[238,1350,414,1853]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 4. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Male habitus. Bar = 20mm." captionText-1="FIGURES 5–9. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9)." captionText-2="FIGURE 10–16. Gomphoides infumata, male from REGUA, Rio de Janeiro, Brazil (10), cerci and epiproct in lateral view (11) (© Tom Kompier); G. perdita, male from Uberlândia, Minas Gerais, Brazil (12), cerci and epiproct in lateral view (13) (© Diogo Vilela); G. praevia, male holotype, habitus in dorsal view (14), cerci and epiproct in lateral (15) and dorsal (16) views (© NHMW NoaS Image Collection / Harald Bruckner)." figureDoi-0="http://doi.org/10.5281/zenodo.6831872" figureDoi-1="http://doi.org/10.5281/zenodo.6831877" figureDoi-2="http://doi.org/10.5281/zenodo.6831881" httpUri-0="https://zenodo.org/record/6831872/files/figure.png" httpUri-1="https://zenodo.org/record/6831877/files/figure.png" httpUri-2="https://zenodo.org/record/6831881/files/figure.png" pageId="2" pageNumber="289">Figures 4–10</figureCitation>
).
</paragraph>
<paragraph id="8BA73695FF8FAC4868CFFE0D03A7FD0A" blockId="2.[151,1436,474,681]" pageId="2" pageNumber="289">
<emphasis id="B96CEA87FF8FAC4868CFFE0D000EFD26" box="[199,260,619,645]" italics="true" pageId="2" pageNumber="289">Head</emphasis>
. Labrum, labium, clypeus and antefrons yellow. Postfrons and vertex black. Occipital bar yellow, with long black setae in a posterior longitudinal row.
</paragraph>
<caption id="DF67661DFF8FAC48689FFBE205CFF83F" ID-DOI="http://doi.org/10.5281/zenodo.6831872" ID-Zenodo-Dep="6831872" box="[151,1221,1924,1948]" httpUri="https://zenodo.org/record/6831872/files/figure.png" pageId="2" pageNumber="289" startId="2.[151,250,1924,1948]" targetBox="[151,1436,702,1898]" targetPageId="2">
<paragraph id="8BA73695FF8FAC48689FFBE205CFF83F" blockId="2.[151,1221,1924,1948]" box="[151,1221,1924,1948]" pageId="2" pageNumber="289">
<emphasis id="B96CEA87FF8FAC48689FFBE20007F83F" bold="true" box="[151,269,1924,1948]" pageId="2" pageNumber="289">FIGURE 4</emphasis>
.
<taxonomicName id="4C184D16FF8FAC486911FBE20246F83F" authority="Datto-Liberato &amp; Guillermo-Ferreira" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" box="[281,844,1924,1948]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="289" phylum="Arthropoda" rank="species" species="davi" status="sp. nov.">
<emphasis id="B96CEA87FF8FAC486911FBE200C6F83F" box="[281,460,1924,1948]" italics="true" pageId="2" pageNumber="289">Gomphoides davi</emphasis>
Datto-Liberato &amp; Guillermo-Ferreira
</taxonomicName>
,
<emphasis id="B96CEA87FF8FAC486B5EFBE202ADF83F" bold="true" box="[854,935,1924,1948]" pageId="2" pageNumber="289">
<taxonomicNameLabel id="A25F57FCFF8FAC486B5EFBE202ADF83F" box="[854,935,1924,1948]" pageId="2" pageNumber="289" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male habitus. Bar = 20mm.
</paragraph>
</caption>
<paragraph id="8BA73695FF8EAC4968CFFCF105D1FEC6" blockId="3.[151,1437,151,753]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFCF10012FF12" box="[199,280,151,177]" italics="true" pageId="3" pageNumber="290">Thorax</emphasis>
. Prothorax black, with long black hair-like setae on posterior margin. Pterothorax dark brown. Mesepisternum with two yellow stripes, yellow on anterior margin, with long black hair-like setae (
<figureCitation id="13232A10FF8EAC496C98FCDD05FAFF76" box="[1168,1264,186,213]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[158,1427,784,1516]" captionTargetId="figure-280@3.[151,1436,777,1523]" captionTargetPageId="3" captionText="FIGURES 5–9. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9)." figureDoi="http://doi.org/10.5281/zenodo.6831877" httpUri="https://zenodo.org/record/6831877/files/figure.png" pageId="3" pageNumber="290">Figure 5</figureCitation>
); mesepimeron with yellow stripe and small setae; metepisternum with yellow stripe, with small black setae; metepimeron with a wide yellow stripe. Legs mostly yellowish brown and black; coxae and trochanters yellowish brown; dorsal-anterior surface of femora yellowish brown and ventroposterior (extensor) surface black; tibiae black; tarsi black; hind femur with several short spines and tibiae with 13 long spines on ventrolateral (extensor) surface.
</paragraph>
<paragraph id="8BA73695FF8EAC4968CFFD0905D9FE0E" blockId="3.[151,1437,151,753]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFD090001FE2B" box="[199,267,367,392]" italics="true" pageId="3" pageNumber="290">Wings</emphasis>
: hyaline (
<figureCitation id="13232A10FF8EAC496973FD0900D7FE2A" box="[379,477,367,393]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[158,1427,784,1516]" captionTargetId="figure-280@3.[151,1436,777,1523]" captionTargetPageId="3" captionText="FIGURES 5–9. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9)." figureDoi="http://doi.org/10.5281/zenodo.6831877" httpUri="https://zenodo.org/record/6831877/files/figure.png" pageId="3" pageNumber="290">Figure 6</figureCitation>
); venation brown to black, pt yellowish brown.
<emphasis id="B96CEA87FF8EAC496BFEFD090552FE2B" box="[1014,1112,367,392]" italics="true" pageId="3" pageNumber="290">Venation</emphasis>
: 17 Ax in Fw; 14 Ax in HW; Fw triangle with three cells, subtriangle with four cells; Hw triangle and subtriangle with 3 cells.
</paragraph>
<paragraph id="8BA73695FF8EAC4968CFFDD10564FE72" blockId="3.[151,1437,151,753]" box="[199,1134,439,465]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFDD10089FE72" box="[199,387,439,465]" italics="true" pageId="3" pageNumber="290">Vesica spermalis</emphasis>
: vesica spermalis distal segment with two long flagella (
<figureCitation id="13232A10FF8EAC496BF6FDD1056AFE72" box="[1022,1120,439,465]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[158,1427,784,1516]" captionTargetId="figure-280@3.[151,1436,777,1523]" captionTargetPageId="3" captionText="FIGURES 5–9. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9)." figureDoi="http://doi.org/10.5281/zenodo.6831877" httpUri="https://zenodo.org/record/6831877/files/figure.png" pageId="3" pageNumber="290">Figure 7</figureCitation>
).
</paragraph>
<paragraph id="8BA73695FF8EAC4968CFFDBD0551FD9E" blockId="3.[151,1437,151,753]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFDBD003BFE56" box="[199,305,475,501]" italics="true" pageId="3" pageNumber="290">Abdomen</emphasis>
. S1–3 mostly yellow; S3–9 yellow anteriorly and black posteriorly; S10 black with a dorsal yellow spot, longer than S9. Hamuli black (
<figureCitation id="13232A10FF8EAC496A27FD990398FDBA" box="[559,658,511,537]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[158,1427,784,1516]" captionTargetId="figure-280@3.[151,1436,777,1523]" captionTargetPageId="3" captionText="FIGURES 5–9. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9)." figureDoi="http://doi.org/10.5281/zenodo.6831877" httpUri="https://zenodo.org/record/6831877/files/figure.png" pageId="3" pageNumber="290">Figure 8</figureCitation>
). Cerci mostly yellow with black anterior margin and ventral projection, lanceolate posteriorly, with black apical blunted tooth (
<figureCitation id="13232A10FF8EAC496B30FE450290FD9E" box="[824,922,547,573]" captionStart="FIGURES 5–9" captionStartId="3.[151,264,1546,1570]" captionTargetBox="[158,1427,784,1516]" captionTargetId="figure-280@3.[151,1436,777,1523]" captionTargetPageId="3" captionText="FIGURES 5–9. Gomphoides davi Datto-Liberato &amp; Guillermo-Ferreira, sp. nov. Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9)." figureDoi="http://doi.org/10.5281/zenodo.6831877" httpUri="https://zenodo.org/record/6831877/files/figure.png" pageId="3" pageNumber="290">Figure 9</figureCitation>
). Epiproct black.
</paragraph>
<paragraph id="8BA73695FF8EAC4968CFFE2102E5FD6E" blockId="3.[151,1437,151,753]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFE210063FDC3" box="[199,361,583,608]" italics="true" pageId="3" pageNumber="290">Measurements</emphasis>
. Total length (including caudal appendages) (60.0); abdomen length (excluding caudal appendages) (38.0); head maximum width (8.15); Fw length (37.0); Hw length (36.30); Hw width (proximal to costal nodus) (9.8); pt length (5.5) in Fw, (5.5) in Hw; length of metathoracic femur (6.0); metathoracic tibia (4.0); total length of cercus in lateral view (1.9); length of epiproct in lateral view (0.8).
</paragraph>
<paragraph id="8BA73695FF8EAC4968CFFEB10094FD52" blockId="3.[151,1437,151,753]" box="[199,414,727,753]" pageId="3" pageNumber="290">Female. Unknown.</paragraph>
</subSubSection>
<caption id="DF67661DFF8EAC49689FFA6C004BF9C9" ID-DOI="http://doi.org/10.5281/zenodo.6831877" ID-Zenodo-Dep="6831877" httpUri="https://zenodo.org/record/6831877/files/figure.png" pageId="3" pageNumber="290" startId="3.[151,264,1546,1570]" targetBox="[158,1427,784,1516]" targetPageId="3">
<paragraph id="8BA73695FF8EAC49689FFA6C004BF9C9" blockId="3.[151,1437,1546,1642]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC49689FFA6C003FF981" bold="true" box="[151,309,1546,1570]" pageId="3" pageNumber="290">FIGURES 5–9</emphasis>
.
<taxonomicName id="4C184D16FF8EAC49694CFA6C0276F981" authority="Datto-Liberato &amp; Guillermo-Ferreira" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" box="[324,892,1546,1570]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="davi" status="sp. nov.">
<emphasis id="B96CEA87FF8EAC49694CFA6C00F2F981" box="[324,504,1546,1570]" italics="true" pageId="3" pageNumber="290">Gomphoides davi</emphasis>
Datto-Liberato &amp; Guillermo-Ferreira
</taxonomicName>
,
<emphasis id="B96CEA87FF8EAC496B80FA6C02D0F981" bold="true" box="[904,986,1546,1570]" pageId="3" pageNumber="290">
<taxonomicNameLabel id="A25F57FCFF8EAC496B80FA6C02D0F981" box="[904,986,1546,1570]" pageId="3" pageNumber="290" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Coloration pattern of thorax in dorsal view (5); Fw and Hw (6); vesica spermalis in ventral and lateral views (7); hamuli in lateral view (8); male appendages in lateral and dorsal views (9).
</paragraph>
</caption>
<subSubSection id="C302651EFF8EAC4968CFFAFE021EF959" pageId="3" pageNumber="290" type="discussion">
<paragraph id="8BA73695FF8EAC4968CFFAFE021EF959" blockId="3.[151,1437,1688,2039]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFAFE0031F911" bold="true" box="[199,315,1688,1714]" pageId="3" pageNumber="290">Remarks.</emphasis>
The male was collected basking at a sunny spot on a tree, with sandy soil below, around dusk (between 17–18h). An intensive survey during twelve days of fieldwork was conducted on the
<typeStatus id="54A38837FF8EAC496CB0FADB05E2F974" box="[1208,1256,1725,1751]" pageId="3" pageNumber="290">type</typeStatus>
locality and all around the EPA-UR and no other specimens were found.
</paragraph>
</subSubSection>
<subSubSection id="C302651EFF8EAC4F68CFFB620472FF76" lastPageId="5" lastPageNumber="292" pageId="3" pageNumber="290" type="diagnosis">
<paragraph id="8BA73695FF8EAC4E68CFFB6201C0FF76" blockId="3.[151,1437,1688,2039]" lastBlockId="4.[151,1437,151,393]" lastPageId="4" lastPageNumber="291" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC4968CFFB6200C5F8BD" bold="true" box="[199,463,1796,1822]" pageId="3" pageNumber="290">Differential diagnosis.</emphasis>
This species can be distinguished from other
<taxonomicName id="4C184D16FF8EAC496BDEFB6205B6F8BD" box="[982,1212,1796,1822]" pageId="3" pageNumber="290">
<emphasis id="B96CEA87FF8EAC496BDEFB62056EF8BD" box="[982,1124,1796,1822]" italics="true" pageId="3" pageNumber="290">Gomphoides</emphasis>
species
</taxonomicName>
(
<figureCitation id="13232A10FF8EAC496CC4FB62047AF8BD" box="[1228,1392,1796,1822]" captionStart="FIGURE 10–16" captionStartId="4.[151,250,1879,1903]" captionTargetBox="[238,1350,414,1843]" captionTargetId="figure-125@4.[238,1350,414,1853]" captionTargetPageId="4" captionText="FIGURE 10–16. Gomphoides infumata, male from REGUA, Rio de Janeiro, Brazil (10), cerci and epiproct in lateral view (11) (© Tom Kompier); G. perdita, male from Uberlândia, Minas Gerais, Brazil (12), cerci and epiproct in lateral view (13) (© Diogo Vilela); G. praevia, male holotype, habitus in dorsal view (14), cerci and epiproct in lateral (15) and dorsal (16) views (© NHMW NoaS Image Collection / Harald Bruckner)." figureDoi="http://doi.org/10.5281/zenodo.6831881" httpUri="https://zenodo.org/record/6831881/files/figure.png" pageId="3" pageNumber="290">Figures 10–16</figureCitation>
) by the shape of epiproct and the characters summarized in
<tableCitation id="C69A032EFF8EAC496B01FB4E0254F8E1" box="[777,862,1832,1858]" captionStart="TABLE 1" captionStartId="5.[151,239,260,284]" captionTargetPageId="5" captionText="TABLE 1. Characters that may be used to distinguish between males of Gomphoides species." httpUri="http://table.plazi.org/id/DF67661DFF88AC4F689FFD620575FEBF" pageId="3" pageNumber="290" tableUuid="DF67661DFF88AC4F689FFD620575FEBF">Table 1</tableCitation>
.
<taxonomicName id="4C184D16FF8EAC496B63FB4E053AF8E1" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" box="[875,1072,1832,1858]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="davi">
<emphasis id="B96CEA87FF8EAC496B63FB4E053AF8E1" box="[875,1072,1832,1858]" italics="true" pageId="3" pageNumber="290">Gomphoides davi</emphasis>
</taxonomicName>
has the shortest epiproct among its congeners, and the unique combination of four cells in Fw subtriangle and four cells in the Hw anal loop.
<taxonomicName id="4C184D16FF8EAC496D54FB2B002DF829" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="davi">
<emphasis id="B96CEA87FF8EAC496D54FB2B002DF829" italics="true" pageId="3" pageNumber="290">Gomphoides davi</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C184D16FF8EAC4969A2FB170314F829" baseAuthorityName="Foerster" baseAuthorityYear="1914" box="[426,542,1904,1931]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="perdita">
<emphasis id="B96CEA87FF8EAC4969A2FB170314F829" box="[426,542,1904,1931]" italics="true" pageId="3" pageNumber="290">G. perdita</emphasis>
</taxonomicName>
, but it is distinguished from the latter by the S10 black, yellow in
<taxonomicName id="4C184D16FF8EAC496D28FB17049FF829" baseAuthorityName="Foerster" baseAuthorityYear="1914" box="[1312,1429,1904,1931]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="perdita">
<emphasis id="B96CEA87FF8EAC496D28FB17049FF829" box="[1312,1429,1904,1931]" italics="true" pageId="3" pageNumber="290">G. perdita</emphasis>
</taxonomicName>
; four cells in the subtriangle of Fw, there are three cells in
<taxonomicName id="4C184D16FF8EAC496B31FBF302A4F80D" baseAuthorityName="Foerster" baseAuthorityYear="1914" box="[825,942,1940,1967]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="perdita">
<emphasis id="B96CEA87FF8EAC496B31FBF302A4F80D" box="[825,942,1940,1967]" italics="true" pageId="3" pageNumber="290">G. perdita</emphasis>
</taxonomicName>
; epiproct shorter than half of the length of cerci, while epiproct is more than half the length of the cerci in
<taxonomicName id="4C184D16FF8EAC496B7DFBDF02E3F871" baseAuthorityName="Foerster" baseAuthorityYear="1914" box="[885,1001,1976,2003]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="perdita">
<emphasis id="B96CEA87FF8EAC496B7DFBDF02E3F871" box="[885,1001,1976,2003]" italics="true" pageId="3" pageNumber="290">G. perdita</emphasis>
</taxonomicName>
. Dorsal projections on S8–9 are more pronounced in
<taxonomicName id="4C184D16FF8EAC496936FBBB009AF855" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" box="[318,400,2012,2039]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="davi">
<emphasis id="B96CEA87FF8EAC496936FBBB009AF855" box="[318,400,2012,2039]" italics="true" pageId="3" pageNumber="290">G. davi</emphasis>
</taxonomicName>
are similar to
<taxonomicName id="4C184D16FF8EAC496A38FBBB03A8F855" baseAuthorityName="Foerster" baseAuthorityYear="1914" box="[560,674,2012,2039]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="perdita">
<emphasis id="B96CEA87FF8EAC496A38FBBB03A8F855" box="[560,674,2012,2039]" italics="true" pageId="3" pageNumber="290">G. perdita</emphasis>
</taxonomicName>
, more pronounced in
<taxonomicName id="4C184D16FF8EAC496B9EFBBB0516F855" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[918,1052,2012,2039]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="290" phylum="Arthropoda" rank="species" species="infumata">
<emphasis id="B96CEA87FF8EAC496B9EFBBB0516F855" box="[918,1052,2012,2039]" italics="true" pageId="3" pageNumber="290">G. infumata</emphasis>
</taxonomicName>
. Lateral flanges well developed in
<taxonomicName id="4C184D16FF89AC4E689FFCF1001AFF13" authorityName="St Quentin" authorityYear="1967" box="[151,272,151,177]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="291" phylum="Arthropoda" rank="species" species="praevia">
<emphasis id="B96CEA87FF89AC4E689FFCF1001AFF13" box="[151,272,151,177]" italics="true" pageId="4" pageNumber="291">G. praevia</emphasis>
</taxonomicName>
, when compared to the other three species. Vesica spermalis of
<taxonomicName id="4C184D16FF89AC4E6BD6FCF1053AFF12" authorityName="Datto-Liberato &amp; Guillermo-Ferreira" authorityYear="2022" box="[990,1072,151,177]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="291" phylum="Arthropoda" rank="species" species="davi">
<emphasis id="B96CEA87FF89AC4E6BD6FCF1053AFF12" box="[990,1072,151,177]" italics="true" pageId="4" pageNumber="291">G. davi</emphasis>
</taxonomicName>
is similar to the other three species.
</paragraph>
<paragraph id="8BA73695FF89AC4E68CFFCB9006FFE2A" blockId="4.[151,1437,151,393]" pageId="4" pageNumber="291">
The fact that this species was found in an area under environmental protection gives hope for the conservation of its biodiversity, but at the same time raises an alarming concern due to the increasing impacts and threats to the rivers and streams of the Uberaba River basin (
<bibRefCitation id="EF894B64FF89AC4E6AB9FD410288FEE2" author="Valera, C. A. &amp; Valle Junior, R. F. &amp; Varandas, S. G. P. &amp; Sanches Fernandes, L. F. &amp; Pacheco, F. A. L." box="[689,898,295,321]" pageId="4" pageNumber="291" pagination="463 - 473" refId="ref3193" refString="Valera, C. A., Valle Junior, R. F., Varandas, S. G. P., Sanches Fernandes, L. F. &amp; Pacheco, F. A. L. (2016) The role of environmental land use conflicts in soil fertility: A study on the Uberaba River basin, Brazil. Science of the Total Environment, 562, 463 - 473. https: // doi. org / 10.1016 / j. scitotenv. 2016.04.046" type="journal article" year="2016">
Valera
<emphasis id="B96CEA87FF89AC4E6B0BFD41023CFEE2" box="[771,822,295,321]" italics="true" pageId="4" pageNumber="291">et al</emphasis>
. 2016
</bibRefCitation>
). The EPA-UR has areas occupied by degraded pastures (
<bibRefCitation id="EF894B64FF89AC4E6908FD2D00CEFEC6" author="Vieira, A. S. &amp; do Valle Junior, R. F. &amp; Rodrigues, V. S. &amp; da Silva Quinaia, T. L. &amp; Mendes, R. G. &amp; Valera, C. A. &amp; Fernandes, L. F. S. &amp; Pacheco, F. A. L." box="[256,452,331,357]" pageId="4" pageNumber="291" pagination="146019" refId="ref3357" refString="Vieira, A. S., do Valle Junior, R. F., Rodrigues, V. S., da Silva Quinaia, T. L., Mendes, R. G., Valera, C. A., Fernandes, L. F. S. &amp; Pacheco, F. A. L. (2021) Estimating water erosion from the brightness index of orbital images: A framework for the prognosis of degraded pastures. Science of the Total Environment, 776, 146019. https: // doi. org / 10.1016 / j. scitotenv. 2021.146019" type="journal article" year="2021">
Vieira
<emphasis id="B96CEA87FF89AC4E6944FD2D0076FEC6" box="[332,380,331,357]" italics="true" pageId="4" pageNumber="291">et al</emphasis>
. 2021
</bibRefCitation>
), which increase soil erosion and impact water resources and aquatic biodiversity (
<bibRefCitation id="EF894B64FF89AC4E6D5BFD2D001BFE2A" author="Valera, C. A. &amp; Valle Junior, R. F. &amp; Varandas, S. G. P. &amp; Sanches Fernandes, L. F. &amp; Pacheco, F. A. L." pageId="4" pageNumber="291" pagination="463 - 473" refId="ref3193" refString="Valera, C. A., Valle Junior, R. F., Varandas, S. G. P., Sanches Fernandes, L. F. &amp; Pacheco, F. A. L. (2016) The role of environmental land use conflicts in soil fertility: A study on the Uberaba River basin, Brazil. Science of the Total Environment, 562, 463 - 473. https: // doi. org / 10.1016 / j. scitotenv. 2016.04.046" type="journal article" year="2016">
Valera
<emphasis id="B96CEA87FF89AC4E689FFD0901C5FE2A" box="[151,207,367,393]" italics="true" pageId="4" pageNumber="291">et al.</emphasis>
2016
</bibRefCitation>
,
<bibRefCitation id="EF894B64FF89AC4E6914FD090053FE2A" author="Valera, C. A. &amp; Pissarra, T. C. T. &amp; Filho, M. V. M. &amp; do Valle Junior, R. F. &amp; Oliveira, C. F. &amp; Moura, J. P. &amp; Fernandes, L. F. S. &amp; Pacheco, F. A. L." box="[284,345,367,393]" pageId="4" pageNumber="291" pagination="549" refId="ref3077" refString="Valera, C. A., Pissarra, T. C. T., Filho, M. V. M., do Valle Junior, R. F., Oliveira, C. F., Moura, J. P., Fernandes, L. F. S. &amp; Pacheco, F. A. L. (2019) The buffer capacity of riparian vegetation to control water quality in anthropogenic catchments from a legally protected area: A critical view over the Brazilian new forest code. Water, Switzerland, 11, 549. https: // doi. org / 10.3390 / w 11030549" type="journal article" year="2019">2019</bibRefCitation>
).
</paragraph>
<caption id="DF67661DFF89AC4E689FFB3103A1F878" ID-DOI="http://doi.org/10.5281/zenodo.6831881" ID-Zenodo-Dep="6831881" httpUri="https://zenodo.org/record/6831881/files/figure.png" pageId="4" pageNumber="291" startId="4.[151,250,1879,1903]" targetBox="[238,1350,414,1843]" targetPageId="4">
<paragraph id="8BA73695FF89AC4E689FFB3103A1F878" blockId="4.[151,1437,1879,2012]" pageId="4" pageNumber="291">
<emphasis id="B96CEA87FF89AC4E689FFB310048F8CC" bold="true" box="[151,322,1879,1903]" pageId="4" pageNumber="291">FIGURE 10–16</emphasis>
.
<taxonomicName id="4C184D16FF89AC4E6959FB31033EF8CC" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[337,564,1879,1903]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="291" phylum="Arthropoda" rank="species" species="infumata">
<emphasis id="B96CEA87FF89AC4E6959FB31033EF8CC" box="[337,564,1879,1903]" italics="true" pageId="4" pageNumber="291">Gomphoides infumata</emphasis>
</taxonomicName>
, male from REGUA, Rio de Janeiro, Brazil (10), cerci and epiproct in lateral view (11) (© Tom Kompier);
<taxonomicName id="4C184D16FF89AC4E6985FB1A00FEF830" baseAuthorityName="Foerster" baseAuthorityYear="1914" box="[397,500,1915,1939]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="291" phylum="Arthropoda" rank="species" species="perdita">
<emphasis id="B96CEA87FF89AC4E6985FB1A00FEF830" box="[397,500,1915,1939]" italics="true" pageId="4" pageNumber="291">G. perdita</emphasis>
</taxonomicName>
, male from Uberlândia, Minas Gerais, Brazil (12), cerci and epiproct in lateral view (13) (© Diogo Vilela);
<taxonomicName id="4C184D16FF89AC4E6925FBC60093F814" authorityName="St Quentin" authorityYear="1967" box="[301,409,1952,1975]" class="Insecta" family="Gomphidae" genus="Gomphoides" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="291" phylum="Arthropoda" rank="species" species="praevia">
<emphasis id="B96CEA87FF89AC4E6925FBC60093F814" box="[301,409,1952,1975]" italics="true" pageId="4" pageNumber="291">G. praevia</emphasis>
</taxonomicName>
, male holotype, habitus in dorsal view (14), cerci and epiproct in lateral (15) and dorsal (16) views (© NHMW NoaS Image Collection / Harald Bruckner).
</paragraph>
</caption>
<paragraph id="8BA73695FF88AC4F68CFFCF10472FF76" blockId="5.[151,1436,151,213]" pageId="5" pageNumber="292">
The distribution of all four
<taxonomicName id="4C184D16FF88AC4F6A00FCF103FAFF12" box="[520,752,151,177]" pageId="5" pageNumber="292">
<emphasis id="B96CEA87FF88AC4F6A00FCF1039CFF12" box="[520,662,151,177]" italics="true" pageId="5" pageNumber="292">Gomphoides</emphasis>
species
</taxonomicName>
and the recent new records, including the species here described, shows that collection gaps for Neotropical gomphids is still an issue that requires attention (
<figureCitation id="13232A10FF88AC4F6CF2FCDD0461FF76" box="[1274,1387,187,213]" captionStart="FIGURE 17" captionStartId="5.[151,250,1575,1599]" captionTargetBox="[160,1430,624,1551]" captionTargetId="figure-142@5.[151,1436,624,1551]" captionTargetPageId="5" captionText="FIGURE 17. Distribution of the hitherto known species of Gomphoides in South America, based on minimal convex polygons using sites of occurrence." figureDoi="http://doi.org/10.5281/zenodo.6831883" httpUri="https://zenodo.org/record/6831883/files/figure.png" pageId="5" pageNumber="292">Figure 17</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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